299 lines
32 KiB
XML
299 lines
32 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.e84187" ID-Pensoft-Pub="2747-8106-96-179" ID-Pensoft-UUID="FE0861D71BB454999EE6637C0D9B2B0C" ID-ZooBank="35B619086E6548B09A177281C2253391" ModsDocID="2747-8106-96-179" checkinTime="1670952513976" checkinUser="pensoft" docAuthor="Gruendel, Joachim, Keupp, Helmut, Lang, Fritz & Nuetzel, Alexander" docDate="2022" docId="03C8F07BEC87525E868F3FA2C68BE3BE" docLanguage="en" docName="Zitteliana 96: 179-221" docOrigin="Zitteliana 96" docPubDate="2022-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.96.e84187" docTitle="Itieroptygmatis cylindrata Gründel & Keupp & Lang & Nützel 2022, sp. nov." docType="treatment" docUuid="05FC3FC9-3FBE-4EE3-8FA8-0B64093E42EA" docUuidSource="ZooBank" docVersion="1" id="FE0861D71BB454999EE6637C0D9B2B0C" lastPageNumber="179" masterDocId="FE0861D71BB454999EE6637C0D9B2B0C" masterDocTitle="Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany)" masterLastPageNumber="221" masterPageNumber="179" pageNumber="179" updateTime="1670952513976" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gruendel, Joachim</mods:namePart>
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<mods:affiliation>Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">joachim.gruendel@lingua-pura.de</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Keupp, Helmut</mods:namePart>
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<mods:affiliation>Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lang, Fritz</mods:namePart>
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<mods:affiliation>Drosselweg 16, 96114 Hirschaid, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nuetzel, Alexander</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8852-7688</mods:nameIdentifier>
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<mods:affiliation>SNSB-Bayerische Staatssammlung fuer Palaeontologie und Geologie, Richard-Wagner-Str. 10, 80333 Muenchen, Germany & Department of Earth and Environmental Sciences, Paleontology and Geobiology, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zitteliana</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>96</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>179</mods:start>
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<mods:end>221</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zitteliana.96.e84187</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zitteliana.96.e84187</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2747-8106-96-179</mods:identifier>
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<mods:identifier type="ZooBank">35B619086E6548B09A177281C2253391</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FE0861D71BB454999EE6637C0D9B2B0C</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:05FC3FC9-3FBE-4EE3-8FA8-0B64093E42EA" httpUri="http://treatment.plazi.org/id/03C8F07BEC87525E868F3FA2C68BE3BE" lastPageNumber="179" pageId="0" pageNumber="179">
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<subSubSection pageId="0" pageNumber="179" type="nomenclature">
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName LSID="https://zoobank.org/05FC3FC9-3FBE-4EE3-8FA8-0B64093E42EA" authority="Gründel & Keupp & Lang & Nützel, 2022" authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" family="Itieriidae" genus="Itieroptygmatis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieroptygmatis cylindrata" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="cylindrata" status="sp. nov.">Itieroptygmatis cylindrata</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="179">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="description">
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<paragraph pageId="0" pageNumber="179">
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<figureCitation captionStart="Plate 18" captionStartId="F18" captionText="Plate 18. (1 - 10) Itieroptygmatis cylindrata sp. nov., morphotype 1. (1 - 2) Holotype, SNSB-BSPG 2021 XV 80, Saal (collection Lang); (1) lateral view, height 15 mm; (2) oblique view, width 5.5 mm (Gruendel 2017, fig. 15 B). (3 - 4) Paratype, SNSB-BSPG 2021 XV 81, Saal (collection Lang), apical and lateral views, width 14 mm. (5 - 6) Paratype, SNSB-BSPG 2021 XV 82, Saal (collection Lang), apical and lateral views, width 16 mm (original Gruendel 2017, fig. 15 C). (7 - 8) Paratype, SNSB-BSPG 2021 XV 83, Saal (collection Lang), basal and lateral views, width 15 mm. (9 - 10) Paratype, SNSB-BSPG 2021 XV 84, Saal (collection Lang); (9) columellar section, height 16 mm; (10) detail of (9), height 7.5 mm. (11) Itieroptygmatis cylindrata sp. nov., transition from morphotype 1 to morphotype 2, paratype, SNSB-BSPG 2021 XV 85, Saal (collection Schaefer), lateral view, height 33 mm. (12 - 14) Itieroptygmatis cylindrata sp. nov., morphotype 2, paratype, SNSB-BSPG 2021 XV 86, Saal (collection Lang), (12) lateral view, height 27 mm; (13) apex in lateral view, height 12 mm; (14) apical, width 13 mm. (15 - 16) Itieroptygmatis sp. 1. (15) SNSB-BSPG 2021 XV 88, Saal (collection Lang), lateral view, height 47 mm. (16) SNSB-BSPG 2021 XV 89, Saal (collection Lang), base, width 21 mm." figureDoi="10.3897/zitteliana.96.e84187.plate18" httpUri="https://binary.pensoft.net/fig/780396" pageId="0" pageNumber="179">Plate 18: figs 1-14</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="reference_group">
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName authority="Gründel & Keupp & Lang & Nützel, 2022" authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" family="Itieriidae" genus="Itieria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieria austriaca" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="austriaca" status="sp. nov.">?1882 - Itieria austriaca</taxonomicName>
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Zitt. - Schlosser: 84, pl. 12, figs 8, 9.
