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<document id="F1120AF220B3D2BAF0118E5AC19F2358" ID="10.11646/zootaxa.4139.2.1" ID-DOI="10.11646/zootaxa.4139.2.1" ID-GBIF-Dataset="aeecbbb4-4764-41dd-9562-b25f91f9f70e" ID-ISSN="1175-5326" ID-Zenodo-Dep="265016" ID-ZooBank="2EAFF1B9-1FA2-44B1-89A3-476359314C5E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468994861279" checkinUser="plazi" docAuthor="Nguyen, Anh D. &amp; Golovatch, Sergei I." docDate="2016" docId="03FECD21FFA3FFF9FF4F3DBCFA35FDDE" docLanguage="en" docName="zootaxa.4139.2.1.pdf" docOrigin="Zootaxa 4139 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enghoffosoma" docType="key" docVersion="11" lastPageNumber="166" masterDocId="FFC7B559FFACFFF6FFD83D24FFAEFF84" masterDocTitle="The millipede genus Enghoffosoma Golovatch, 1993 recorded in Vietnam for the first time, with descriptions of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="166" masterPageNumber="151" pageNumber="166" updateTime="1698365308467" updateUser="plazi">
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<mods:title id="83005B21B0319480B98C0060E21BEC3E">The millipede genus Enghoffosoma Golovatch, 1993 recorded in Vietnam for the first time, with descriptions of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:name id="63717145E3B04634CCDA4F36D48BC458" type="personal">
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<mods:namePart id="39930F6E50CF76A4749786DB55F30AEE">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="709535BCA6904F34849F11CDB0D781C3">Golovatch, Sergei I.</mods:namePart>
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<mods:title id="0CE4E59D8676BAD96324ADA28DDD9913">Zootaxa</mods:title>
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<mods:part id="78AAE65540DF07495E4B155AC0411DB6">
<mods:date id="2C71BA10223251BD4CA966DB46899487">2016</mods:date>
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<mods:number id="D87373C2061E25EBBB2DEFA18FB0A9F9">4139</mods:number>
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<mods:identifier id="08024F9B38D6107381FFFC5621019334" type="DOI">10.11646/zootaxa.4139.2.1</mods:identifier>
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<treatment id="03FECD21FFA3FFF9FF4F3DBCFA35FDDE" ID-DOI="http://doi.org/10.5281/zenodo.6067539" ID-GBIF-Taxon="123198409" ID-Zenodo-Dep="6067539" LSID="urn:lsid:plazi:treatment:03FECD21FFA3FFF9FF4F3DBCFA35FDDE" httpUri="http://treatment.plazi.org/id/03FECD21FFA3FFF9FF4F3DBCFA35FDDE" lastPageNumber="166" pageId="15" pageNumber="166">
<subSubSection id="C34D2FBCFFA3FFF9FF4F3DBCFD9CFF57" pageId="15" pageNumber="166" type="nomenclature">
<paragraph id="8BE87C37FFA3FFF9FF4F3DBCFCD4FF36" blockId="15.[151,890,151,211]" box="[151,890,151,178]" pageId="15" pageNumber="166">
<heading id="D0A0CB5BFFA3FFF9FF4F3DBCFCD4FF36" bold="true" box="[151,890,151,178]" fontSize="11" level="1" pageId="15" pageNumber="166" reason="1">
<emphasis id="B923A025FFA3FFF9FF4F3DBCFCD4FF36" bold="true" box="[151,890,151,178]" pageId="15" pageNumber="166">
Key to species of
<taxonomicName id="4C5707B4FFA3FFF9FEB53DB3FD8EFF35" ID-CoL="7PD8X" box="[365,544,151,177]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B923A025FFA3FFF9FEB53DB3FD8EFF35" bold="true" box="[365,544,151,177]" italics="true" pageId="15" pageNumber="166">Enghoffosoma</emphasis>
