treatments-xml/data/14/7B/C4/147BC44BC44C2D5EFF69F952C7547C0B.xml
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<document id="769414A6C294C6517AF3F1263C258C0D" ID-CLB-Dataset="9446" ID-DOI="10.11646/zootaxa.4885.4.6" ID-GBIF-Dataset="c7579c32-46c4-4a34-86ee-cd06de94a790" ID-ISSN="1175-5326" ID-Zenodo-Dep="4296955" ID-ZooBank="42DEF6F1-9C16-48AD-BD61-8B156111C20D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606684780813" checkinUser="plazi" docAuthor="Salazar-Vallejo, Sergio I." docDate="2020" docId="147BC44BC44C2D5EFF69F952C7547C0B" docLanguage="en" docName="zootaxa.4885.4.6.pdf" docOrigin="Zootaxa 4885 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Saphobranchia omorpha Salazar-Vallejo 2020, n. sp." docType="treatment" docVersion="9" lastPageNumber="574" masterDocId="E842BC33C4472D50FFFEFFB8C4647F32" masterDocTitle="Four new deep-water flabelligerid species from Pacific Costa Rica (Annelida Sedentaria, Flabelligeridae)" masterLastPageNumber="578" masterPageNumber="560" pageNumber="571" updateTime="1698846847864" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E4C89E5126A85A4CB29ABA38063C4F05">Four new deep-water flabelligerid species from Pacific Costa Rica (Annelida Sedentaria, Flabelligeridae)</mods:title>
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<mods:namePart id="94D0026BC9834AB945CC00971C665DA5">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:date id="2C4AF6E6DCDFE92420A137BC859B3A24">2020</mods:date>
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<mods:number id="A2304F28BA645C85B85B4DA6FE805105">2020-11-27</mods:number>
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<treatment id="147BC44BC44C2D5EFF69F952C7547C0B" ID-DOI="http://doi.org/10.5281/zenodo.4329990" ID-GBIF-Taxon="170618220" ID-Zenodo-Dep="4329990" LSID="urn:lsid:plazi:treatment:147BC44BC44C2D5EFF69F952C7547C0B" httpUri="http://treatment.plazi.org/id/147BC44BC44C2D5EFF69F952C7547C0B" lastPageId="14" lastPageNumber="574" pageId="11" pageNumber="571">
<subSubSection id="D4C826D6C44C2D5BFF69F952C7AE781B" pageId="11" pageNumber="571" type="nomenclature">
<paragraph id="9C6D755DC44C2D5BFF69F952C66B7837" blockId="11.[151,970,1770,1869]" box="[151,527,1770,1797]" pageId="11" pageNumber="571">
<heading id="C725C231C44C2D5BFF69F952C66B7837" bold="true" box="[151,527,1770,1797]" fontSize="11" level="1" pageId="11" pageNumber="571" reason="1">
<emphasis id="AEA6A94FC44C2D5BFF69F952C66B7837" bold="true" box="[151,527,1770,1797]" pageId="11" pageNumber="571">
<taxonomicName id="5BD20EDEC44C2D5BFF69F952C5A17837" ID-CoL="85RYN" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[151,453,1770,1797]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="571" phylum="Annelida" rank="species" species="omorpha" status="sp. nov.">
<emphasis id="AEA6A94FC44C2D5BFF69F952C5A17837" bold="true" box="[151,453,1770,1797]" italics="true" pageId="11" pageNumber="571">Saphobranchia omorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C44C2D5BFE33F953C66B7837" box="[461,527,1771,1797]" pageId="11" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9C6D755DC44C2D5BFF69F8B6C7AE781B" blockId="11.[151,970,1770,1869]" box="[151,970,1806,1833]" pageId="11" pageNumber="571">
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urn:lsid:zoobank.org:act:
<uuid id="E8744F88C44C2D5BFE54F8B6C7AE781B" box="[426,970,1806,1833]" pageId="11" pageNumber="571">C97609B8-AF2E-413B-80C9-F45CC68CF5A2</uuid>
</uri>
</paragraph>
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<subSubSection id="D4C826D6C44C2D5BFF69F88BC507787F" box="[151,355,1843,1869]" pageId="11" pageNumber="571" type="description">
<paragraph id="9C6D755DC44C2D5BFF69F88BC507787F" blockId="11.[151,970,1770,1869]" box="[151,355,1843,1869]" pageId="11" pageNumber="571">
<figureCitation id="04E969D8C44C2D5BFF69F88BC566787F" box="[151,258,1843,1869]" captionStart="FIGURE 6" captionStartId="12.[151,250,1171,1195]" captionTargetBox="[172,1415,182,1146]" captionTargetId="figure-18@12.[172,1416,181,1147]" captionTargetPageId="12" captionText="FIGURE 6. Saphobranchia omorpha n. sp., holotype (SIO A9595). A. Dorsal view. B. Anterior region, dorsal view. C. Last chaetigers, dorsal view. D. Anterior region, ventral view. E. Last chaetiger, ventral view. Scale bars.- A: 1.5 mm, B: 0.90 mm, C: 0.60 mm, D: 0.95 mm, E: 0.52 mm." figureDoi="http://doi.org/10.5281/zenodo.4296967" httpUri="https://zenodo.org/record/4296967/files/figure.png" pageId="11" pageNumber="571">Figures 6</figureCitation>
,