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</paragraph>
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName authority="Gründel & Keupp & Lang & Nützel, 2022" authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" pageId="0" pageNumber="179" rank="superFamily" status="sp. nov." superFamily="Nerineoidea">2017 - Nerineoidea</taxonomicName>
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Nr. 9a, b -
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<normalizedToken originalValue="Gründel">Gruendel</normalizedToken>
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: 33, pl. 15B-C.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="etymology">
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<paragraph pageId="0" pageNumber="179">Derivatio nominis.</paragraph>
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<paragraph pageId="0" pageNumber="179">
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Latin
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<emphasis italics="true" pageId="0" pageNumber="179">cylindratus</emphasis>
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- cylindrical; according to the approximately cylindrical shell shape of the late teleoconch of this species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="holotype">
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<paragraph pageId="0" pageNumber="179">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="179">SNSB-BSPG 2021 XV 80, collection Lang.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="locus typicus">
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<paragraph pageId="0" pageNumber="179">Locus typicus.</paragraph>
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<paragraph pageId="0" pageNumber="179">Saal Quarry near Kelheim.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="stratum typicum">
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<paragraph pageId="0" pageNumber="179">Stratum typicum.</paragraph>
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<paragraph pageId="0" pageNumber="179">Upper Kimmeridgian.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="paratypes">
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<paragraph pageId="0" pageNumber="179">Paratypes.</paragraph>
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<paragraph pageId="0" pageNumber="179">
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45 specimens from Saal (43 specimens collection Lang, BSPG (SNSB-BSPG 2021 XV 81-84, 86, 109-146); one specimen collection Keupp, SNSB-BSPG 2021 XV 87, one specimen collection
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<normalizedToken originalValue="Schäfer">Schaefer</normalizedToken>
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, BSPG SNSB-BSPG 2021 XV 85; additional specimens on block with
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<taxonomicName class="Gastropoda" family="Neritopsidae" genus="Cassianopsis" kingdom="Animalia" lsidName="Cassianopsis quenstedti" order="Cycloneritida" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="quenstedti">
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<emphasis italics="true" pageId="0" pageNumber="179">Cassianopsis quenstedti</emphasis>
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</taxonomicName>
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SNSB-BSPG 2015 VII 58.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="materials_examined">
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<paragraph pageId="0" pageNumber="179">Material.</paragraph>
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<paragraph pageId="0" pageNumber="179">
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A total of
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<specimenCount type="generic">56 specimens</specimenCount>
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from Saal,
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<specimenCount type="generic" typeStatus="types">46 types</specimenCount>
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(see above) and
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<specimenCount type="generic">10 specimens</specimenCount>
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without
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<typeStatus>type</typeStatus>
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status from private collections:
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<specimenCount type="generic">6 specimens</specimenCount>
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collection Lang,
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<specimenCount type="generic">4 specimens</specimenCount>
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collection
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<normalizedToken originalValue="Schäfer">Schaefer</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="diagnosis">
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<paragraph pageId="0" pageNumber="179">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="179">The adapical part of the shell is slender with many whorls lacking a ramp. Later whorls are broad in relation to their height and have a ramp which becomes broader from whorl to whorl in most specimens. Below the ramp, a weak concavity (lateral view) is developed; rarely, the ramp remains narrow and the concavity is lacking. The last whorls of large specimens increase only slowly in width and this part of the shell is more or less cylindrical. The transition from whorl face to base is evenly rounded without edge or spiral cord. Aperture with two columellar and palatal plaits and one parietal plait.</paragraph>
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<caption doi="10.3897/zitteliana.96.e84187.plate18" httpUri="https://binary.pensoft.net/fig/780396" pageId="0" pageNumber="179" start="Plate 18" startId="F18">
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<paragraph pageId="0" pageNumber="179">
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<emphasis bold="true" pageId="0" pageNumber="179">Plate 18.</emphasis>
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<emphasis bold="true" pageId="0" pageNumber="179">(1-10)</emphasis>
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<taxonomicName authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" family="Itieriidae" genus="Itieroptygmatis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieroptygmatis cylindrata" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="cylindrata">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieroptygmatis cylindrata</emphasis>
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</taxonomicName>
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sp. nov., morphotype 1.