</taxonomicName>
known to occur in
<collectingCountry id="F3403CA7FFA3FFF9FCC83DBCFCD4FF36" box="[784,890,152,178]" name="Vietnam" pageId="15" pageNumber="166">Vietnam</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE87C37FFA3FFF9FF4F3D9EFD9CFF57" blockId="15.[151,890,151,211]" box="[151,562,186,211]" pageId="15" pageNumber="166">(based on gonopod characters alone)</paragraph>
</subSubSection>
<subSubSection id="C34D2FBCFFA3FFF9FF4F3C25FA35FDDE" pageId="15" pageNumber="166" type="key">
<keyStep id="30A36492FFA3FFF9FF4F3C25FA35FEB1" pageId="15" pageNumber="166">
<paragraph id="8BE87C37FFA3FFF9FF4F3C25FA35FE93" blockId="15.[151,1436,257,602]" box="[151,1435,257,280]" pageId="15" pageNumber="166">
<keyLead id="30A6DF02FFA3FFF9FF4F3C25FA35FE93" box="[151,1435,257,280]" pageId="15" pageNumber="166">
1 Gonopod postfemoral region consisting only of a solenomere and a process
<emphasis id="B923A025FFA3FFF9FC623C26FC6BFE9C" bold="true" box="[954,965,258,280]" pageId="15" pageNumber="166">e</emphasis>
.................................... 2
</keyLead>
</paragraph>
<paragraph id="8BE87C37FFA3FFF9FF4F3C3BFA35FEB1" blockId="15.[151,1436,257,602]" box="[151,1435,287,309]" pageId="15" pageNumber="166">
<keyLead id="30A6DF02FFA3FFF9FF4F3C3BFA35FEB1" box="[151,1435,287,309]" pageId="15" pageNumber="166">
- Gonopod postfemoral region consisting of a solenomere and processes
<emphasis id="B923A025FFA3FFF9FC543C3BFC39FEB1" bold="true" box="[908,919,287,309]" pageId="15" pageNumber="166">e</emphasis>
and
<emphasis id="B923A025FFA3FFF9FC1D3C3BFC7CFEB1" bold="true" box="[965,978,287,309]" pageId="15" pageNumber="166">p</emphasis>
.................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A36492FFA3FFF9FF4F3C18FA35FEEB" pageId="15" pageNumber="166">
<paragraph id="8BE87C37FFA3FFF9FF4F3C18FA32FED6" blockId="15.[151,1436,257,602]" box="[151,1436,316,338]" pageId="15" pageNumber="166">
<keyLead id="30A6DF02FFA3FFF9FF4F3C18FA32FED6" box="[151,1436,316,338]" pageId="15" pageNumber="166">
2 Process
<emphasis id="B923A025FFA3FFF9FEE83C18FE95FED6" bold="true" box="[304,315,316,338]" pageId="15" pageNumber="166">e</emphasis>
small, triangular (
<figureCitation id="136C60B2FFA3FFF9FE333C18FDE8FED6" box="[491,582,316,338]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="7.[151,250,1997,2019]" captionStartId-1="8.[151,250,1431,1453]" captionTargetBox-0="[161,1425,956,1967]" captionTargetBox-1="[151,1436,193,1409]" captionTargetId-0="figure@7.[151,1435,949,1976]" captionTargetId-1="figure@8.[151,1436,193,1410]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 7. Enghoffosoma triangulare sp. nov. Holotype, right gonopod, lateral view (A); mesal view (B). Scale = 1 mm." captionText-1="FIGURE 8. Enghoffosoma triangulare sp. nov. Holotype, right gonopod, lateroventral view (A); postfemoral region, lateroventral view (B)." httpUri-0="https://zenodo.org/record/265023/files/figure.png" httpUri-1="https://zenodo.org/record/265024/files/figure.png" pageId="15" pageNumber="166">Figs. 78</figureCitation>
)...................................................
<taxonomicName id="4C5707B4FFA3FFF9FB1A3C19FAE7FED6" ID-CoL="39T4L" box="[1218,1353,317,338]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="triangulare" status="sp. nov.">
<emphasis id="B923A025FFA3FFF9FB1A3C19FAE7FED6" box="[1218,1353,317,338]" italics="true" pageId="15" pageNumber="166">E. triangulare</emphasis>
</taxonomicName>
<emphasis id="B923A025FFA3FFF9FA973C18FA32FED6" bold="true" box="[1359,1436,316,338]" pageId="15" pageNumber="166">
<taxonomicNameLabel id="A2101D5EFFA3FFF9FA973C18FA32FED6" box="[1359,1436,316,338]" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE87C37FFA3FFF9FF4F3C7EFA35FEEB" blockId="15.[151,1436,257,602]" box="[151,1435,345,368]" pageId="15" pageNumber="166">
<keyLead id="30A6DF02FFA3FFF9FF4F3C7EFA35FEEB" box="[151,1435,345,368]" pageId="15" pageNumber="166">
- Process
<emphasis id="B923A025FFA3FFF9FEE83C7EFE95FEF4" bold="true" box="[304,315,346,368]" pageId="15" pageNumber="166">e</emphasis>
large, shaped otherwise....................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A36492FFA3FFF9FF4F3C53FA32FE2E" pageId="15" pageNumber="166">
<paragraph id="8BE87C37FFA3FFF9FF4F3C53FA38FE09" blockId="15.[151,1436,257,602]" box="[151,1430,375,397]" pageId="15" pageNumber="166">
<keyLead id="30A6DF02FFA3FFF9FF4F3C53FA38FE09" box="[151,1430,375,397]" pageId="15" pageNumber="166">
3 Process
<emphasis id="B923A025FFA3FFF9FEE83C53FE95FE09" bold="true" box="[304,315,375,397]" pageId="15" pageNumber="166">e</emphasis>
stout, straight, slightly shorter than solenomere (
<figureCitation id="136C60B2FFA3FFF9FCC83C53FC2DFE09" box="[784,899,375,397]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="13.[151,250,1037,1059]" captionStartId-1="13.[151,250,1969,1991]" captionTargetBox-0="[165,1421,202,1004]" captionTargetBox-1="[151,1436,1108,1947]" captionTargetId-0="figure@13.[151,1436,193,1015]" captionTargetId-1="figure@13.[151,1436,1108,1947]" captionTargetPageId-0="13" captionTargetPageId-1="13" captionText-0="FIGURE 13. Enghoffosoma digitatum sp. nov. Holotype, right gonopod, mesal view (A); lateral view (B). Scale = 1 mm." captionText-1="FIGURE 14. Enghoffosoma digitatum sp. nov. Holotype, right gonopod, mesal view (A); postfemoral region, mesal view (B)." httpUri-0="https://zenodo.org/record/265029/files/figure.png" httpUri-1="https://zenodo.org/record/265030/files/figure.png" pageId="15" pageNumber="166">Figs. 1314</figureCitation>
).........................
<taxonomicName id="4C5707B4FFA3FFF9FB143C53FAEDFE08" ID-CoL="39T4B" box="[1228,1347,375,396]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="digitatum" status="sp. nov.">
<emphasis id="B923A025FFA3FFF9FB143C53FAEDFE08" box="[1228,1347,375,396]" italics="true" pageId="15" pageNumber="166">E. digitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2101D5EFFA3FFF9FA913C53FA38FE09" box="[1353,1430,375,397]" pageId="15" pageNumber="166" rank="species">
<emphasis id="B923A025FFA3FFF9FA913C53FA38FE09" bold="true" box="[1353,1430,375,397]" pageId="15" pageNumber="166">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BE87C37FFA3FFF9FF4F3CB0FA32FE2E" blockId="15.[151,1436,257,602]" box="[151,1436,404,426]" pageId="15" pageNumber="166">
<keyLead id="30A6DF02FFA3FFF9FF4F3CB0FA32FE2E" box="[151,1436,404,426]" pageId="15" pageNumber="166">
<emphasis id="B923A025FFA3FFF9FF4F3CB0FF31FE2E" bold="true" box="[151,159,404,426]" pageId="15" pageNumber="166">-</emphasis>
Process
<emphasis id="B923A025FFA3FFF9FEE83CB0FE95FE2E" bold="true" box="[304,315,404,426]" pageId="15" pageNumber="166">e</emphasis>
lobe-shaped, curved ventrad, much shorter than solenomere (
<figureCitation id="136C60B2FFA3FFF9FC523CB0FC71FE2E" box="[906,991,404,426]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="4.[151,250,1963,1985]" captionStartId-1="5.[151,250,1190,1212]" captionTargetBox-0="[197,1389,882,1933]" captionTargetBox-1="[151,1436,193,1168]" captionTargetId-0="figure@4.[189,1398,877,1941]" captionTargetId-1="figure@5.[151,1436,193,1169]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 4. Enghoffosoma fedorenkoi (Golovatch, 2016), comb. nov., male from Chu Yan Sin National Park, Vietnam. Right gonopod, lateral view (A); mesal view (B). Scale = 1 mm." captionText-1="FIGURE 5. Enghoffosoma fedorenkoi (Golovatch, 2016), comb. nov., male from Chu Yan Sin National Park; Vietnam. Left gonopod, lateroventral view (A), mesoventral view (B)." httpUri-0="https://zenodo.org/record/265020/files/figure.png" httpUri-1="https://zenodo.org/record/265021/files/figure.png" pageId="15" pageNumber="166">Figs 45</figureCitation>
).........................
<taxonomicName id="4C5707B4FFA3FFF9FAC33CB1FA32FE2E" ID-CoL="39T4D" box="[1307,1436,405,426]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="fedorenkoi">
<emphasis id="B923A025FFA3FFF9FAC33CB1FA32FE2E" box="[1307,1436,405,426]" italics="true" pageId="15" pageNumber="166">E. fedorenkoi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A36492FFA3FFF9FF4F3C95FA35FE61" pageId="15" pageNumber="166">
<paragraph id="8BE87C37FFA3FFF9FF4F3C95FA32FE4C" blockId="15.[151,1436,257,602]" box="[151,1436,433,456]" pageId="15" pageNumber="166">
<keyLead id="30A6DF02FFA3FFF9FF4F3C95FA32FE4C" box="[151,1436,433,456]" pageId="15" pageNumber="166">
4 Femorite with an apical shelf
<emphasis id="B923A025FFA3FFF9FE263C96FDA5FE4C" bold="true" box="[510,523,434,456]" pageId="15" pageNumber="166">b</emphasis>
. Process
<emphasis id="B923A025FFA3FFF9FDBF3C96FDDAFE4C" bold="true" box="[615,628,434,456]" pageId="15" pageNumber="166">p</emphasis>
small, retrorse and tuberculiform (
<figureCitation id="136C60B2FFA3FFF9FC1A3C95FB9FFE43" box="[962,1073,433,455]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStartId-0="10.[151,250,1397,1419]" captionStartId-1="11.[151,250,1430,1452]" captionTargetBox-0="[162,1425,194,1366]" captionTargetBox-1="[151,1436,193,1408]" captionTargetId-0="figure@10.[151,1436,193,1375]" captionTargetId-1="figure@11.[151,1436,193,1409]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 10. Enghoffosoma retrorsum sp. nov. Holotype, right gonopod, lateral view (A); mesal view (B). Scale = 1 mm." captionText-1="FIGURE 11. Enghoffosoma retrorsum sp. nov. Holotype, right gonopod, mesal view (A); postfemoral region, mesal view (B)." httpUri-0="https://zenodo.org/record/265026/files/figure.png" httpUri-1="https://zenodo.org/record/265027/files/figure.png" pageId="15" pageNumber="166">Figs 1011</figureCitation>
)............
<taxonomicName id="4C5707B4FFA3FFF9FB083C96FAE7FE43" ID-CoL="39T4H" box="[1232,1353,434,455]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="retrorsum" status="sp. nov.">
<emphasis id="B923A025FFA3FFF9FB083C96FAE7FE43" box="[1232,1353,434,455]" italics="true" pageId="15" pageNumber="166">E. retrorsum</emphasis>
</taxonomicName>
<emphasis id="B923A025FFA3FFF9FA973C96FA32FE4C" bold="true" box="[1359,1436,434,456]" pageId="15" pageNumber="166">
<taxonomicNameLabel id="A2101D5EFFA3FFF9FA973C96FA32FE4C" box="[1359,1436,434,456]" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE87C37FFA3FFF9FF4F3CEBFA35FE61" blockId="15.[151,1436,257,602]" box="[151,1435,463,485]" pageId="15" pageNumber="166">
<keyLead id="30A6DF02FFA3FFF9FF4F3CEBFA35FE61" box="[151,1435,463,485]" pageId="15" pageNumber="166">
- Femorite without apical shelf
<emphasis id="B923A025FFA3FFF9FDD83CEBFDA3FE61" bold="true" box="[512,525,463,485]" pageId="15" pageNumber="166">b</emphasis>
. Process
<emphasis id="B923A025FFA3FFF9FDB13CEBFDD8FE61" bold="true" box="[617,630,463,485]" pageId="15" pageNumber="166">p</emphasis>
large, shaped otherwise............................................. 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A36492FFA3FFF9FF4F3CC8FA35FDDE" pageId="15" pageNumber="166">
<paragraph id="8BE87C37FFA3FFF9FF4F3CC8FA35FD9B" blockId="15.[151,1436,257,602]" pageId="15" pageNumber="166">
<keyLead id="30A6DF02FFA3FFF9FF4F3CC8FA35FD9B" pageId="15" pageNumber="166">
5 Solenomere large, twisted in distal part; process
<emphasis id="B923A025FFA3FFF9FD1D3CC8FD7EFD86" bold="true" box="[709,720,492,514]" pageId="15" pageNumber="166">e</emphasis>
large, serrate at anterior margin; process
<emphasis id="B923A025FFA3FFF9FBA83CC8FBD3FD86" bold="true" box="[1136,1149,492,514]" pageId="15" pageNumber="166">p</emphasis>
anchor-shaped; apical self
<emphasis id="B923A025FFA3FFF9FA563CC8FA35FD86" bold="true" box="[1422,1435,492,514]" pageId="15" pageNumber="166">b</emphasis>
absent (
<figureCitation id="136C60B2FFA3FFF9FEF33F2DFECBFD9B" box="[299,357,521,543]" captionStart="FIGURE 2" captionStartId="2.[151,250,1891,1913]" captionTargetBox="[151,1435,982,1869]" captionTargetId="figure@2.[151,1435,982,1869]" captionTargetPageId="2" captionText="FIGURE 2. Enghoffosoma anchoriforme Likhitrakarn, Golovatch &amp; Panha, 2014, male from Cat Tien National Park, Vietnam. Right gonopod, mesal view (A); postfemoral region, mesal view (B)." httpUri="https://zenodo.org/record/265018/files/figure.png" pageId="15" pageNumber="166">Fig. 2</figureCitation>
)...........................................................................
<taxonomicName id="4C5707B4FFA3FFF9FB263F2EFA35FD9B" ID-CoL="39T48" box="[1278,1435,522,543]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="anchoriforme">
<emphasis id="B923A025FFA3FFF9FB263F2EFA35FD9B" box="[1278,1435,522,543]" italics="true" pageId="15" pageNumber="166">E. anchoriforme</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE87C37FFA3FFF9FF4F3F03FA35FDDE" blockId="15.[151,1436,257,602]" pageId="15" pageNumber="166">
<keyLead id="30A6DF02FFA3FFF9FF4F3F03FA35FDDE" pageId="15" pageNumber="166">
- Solenomere large, spiralled; distal part with a tooth terminating the seminal groove; process
<emphasis id="B923A025FFA3FFF9FB853F03FBC6FDB9" bold="true" box="[1117,1128,551,573]" pageId="15" pageNumber="166">e</emphasis>
small, tuberculiform; process
<emphasis id="B923A025FFA3FFF9FA563F03FA35FDB9" bold="true" box="[1422,1435,551,573]" pageId="15" pageNumber="166">p</emphasis>
small, erect, subrectangular (
<figureCitation id="136C60B2FFA3FFF9FE2E3F60FD87FDDE" box="[502,553,580,602]" captionStart="FIGURE 1" captionStartId="2.[151,250,880,902]" captionTargetBox="[153,1433,193,858]" captionTargetId="figure@2.[153,1433,193,859]" captionTargetPageId="2" captionText="FIGURE 1. Enghoffosoma lanceolatum Likhitrakarn, Golovatch &amp; Panha, 2014, male from Yok Don National Park, Vietnam. Right gonopod, mesal view (A), ventral view (B), and lateral view (C)." httpUri="https://zenodo.org/record/265017/files/figure.png" pageId="15" pageNumber="166">Fig.1</figureCitation>
)............................................................
<taxonomicName id="4C5707B4FFA3FFF9FAD33F61FA35FDDE" ID-CoL="39T4F" box="[1291,1435,581,602]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="lanceolatum">
<emphasis id="B923A025FFA3FFF9FAD33F61FA35FDDE" box="[1291,1435,581,602]" italics="true" pageId="15" pageNumber="166">E. lanceolatum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>