<figureCitation id="04E969D8C44C2D5BFEF0F88BC57A787F" box="[270,286,1843,1869]" captionStart="FIGURE 7" captionStartId="13.[151,250,1632,1656]" captionTargetBox="[159,1429,417,1605]" captionTargetId="figure-136@13.[157,1430,417,1608]" captionTargetPageId="13" captionText="FIGURE 7. Saphobranchia omorpha n. sp., holotype (SIO A9595).A. Chaetiger 14, right parapodium, anterior view. B. Close up of parapodium. C. Central notochaetae, basal, medial and distal regions. D. Central neurochaetae, basal, medial and distal regions (inset: neurochaetal tip). Scale bars.- A: 0.36 mm, B: 0.21 mm, C: 35 μm, D: 65 μm." figureDoi="http://doi.org/10.5281/zenodo.4296969" httpUri="https://zenodo.org/record/4296969/files/figure.png" pageId="11" pageNumber="571">7</figureCitation>
,
<figureCitation id="04E969D8C44C2D5BFED4F88BC507787F" box="[298,355,1843,1869]" captionStart="FIGURE 8" captionStartId="14.[151,250,1797,1821]" captionTargetBox="[157,1430,849,1774]" captionTargetId="figure-328@14.[157,1430,849,1774]" captionTargetPageId="14" captionText="FIGURE 8. Living specimens. A. Lamispina polycerata n. sp., holotype (SIO A9842), anterior fragment, left lateral view. B. Same, posterior fragment, dorsal view. C. Saphobranchia canela n. sp., holotype (SIO A1332), dorsal view. D. Same, ventral view. E. Same, anterior end, frontal view (Br: branchia, Pa: palp). F. S. ilys n. sp., holotype (SIO A 9605), dorsal view. G. Same, ventral view. H. S. omorpha n. sp., holotype (SIO A9595), dorsal view. I. Same, ventral view. Scale bars: A, B: 1.0 mm; C, D: 2.2 mm; E: 0.4 mm; F, G: 3.5 mm; H, I: 3.7 mm (Photos: Greg Rouse)." figureDoi="http://doi.org/10.5281/zenodo.4296971" httpUri="https://zenodo.org/record/4296971/files/figure.png" pageId="11" pageNumber="571">8H, I</figureCitation>
</paragraph>
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<subSubSection id="D4C826D6C44C2D5BFF69F8C2C5A378EF" pageId="11" pageNumber="571" type="diagnosis">
<paragraph id="9C6D755DC44C2D5BFF69F8C2C5A378EF" blockId="11.[151,1436,1914,2013]" pageId="11" pageNumber="571">
<emphasis id="AEA6A94FC44C2D5BFF69F8C2C56E78A6" bold="true" box="[151,266,1914,1940]" pageId="11" pageNumber="571">Diagnosis</emphasis>
.
<taxonomicName id="5BD20EDEC44C2D5BFEE6F8C3C5A778A7" authorityName="Chamberlin" authorityYear="1919" box="[280,451,1915,1941]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="571" phylum="Annelida" rank="genus">
<emphasis id="AEA6A94FC44C2D5BFEE6F8C3C5A778A7" box="[280,451,1915,1941]" italics="true" pageId="11" pageNumber="571">Saphobranchia</emphasis>
</taxonomicName>
with tunic without sand particles; median chaetigers with notochaetae longer than body width; neurochaetae with tips straight, basal anchylosed region 1/3 chaetal length, articles 46 times longer than wide; no gonopodial lobes.
</paragraph>
</subSubSection>
<caption id="C8AD25D5C44B2D5CFF69FB2BC6457BC1" ID-DOI="http://doi.org/10.5281/zenodo.4296967" ID-Zenodo-Dep="4296967" httpUri="https://zenodo.org/record/4296967/files/figure.png" pageId="12" pageNumber="572" startId="12.[151,250,1171,1195]" targetBox="[172,1415,182,1146]" targetPageId="12">
<paragraph id="9C6D755DC44B2D5CFF69FB2BC6457BC1" blockId="12.[151,1437,1171,1268]" pageId="12" pageNumber="572">
<emphasis id="AEA6A94FC44B2D5CFF69FB2BC5707B99" bold="true" box="[151,276,1171,1195]" pageId="12" pageNumber="572">FIGURE 6.</emphasis>
<taxonomicName id="5BD20EDEC44B2D5CFEE2FB2BC67E7B99" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[284,538,1171,1195]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="12" pageNumber="572" phylum="Annelida" rank="species" species="omorpha" status="sp. nov.">
<emphasis id="AEA6A94FC44B2D5CFEE2FB2BC67E7B99" box="[284,538,1171,1195]" italics="true" pageId="12" pageNumber="572">Saphobranchia omorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C44B2D5CFDDCFB2CC6387B99" box="[546,604,1172,1195]" pageId="12" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
, holotype (SIO A9595). A. Dorsal view. B. Anterior region, dorsal view. C. Last chaetigers, dorsal view. D. Anterior region, ventral view. E. Last chaetiger, ventral view. Scale bars.- A: 1.5 mm, B: 0.90 mm, C: 0.60 mm, D: 0.95 mm, E: 0.52 mm.
</paragraph>
</caption>
<subSubSection id="D4C826D6C44B2D5CFF39FA9AC7C67A52" pageId="12" pageNumber="572" type="materials_examined">
<paragraph id="9C6D755DC44B2D5CFF39FA9AC7C67A52" blockId="12.[151,1437,1313,2024]" pageId="12" pageNumber="572">
<emphasis id="AEA6A94FC44B2D5CFF39FA9AC5087A09" bold="true" box="[199,364,1313,1340]" pageId="12" pageNumber="572">Type material</emphasis>
.
<materialsCitation id="2CBA7F00C44B2D5CFE86FA9AC7C67A52" ID-GBIF-Occurrence="2988754308" accessionNumber="MW172264" collectingDate="2018-10-18" collectionCode="SIO, MW" collectorName="G. Rouse &amp; A. Hatch" elevation="1784" latitude="9.117001" longLatPrecision="1" longitude="-84.840004" pageId="12" pageNumber="572" specimenCode="A9595" specimenCount="1" typeStatus="holotype">
<typeStatus id="4369CBFFC44B2D5CFE86FA9AC5847A0E" box="[376,480,1314,1340]" pageId="12" pageNumber="572" type="holotype">Holotype</typeStatus>
(
<collectionCode id="FAC3ED98C44B2D5CFE13FA9AC6787A0E" box="[493,540,1314,1340]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" lsid="urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="12" pageNumber="572" type="University or college">SIO</collectionCode>
<specimenCode id="CC74DD26C44B2D5CFDE1FA99C60B7A0E" box="[543,623,1313,1340]" collectionCode="SIO" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" lsid="urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="12" pageNumber="572" type="University or college">A9595</specimenCode>
), CR 18-0013,
<collectorName id="3127108BC44B2D5CFCECFA9AC7367A0E" box="[786,850,1314,1340]" pageId="12" pageNumber="572">Alvin</collectorName>
dive 4972,
<collectorName id="3127108BC44B2D5CFC2EFA9AC05D7A0E" box="[976,1081,1314,1340]" pageId="12" pageNumber="572">Jaco Scar</collectorName>
(
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,
<geoCoordinate id="F9E6139AC44B2D5CFAFAFA99C4D57A52" degrees="84" direction="west" minutes="50" orientation="longitude" pageId="12" pageNumber="572" precision="1" seconds="24.00" value="-84.840004">84°5024.00” W</geoCoordinate>
),
<quantity id="5B2AD8B8C44B2D5CFF37FAFEC5447A52" box="[201,288,1350,1376]" metricMagnitude="3" metricUnit="m" metricValue="1.784" pageId="12" pageNumber="572" unit="m" value="1784.0">
<elevation id="17FF926EC44B2D5CFF37FAFEC5447A52" box="[201,288,1350,1376]" metricMagnitude="3" metricUnit="m" metricValue="1.784" pageId="12" pageNumber="572" unit="m" value="1784.0">1784 m</elevation>
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,
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,
<collectorName id="3127108BC44B2D5CFE35FAFEC6577A52" box="[459,563,1350,1376]" pageId="12" pageNumber="572">G. Rouse</collectorName>
&amp;
<collectorName id="3127108BC44B2D5CFDA8FAFEC6DA7A52" box="[598,702,1350,1376]" pageId="12" pageNumber="572">A. Hatch</collectorName>
, coll. [
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].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C44B2D5EFF39FAD1C48F7FE7" lastPageId="14" lastPageNumber="574" pageId="12" pageNumber="572" type="description">
<paragraph id="9C6D755DC44B2D5CFF39FAD1C4B9790B" blockId="12.[151,1437,1313,2024]" pageId="12" pageNumber="572">
<emphasis id="AEA6A94FC44B2D5CFF39FAD1C5357AB1" bold="true" box="[199,337,1385,1411]" pageId="12" pageNumber="572">Description</emphasis>
.
<typeStatus id="4369CBFFC44B2D5CFE9FFAD2C5AD7AB6" box="[353,457,1386,1412]" pageId="12" pageNumber="572" type="holotype">Holotype</typeStatus>
(SIO A9595) originally with 32 chaetigers (
<figureCitation id="04E969D8C44B2D5CFC3FFAD2C0787AB6" box="[961,1052,1386,1412]" captionStart="FIGURE 8" captionStartId="14.[151,250,1797,1821]" captionTargetBox="[157,1430,849,1774]" captionTargetId="figure-328@14.[157,1430,849,1774]" captionTargetPageId="14" captionText="FIGURE 8. Living specimens. A. Lamispina polycerata n. sp., holotype (SIO A9842), anterior fragment, left lateral view. B. Same, posterior fragment, dorsal view. C. Saphobranchia canela n. sp., holotype (SIO A1332), dorsal view. D. Same, ventral view. E. Same, anterior end, frontal view (Br: branchia, Pa: palp). F. S. ilys n. sp., holotype (SIO A 9605), dorsal view. G. Same, ventral view. H. S. omorpha n. sp., holotype (SIO A9595), dorsal view. I. Same, ventral view. Scale bars: A, B: 1.0 mm; C, D: 2.2 mm; E: 0.4 mm; F, G: 3.5 mm; H, I: 3.7 mm (Photos: Greg Rouse)." figureDoi="http://doi.org/10.5281/zenodo.4296971" httpUri="https://zenodo.org/record/4296971/files/figure.png" pageId="12" pageNumber="572">Fig. 8H</figureCitation>
), 12 posterior chaetigers removed for molecular studies; now without posterior region. Body pale, anteriorly swollen, slightly bent laterally, broken along left margins of chaetigers 58, tapered medially and posteriorly, anterior chaetigers with notochaetae up to 9 times longer than body width; body wall broken along left size in chaetigers 58 (
<figureCitation id="04E969D8C44B2D5CFBAFFA6EC0C87AC2" box="[1105,1196,1493,1520]" captionStart="FIGURE 6" captionStartId="12.[151,250,1171,1195]" captionTargetBox="[172,1415,182,1146]" captionTargetId="figure-18@12.[172,1416,181,1147]" captionTargetPageId="12" captionText="FIGURE 6. Saphobranchia omorpha n. sp., holotype (SIO A9595). A. Dorsal view. B. Anterior region, dorsal view. C. Last chaetigers, dorsal view. D. Anterior region, ventral view. E. Last chaetiger, ventral view. Scale bars.- A: 1.5 mm, B: 0.90 mm, C: 0.60 mm, D: 0.95 mm, E: 0.52 mm." figureDoi="http://doi.org/10.5281/zenodo.4296967" httpUri="https://zenodo.org/record/4296967/files/figure.png" pageId="12" pageNumber="572">Fig. 6A</figureCitation>
);
<quantity id="5B2AD8B8C44B2D5CFB3CFA6EC1767AC2" box="[1218,1298,1494,1520]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="572" unit="mm" value="13.0">13 mm</quantity>
long,
<quantity id="5B2AD8B8C44B2D5CFAA5FA6EC1F97AC2" box="[1371,1437,1494,1520]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="572" unit="mm" value="1.0">1 mm</quantity>
wide (widest by chaetigers 78,
<quantity id="5B2AD8B8C44B2D5CFDFBFA42C6047926" box="[517,608,1530,1556]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="572" unit="mm" value="2.8">2.8 mm</quantity>
), cephalic cage chaetae
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long, 20 chaetigers (last one removed for a slide).
</paragraph>
<paragraph id="9C6D755DC44B2D5CFF39F9FAC7D57996" blockId="12.[151,1437,1313,2024]" pageId="12" pageNumber="572">
Tunic thin, without sand particles; papillae long, in about three different lengths, up to 10 times longer than wide, sparse, variably eroded along body, dorsally and ventrally of similar shape, shorter and more abundant ventrally (
<figureCitation id="04E969D8C44B2D5CFF1FF932C5247991" box="[225,320,1673,1700]" captionStart="FIGURE 6" captionStartId="12.[151,250,1171,1195]" captionTargetBox="[172,1415,182,1146]" captionTargetId="figure-18@12.[172,1416,181,1147]" captionTargetPageId="12" captionText="FIGURE 6. Saphobranchia omorpha n. sp., holotype (SIO A9595). A. Dorsal view. B. Anterior region, dorsal view. C. Last chaetigers, dorsal view. D. Anterior region, ventral view. E. Last chaetiger, ventral view. Scale bars.- A: 1.5 mm, B: 0.90 mm, C: 0.60 mm, D: 0.95 mm, E: 0.52 mm." figureDoi="http://doi.org/10.5281/zenodo.4296967" httpUri="https://zenodo.org/record/4296967/files/figure.png" pageId="12" pageNumber="572">Figs 6D</figureCitation>
;
<figureCitation id="04E969D8C44B2D5CFEB5F932C5007996" box="[331,356,1674,1700]" captionStart="FIGURE 8" captionStartId="14.[151,250,1797,1821]" captionTargetBox="[157,1430,849,1774]" captionTargetId="figure-328@14.[157,1430,849,1774]" captionTargetPageId="14" captionText="FIGURE 8. Living specimens. A. Lamispina polycerata n. sp., holotype (SIO A9842), anterior fragment, left lateral view. B. Same, posterior fragment, dorsal view. C. Saphobranchia canela n. sp., holotype (SIO A1332), dorsal view. D. Same, ventral view. E. Same, anterior end, frontal view (Br: branchia, Pa: palp). F. S. ilys n. sp., holotype (SIO A 9605), dorsal view. G. Same, ventral view. H. S. omorpha n. sp., holotype (SIO A9595), dorsal view. I. Same, ventral view. Scale bars: A, B: 1.0 mm; C, D: 2.2 mm; E: 0.4 mm; F, G: 3.5 mm; H, I: 3.7 mm (Photos: Greg Rouse)." figureDoi="http://doi.org/10.5281/zenodo.4296971" httpUri="https://zenodo.org/record/4296971/files/figure.png" pageId="12" pageNumber="572">8I</figureCitation>
), and surrounding chaetal bundle bases (
<figureCitation id="04E969D8C44B2D5CFCD3F932C7C17996" box="[813,933,1673,1700]" captionStart="FIGURE 6" captionStartId="12.[151,250,1171,1195]" captionTargetBox="[172,1415,182,1146]" captionTargetId="figure-18@12.[172,1416,181,1147]" captionTargetPageId="12" captionText="FIGURE 6. Saphobranchia omorpha n. sp., holotype (SIO A9595). A. Dorsal view. B. Anterior region, dorsal view. C. Last chaetigers, dorsal view. D. Anterior region, ventral view. E. Last chaetiger, ventral view. Scale bars.- A: 1.5 mm, B: 0.90 mm, C: 0.60 mm, D: 0.95 mm, E: 0.52 mm." figureDoi="http://doi.org/10.5281/zenodo.4296967" httpUri="https://zenodo.org/record/4296967/files/figure.png" pageId="12" pageNumber="572">Fig. 6C, E</figureCitation>
).
</paragraph>
<paragraph id="9C6D755DC44B2D5CFF39F916C10C79FA" blockId="12.[151,1437,1313,2024]" box="[199,1384,1710,1736]" pageId="12" pageNumber="572">Anterior end features not seen; one palp barely exposed, brownish; not dissected to avoid further damage.</paragraph>
<paragraph id="9C6D755DC44B2D5CFF39F96AC6B67806" blockId="12.[151,1437,1313,2024]" pageId="12" pageNumber="572">Cephalic cage chaetae 23 times longer than body width. Chaetigers 12 forming cephalic cage, chaetae directed anteriorly; chaetiger 3 with notochaetae directed anteriorly, neurochaetae directed laterally. Chaetae arranged in short rows, notochaetae dorsolateral along body.</paragraph>
<paragraph id="9C6D755DC44B2D5CFF39F886C7E078DA" blockId="12.[151,1437,1313,2024]" pageId="12" pageNumber="572">Chaetiger 1 with 78 notochaetae, 34 neurochaetae, longest ones basally to medially anchylosed, shorter ones completely articulated, articles 34 times longer than wide medially, progressively longer distally. Chaetiger 2 with 7 thinner notochaetae, 4/5 as long as those of chaetiger 1 or of similar size, anchylosed region progressively shorter in smaller chaetae; neurochaetae thicker, 1112 per bundle, some with adsorbed brownish particles along their length, articles 23 times longer than wide, slightly longer distally.</paragraph>
<paragraph id="9C6D755DC44A2D5DFF39FF2FC03D7E2F" blockId="13.[151,1437,151,393]" pageId="13" pageNumber="573">
Anterior dorsal margin of chaetiger 1 with several papillae and two small blunt conical projections internal to notochaetae (
<figureCitation id="04E969D8C44A2D5DFEB2FF03C5C27FE4" box="[332,422,187,214]" captionStart="FIGURE 6" captionStartId="12.[151,250,1171,1195]" captionTargetBox="[172,1415,182,1146]" captionTargetId="figure-18@12.[172,1416,181,1147]" captionTargetPageId="12" captionText="FIGURE 6. Saphobranchia omorpha n. sp., holotype (SIO A9595). A. Dorsal view. B. Anterior region, dorsal view. C. Last chaetigers, dorsal view. D. Anterior region, ventral view. E. Last chaetiger, ventral view. Scale bars.- A: 1.5 mm, B: 0.90 mm, C: 0.60 mm, D: 0.95 mm, E: 0.52 mm." figureDoi="http://doi.org/10.5281/zenodo.4296967" httpUri="https://zenodo.org/record/4296967/files/figure.png" pageId="13" pageNumber="573">Fig. 6B</figureCitation>
), also visible in chaetiger 2. Chaetigers 14 of similar length. No chaetal transition from cephalic cage to body chaetae; all neurochaetae multiarticulated. Gonopodial lobes not seen (
<figureCitation id="04E969D8C44A2D5DFB4DFF67C16A7FC8" box="[1203,1294,223,250]" captionStart="FIGURE 6" captionStartId="12.[151,250,1171,1195]" captionTargetBox="[172,1415,182,1146]" captionTargetId="figure-18@12.[172,1416,181,1147]" captionTargetPageId="12" captionText="FIGURE 6. Saphobranchia omorpha n. sp., holotype (SIO A9595). A. Dorsal view. B. Anterior region, dorsal view. C. Last chaetigers, dorsal view. D. Anterior region, ventral view. E. Last chaetiger, ventral view. Scale bars.- A: 1.5 mm, B: 0.90 mm, C: 0.60 mm, D: 0.95 mm, E: 0.52 mm." figureDoi="http://doi.org/10.5281/zenodo.4296967" httpUri="https://zenodo.org/record/4296967/files/figure.png" pageId="13" pageNumber="573">Fig. 6D</figureCitation>
). Testis seen through broken body wall, confirmed by abundant spherical spermatids within gonad.
</paragraph>
<paragraph id="9C6D755DC44A2D5DFF39FE9FC16F7EBB" blockId="13.[151,1437,151,393]" pageId="13" pageNumber="573">
Parapodia laterally projected from body wall; parapodia lateral, median neuropodia ventrolateral. Notopodia and neuropodia surrounded by several cylindrical papillae (
<figureCitation id="04E969D8C44A2D5DFCD4FEF3C7E77E57" box="[810,899,331,357]" captionStart="FIGURE 7" captionStartId="13.[151,250,1632,1656]" captionTargetBox="[159,1429,417,1605]" captionTargetId="figure-136@13.[157,1430,417,1608]" captionTargetPageId="13" captionText="FIGURE 7. Saphobranchia omorpha n. sp., holotype (SIO A9595).A. Chaetiger 14, right parapodium, anterior view. B. Close up of parapodium. C. Central notochaetae, basal, medial and distal regions. D. Central neurochaetae, basal, medial and distal regions (inset: neurochaetal tip). Scale bars.- A: 0.36 mm, B: 0.21 mm, C: 35 μm, D: 65 μm." figureDoi="http://doi.org/10.5281/zenodo.4296969" httpUri="https://zenodo.org/record/4296969/files/figure.png" pageId="13" pageNumber="573">Fig. 7A</figureCitation>
); papillae covered by a very thin tunic (
<figureCitation id="04E969D8C44A2D5DFAC6FEF3C1EA7E57" box="[1336,1422,331,357]" captionStart="FIGURE 7" captionStartId="13.[151,250,1632,1656]" captionTargetBox="[159,1429,417,1605]" captionTargetId="figure-136@13.[157,1430,417,1608]" captionTargetPageId="13" captionText="FIGURE 7. Saphobranchia omorpha n. sp., holotype (SIO A9595).A. Chaetiger 14, right parapodium, anterior view. B. Close up of parapodium. C. Central notochaetae, basal, medial and distal regions. D. Central neurochaetae, basal, medial and distal regions (inset: neurochaetal tip). Scale bars.- A: 0.36 mm, B: 0.21 mm, C: 35 μm, D: 65 μm." figureDoi="http://doi.org/10.5281/zenodo.4296969" httpUri="https://zenodo.org/record/4296969/files/figure.png" pageId="13" pageNumber="573">Fig. 7B</figureCitation>
), longest papillae about as 1/13 as long as longest notochaetae, or 1/11 as long as longest neurochaetae.
</paragraph>
<caption id="C8AD25D5C44A2D5DFF69F9D8C05B79F2" ID-DOI="http://doi.org/10.5281/zenodo.4296969" ID-Zenodo-Dep="4296969" httpUri="https://zenodo.org/record/4296969/files/figure.png" pageId="13" pageNumber="573" startId="13.[151,250,1632,1656]" targetBox="[159,1429,417,1605]" targetPageId="13">
<paragraph id="9C6D755DC44A2D5DFF69F9D8C05B79F2" blockId="13.[151,1437,1632,1728]" pageId="13" pageNumber="573">
<emphasis id="AEA6A94FC44A2D5DFF69F9D8C577794A" bold="true" box="[151,275,1632,1656]" pageId="13" pageNumber="573">FIGURE 7.</emphasis>
<taxonomicName id="5BD20EDEC44A2D5DFEE7F9D8C671794A" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[281,533,1632,1656]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="13" pageNumber="573" phylum="Annelida" rank="species" species="omorpha" status="sp. nov.">
<emphasis id="AEA6A94FC44A2D5DFEE7F9D8C671794A" box="[281,533,1632,1656]" italics="true" pageId="13" pageNumber="573">Saphobranchia omorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C44A2D5DFDE2F9D8C6307945" box="[540,596,1632,1655]" pageId="13" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
, holotype (SIO A9595). A. Chaetiger 14, right parapodium, anterior view. B. Close up of parapodium. C. Central notochaetae, basal, medial and distal regions. D. Central neurochaetae, basal, medial and distal regions (inset: neurochaetal tip). Scale bars.- A: 0.36 mm, B: 0.21 mm, C: 35 μm, D: 65 μm.
</paragraph>
</caption>
<paragraph id="9C6D755DC44A2D5DFF39F956C4B978D3" blockId="13.[151,1437,1774,2017]" pageId="13" pageNumber="573">
Median notochaetae arranged in short transverse rows, chaetal fascicles fan-shaped, directed dorsally, at least 5 times longer than body width. All notochaetae multiarticulate capillaries; 14 per bundle, central notochaetae with basally anchylosed region about 1/3 chaetal length (
<figureCitation id="04E969D8C44A2D5DFD1BF88EC75A7862" box="[741,830,1846,1872]" captionStart="FIGURE 7" captionStartId="13.[151,250,1632,1656]" captionTargetBox="[159,1429,417,1605]" captionTargetId="figure-136@13.[157,1430,417,1608]" captionTargetPageId="13" captionText="FIGURE 7. Saphobranchia omorpha n. sp., holotype (SIO A9595).A. Chaetiger 14, right parapodium, anterior view. B. Close up of parapodium. C. Central notochaetae, basal, medial and distal regions. D. Central neurochaetae, basal, medial and distal regions (inset: neurochaetal tip). Scale bars.- A: 0.36 mm, B: 0.21 mm, C: 35 μm, D: 65 μm." figureDoi="http://doi.org/10.5281/zenodo.4296969" httpUri="https://zenodo.org/record/4296969/files/figure.png" pageId="13" pageNumber="573">Fig. 7C</figureCitation>
), basal notochaetae with a shorter anchylosed region, not fully articulated; articles 46 times longer than wide medially, progressively longer distally. Neurochaetae twice longer than body width, nine per bundle, central neurochaetae with anchylosed region 1/3 as long as chaetal length, medially with articles 34 times longer than wide, progressively longer distally (
<figureCitation id="04E969D8C44A2D5DFBCAF81AC0F4788F" box="[1076,1168,1954,1981]" captionStart="FIGURE 7" captionStartId="13.[151,250,1632,1656]" captionTargetBox="[159,1429,417,1605]" captionTargetId="figure-136@13.[157,1430,417,1608]" captionTargetPageId="13" captionText="FIGURE 7. Saphobranchia omorpha n. sp., holotype (SIO A9595).A. Chaetiger 14, right parapodium, anterior view. B. Close up of parapodium. C. Central notochaetae, basal, medial and distal regions. D. Central neurochaetae, basal, medial and distal regions (inset: neurochaetal tip). Scale bars.- A: 0.36 mm, B: 0.21 mm, C: 35 μm, D: 65 μm." figureDoi="http://doi.org/10.5281/zenodo.4296969" httpUri="https://zenodo.org/record/4296969/files/figure.png" pageId="13" pageNumber="573">Fig. 7D</figureCitation>
), tips straight (
<figureCitation id="04E969D8C44A2D5DFAC3F81AC1F3788F" box="[1341,1431,1954,1981]" captionStart="FIGURE 7" captionStartId="13.[151,250,1632,1656]" captionTargetBox="[159,1429,417,1605]" captionTargetId="figure-136@13.[157,1430,417,1608]" captionTargetPageId="13" captionText="FIGURE 7. Saphobranchia omorpha n. sp., holotype (SIO A9595).A. Chaetiger 14, right parapodium, anterior view. B. Close up of parapodium. C. Central notochaetae, basal, medial and distal regions. D. Central neurochaetae, basal, medial and distal regions (inset: neurochaetal tip). Scale bars.- A: 0.36 mm, B: 0.21 mm, C: 35 μm, D: 65 μm." figureDoi="http://doi.org/10.5281/zenodo.4296969" httpUri="https://zenodo.org/record/4296969/files/figure.png" pageId="13" pageNumber="573">Fig. 7D</figureCitation>
, inset).
</paragraph>
<paragraph id="9C6D755DC4492D5EFF39FF2FC48F7FE7" blockId="14.[151,1437,151,825]" pageId="14" pageNumber="574">Posterior region cylindrical, tapered (seen in photos before cropping for molecular studies); pygidium unknown.</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C4492D5EFF39FF67C5357E73" pageId="14" pageNumber="574" type="etymology">
<paragraph id="9C6D755DC4492D5EFF39FF67C5357E73" blockId="14.[151,1437,151,825]" pageId="14" pageNumber="574">
<emphasis id="AEA6A94FC4492D5EFF39FF67C52C7FCB" bold="true" box="[199,328,223,249]" pageId="14" pageNumber="574">Etymology</emphasis>
. The specific name is from the Greek
<emphasis id="AEA6A94FC4492D5EFCF2FF67C7187FCB" box="[780,892,223,249]" italics="true" pageId="14" pageNumber="574">ómorphos</emphasis>
, beautiful, masculine, latinised and declined in feminine, for referring to the beauty of the specimen. The specific name is regarded as a noun in apposition (
<bibRefCitation id="F84308ACC4492D5EFAA7FEBBC4B67E73" author="ICZN &amp; International Commission on Zoological Nomenclature" pageId="14" pageNumber="574" refId="ref15696" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature in association with the British Museum (Natural History), London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
, Art. 31.2).
</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C4492D5EFF39FEF3C11B7C27" pageId="14" pageNumber="574" type="discussion">
<paragraph id="9C6D755DC4492D5EFF39FEF3C6C97D2B" blockId="14.[151,1437,151,825]" pageId="14" pageNumber="574">
<emphasis id="AEA6A94FC4492D5EFF39FEF3C5507E57" bold="true" box="[199,308,331,357]" pageId="14" pageNumber="574">Remarks</emphasis>
.
<taxonomicName id="5BD20EDEC4492D5EFEBDFEF3C63F7E57" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[323,603,331,357]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="14" pageNumber="574" phylum="Annelida" rank="species" species="omorpha" status="sp. nov.">
<emphasis id="AEA6A94FC4492D5EFEBDFEF3C63F7E57" box="[323,603,331,357]" italics="true" pageId="14" pageNumber="574">Saphobranchia omorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C4492D5EFD9DFEF3C6C47E57" box="[611,672,331,357]" pageId="14" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
groups with
<taxonomicName id="5BD20EDEC4492D5EFCC8FEF3C00E7E57" authority="(Fauchald, 1972)" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Fauchald" baseAuthorityYear="1972" box="[822,1130,331,357]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="14" pageNumber="574" phylum="Annelida" rank="species" species="micans" status="comb. nov.">
<emphasis id="AEA6A94FC4492D5EFCC8FEF3C7C67E56" box="[822,930,331,357]" italics="true" pageId="14" pageNumber="574">S. micans</emphasis>
(
<bibRefCitation id="F84308ACC4492D5EFC4CFEF3C0067E57" author="Fauchald, K." box="[946,1122,331,357]" pageId="14" pageNumber="574" pagination="1 - 575" refId="ref15175" refString="Fauchald, K. (1972) Benthic polychaetous annelids from deep waters off Western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1 - 575." type="journal article" year="1972">Fauchald, 1972</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="B5951434C4492D5EFB8CFEF3C0B37E57" box="[1138,1239,331,357]" pageId="14" pageNumber="574" rank="species">n. comb.</taxonomicNameLabel>
, and
<taxonomicName id="5BD20EDEC4492D5EFAEAFEF3C13C7E57" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[1300,1368,331,357]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="14" pageNumber="574" phylum="Annelida" rank="species" species="ilys" status="sp. nov.">
<emphasis id="AEA6A94FC4492D5EFAEAFEF3C13C7E57" box="[1300,1368,331,357]" italics="true" pageId="14" pageNumber="574">S. ilys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C4492D5EFAA1FEF3C1F87E57" box="[1375,1436,331,357]" pageId="14" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
because their bodies do not have abundant sand particles adhered on tunic, and their notochaetae are longer than body width. Their main difference after the key above is the
<typeStatus id="4369CBFFC4492D5EFCC4FE2BC70E7E9F" box="[826,874,403,429]" pageId="14" pageNumber="574">type</typeStatus>
of neurochaetal tips. In
<taxonomicName id="5BD20EDEC4492D5EFB87FE2BC09E7E9F" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[1145,1274,403,429]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="14" pageNumber="574" phylum="Annelida" rank="species" species="omorpha" status="sp. nov.">
<emphasis id="AEA6A94FC4492D5EFB87FE2BC09E7E9F" box="[1145,1274,403,429]" italics="true" pageId="14" pageNumber="574">S. omorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C4492D5EFAFFFE2BC1597E9F" box="[1281,1341,403,429]" pageId="14" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
they are straight whereas in
<taxonomicName id="5BD20EDEC4492D5EFE8CFE0FC5B97EE2" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Fauchald" baseAuthorityYear="1972" box="[370,477,439,465]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="14" pageNumber="574" phylum="Annelida" rank="species" species="micans">
<emphasis id="AEA6A94FC4492D5EFE8CFE0FC5B97EE2" box="[370,477,439,465]" italics="true" pageId="14" pageNumber="574">S. micans</emphasis>
</taxonomicName>
and
<taxonomicName id="5BD20EDEC4492D5EFDEAFE0FC6337EE3" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[532,599,439,465]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="14" pageNumber="574" phylum="Annelida" rank="species" species="ilys" status="sp. nov.">
<emphasis id="AEA6A94FC4492D5EFDEAFE0FC6337EE3" box="[532,599,439,465]" italics="true" pageId="14" pageNumber="574">S. ilys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C4492D5EFDA0FE0FC6FF7EE3" box="[606,667,439,465]" pageId="14" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
, they are falcate. Another conspicuous difference is the size of chaetae, because in
<taxonomicName id="5BD20EDEC4492D5EFEBEFE63C5A57EC7" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[320,449,475,501]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="14" pageNumber="574" phylum="Annelida" rank="species" species="omorpha" status="sp. nov.">
<emphasis id="AEA6A94FC4492D5EFEBEFE63C5A57EC7" box="[320,449,475,501]" italics="true" pageId="14" pageNumber="574">S. omorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C4492D5EFE39FE63C6677EC7" box="[455,515,475,501]" pageId="14" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
they are very long, more than 10 times longer than body width, whereas in the two other species, they can be up to 5 times as long.
</paragraph>
<paragraph id="9C6D755DC4492D5EFF39FD9BC11B7C27" blockId="14.[151,1437,151,825]" pageId="14" pageNumber="574">
An additional comparison might be needed because
<taxonomicName id="5BD20EDEC4492D5EFCDAFD9BC7C37D0F" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[804,935,547,573]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="14" pageNumber="574" phylum="Annelida" rank="species" species="omorpha" status="sp. nov.">
<emphasis id="AEA6A94FC4492D5EFCDAFD9BC7C37D0F" box="[804,935,547,573]" italics="true" pageId="14" pageNumber="574">S. omorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C4492D5EFC4FFD9BC7947D0F" box="[945,1008,547,573]" pageId="14" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="5BD20EDEC4492D5EFBD2FD9BC0157D0F" authorityName="Salazar-Vallejo" authorityYear="2020" box="[1068,1137,547,573]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="14" pageNumber="574" phylum="Annelida" rank="species" species="ilys">
<emphasis id="AEA6A94FC4492D5EFBD2FD9BC0157D0F" box="[1068,1137,547,573]" italics="true" pageId="14" pageNumber="574">S. ilys</emphasis>
</taxonomicName>
share pale body wall and brownish chaetae. As indicated above, their main differences are in the development of tunic papillae, associated sediment particles, parapodial development, and chaetal size in median chaetigers.
<taxonomicName id="5BD20EDEC4492D5EFBBEFDD3C13C7DB7" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[1088,1368,619,645]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="14" pageNumber="574" phylum="Annelida" rank="species" species="omorpha" status="sp. nov.">
<emphasis id="AEA6A94FC4492D5EFBBEFDD3C13C7DB7" box="[1088,1368,619,645]" italics="true" pageId="14" pageNumber="574">Saphobranchia omorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C4492D5EFA9EFDD3C1F87DB7" box="[1376,1436,619,645]" pageId="14" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
has a pilose body wall because the scarce dorsal papillae lack sediment particles, or have fine particles forming a thin layer, parapodia are clearly projected laterally, and chaetae are up to 4 times longer than body width. On the contrary,
<taxonomicName id="5BD20EDEC4492D5EFEFEFD6FC5267DC3" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[256,322,727,753]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="14" pageNumber="574" phylum="Annelida" rank="species" species="ilys" status="sp. nov.">
<emphasis id="AEA6A94FC4492D5EFEFEFD6FC5267DC3" box="[256,322,727,753]" italics="true" pageId="14" pageNumber="574">S. ilys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C4492D5EFEB6FD6FC5E07DC3" box="[328,388,727,753]" pageId="14" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
has a rugose body wall because dorsal papillae are short, blunt mainly with fine sediment, and a few sand particles, parapodia are slightly projected from the body wall, and chaetae are as long as body width.
</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C4492D5EFF39FCA6C7547C0B" box="[199,816,798,825]" pageId="14" pageNumber="574" type="distribution">
<paragraph id="9C6D755DC4492D5EFF39FCA6C7547C0B" blockId="14.[151,1437,151,825]" box="[199,816,798,825]" pageId="14" pageNumber="574">
<emphasis id="AEA6A94FC4492D5EFF39FCA6C53D7C0A" bold="true" box="[199,345,798,824]" pageId="14" pageNumber="574">Distribution</emphasis>
. Off Pacific
<collectingCountry id="E4C535CDC4492D5EFE14FCA7C6027C0B" box="[490,614,799,825]" name="Costa Rica" pageId="14" pageNumber="574">Costa Rica</collectingCountry>
, in
<quantity id="5B2AD8B8C4492D5EFD70FCA7C6877C0B" box="[654,739,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.784" pageId="14" pageNumber="574" unit="m" value="1784.0">1784 m</quantity>
depth.
</paragraph>
</subSubSection>
</treatment>
</document>