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<document id="5EAF63B7A4C0EC1F13ED79F3E2B12544" ID-DOI="10.5324/cjcr.v0i30.2240" ID-ISSN="2387-5372" ID-Zenodo-Dep="7987298" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1685469670408" checkinUser="felipe" docAuthor="Epler, John H." docDate="2017" docId="03E2824B3D78B920143C6DA7FD9BFAB7" docLanguage="en" docName="ChiroJChiroRes.30.4-18.pdf" docOrigin="CHIRONOMUS Journal of Chironomidae Research 30 (30)" docSource="https://www.ntnu.no/ojs/index.php/chironomus/article/view/2240" docStyle="DocumentStyle:B21B57A70ED88EDAB7971C4BDED0DA62.1:ChiroJChiroRes.2015-.journal_article" docStyleId="B21B57A70ED88EDAB7971C4BDED0DA62" docStyleName="ChiroJChiroRes.2015-.journal_article" docStyleVersion="1" docTitle="Parametriocnemus Goetghebuer 1931" docType="treatment" docVersion="1" lastPageNumber="8" masterDocId="FFDBFA333D7BB92417046A28FFD5FF8F" masterDocTitle="AN ANNOTATED PRELIMINARY LIST OF THE CHIRONOMIDAE (DIPTERA) OF ZURQUÍ, COSTA RICA Abstract Introduction" masterLastPageNumber="18" masterPageNumber="4" pageNumber="7" updateTime="1685624128335" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="07F6D07C8DCE5A321859F8F338166622">AN ANNOTATED PRELIMINARY LIST OF THE CHIRONOMIDAE (DIPTERA) OF ZURQUÍ, COSTA RICA Abstract Introduction</mods:title>
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<mods:namePart id="3BAEF981CC575B8ECA80D64558A069BF">Epler, John H.</mods:namePart>
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<treatment id="03E2824B3D78B920143C6DA7FD9BFAB7" LSID="urn:lsid:plazi:treatment:03E2824B3D78B920143C6DA7FD9BFAB7" httpUri="http://treatment.plazi.org/id/03E2824B3D78B920143C6DA7FD9BFAB7" lastPageId="4" lastPageNumber="8" pageId="3" pageNumber="7">
<subSubSection id="C35160D63D78B927143C6DA7FBDCF828" box="[824,1033,1935,1959]" pageId="3" pageNumber="7" type="nomenclature">
<paragraph id="8BF4335D3D78B927143C6DA7FBDCF828" blockId="3.[824,1361,1935,2023]" box="[824,1033,1935,1959]" pageId="3" pageNumber="7">
<taxonomicName id="4C4B48DE3D78B927143C6DA7FBDCF828" ID-CoL="6FJS" ID-ENA="611373" authorityName="Goetghebuer" authorityYear="1931" box="[824,1033,1935,1959]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEF4F3D78B927143C6DA7FBDCF828" bold="true" box="[824,1033,1935,1959]" italics="true" pageId="3" pageNumber="7">Parametriocnemus</emphasis>
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</paragraph>
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<paragraph id="8BF4335D3D78B92013146DA7FDFEFEF3" blockId="3.[824,1361,1935,2023]" lastBlockId="4.[227,764,227,380]" lastPageId="4" lastPageNumber="8" pageId="3" pageNumber="7">
- Four species were collected; all appear to be undescribed.
<taxonomicName id="4C4B48DE3D78B92713826D87FC82F869" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93FEF4F3D78B92713826D87FA84F849" box="[1158,1361,1967,1990]" italics="true" pageId="3" pageNumber="7">Parametriocnemus</emphasis>
sp.
</taxonomicName>
ZUR-1 and
<emphasis id="B93FEF4F3D78B92714E46DE7FC25F869" box="[992,1008,1999,2022]" italics="true" pageId="3" pageNumber="7">P</emphasis>
. sp. ZUR-2 resemble the Nearctic
<taxonomicName id="4C4B48DE3D7FB92016006ACCFDC1FF74" authority="(Johannsen)" baseAuthorityName="Johannsen" baseAuthorityYear="1905" box="[260,532,227,251]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="8" phylum="Arthropoda" rank="species" species="lundbeckii">
<emphasis id="B93FEF4F3D7FB92016006ACCFE58FF74" box="[260,397,227,251]" italics="true" pageId="4" pageNumber="8">P. lundbeckii</emphasis>
(Johannsen)
</taxonomicName>
but have a lower AR;
<emphasis id="B93FEF4F3D7FB92017E76B2CFF26FE94" box="[227,243,260,283]" italics="true" pageId="4" pageNumber="8">P</emphasis>
. sp. ZUR-1 was 0.670.72;
<emphasis id="B93FEF4F3D7FB92015366B2CFD97FE94" box="[562,578,260,283]" italics="true" pageId="4" pageNumber="8">P</emphasis>
. sp. ZUR-2 was 0.52-0.68 (for
<taxonomicName id="4C4B48DE3D7FB92016846B0CFDDCFEB4" baseAuthorityName="Johannsen" baseAuthorityYear="1905" box="[384,521,291,315]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="8" phylum="Arthropoda" rank="species" species="lundbeckii">
<emphasis id="B93FEF4F3D7FB92016846B0CFDDCFEB4" box="[384,521,291,315]" italics="true" pageId="4" pageNumber="8">P. lundbeckii</emphasis>
</taxonomicName>
see below). Both may represent a single species; more measurements of other body parts are necessary.
</paragraph>
<paragraph id="8BF4335D3D7FB92017E76BBAFD96FDC5" blockId="4.[227,764,402,586]" pageId="4" pageNumber="8">
<bibRefCitation id="EFDA4EAC3D7FB92017E76BBAFE6BFE25" author="Sublette, J. E." box="[227,446,402,426]" pageId="4" pageNumber="8" pagination="477 - 564" refId="ref10537" refString="Sublette, J. E. 1967. Type specimens of Chironomidae (Diptera) in the Cornell University Collection. - Journal of the Kansas Entomological Society 40: 477 - 564." type="journal article" year="1967">Sublette (1967: 539)</bibRefCitation>
gave an AR of 1.40 based on a specimen (a “metatype”) from the
<typeStatus id="54F08DFF3D7FB920157B6B9BFD78FE44" box="[639,685,435,459]" pageId="4" pageNumber="8">type</typeStatus>
series; he also gave ARs of four Johannsen-determined specimens that ranged from 1.261.60, mean 1.36.
<bibRefCitation id="EFDA4EAC3D7FB92017E7683AFE55FDA5" author="Saether, O. A." box="[227,384,530,554]" pageId="4" pageNumber="8" pagination="1 - 154" refId="ref9993" refString="Saether, O. A. 1969. Some Nearctic Podonominae, Diamesinae, and Orthocladiinae (Diptera: Chironomidae). - Fisheries Research Board of Canada Bulletin 170: 1 - 154." type="journal article" year="1969">Saether (1969)</bibRefCitation>
gave a range of 0.90-1.30, mean 1.12, for the AR of
<taxonomicName id="4C4B48DE3D7FB92016B7681BFDE9FDC5" baseAuthorityName="Johannsen" baseAuthorityYear="1905" box="[435,572,562,586]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="8" phylum="Arthropoda" rank="species" species="lundbeckii">
<emphasis id="B93FEF4F3D7FB92016B7681BFDE9FDC5" box="[435,572,562,586]" italics="true" pageId="4" pageNumber="8">P. lundbeckii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF4335D3D7FB92017E76849FE6AFC96" blockId="4.[227,764,609,793]" pageId="4" pageNumber="8">
<materialsCitation id="3B2339003D7FB92017E76849FE6DFC96" country="Guatemala" location="Guatemala" pageId="4" pageNumber="8" specimenCount="5" specimenCount-female="5">
<bibRefCitation id="EFDA4EAC3D7FB92017E76849FE08FDF6" author="Sublette, J. E. &amp; Sasa, M." box="[227,477,609,633]" pageId="4" pageNumber="8" pagination="1 - 60" refId="ref10571" refString="Sublette, J. E. &amp; Sasa, M. 1994. Chironomidae collected in Onchocerciasis endemic areas of Guatemala. - Spixiana Supplement 20: 1 - 60." type="journal article" year="1994">Sublette &amp; Sasa (1994)</bibRefCitation>
recorded larval, pupal and adult
<taxonomicName id="4C4B48DE3D7FB920162468AAFE7FFD16" baseAuthorityName="Johannsen" baseAuthorityYear="1905" box="[288,426,641,665]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="8" phylum="Arthropoda" rank="species" species="lundbeckii">
<emphasis id="B93FEF4F3D7FB920162468AAFE7FFD16" box="[288,426,641,665]" italics="true" pageId="4" pageNumber="8">P. lundbeckii</emphasis>
</taxonomicName>
from
<collectingCountry id="F35C73CD3D7FB92016E968A9FDB1FD16" box="[493,612,641,665]" name="Guatemala" pageId="4" pageNumber="8">Guatemala</collectingCountry>
. I examined
<specimenCount id="9D4DF8D43D7FB92015EA68A9FEE3FD36" pageId="4" pageNumber="8" type="female">4 females</specimenCount>
and
<specimenCount id="9D4DF8D43D7FB9201675688AFE3FFD36" box="[369,490,673,697]" pageId="4" pageNumber="8" type="female">one female</specimenCount>
pupal exuviae from this Guatemalan material (no males were available). None of these specimens could be reliably identified as
<taxonomicName id="4C4B48DE3D7FB920162A692AFE6DFC96" baseAuthorityName="Johannsen" baseAuthorityYear="1905" box="[302,440,769,793]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="8" phylum="Arthropoda" rank="species" species="lundbeckii">
<emphasis id="B93FEF4F3D7FB920162A692AFE6DFC96" box="[302,440,769,793]" italics="true" pageId="4" pageNumber="8">P. lundbeckii</emphasis>
</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF4335D3D7FB92017E76918FD82FBC7" blockId="4.[227,764,816,1096]" pageId="4" pageNumber="8">
<bibRefCitation id="EFDA4EAC3D7FB92017E76918FEB6FCC6" author="Epler, J. H." box="[227,355,816,841]" pageId="4" pageNumber="8" refId="ref8808" refString="Epler, J. H. 2001. Identification Manual for the larval Chironomidae (Diptera) of North and South Carolina. A guide to the taxonomy of the midges of the southeastern United States, including Florida. Special Publication SJ 2001 - SP 13. - North Carolina Department of Environment and Natural Resources, Raleigh, NC, and St. Johns River Water Management District, Palatka, FL. 526 pp." type="book" year="2001">Epler (2001</bibRefCitation>
: 7.117) discussed a
<taxonomicName id="4C4B48DE3D7FB92015356919FD29FCC7" authorityName="Goetghebuer" authorityYear="1931" box="[561,764,817,840]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEF4F3D7FB92015356919FD29FCC7" box="[561,764,817,840]" italics="true" pageId="4" pageNumber="8">Parametriocnemus</emphasis>
</taxonomicName>
from Great Smoky National Park that had genitalia similar to
<taxonomicName id="4C4B48DE3D7FB92016496959FE00FC07" baseAuthorityName="Johannsen" baseAuthorityYear="1905" box="[333,469,880,904]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="8" phylum="Arthropoda" rank="species" species="lundbeckii">
<emphasis id="B93FEF4F3D7FB92016496959FE00FC07" box="[333,469,880,904]" italics="true" pageId="4" pageNumber="8">P. lundbeckii</emphasis>
</taxonomicName>
but had an AR of 0.40. Two other specimens resembling
<taxonomicName id="4C4B48DE3D7FB920151669B9FD4EFC27" baseAuthorityName="Johannsen" baseAuthorityYear="1905" box="[530,667,912,936]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="8" phylum="Arthropoda" rank="species" species="lundbeckii">
<emphasis id="B93FEF4F3D7FB920151669B9FD4EFC27" box="[530,667,912,936]" italics="true" pageId="4" pageNumber="8">P. lundbeckii</emphasis>
</taxonomicName>
from the same area have an AR of around 0.84, and Ive also examined
<taxonomicName id="4C4B48DE3D7FB920168F69F9FDCDFC67" baseAuthorityName="Johannsen" baseAuthorityYear="1905" box="[395,536,976,1000]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="8" phylum="Arthropoda" rank="species" species="lundbeckii">
<emphasis id="B93FEF4F3D7FB920168F69F9FDCDFC67" box="[395,536,976,1000]" italics="true" pageId="4" pageNumber="8">P. lundbeckii</emphasis>
</taxonomicName>
from the same area with “typical” ARs (~1.40). It appears obvious that a thorough study of
<taxonomicName id="4C4B48DE3D7FB92016C16E39FD84FBA7" baseAuthorityName="Johannsen" baseAuthorityYear="1905" box="[453,593,1040,1064]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="8" phylum="Arthropoda" rank="species" species="lundbeckii">
<emphasis id="B93FEF4F3D7FB92016C16E39FD84FBA7" box="[453,593,1040,1064]" italics="true" pageId="4" pageNumber="8">P. lundbeckii</emphasis>
</taxonomicName>
is necessary to determine just what that species is.
</paragraph>
<paragraph id="8BF4335D3D7FB92017E76E48FD9BFAB7" blockId="4.[227,764,1120,1336]" pageId="4" pageNumber="8">
According to my notes,
<emphasis id="B93FEF4F3D7FB92016EC6E48FE28FBF8" box="[488,509,1120,1143]" italics="true" pageId="4" pageNumber="8">P.</emphasis>
sp. ZUR-4 resembles a species I have seen from northwestern
<collectingCountry id="F35C73CD3D7FB92015826EA8FD29FB17" box="[646,764,1152,1176]" name="Costa Rica" pageId="4" pageNumber="8">Costa Rica</collectingCountry>
that I had designated
<emphasis id="B93FEF4F3D7FB92016DD6E88FE3CFB38" box="[473,489,1184,1207]" italics="true" pageId="4" pageNumber="8">P</emphasis>
. sp. CR-3. Those specimens, and other Costa Rican
<taxonomicName id="4C4B48DE3D7FB92015356EE8FEE6FB77" pageId="4" pageNumber="8">
<emphasis id="B93FEF4F3D7FB92015356EE8FD29FB58" box="[561,764,1216,1239]" italics="true" pageId="4" pageNumber="8">Parametriocnemus</emphasis>
species
</taxonomicName>
, are out on loan and were not available for comparison. All of the Zurquí
<taxonomicName id="4C4B48DE3D7FB92015356F28FD29FA98" authorityName="Goetghebuer" authorityYear="1931" box="[561,764,1280,1303]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEF4F3D7FB92015356F28FD29FA98" box="[561,764,1280,1303]" italics="true" pageId="4" pageNumber="8">Parametriocnemus</emphasis>
</taxonomicName>
specimens had an AR below 1.00.
</paragraph>
</subSubSection>
</treatment>
</document>