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<emphasis bold="true" pageId="0" pageNumber="179">(1-2)</emphasis>
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Holotype, SNSB-BSPG 2021 XV 80, Saal (collection Lang);
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<emphasis bold="true" pageId="0" pageNumber="179">(1)</emphasis>
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lateral view, height 15 mm;
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<emphasis bold="true" pageId="0" pageNumber="179">(2)</emphasis>
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oblique view, width 5.5 mm (
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<bibRefCitation author="Gruendel, J" journalOrPublisher="Der Steinkern" pageId="0" pageNumber="179" pagination="20 - 47" refId="B30" refString="Gruendel, J, 2017. Die Gastropoden-Fauna von Saal bei Kelheim. Der Steinkern 30: 20 - 47" title="Die Gastropoden-Fauna von Saal bei Kelheim." volume="30" year="2017">
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<normalizedToken originalValue="Gründel">Gruendel</normalizedToken>
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2017
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</bibRefCitation>
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, fig. 15 B).
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<emphasis bold="true" pageId="0" pageNumber="179">(3-4)</emphasis>
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Paratype, SNSB-BSPG 2021 XV 81, Saal (collection Lang), apical and lateral views, width 14 mm.
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<emphasis bold="true" pageId="0" pageNumber="179">(5-6)</emphasis>
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Paratype, SNSB-BSPG 2021 XV 82, Saal (collection Lang), apical and lateral views, width 16 mm (original
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<bibRefCitation author="Gruendel, J" journalOrPublisher="Der Steinkern" pageId="0" pageNumber="179" pagination="20 - 47" refId="B30" refString="Gruendel, J, 2017. Die Gastropoden-Fauna von Saal bei Kelheim. Der Steinkern 30: 20 - 47" title="Die Gastropoden-Fauna von Saal bei Kelheim." volume="30" year="2017">
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<normalizedToken originalValue="Gründel">Gruendel</normalizedToken>
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2017
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</bibRefCitation>
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, fig. 15 C).
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<emphasis bold="true" pageId="0" pageNumber="179">(7-8)</emphasis>
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Paratype, SNSB-BSPG 2021 XV 83, Saal (collection Lang), basal and lateral views, width 15 mm.
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<emphasis bold="true" pageId="0" pageNumber="179">(9-10)</emphasis>
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Paratype, SNSB-BSPG 2021 XV 84, Saal (collection Lang);
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<emphasis bold="true" pageId="0" pageNumber="179">(9)</emphasis>
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columellar section, height 16 mm;
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<emphasis bold="true" pageId="0" pageNumber="179">(10)</emphasis>
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detail of (9), height 7.5 mm.
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<emphasis bold="true" pageId="0" pageNumber="179">(11)</emphasis>
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<taxonomicName authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" family="Itieriidae" genus="Itieroptygmatis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieroptygmatis cylindrata" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="cylindrata">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieroptygmatis cylindrata</emphasis>
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</taxonomicName>
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sp. nov., transition from morphotype 1 to morphotype 2, paratype, SNSB-BSPG 2021 XV 85, Saal (collection
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<normalizedToken originalValue="Schäfer">Schaefer</normalizedToken>
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), lateral view, height 33 mm.
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<emphasis bold="true" pageId="0" pageNumber="179">(12-14)</emphasis>
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<taxonomicName authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" family="Itieriidae" genus="Itieroptygmatis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieroptygmatis cylindrata" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="cylindrata">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieroptygmatis cylindrata</emphasis>
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</taxonomicName>
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sp. nov., morphotype 2, paratype, SNSB-BSPG 2021 XV 86, Saal (collection Lang),
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<emphasis bold="true" pageId="0" pageNumber="179">(12)</emphasis>
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lateral view, height 27 mm;
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<emphasis bold="true" pageId="0" pageNumber="179">(13)</emphasis>
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apex in lateral view, height 12 mm;
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<emphasis bold="true" pageId="0" pageNumber="179">(14)</emphasis>
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apical, width 13 mm.
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<emphasis bold="true" pageId="0" pageNumber="179">(15-16)</emphasis>
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<taxonomicName authorityName="Charvet & Termier" authorityYear="1971" class="Gastropoda" family="Itieriidae" genus="Itieroptygmatis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieroptygmatis" pageId="0" pageNumber="179" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieroptygmatis</emphasis>
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</taxonomicName>
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sp. 1.
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<emphasis bold="true" pageId="0" pageNumber="179">(15)</emphasis>
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SNSB-BSPG 2021 XV 88, Saal (collection Lang), lateral view, height 47 mm.
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<emphasis bold="true" pageId="0" pageNumber="179">(16)</emphasis>
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SNSB-BSPG 2021 XV 89, Saal (collection Lang), base, width 21 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="179">This species is present in two morphotypes and specimens that are intermediate:</paragraph>
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<paragraph pageId="0" pageNumber="179">
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Morphotype 1: Plate
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<figureCitation captionStart="Plate 18" captionStartId="F18" captionText="Plate 18. (1 - 10) Itieroptygmatis cylindrata sp. nov., morphotype 1. (1 - 2) Holotype, SNSB-BSPG 2021 XV 80, Saal (collection Lang); (1) lateral view, height 15 mm; (2) oblique view, width 5.5 mm (Gruendel 2017, fig. 15 B). (3 - 4) Paratype, SNSB-BSPG 2021 XV 81, Saal (collection Lang), apical and lateral views, width 14 mm. (5 - 6) Paratype, SNSB-BSPG 2021 XV 82, Saal (collection Lang), apical and lateral views, width 16 mm (original Gruendel 2017, fig. 15 C). (7 - 8) Paratype, SNSB-BSPG 2021 XV 83, Saal (collection Lang), basal and lateral views, width 15 mm. (9 - 10) Paratype, SNSB-BSPG 2021 XV 84, Saal (collection Lang); (9) columellar section, height 16 mm; (10) detail of (9), height 7.5 mm. (11) Itieroptygmatis cylindrata sp. nov., transition from morphotype 1 to morphotype 2, paratype, SNSB-BSPG 2021 XV 85, Saal (collection Schaefer), lateral view, height 33 mm. (12 - 14) Itieroptygmatis cylindrata sp. nov., morphotype 2, paratype, SNSB-BSPG 2021 XV 86, Saal (collection Lang), (12) lateral view, height 27 mm; (13) apex in lateral view, height 12 mm; (14) apical, width 13 mm. (15 - 16) Itieroptygmatis sp. 1. (15) SNSB-BSPG 2021 XV 88, Saal (collection Lang), lateral view, height 47 mm. (16) SNSB-BSPG 2021 XV 89, Saal (collection Lang), base, width 21 mm." figureDoi="10.3897/zitteliana.96.e84187.plate18" httpUri="https://binary.pensoft.net/fig/780396" pageId="0" pageNumber="179">18</figureCitation>
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: figs 1-10;
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</paragraph>
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<paragraph pageId="0" pageNumber="179">
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Transition from morphotype 1 to morphotype 2: Plate
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<figureCitation captionStart="Plate 18" captionStartId="F18" captionText="Plate 18. (1 - 10) Itieroptygmatis cylindrata sp. nov., morphotype 1. (1 - 2) Holotype, SNSB-BSPG 2021 XV 80, Saal (collection Lang); (1) lateral view, height 15 mm; (2) oblique view, width 5.5 mm (Gruendel 2017, fig. 15 B). (3 - 4) Paratype, SNSB-BSPG 2021 XV 81, Saal (collection Lang), apical and lateral views, width 14 mm. (5 - 6) Paratype, SNSB-BSPG 2021 XV 82, Saal (collection Lang), apical and lateral views, width 16 mm (original Gruendel 2017, fig. 15 C). (7 - 8) Paratype, SNSB-BSPG 2021 XV 83, Saal (collection Lang), basal and lateral views, width 15 mm. (9 - 10) Paratype, SNSB-BSPG 2021 XV 84, Saal (collection Lang); (9) columellar section, height 16 mm; (10) detail of (9), height 7.5 mm. (11) Itieroptygmatis cylindrata sp. nov., transition from morphotype 1 to morphotype 2, paratype, SNSB-BSPG 2021 XV 85, Saal (collection Schaefer), lateral view, height 33 mm. (12 - 14) Itieroptygmatis cylindrata sp. nov., morphotype 2, paratype, SNSB-BSPG 2021 XV 86, Saal (collection Lang), (12) lateral view, height 27 mm; (13) apex in lateral view, height 12 mm; (14) apical, width 13 mm. (15 - 16) Itieroptygmatis sp. 1. (15) SNSB-BSPG 2021 XV 88, Saal (collection Lang), lateral view, height 47 mm. (16) SNSB-BSPG 2021 XV 89, Saal (collection Lang), base, width 21 mm." figureDoi="10.3897/zitteliana.96.e84187.plate18" httpUri="https://binary.pensoft.net/fig/780396" pageId="0" pageNumber="179">18</figureCitation>
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: fig. 11;
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</paragraph>
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<paragraph pageId="0" pageNumber="179">
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Morphotype 2: Plate
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<figureCitation captionStart="Plate 18" captionStartId="F18" captionText="Plate 18. (1 - 10) Itieroptygmatis cylindrata sp. nov., morphotype 1. (1 - 2) Holotype, SNSB-BSPG 2021 XV 80, Saal (collection Lang); (1) lateral view, height 15 mm; (2) oblique view, width 5.5 mm (Gruendel 2017, fig. 15 B). (3 - 4) Paratype, SNSB-BSPG 2021 XV 81, Saal (collection Lang), apical and lateral views, width 14 mm. (5 - 6) Paratype, SNSB-BSPG 2021 XV 82, Saal (collection Lang), apical and lateral views, width 16 mm (original Gruendel 2017, fig. 15 C). (7 - 8) Paratype, SNSB-BSPG 2021 XV 83, Saal (collection Lang), basal and lateral views, width 15 mm. (9 - 10) Paratype, SNSB-BSPG 2021 XV 84, Saal (collection Lang); (9) columellar section, height 16 mm; (10) detail of (9), height 7.5 mm. (11) Itieroptygmatis cylindrata sp. nov., transition from morphotype 1 to morphotype 2, paratype, SNSB-BSPG 2021 XV 85, Saal (collection Schaefer), lateral view, height 33 mm. (12 - 14) Itieroptygmatis cylindrata sp. nov., morphotype 2, paratype, SNSB-BSPG 2021 XV 86, Saal (collection Lang), (12) lateral view, height 27 mm; (13) apex in lateral view, height 12 mm; (14) apical, width 13 mm. (15 - 16) Itieroptygmatis sp. 1. (15) SNSB-BSPG 2021 XV 88, Saal (collection Lang), lateral view, height 47 mm. (16) SNSB-BSPG 2021 XV 89, Saal (collection Lang), base, width 21 mm." figureDoi="10.3897/zitteliana.96.e84187.plate18" httpUri="https://binary.pensoft.net/fig/780396" pageId="0" pageNumber="179">18</figureCitation>
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: figs 12-14.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="description">
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<paragraph pageId="0" pageNumber="179">Description.</paragraph>
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<paragraph pageId="0" pageNumber="179">A large specimen (early whorl missing) is 33 mm high. The early shell is very slender consisting of at least eight smooth whorls having a weakly convex whorl face and somewhat impressed sutures. The following 2-3 whorls increase rapidly in width and are very wide in relation to their height. As a result, the spire is coeloconoid. In the mentioned 2-3 whorls following the slender spire whorls, a broad, distinctly concave, funnel-shaped ramp is formed. From now on, the ontogenetic evolution of the shell follows different pathways leading to two varieties (morphotype 1 and 2 as well as to intermediate forms).</paragraph>
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<paragraph pageId="0" pageNumber="179">In morphotype 1 (much more abundant than morphotype 2), the wide ramp continues and becomes wider. It is delimited from whorl face by a broad concavity with indistinct borders. Initially, the whorls continue to rapidly increase in width. In large, more or less fully grown specimens, the increase in width decelerates, the last whorls are high and almost cylindrical in shape. The edge of the ramp is projecting abaxially and forms the whorl periphery.</paragraph>
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<paragraph pageId="0" pageNumber="179">
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In the other extreme form, morphotype 2, the rapid increase of the width of the whorls and the formation of a broad ramp is confined to 2-3 whorls. Afterwards, the whorls become wider and increase in width only slowly, but considerably increase in height. The ramp forms only a narrow band. The whorl face is straight and lacks a concavity below the ramp so that this part of the shell is almost cylindrical. As mentioned, there are transitional forms between varieties 1 and 2 (Plate
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<figureCitation captionStart="Plate 18" captionStartId="F18" captionText="Plate 18. (1 - 10) Itieroptygmatis cylindrata sp. nov., morphotype 1. (1 - 2) Holotype, SNSB-BSPG 2021 XV 80, Saal (collection Lang); (1) lateral view, height 15 mm; (2) oblique view, width 5.5 mm (Gruendel 2017, fig. 15 B). (3 - 4) Paratype, SNSB-BSPG 2021 XV 81, Saal (collection Lang), apical and lateral views, width 14 mm. (5 - 6) Paratype, SNSB-BSPG 2021 XV 82, Saal (collection Lang), apical and lateral views, width 16 mm (original Gruendel 2017, fig. 15 C). (7 - 8) Paratype, SNSB-BSPG 2021 XV 83, Saal (collection Lang), basal and lateral views, width 15 mm. (9 - 10) Paratype, SNSB-BSPG 2021 XV 84, Saal (collection Lang); (9) columellar section, height 16 mm; (10) detail of (9), height 7.5 mm. (11) Itieroptygmatis cylindrata sp. nov., transition from morphotype 1 to morphotype 2, paratype, SNSB-BSPG 2021 XV 85, Saal (collection Schaefer), lateral view, height 33 mm. (12 - 14) Itieroptygmatis cylindrata sp. nov., morphotype 2, paratype, SNSB-BSPG 2021 XV 86, Saal (collection Lang), (12) lateral view, height 27 mm; (13) apex in lateral view, height 12 mm; (14) apical, width 13 mm. (15 - 16) Itieroptygmatis sp. 1. (15) SNSB-BSPG 2021 XV 88, Saal (collection Lang), lateral view, height 47 mm. (16) SNSB-BSPG 2021 XV 89, Saal (collection Lang), base, width 21 mm." figureDoi="10.3897/zitteliana.96.e84187.plate18" httpUri="https://binary.pensoft.net/fig/780396" pageId="0" pageNumber="179">18</figureCitation>
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: fig. 11).
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</paragraph>
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<paragraph pageId="0" pageNumber="179">All individuals have a continuous, rounded transition from whorl face to the strongly convex base. Base and whorl face are smooth. The base has a distinct umbilicus. The growth lines run straight and somewhat opisthocline on the whorl face and curve strongly backward below the ramp. The aperture is very narrow, adapically acute and has a weakly developed siphonal canal. The inner lip is broadened and detached in the columellar area; it partly covers the umbilicus. The inner lip bears a strong parietal and two columellar plaits. The adapical columellar plait is weaker than the abapical one. The aperture has two palatal plaits which are, however, rarely recognizable.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="remarks">
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<paragraph pageId="0" pageNumber="179">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" family="Itieriidae" genus="Itieria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieria austriaca" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="austriaca">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieria austriaca</emphasis>
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</taxonomicName>
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Zittel as described by
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<bibRefCitation author="Schlosser, M" journalOrPublisher="Palaeontographica" pageId="0" pageNumber="179" pagination="41 - 110" refId="B88" refString="Schlosser, M, 1882. Die Fauna des Kelheimer Diceras-Kalkes. I. Vertebrata, Crustacea, Cephalopoda und Gastropoda. Palaeontographica 28: 41 - 110" title="Die Fauna des Kelheimer Diceras-Kalkes. I. Vertebrata, Crustacea, Cephalopoda und Gastropoda." volume="28" year="1882">Schlosser (1882)</bibRefCitation>
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closely resembles morphotype 2 of
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<taxonomicName authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" family="Itieriidae" genus="Itieroptygmatis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieroptygmatis cylindrata" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="cylindrata">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieroptygmatis cylindrata</emphasis>
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</taxonomicName>
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sp. nov. However, it has a distinctly lower last whorl in relation to its spire height. The real
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<taxonomicName authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" family="Itieriidae" genus="Itieria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieria austriaca" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="austriaca">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieria austriaca</emphasis>
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</taxonomicName>
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Zittel, 1873 differs distinctly from
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<taxonomicName authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" family="Itieriidae" genus="Itieroptygmatis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieroptygmatis cylindrata" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="cylindrata">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieroptygmatis cylindrata</emphasis>
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</taxonomicName>
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sp. nov. by lacking a ramp and by having a subsutural row of knobs.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="relationships">
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<paragraph pageId="0" pageNumber="179">Relationships.</paragraph>
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName class="Gastropoda" family="Itieriidae" genus="Itieria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieria (Campichia) pellati" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="pellati" subGenus="Campichia">Itieria (Campichia) pellati</taxonomicName>
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Cossmann and
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<taxonomicName class="Gastropoda" family="Itieriidae" genus="Itieria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieria (Campichia) truncata" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="truncata" subGenus="Campichia">Itieria (Campichia) truncata</taxonomicName>
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Pictet and Campiche, both sensu
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<bibRefCitation author="Cossmann, M" journalOrPublisher="The author, Paris" pageId="0" pageNumber="179" refId="B13" refString="Cossmann, M, 1916. Essais de paleoconchologie comparee 10. The author, Paris" title="Essais de paleoconchologie comparee 10." year="1916">Cossmann (1916)</bibRefCitation>
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, resemble morphotype 1 of
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<taxonomicName authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" family="Itieriidae" genus="Itieroptygmatis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieroptygmatis cylindrata" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="cylindrata">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieroptygmatis cylindrata</emphasis>
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</taxonomicName>
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sp. nov. However, their spire is lower and has not as many whorls, the last whorl is much broader than the spire (having wide ramp only on a single whorl).
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<taxonomicName class="Gastropoda" family="Itieriidae" genus="Itieria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieria obtusiceps" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="obtusiceps">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieria obtusiceps</emphasis>
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</taxonomicName>
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Zittel, 1873 resembles morphotype 2 of
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<taxonomicName class="Gastropoda" family="Itieriidae" genus="Itieroptygmatis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieroptygmatis cylindrata." pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="cylindrata.">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieroptygmatis cylindrata.</emphasis>
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</taxonomicName>
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However,
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<taxonomicName lsidName="I. obtusiceps" pageId="0" pageNumber="179" rank="species" species="obtusiceps">
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<emphasis italics="true" pageId="0" pageNumber="179">I. obtusiceps</emphasis>
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</taxonomicName>
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has a less distinct ramp, the spire is blunter, and the shell outline is more oval-shaped.
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<taxonomicName class="Gastropoda" family="Nerineidae" genus="Phaneroptyxis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phaneroptyxis nogreti" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nogreti">
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<emphasis italics="true" pageId="0" pageNumber="179">Phaneroptyxis nogreti</emphasis>
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</taxonomicName>
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Guir. and
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<normalizedToken originalValue="Ogérien">Ogerien</normalizedToken>
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sensu
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<bibRefCitation author="Cossmann, M" journalOrPublisher="The author and Comptoir Geologique, Paris" pageId="0" pageNumber="179" refId="B12" refString="Cossmann, M, 1898. Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1-179." title="Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1 - 179." year="1898">Cossmann (1898)</bibRefCitation>
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differs from morphotype 1 of
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<taxonomicName authorityName="Gründel & Keupp & Lang & Nützel" authorityYear="2022" class="Gastropoda" family="Itieriidae" genus="Itieroptygmatis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itieroptygmatis cylindrata" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="cylindrata">
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<emphasis italics="true" pageId="0" pageNumber="179">Itieroptygmatis cylindrata</emphasis>
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</taxonomicName>
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sp. nov. by having a lower last whorl (in relation to spire height) in large specimens, lacking a wide ramp, and not having a cylindrical shape of the last whorl.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |