233 lines
29 KiB
XML
233 lines
29 KiB
XML
<document id="D2A792A289EAD30C0AF722B221D2C8B8" ID="10.11646/zootaxa.4139.2.1" ID-DOI="10.11646/zootaxa.4139.2.1" ID-GBIF-Dataset="aeecbbb4-4764-41dd-9562-b25f91f9f70e" ID-ISSN="1175-5326" ID-Zenodo-Dep="265016" ID-ZooBank="2EAFF1B9-1FA2-44B1-89A3-476359314C5E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468994861279" checkinUser="plazi" docAuthor="Nguyen, Anh D. & Golovatch, Sergei I." docDate="2016" docId="03FECD21FFA9FFFEFF4F3BDBFE28F9B1" docLanguage="en" docName="zootaxa.4139.2.1.pdf" docOrigin="Zootaxa 4139 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enghoffosoma triangulare Nguyen & Golovatch, 2016, sp. nov." docType="treatment" docVersion="11" lastPageNumber="159" masterDocId="FFC7B559FFACFFF6FFD83D24FFAEFF84" masterDocTitle="The millipede genus Enghoffosoma Golovatch, 1993 recorded in Vietnam for the first time, with descriptions of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="166" masterPageNumber="151" pageNumber="156" updateTime="1698365308467" updateUser="plazi">
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<mods:mods id="DCA4BA212968819729D23B980308557F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B7860AA0285CE34ED26C087153B77F53">
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<mods:title id="6723ECB06B8CCDD55A736747348939F0">The millipede genus Enghoffosoma Golovatch, 1993 recorded in Vietnam for the first time, with descriptions of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="3C9DA0EB78EB7AB9D1EDA89F5DBC887F" type="personal">
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<mods:roleTerm id="ABDCA7792354FE170FE6B922AABBB3AA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A02B47D5C338D73FE74ECA907F3CDCD0">Nguyen, Anh D.</mods:namePart>
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</mods:name>
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<mods:name id="961A1F1FE1F8F1E84710661842B4ADA3" type="personal">
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<mods:role id="8CDC8674444108D0FE2ECE955130E3FB">
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<mods:roleTerm id="1BCE5F93982FC9056F0152200E2FDF28">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F41B869E4478A0BC404F8C4CDB06DC23">Golovatch, Sergei I.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="17DEF9E9EAD4DB5CB3FE75640B2451E0">text</mods:typeOfResource>
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<mods:title id="24385F77EF5234BB87A838EAD27EE859">Zootaxa</mods:title>
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<mods:part id="5A30101DCB0F3497502947A13BD01882">
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<mods:date id="4E1E455BD50C91D9FFFE3D261A521131">2016</mods:date>
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<mods:detail id="1CBCEACDC9A097BB58AECCC10AC15E50" type="volume">
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<mods:number id="4DF5F191D7ACC60840BE06AA1E20D91F">4139</mods:number>
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<mods:detail id="225F4BA6D71605212BEF1AD915844FC1" type="issue">
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<mods:number id="C38F61CEA9DED493A0F23374EEA23D4F">2</mods:number>
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<mods:classification id="73AD6977CD543911BEE96DB37C82BAF9">journal article</mods:classification>
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<mods:identifier id="5426418660298C7B4AB675CB73BAF5BD" type="DOI">10.11646/zootaxa.4139.2.1</mods:identifier>
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<mods:identifier id="6E73CE9331AF76EC36071853683E56F7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="852AAD9154E6797FFA2425A8C184DD40" type="Zenodo-Dep">265016</mods:identifier>
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<treatment id="03FECD21FFA9FFFEFF4F3BDBFE28F9B1" ID-DOI="http://doi.org/10.5281/zenodo.6067531" ID-GBIF-Taxon="123198413" ID-Zenodo-Dep="6067531" LSID="urn:lsid:plazi:treatment:03FECD21FFA9FFFEFF4F3BDBFE28F9B1" httpUri="http://treatment.plazi.org/id/03FECD21FFA9FFFEFF4F3BDBFE28F9B1" lastPageId="8" lastPageNumber="159" pageId="5" pageNumber="156">
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<subSubSection id="C34D2FBCFFA9FFF3FF4F3BDBFEA4F8BF" pageId="5" pageNumber="156" type="nomenclature">
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<paragraph id="8BE87C37FFA9FFF3FF4F3BDBFD91F89D" blockId="5.[151,575,1791,1851]" box="[151,575,1791,1817]" pageId="5" pageNumber="156">
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<heading id="D0A0CB5BFFA9FFF3FF4F3BDBFD91F89D" bold="true" box="[151,575,1791,1817]" fontSize="11" level="1" pageId="5" pageNumber="156" reason="1">
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<emphasis id="B923A025FFA9FFF3FF4F3BDBFD91F89D" bold="true" box="[151,575,1791,1817]" pageId="5" pageNumber="156">
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<taxonomicName id="4C5707B4FFA9FFF3FF4F3BDBFE74F89D" ID-CoL="39T4L" box="[151,474,1791,1817]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="triangulare" status="sp. nov.">
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<emphasis id="B923A025FFA9FFF3FF4F3BDBFE74F89D" bold="true" box="[151,474,1791,1817]" italics="true" pageId="5" pageNumber="156">Enghoffosoma triangulare</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2101D5EFFA9FFF3FE393BDBFD91F89D" box="[481,575,1791,1817]" pageId="5" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE87C37FFA9FFF3FF4F3A06FEA4F8BF" blockId="5.[151,575,1791,1851]" box="[151,266,1826,1851]" pageId="5" pageNumber="156">
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(
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<figureCitation id="136C60B2FFA9FFF3FF473A06FEADF8BF" box="[159,259,1826,1851]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="6.[151,250,1768,1790]" captionStartId-1="7.[151,250,1997,2019]" captionStartId-2="8.[151,250,1431,1453]" captionTargetBox-0="[153,1423,661,1737]" captionTargetBox-1="[161,1425,956,1967]" captionTargetBox-2="[151,1436,193,1409]" captionTargetId-0="figure@6.[151,1435,661,1747]" captionTargetId-1="figure@7.[151,1435,949,1976]" captionTargetId-2="figure@8.[151,1436,193,1410]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 6. Enghoffosoma triangulare sp. nov. Holotype, anterior segments, lateral view (A), dorsal view (B), ventral view (C); midbody segments, dorsal view (D), lateral view (E); posterior segments, lateral view (F); sternum between coxae 4, ventral view (G); hypoproct and paraprocts, ventral view (H). Scale = 1 mm." captionText-1="FIGURE 7. Enghoffosoma triangulare sp. nov. Holotype, right gonopod, lateral view (A); mesal view (B). Scale = 1 mm." captionText-2="FIGURE 8. Enghoffosoma triangulare sp. nov. Holotype, right gonopod, lateroventral view (A); postfemoral region, lateroventral view (B)." httpUri-0="https://zenodo.org/record/265022/files/figure.png" httpUri-1="https://zenodo.org/record/265023/files/figure.png" httpUri-2="https://zenodo.org/record/265024/files/figure.png" pageId="5" pageNumber="156">Figs 6–8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D2FBCFFA9FFF3FF4F3A4EFC48F84F" pageId="5" pageNumber="156" type="materials_examined">
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<paragraph id="8BE87C37FFA9FFF3FF4F3A4EFCEDF823" blockId="5.[151,1436,1898,2031]" pageId="5" pageNumber="156">
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<emphasis id="B923A025FFA9FFF3FF4F3A4EFE2EF807" bold="true" box="[151,384,1898,1923]" pageId="5" pageNumber="156">Material examined.</emphasis>
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<emphasis id="B923A025FFA9FFF3FE5F3A4EFE40F806" box="[391,494,1898,1922]" italics="true" pageId="5" pageNumber="156">
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<typeStatus id="54ECC295FFA9FFF3FE5F3A4EFE40F806" box="[391,494,1898,1922]" pageId="5" pageNumber="156" type="holotype">Holotype</typeStatus>
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</emphasis>
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male (
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<emphasis id="B923A025FFA9FFF3FDE33A4EFCABF807" bold="true" box="[571,773,1898,1923]" pageId="5" pageNumber="156">IEBR-Myr 123H</emphasis>
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),
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<collectingCountry id="F3403CA7FFA9FFF3FCC43A4EFCD0F807" box="[796,894,1898,1923]" name="Vietnam" pageId="5" pageNumber="156">Vietnam</collectingCountry>
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,
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<collectingRegion id="4993B2D5FFA9FFF3FC503A4FFC58F807" box="[904,1014,1898,1923]" country="Vietnam" name="Dong Nai" pageId="5" pageNumber="156">Dong Nai</collectingRegion>
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Prov., Cat Tien National Park, forest,
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<date id="FFE95AF7FFA9FFF3FF4F3AABFEDAF823" box="[151,372,1934,1959]" pageId="5" pageNumber="156" value="2004-09-24">24 September 2004</date>
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, leg. Anh Duc Nguyen & S.I. Golovatch.
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</paragraph>
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<paragraph id="8BE87C37FFA9FFF3FF1F3A97FC48F84F" blockId="5.[151,1436,1898,2031]" box="[199,998,1970,1995]" pageId="5" pageNumber="156">
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<emphasis id="B923A025FFA9FFF3FF1F3A97FE97F84E" box="[199,313,1971,1994]" italics="true" pageId="5" pageNumber="156">
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<typeStatus id="54ECC295FFA9FFF3FF1F3A97FE97F84E" box="[199,313,1971,1994]" pageId="5" pageNumber="156" type="paratype">Paratypes</typeStatus>
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</emphasis>
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:
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<specimenCount id="9D51B7BEFFA9FFF3FE9F3A97FE05F84F" box="[327,427,1970,1995]" pageId="5" pageNumber="156" type="female">1 female</specimenCount>
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,
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<specimenCount id="9D51B7BEFFA9FFF3FE6E3A97FE5BF84F" box="[438,501,1970,1995]" pageId="5" pageNumber="156" type="generic">1 juv.</specimenCount>
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(
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<emphasis id="B923A025FFA9FFF3FDDC3A96FD67F84F" bold="true" box="[516,713,1970,1995]" pageId="5" pageNumber="156">IEBR-Myr 123P</emphasis>
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), together with
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<typeStatus id="54ECC295FFA9FFF3FCA43A96FC4CF84F" box="[892,994,1970,1995]" pageId="5" pageNumber="156" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D2FBCFFA9FFF0FF1F3AF2FEDDFEC4" lastPageId="6" lastPageNumber="157" pageId="5" pageNumber="156" type="diagnosis">
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<paragraph id="8BE87C37FFA9FFF0FF1F3AF2FEA4FF7C" blockId="5.[151,1436,1898,2031]" lastBlockId="6.[151,1436,151,608]" lastPageId="6" lastPageNumber="157" pageId="5" pageNumber="156">
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<emphasis id="B923A025FFA9FFF3FF1F3AF2FEEFF86B" bold="true" box="[199,321,2006,2031]" pageId="5" pageNumber="156">Diagnosis.</emphasis>
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The species is easily recognized by the paraterga being well-developed; the gonopod femorite cylindrical and slender, but somewhat curved in the middle, without processes; the solenomere is slightly spiralled, as large as the femorite at base, then tapering towards tip; distal part with a small pointed tooth; process
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<emphasis id="B923A025FFAAFFF0FAC53D98FA87FF51" bold="true" box="[1309,1321,188,213]" pageId="6" pageNumber="157">e</emphasis>
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small and triangular.
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</paragraph>
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<paragraph id="8BE87C37FFAAFFF0FF1F3C20FEDDFEC4" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="157">
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<emphasis id="B923A025FFAAFFF0FF1F3C20FEBAFE99" bold="true" box="[199,276,260,285]" pageId="6" pageNumber="157">Name.</emphasis>
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“
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<taxonomicName id="4C5707B4FFAAFFF0FEF43C20FE04FE98" box="[300,426,260,284]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="triangulare">
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<emphasis id="B923A025FFAAFFF0FEF43C20FE04FE98" box="[300,426,260,284]" italics="true" pageId="6" pageNumber="157">triangulare</emphasis>
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</taxonomicName>
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”, an adjective in neuter gender to emphasize the triangular process
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<emphasis id="B923A025FFAAFFF0FB0A3C20FB70FE99" bold="true" box="[1234,1246,260,285]" pageId="6" pageNumber="157">e</emphasis>
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of the gonopod postfemoral region.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D2FBCFFAAFFF1FF1F3C68FB53FD74" lastPageId="7" lastPageNumber="158" pageId="6" pageNumber="157" type="description">
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<paragraph id="8BE87C37FFAAFFF0FF1F3C68FF47FE54" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="157">
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<emphasis id="B923A025FFAAFFF0FF1F3C68FEF7FEE1" bold="true" box="[199,345,332,357]" pageId="6" pageNumber="157">Description.</emphasis>
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Body length about
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<quantity id="4CAFD1D2FFAAFFF0FDEA3C68FD38FEE0" box="[562,662,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="3.28" pageId="6" pageNumber="157" unit="mm" value="32.8">32.8 mm</quantity>
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(male),
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<quantity id="4CAFD1D2FFAAFFF0FD2A3C68FCF8FEE0" box="[754,854,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.67" pageId="6" pageNumber="157" unit="mm" value="26.7">26.7 mm</quantity>
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(female); width of midbody pro- and metazonae
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<quantity id="4CAFD1D2FFAAFFF0FAA13C68FF6DFE03" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="157" unit="mm" value="2.4">2.4 mm</quantity>
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(male), 2.0 mm (female) and 3.0 mm (male),
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<quantity id="4CAFD1D2FFAAFFF0FD1A3C54FCB6FE03" box="[706,792,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="157" unit="mm" value="2.9">2.9 mm</quantity>
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(female), respectively. Body generally shining, yellowish brown; four anteriolateral and posterolateral areas on metaterga darker. Antennae, legs, sterna and paraterga yellow.
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</paragraph>
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<paragraph id="8BE87C37FFAAFFF0FF1F3CF8FACDFD9C" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="157">
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Clypeolabral region densely setose. Epicranial suture evident, with 2+2 setae on vertex. Antennae long and slender, nearly reaching segment 4 when stretched laterally. In length, antennomere 6>2=5>1–7 (
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<figureCitation id="136C60B2FFAAFFF0FB0E3CDBFAB3FD9C" box="[1238,1309,511,536]" captionStart="FIGURE 6" captionStartId="6.[151,250,1768,1790]" captionTargetBox="[153,1423,661,1737]" captionTargetId="figure@6.[151,1435,661,1747]" captionTargetPageId="6" captionText="FIGURE 6. Enghoffosoma triangulare sp. nov. Holotype, anterior segments, lateral view (A), dorsal view (B), ventral view (C); midbody segments, dorsal view (D), lateral view (E); posterior segments, lateral view (F); sternum between coxae 4, ventral view (G); hypoproct and paraprocts, ventral view (H). Scale = 1 mm." httpUri="https://zenodo.org/record/265022/files/figure.png" pageId="6" pageNumber="157">Figs 6</figureCitation>
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A–C).
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</paragraph>
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<paragraph id="8BE87C37FFAAFFF0FF1F3F00FD50FDE4" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="157">
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Collum (
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<figureCitation id="136C60B2FFAAFFF0FEE93F00FED4FDB9" box="[305,378,548,573]" captionStart="FIGURE 6" captionStartId="6.[151,250,1768,1790]" captionTargetBox="[153,1423,661,1737]" captionTargetId="figure@6.[151,1435,661,1747]" captionTargetPageId="6" captionText="FIGURE 6. Enghoffosoma triangulare sp. nov. Holotype, anterior segments, lateral view (A), dorsal view (B), ventral view (C); midbody segments, dorsal view (D), lateral view (E); posterior segments, lateral view (F); sternum between coxae 4, ventral view (G); hypoproct and paraprocts, ventral view (H). Scale = 1 mm." httpUri="https://zenodo.org/record/265022/files/figure.png" pageId="6" pageNumber="157">Fig. 6</figureCitation>
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B) surface shining and smooth, with two rows of 4+4 and 2+2 setae; paraterga poorlydeveloped, rounded, not extending behind rear margin.
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</paragraph>
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<caption id="DF282CBFFFAAFFF0FF4F3BCCFC0EF8BF" httpUri="https://zenodo.org/record/265022/files/figure.png" pageId="6" pageNumber="157" targetBox="[153,1423,661,1737]" targetPageId="6">
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<paragraph id="8BE87C37FFAAFFF0FF4F3BCCFC0EF8BF" blockId="6.[151,1435,1768,1851]" pageId="6" pageNumber="157">
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<emphasis id="B923A025FFAAFFF0FF4F3BCCFEBAF97A" bold="true" box="[151,276,1768,1790]" pageId="6" pageNumber="157">FIGURE 6.</emphasis>
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<taxonomicName id="4C5707B4FFAAFFF0FEC43BCCFD87F979" box="[284,553,1768,1789]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="triangulare" status="sp. nov.">
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<emphasis id="B923A025FFAAFFF0FEC43BCCFD87F979" box="[284,553,1768,1789]" italics="true" pageId="6" pageNumber="157">Enghoffosoma triangulare</emphasis>
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</taxonomicName>
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<emphasis id="B923A025FFAAFFF0FDE93BCCFD2DF97A" bold="true" box="[561,643,1768,1790]" pageId="6" pageNumber="157">
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<taxonomicNameLabel id="A2101D5EFFAAFFF0FDE93BCCFD2DF97A" box="[561,643,1768,1790]" pageId="6" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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Holotype, anterior segments, lateral view (A), dorsal view (B), ventral view (C); midbody segments, dorsal view (D), lateral view (E); posterior segments, lateral view (F); sternum between coxae 4, ventral view (G); hypoproct and paraprocts, ventral view (H). Scale = 1 mm.
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</paragraph>
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</caption>
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<paragraph id="8BE87C37FFAAFFF0FF1F3A40FA33F86C" blockId="6.[151,1437,1891,2024]" pageId="6" pageNumber="157">In width, head<segment 2=3<4<2=5–16, thereafter body gradually tapering towards telson. Prozonae shining and smooth. Metaterga finely granulated, slightly wrinkled in posterolateral areas, with a row of 2+2 setae and traces of a row of 3+3 setae before and after transverse sulcus, respectively. Transverse sulcus present on segments 5–19, thin and deep, reaching to base of paraterga, striolate at bottom, especially on segments of rear half of body.</paragraph>
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<paragraph id="8BE87C37FFABFFF1FF4F3DB3FB2BFF7C" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="158">Sides below paraterga rugose, with several transverse wrinkles. Pleurosternal carinae well-developed, ridgeshaped; caudal corner spiniform and drawn behind posterior margin, present until segment 16, thereafter missing. Stricture between pro- and metazonae narrow, striolate at bottom. Axial line distinct, thin.</paragraph>
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<paragraph id="8BE87C37FFABFFF1FF1F3C21FAD6FEBB" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="158">
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Paraterga (
|
||
<figureCitation id="136C60B2FFABFFF1FE993C20FE26FE99" box="[321,392,260,285]" captionStart="FIGURE 6" captionStartId="6.[151,250,1768,1790]" captionTargetBox="[153,1423,661,1737]" captionTargetId="figure@6.[151,1435,661,1747]" captionTargetPageId="6" captionText="FIGURE 6. Enghoffosoma triangulare sp. nov. Holotype, anterior segments, lateral view (A), dorsal view (B), ventral view (C); midbody segments, dorsal view (D), lateral view (E); posterior segments, lateral view (F); sternum between coxae 4, ventral view (G); hypoproct and paraprocts, ventral view (H). Scale = 1 mm." httpUri="https://zenodo.org/record/265022/files/figure.png" pageId="7" pageNumber="158">Figs 6</figureCitation>
|
||
D–E) well-developed, beak-like, drawn behind posterior margin. Caudolateral corner acute, even pointed in most of caudal segments. Calluses small on poreless paraterga, but larger on pore-bearing ones.
|
||
</paragraph>
|
||
<paragraph id="8BE87C37FFABFFF1FF1F3C68FBECFE0C" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="158">
|
||
Epiproct (
|
||
<figureCitation id="136C60B2FFABFFF1FEE03C68FE2FFEE1" box="[312,385,332,357]" captionStart="FIGURE 6" captionStartId="6.[151,250,1768,1790]" captionTargetBox="[153,1423,661,1737]" captionTargetId="figure@6.[151,1435,661,1747]" captionTargetPageId="6" captionText="FIGURE 6. Enghoffosoma triangulare sp. nov. Holotype, anterior segments, lateral view (A), dorsal view (B), ventral view (C); midbody segments, dorsal view (D), lateral view (E); posterior segments, lateral view (F); sternum between coxae 4, ventral view (G); hypoproct and paraprocts, ventral view (H). Scale = 1 mm." httpUri="https://zenodo.org/record/265022/files/figure.png" pageId="7" pageNumber="158">Figs 6</figureCitation>
|
||
F, H) long and broadly truncated, with 2 small terminal tubercles. Tip with four spinnerets. Hypoproct subtriangular, with two small, distolateral, setiferous knobs (
|
||
<figureCitation id="136C60B2FFABFFF1FC633C4BFBAEFE0C" box="[955,1024,367,392]" captionStart="FIGURE 6" captionStartId="6.[151,250,1768,1790]" captionTargetBox="[153,1423,661,1737]" captionTargetId="figure@6.[151,1435,661,1747]" captionTargetPageId="6" captionText="FIGURE 6. Enghoffosoma triangulare sp. nov. Holotype, anterior segments, lateral view (A), dorsal view (B), ventral view (C); midbody segments, dorsal view (D), lateral view (E); posterior segments, lateral view (F); sternum between coxae 4, ventral view (G); hypoproct and paraprocts, ventral view (H). Scale = 1 mm." httpUri="https://zenodo.org/record/265022/files/figure.png" pageId="7" pageNumber="158">Figs 6</figureCitation>
|
||
F, H).
|
||
</paragraph>
|
||
<paragraph id="8BE87C37FFABFFF1FF1F3CB0FBF1FE54" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="158">
|
||
Sterna densely setose; longitudinal groove vague, whereas transverse one distinct, without modifications except for a densely setose, bifid, trapeziform lamina between male coxae 4 (
|
||
<figureCitation id="136C60B2FFABFFF1FC203C93FB95FE54" box="[1016,1083,439,464]" captionStart="FIGURE 6" captionStartId="6.[151,250,1768,1790]" captionTargetBox="[153,1423,661,1737]" captionTargetId="figure@6.[151,1435,661,1747]" captionTargetPageId="6" captionText="FIGURE 6. Enghoffosoma triangulare sp. nov. Holotype, anterior segments, lateral view (A), dorsal view (B), ventral view (C); midbody segments, dorsal view (D), lateral view (E); posterior segments, lateral view (F); sternum between coxae 4, ventral view (G); hypoproct and paraprocts, ventral view (H). Scale = 1 mm." httpUri="https://zenodo.org/record/265022/files/figure.png" pageId="7" pageNumber="158">Fig. 6</figureCitation>
|
||
G).
|
||
</paragraph>
|
||
<paragraph id="8BE87C37FFABFFF1FF1F3CF9FAC2FD93" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="158">
|
||
Legs (
|
||
<figureCitation id="136C60B2FFABFFF1FED53CF8FEFDFE71" box="[269,339,476,501]" captionStart="FIGURE 6" captionStartId="6.[151,250,1768,1790]" captionTargetBox="[153,1423,661,1737]" captionTargetId="figure@6.[151,1435,661,1747]" captionTargetPageId="6" captionText="FIGURE 6. Enghoffosoma triangulare sp. nov. Holotype, anterior segments, lateral view (A), dorsal view (B), ventral view (C); midbody segments, dorsal view (D), lateral view (E); posterior segments, lateral view (F); sternum between coxae 4, ventral view (G); hypoproct and paraprocts, ventral view (H). Scale = 1 mm." httpUri="https://zenodo.org/record/265022/files/figure.png" pageId="7" pageNumber="158">Figs 6</figureCitation>
|
||
D–E) thin and slender, about 1.5 times as long as midbody height (male). Prefemora not swollen. Femora without modifications. Tarsal brushes present on male leg-pairs 1–9, thereafter gradually thinning out.
|
||
</paragraph>
|
||
<paragraph id="8BE87C37FFABFFF1FF1F3F00FB53FD74" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="158">
|
||
Gonopod (
|
||
<figureCitation id="136C60B2FFABFFF1FE9C3F00FE01FDB8" box="[324,431,548,573]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="7.[151,250,1997,2019]" captionStartId-1="8.[151,250,1431,1453]" captionTargetBox-0="[161,1425,956,1967]" captionTargetBox-1="[151,1436,193,1409]" captionTargetId-0="figure@7.[151,1435,949,1976]" captionTargetId-1="figure@8.[151,1436,193,1410]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 7. Enghoffosoma triangulare sp. nov. Holotype, right gonopod, lateral view (A); mesal view (B). Scale = 1 mm." captionText-1="FIGURE 8. Enghoffosoma triangulare sp. nov. Holotype, right gonopod, lateroventral view (A); postfemoral region, lateroventral view (B)." httpUri-0="https://zenodo.org/record/265023/files/figure.png" httpUri-1="https://zenodo.org/record/265024/files/figure.png" pageId="7" pageNumber="158">Figs 7–8</figureCitation>
|
||
) relatively simple. Coxite large, stout; distoventral part sparsely setose. Prefemorite densely setose, separated laterally from femorite by a transverse sulcus. Femorite slender, cylindrical, but somewhat curved ventrad, with neither processes nor other modifications. Postfemoral region consisting only of a solenomere, clearly separated from femorite by a cingulum. Solenomere slightly spiralled, as large as femorite at base, thereafter tapering towards tip; distal part with a small pointed tooth; process
|
||
<emphasis id="B923A025FFABFFF1FBB83F90FBC2FD49" bold="true" box="[1120,1132,692,717]" pageId="7" pageNumber="158">e</emphasis>
|
||
small, triangular. Seminal groove running distodorsad entirely mesally along femorite before entering onto a thick solenomere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D2FBCFFABFFFEFF1F3FD8FE28F9B1" lastPageId="8" lastPageNumber="159" pageId="7" pageNumber="158" type="discussion">
|
||
<paragraph id="8BE87C37FFABFFF1FF1F3FD8FCFCFCFB" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="158">
|
||
<emphasis id="B923A025FFABFFF1FF1F3FD8FE95FC91" bold="true" box="[199,315,764,789]" pageId="7" pageNumber="158">Remarks.</emphasis>
|
||
The species is clearly distinguished from other known
|
||
<taxonomicName id="4C5707B4FFABFFF1FC043FD8FBD0FC90" box="[988,1150,764,788]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923A025FFABFFF1FC043FD8FBD0FC90" box="[988,1150,764,788]" italics="true" pageId="7" pageNumber="158">Enghoffosoma</emphasis>
|
||
</taxonomicName>
|
||
species by its gonopod conformation. It seems to be more similar to
|
||
<taxonomicName id="4C5707B4FFABFFF1FD6D3E05FB20FCBC" authority="Likhitrakarn et al., 2014" authorityName="Likhitrakarn et al." authorityYear="2014" box="[693,1166,799,824]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="lanceolatum">
|
||
<emphasis id="B923A025FFABFFF1FD6D3E05FCCDFCBC" box="[693,867,800,824]" italics="true" pageId="7" pageNumber="158">E. lanceolatum</emphasis>
|
||
<bibRefCitation id="EFC601C6FFABFFF1FCB73E3BFB20FCBC" author="Likhitrakarn" box="[879,1166,799,824]" pageId="7" pageNumber="158" refString="Likhitrakarn, K., Golovatch, S. I. & Panha, S. (2014) Review of the Southeast Asian millipede genus Enghoffosoma Golovatch, 1993 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species. Zootaxa, 3811 (4), 491 - 514. http: // dx. doi. org / 10.11646 / zootaxa. 3811.4.4" type="journal article" year="2014">
|
||
Likhitrakarn
|
||
<emphasis id="B923A025FFABFFF1FBDE3E05FBEDFCBC" box="[1030,1091,800,824]" italics="true" pageId="7" pageNumber="158">et al.</emphasis>
|
||
, 2014
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
by the well-developed paraterga and the small process
|
||
<emphasis id="B923A025FFABFFF1FDC13E60FD8BFCD9" bold="true" box="[537,549,836,861]" pageId="7" pageNumber="158">e</emphasis>
|
||
.
|
||
<taxonomicName id="4C5707B4FFABFFF1FDE23E61FD46FCD8" box="[570,744,836,860]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="lanceolatum">
|
||
<emphasis id="B923A025FFABFFF1FDE23E61FD46FCD8" box="[570,744,836,860]" italics="true" pageId="7" pageNumber="158">E. lanceolatum</emphasis>
|
||
</taxonomicName>
|
||
differs from the new species in the gonopod showing an apicoventral shelf and a pointed, parabasal subventral process.
|
||
</paragraph>
|
||
<caption id="DF282CBFFFABFFF1FF4F3AE9FADEF867" box="[151,1392,1997,2019]" httpUri="https://zenodo.org/record/265023/files/figure.png" pageId="7" pageNumber="158" targetBox="[161,1425,956,1967]" targetPageId="7">
|
||
<paragraph id="8BE87C37FFABFFF1FF4F3AE9FADEF867" blockId="7.[151,1392,1997,2019]" box="[151,1392,1997,2019]" pageId="7" pageNumber="158">
|
||
<emphasis id="B923A025FFABFFF1FF4F3AE9FEBCF867" bold="true" box="[151,274,1997,2019]" pageId="7" pageNumber="158">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C5707B4FFABFFF1FEC13AE9FD8AF866" box="[281,548,1997,2018]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="triangulare" status="sp. nov.">
|
||
<emphasis id="B923A025FFABFFF1FEC13AE9FD8AF866" box="[281,548,1997,2018]" italics="true" pageId="7" pageNumber="158">Enghoffosoma triangulare</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923A025FFABFFF1FDF23AE9FDD4F867" bold="true" box="[554,634,1997,2019]" pageId="7" pageNumber="158">
|
||
<taxonomicNameLabel id="A2101D5EFFABFFF1FDF23AE9FDD4F867" box="[554,634,1997,2019]" pageId="7" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Holotype, right gonopod, lateral view (A); mesal view (B). Scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF282CBFFFA4FFFEFF4F38B3FED3FA48" httpUri="https://zenodo.org/record/265024/files/figure.png" pageId="8" pageNumber="159" targetBox="[151,1436,193,1409]" targetPageId="8">
|
||
<paragraph id="8BE87C37FFA4FFFEFF4F38B3FED3FA48" blockId="8.[151,1436,1431,1484]" pageId="8" pageNumber="159">
|
||
<emphasis id="B923A025FFA4FFFEFF4F38B3FEB4FA29" bold="true" box="[151,282,1431,1453]" pageId="8" pageNumber="159">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C5707B4FFA4FFFEFEF038BCFD94FA29" box="[296,570,1432,1453]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="triangulare" status="sp. nov.">
|
||
<emphasis id="B923A025FFA4FFFEFEF038BCFD94FA29" box="[296,570,1432,1453]" italics="true" pageId="8" pageNumber="159">Enghoffosoma triangulare</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923A025FFA4FFFEFD9038B3FD0EFA29" bold="true" box="[584,672,1431,1453]" pageId="8" pageNumber="159">
|
||
<taxonomicNameLabel id="A2101D5EFFA4FFFEFD9038B3FD0EFA29" box="[584,672,1431,1453]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Holotype, right gonopod, lateroventral view (A); postfemoral region, lateroventral view (B).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE87C37FFA4FFFEFF1F38DCFE28F9B1" blockId="8.[151,1436,1528,1589]" pageId="8" pageNumber="159">
|
||
The species was found in the Cat Tien National Park, southern
|
||
<collectingCountry id="F3403CA7FFA4FFFEFC5B38DCFC4BF995" box="[899,997,1528,1553]" name="Vietnam" pageId="8" pageNumber="159">Vietnam</collectingCountry>
|
||
, where the best forests in the region are still being conserved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |