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<document id="A69868A9F716F87F67EBFD4028A1D1C8" ID-DOI="10.11646/zootaxa.3636.2.8" ID-GBIF-Dataset="343f524f-f0f7-4ffc-8eea-6db65a9626c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="223767" ID-ZooBank="84E31B3F-9F0F-47CF-A5EF-ADBA8255C66C" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460589225437" checkinUser="plazi" docAuthor="Mendoza, Jose Christopher E." docDate="2013" docId="03C887FAFFDBC724FF344625DADEFB8D" docLanguage="en" docName="zt03636p384.pdf" docOrigin="Zootaxa 3636 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Xanthias joanneae Mendoza, 2013, sp. nov." docType="treatment" docUuid="B372E200-EE6E-4D94-8960-985BA07CD452" docUuidSource="ZooBank" docVersion="9" lastPageNumber="383" masterDocId="FFF1FF82FFDAC72DFFA34519D853FFF1" masterDocTitle="A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines" masterLastPageNumber="384" masterPageNumber="374" pageNumber="375" updateTime="1698325619238" updateUser="plazi">
<mods:mods id="A52F5D9BB682EE164802F2672E3BB606" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="32FEE4EF36DCD59F3ED873AF935B7D4D">
<mods:title id="22BCD46888B818A1D2F8E845A5ED2087">A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines</mods:title>
</mods:titleInfo>
<mods:name id="6561F6D25797620164B261BB642D6A4D" type="personal">
<mods:role id="2C525546EB9527D9C247ACE744937AA2">
<mods:roleTerm id="1622E828BF96D1334F5636C00FE1D744">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3C996B69B50D1A8425C467DEFF29B781">Mendoza, Jose Christopher E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="73C631E829D9161763A1FD98FEC058E8">text</mods:typeOfResource>
<mods:relatedItem id="6B3DBBF497CCE7B90EC013EA4E7FD041" type="host">
<mods:titleInfo id="ABE38554F0DF255693F3079CB6C7F8A1">
<mods:title id="9DD7F75B9E1F9862ED3914593123D894">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DB57ABDEE2E3F631E74496A73286139A">
<mods:date id="16B9361868CA07A3D41C25F1BC79C48A">2013</mods:date>
<mods:detail id="C2D776DCE492A2BE35D64F1EF9BA6B80" type="volume">
<mods:number id="7E80199516D7CB25919994567218707D">3636</mods:number>
</mods:detail>
<mods:detail id="FCD237F6E7314882A86E7AFBD09C19D9" type="issue">
<mods:number id="C5874A62FB580447C88DDCF7CE46BC20">2</mods:number>
</mods:detail>
<mods:extent id="B1A97B60F7368A74C72C66DD12DE570E" unit="page">
<mods:start id="7A242CB186191B2F78012885C4B8C9B6">374</mods:start>
<mods:end id="0738FCD4685CD56324F3621D0F5F460D">384</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="70BDE40E044DA4C09DBB43E02080B266">journal article</mods:classification>
<mods:identifier id="C8B7104D6421C7BCF4622AE0F9973593" type="DOI">10.11646/zootaxa.3636.2.8</mods:identifier>
<mods:identifier id="DAB3FF631E2BF479E0E30199FC2E40BA" type="GBIF-Dataset">343f524f-f0f7-4ffc-8eea-6db65a9626c1</mods:identifier>
<mods:identifier id="03EAF78E5FC35DB81365CF3C45B00819" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="291E6639367D1B3071C12CEBC726823D" type="Zenodo-Dep">223767</mods:identifier>
<mods:identifier id="98ABDD683EA7C2F15A5B4C4F8457B36A" type="ZooBank">84E31B3F-9F0F-47CF-A5EF-ADBA8255C66C</mods:identifier>
</mods:mods>
<treatment id="03C887FAFFDBC724FF344625DADEFB8D" ID-DOI="http://doi.org/10.5281/zenodo.6150383" ID-GBIF-Taxon="119565458" ID-Zenodo-Dep="6150383" LSID="urn:lsid:zoobank.org:act:B372E200-EE6E-4D94-8960-985BA07CD452" httpUri="http://treatment.plazi.org/id/03C887FAFFDBC724FF344625DADEFB8D" lastPageId="9" lastPageNumber="383" pageId="1" pageNumber="375">
<subSubSection id="C37B6567FFDBC72CFF344625D8ACFC87" pageId="1" pageNumber="375" type="nomenclature">
<paragraph id="8BDE36ECFFDBC72CFF344625D98CFCA4" blockId="1.[151,479,827,886]" box="[151,479,827,854]" pageId="1" pageNumber="375">
<heading id="D0968180FFDBC72CFF344625D98CFCA4" bold="true" box="[151,479,827,854]" fontSize="11" level="1" pageId="1" pageNumber="375" reason="1">
<emphasis id="B915EAFEFFDBC72CFF344625D98CFCA4" bold="true" box="[151,479,827,854]" pageId="1" pageNumber="375">
<taxonomicName id="4C614D6FFFDBC72CFF344625D929FCA7" ID-CoL="5C4TN" box="[151,378,828,854]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="375" phylum="Arthropoda" rank="species" species="joanneae" status="sp. nov.">
<emphasis id="B915EAFEFFDBC72CFF344625D929FCA7" bold="true" box="[151,378,828,854]" italics="true" pageId="1" pageNumber="375">Xanthias joanneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2265785FFDBC72CFE224622D98CFCA4" box="[385,479,827,853]" pageId="1" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDE36ECFFDBC72CFF344647D8ACFC87" blockId="1.[151,479,827,886]" box="[151,255,862,886]" pageId="1" pageNumber="375">
<figureCitation id="135A2A69FFDBC72CFF344647D8ACFC87" box="[151,255,862,886]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1895,1918]" captionStartId-1="3.[151,250,1968,1991]" captionStartId-2="4.[151,250,1907,1930]" captionStartId-3="5.[151,250,1920,1943]" captionTargetBox-0="[151,1435,193,1873]" captionTargetBox-1="[292,1282,193,1921]" captionTargetBox-2="[151,1435,193,1886]" captionTargetBox-3="[179,1383,193,1862]" captionTargetId-0="figure@2.[151,1436,193,1874]" captionTargetId-1="figure@3.[292,1283,193,1921]" captionTargetId-2="figure@4.[151,1436,193,1886]" captionTargetId-3="figure@5.[171,1393,193,1873]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Xanthias joanneae sp. nov., live coloration. A, holotype, male, 7.9 × 5.1 mm (NMCR 39080); B, paratype, female, 9.4 × 6.0 mm (NMCR 39081). Photos by (A) Peter K. L. Ng / Tan Swee-Hee (NUS); (B) Chan Tin-Yam / Lin Chia-Wei (NTOU)." captionText-1="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." captionText-2="FIGURE 3. Xanthias joanneae sp. nov., paratype, female, 9.4 × 6.0 mm (NMCR 39081), Pamilacan Is., Bohol Sea (A – C); holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea (D). A, habitus, dorsal view; B, habitus, anterior view; C, D, left chela, external view." captionText-3="FIGURE 4. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080). A, left third maxilliped, external view; B, anterior thoracic sternum, ventral view; C, carpus of right cheliped, dorso-external view; D, abdomen, ventral view; E, right P 5, dorsal view; left G 1 (F, G, I, J), F, external view; G, internal view; I, distal tip, external view; J, distal tip, internal view; H, left G 2, external view. Scale bars: A – E = 1.0 mm; F – H = 0.5 mm; I, J = 0.25 mm." httpUri-0="https://zenodo.org/record/223768/files/figure.png" httpUri-1="https://zenodo.org/record/223769/files/figure.png" httpUri-2="https://zenodo.org/record/223770/files/figure.png" httpUri-3="https://zenodo.org/record/223771/files/figure.png" pageId="1" pageNumber="375">Figs. 1–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B6567FFDBC72CFF3446B8DC66FA08" pageId="1" pageNumber="375" type="materials_examined">
<paragraph id="8BDE36ECFFDBC72CFF3446B8DB41FC2C" blockId="1.[151,1436,929,2033]" pageId="1" pageNumber="375">
<emphasis id="B915EAFEFFDBC72CFF3446B8D9D7FC4B" bold="true" box="[151,388,929,954]" pageId="1" pageNumber="375">Material examined.</emphasis>
<typeStatus id="54DA884EFFDBC72CFE2C46B8D9A9FC48" box="[399,506,929,953]" pageId="1" pageNumber="375" type="holotype">Holotype</typeStatus>
, male, 7.9 ×
<quantity id="4C999B09FFDBC72CFD3846B8DAA6FC48" box="[667,757,929,953]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="375" unit="mm" value="5.1">5.1 mm</quantity>
(NMCR 39080), stn B5, 4 m, reef slope with overhangs, Biking, Panglao Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFD9E46DCDA9FFC2C" box="[573,716,965,989]" pageId="1" pageNumber="375" value="2004-06-02">2004, 2 June</date>
2004.
</paragraph>
<paragraph id="8BDE36ECFFDBC72CFF6446F3DC66FA08" blockId="1.[151,1436,929,2033]" pageId="1" pageNumber="375">
<typeStatus id="54DA884EFFDBC72CFF6446F3D96BFBF3" box="[199,312,1002,1026]" pageId="1" pageNumber="375" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D67FD65FFDBC72CFEEA46F0D9E0FBF0" box="[329,435,1001,1025]" pageId="1" pageNumber="375" type="female">1 female</specimenCount>
, 9.4 × 6.0 mm (NMCR 39081), stn B11, 2–
<quantity id="4C999B09FFDBC72CFC7746F0DC55FBF0" box="[980,1030,1001,1025]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="375" unit="m" value="4.0">4 m</quantity>
, coral rubble, Pamilacan Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFE804114D991FBD4" box="[291,450,1037,1061]" pageId="1" pageNumber="375" value="2004-06-11">2004, 11 June</date>
2004;
<specimenCount id="9D67FD65FFDBC72CFDB74114DA29FBD4" box="[532,634,1037,1061]" pageId="1" pageNumber="375" type="female">1 female</specimenCount>
, 4.1 ×
<quantity id="4C999B09FFDBC72CFD6F4114DB77FBD4" box="[716,804,1037,1061]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="375" unit="mm" value="2.8">2.8 mm</quantity>
(MNHN-IU-2010-8062), stn B24, 38 m, floor of cave, Pamilacan Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFDA04128DAF1FBB8" box="[515,674,1073,1097]" pageId="1" pageNumber="375" value="2004-06-25">2004, 25 June</date>
2004;
<specimenCount id="9D67FD65FFDBC72CFD514128DB11FBB8" box="[754,834,1073,1097]" pageId="1" pageNumber="375" type="male">1 male</specimenCount>
, 4.8 ×
<quantity id="4C999B09FFDBC72CFC314128DBB8FBB8" box="[914,1003,1073,1097]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="375" unit="mm" value="3.2">3.2 mm</quantity>
,
<specimenCount id="9D67FD65FFDBC72CFC5B4128DC0DFBB8" box="[1016,1118,1073,1097]" pageId="1" pageNumber="375" type="female">1 female</specimenCount>
, 5.9 ×
<quantity id="4C999B09FFDBC72CFB0D4128DD56FBB8" box="[1198,1285,1073,1097]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="375" unit="mm" value="3.8">3.8 mm</quantity>
(MNHN-IU- 2010-8061), stn B41, 17–
<quantity id="4C999B09FFDBC72CFE15414CD9BCFB9C" box="[438,495,1109,1133]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="375" unit="m" value="19.0">19 m</quantity>
, floor of large cave, Balicasag Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFB90414CDCEEFB9C" box="[1075,1213,1109,1133]" pageId="1" pageNumber="375" value="2004-07-04">2004, 4 July</date>
2004;
<specimenCount id="9D67FD65FFDBC72CFAA9414CDD04FB9C" box="[1290,1367,1109,1133]" pageId="1" pageNumber="375" type="male">1 male</specimenCount>
, 6.3 ×
<quantity id="4C999B09FFDBC72CFF344160D8BEFB60" box="[151,237,1145,1169]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="375" unit="mm" value="4.2">4.2 mm</quantity>
(MNHN-IU-2010-8063), stn R38, 6–
<quantity id="4C999B09FFDBC72CFD304160DA9EFB60" box="[659,717,1145,1169]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="375" unit="m" value="37.0">37 m</quantity>
, reef slope, Pamilacan Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFB184160DD06FB60" box="[1211,1365,1145,1169]" pageId="1" pageNumber="375" value="2004-06-11">2004, 11 June</date>
2004;
<specimenCount id="9D67FD65FFDBC72CFF344184D8A9FB44" box="[151,250,1181,1205]" pageId="1" pageNumber="375" type="female">1 female</specimenCount>
, 8.9 ×
<quantity id="4C999B09FFDBC72CFEE64184D9C9FB44" box="[325,410,1181,1205]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="375" unit="mm" value="5.5">5.5 mm</quantity>
(ZRC 2013.0433), stn B3, 8 m, base of reef slope, Arco Point, Panglao Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFF3441D8D968FB2B" box="[151,315,1217,1242]" pageId="1" pageNumber="375" value="2004-05-31">2004, 31 May</date>
2004;
<specimenCount id="9D67FD65FFDBC72CFE3141D8D9B7FB28" box="[402,484,1217,1241]" pageId="1" pageNumber="375" type="male">1 male</specimenCount>
, 4.5 × 3.0 mm (ZRC 2013.0434), stn B8, 3 m, subtidal reef platform, Napaling, Panglao Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFE4741FCDA27FB0C" box="[484,628,1253,1277]" pageId="1" pageNumber="375" value="2004-06-07">2004, 7 June</date>
2004;
<specimenCount id="9D67FD65FFDBC72CFD6241FCDB5CFB0C" box="[705,783,1253,1277]" pageId="1" pageNumber="375" type="male">1 male</specimenCount>
, 7.1 ×
<quantity id="4C999B09FFDBC72CFCFF41FCDBE7FB0C" box="[860,948,1253,1277]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="375" unit="mm" value="4.8">4.8 mm</quantity>
,
<specimenCount id="9D67FD65FFDBC72CFC6341FCDC77FB0C" box="[960,1060,1253,1277]" pageId="1" pageNumber="375" type="female">1 female</specimenCount>
, 6.1 ×
<quantity id="4C999B09FFDBC72CFBD241FCDC94FB0C" box="[1137,1223,1253,1277]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="375" unit="mm" value="4.5">4.5 mm</quantity>
(ZRC 2013.0435), stn B10, 3–
<quantity id="4C999B09FFDBC72CFEBF4010D90BFAD0" box="[284,344,1289,1313]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="375" unit="m" value="14.0">14 m</quantity>
, reef wall with small caves, Momo Beach, Panglao Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFB014010DD6EFAD0" box="[1186,1341,1289,1313]" pageId="1" pageNumber="375" value="2004-06-10">2004, 10 Jun.</date>
2004;
<specimenCount id="9D67FD65FFDBC72CFA2D4010D883FAB4" pageId="1" pageNumber="375" type="male">1 male</specimenCount>
, 7.4 ×
<quantity id="4C999B09FFDBC72CFE804034D928FAB4" box="[291,379,1325,1349]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="1" pageNumber="375" unit="mm" value="4.9">4.9 mm</quantity>
(ZRC 2013.0436), stn B12, 24–
<quantity id="4C999B09FFDBC72CFD534034DB7EFAB4" box="[752,813,1325,1349]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="375" unit="m" value="27.0">27 m</quantity>
, reef slope, Doljo Point, Panglao Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFF344048D967FA98" box="[151,308,1361,1385]" pageId="1" pageNumber="375" value="2004-06-14">2004, 14 June</date>
2004;
<specimenCount id="9D67FD65FFDBC72CFE204048D996FA98" box="[387,453,1361,1385]" pageId="1" pageNumber="375" type="generic">2 juv.</specimenCount>
, 3.1 ×
<quantity id="4C999B09FFDBC72CFDB14048DA38FA98" box="[530,619,1361,1385]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="375" unit="mm" value="2.2">2.2 mm</quantity>
, 3.8 ×
<quantity id="4C999B09FFDBC72CFD1B4048DB5CFA98" box="[696,783,1361,1385]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="375" unit="mm" value="2.6">2.6 mm</quantity>
(ZRC 2013.0437), stn B16, 20 m, coral rubble on sand &amp; gravel, Bingag, Panglao Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFD3C406CDB6DFA7C" box="[671,830,1397,1421]" pageId="1" pageNumber="375" value="2004-06-17">2004, 17 June</date>
2004;
<specimenCount id="9D67FD65FFDBC72CFC2D406CDBA7FA7C" box="[910,1012,1397,1421]" pageId="1" pageNumber="375" type="female">1 female</specimenCount>
, 6.4 ×
<quantity id="4C999B09FFDBC72CFBE7406CDCC8FA7C" box="[1092,1179,1397,1421]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="375" unit="mm" value="4.2">4.2 mm</quantity>
(ZRC 2013.0438), stn L41, 90–
<quantity id="4C999B09FFDBC72CFF5E4080D915FA40" box="[253,326,1433,1457]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="375" unit="m" value="100.0">100 m</quantity>
, front of PTA compound, Panglao Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFC184080DC15FA40" box="[955,1094,1433,1457]" pageId="1" pageNumber="375" value="2004-07-01">2004, 1 July</date>
2004;
<specimenCount id="9D67FD65FFDBC72CFB374080DCAAFA40" box="[1172,1273,1433,1457]" pageId="1" pageNumber="375" type="female">1 female</specimenCount>
, 5.6 ×
<quantity id="4C999B09FFDBC72CFAE54080DDCFFA40" box="[1350,1436,1433,1457]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="375" unit="mm" value="3.8">3.8 mm</quantity>
(ZRC 2013.0439), stn L43, 60 m, Pamilacan Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFCD840A4DC57FA24" box="[891,1028,1469,1493]" pageId="1" pageNumber="375" value="2004-07-02">2004, 2 July</date>
2004;
<specimenCount id="9D67FD65FFDBC72CFBF240A4DCE6FA24" box="[1105,1205,1469,1493]" pageId="1" pageNumber="375" type="female">1 female</specimenCount>
, 5.3 ×
<quantity id="4C999B09FFDBC72CFAA340A4DD06FA24" box="[1280,1365,1469,1493]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="375" unit="mm" value="3.5">3.5 mm</quantity>
(ZRC 2013.0440), stn L46, 90–
<quantity id="4C999B09FFDBC72CFE1040F8D9A9FA08" box="[435,506,1505,1529]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="375" unit="m" value="110.0">110 m</quantity>
, Balicasag Is., coll. PANGLAO
<date id="FFDF102CFFDBC72CFCC640F8DBBCFA08" box="[869,1007,1505,1529]" pageId="1" pageNumber="375" value="2004-07-04">2004, 4 July</date>
2004.
</paragraph>
</subSubSection>
<subSubSection id="C37B6567FFDBC72AFF64431CDB1FFF25" lastPageId="7" lastPageNumber="381" pageId="1" pageNumber="375" type="description">
<paragraph id="8BDE36ECFFDBC72CFF64431CD8A9F800" blockId="1.[151,1436,929,2033]" pageId="1" pageNumber="375">
<emphasis id="B915EAFEFFDBC72CFF64431CD90AF9EF" bold="true" box="[199,345,1541,1566]" pageId="1" pageNumber="375">Description.</emphasis>
Carapace (
<figureCitation id="135A2A69FFDBC72CFE43431CDA61F9EC" box="[480,562,1541,1565]" captionStart="FIGURE 1" captionStartId="2.[151,250,1895,1918]" captionTargetBox="[151,1435,193,1873]" captionTargetId="figure@2.[151,1436,193,1874]" captionTargetPageId="2" captionText="FIGURE 1. Xanthias joanneae sp. nov., live coloration. A, holotype, male, 7.9 × 5.1 mm (NMCR 39080); B, paratype, female, 9.4 × 6.0 mm (NMCR 39081). Photos by (A) Peter K. L. Ng / Tan Swee-Hee (NUS); (B) Chan Tin-Yam / Lin Chia-Wei (NTOU)." httpUri="https://zenodo.org/record/223768/files/figure.png" pageId="1" pageNumber="375">Figs. 1</figureCitation>
,
<figureCitation id="135A2A69FFDBC72CFDE6431CDA00F9EC" box="[581,595,1541,1565]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="1" pageNumber="375">2</figureCitation>
A, 3A) transversely ovate, subhexagonal, width about 1.5 times length, dorsal area slightly convex transversely, longitudinally; dorsal surface smooth, glabrous, regions mostly poorly defined, except orbital region, with mesial, posterior limit delineated by deep groove, anterior portions of 2M, 3M feebly indicated; subhepatic, pterygostomian regions smooth, glabrous. Front weakly advanced, broad, about 0.4 times carapace width, slightly deflexed ventrally; separated into two lobes by wide V-shaped notch, continuing posteriorly as shallow median groove on dorsal carapace, as median suture reaching tip of proepistome ventrally; anterior margin of frontal lobes gently convex in dorsal view. Anterolateral margin convex, divided into 4 low, lobiform teeth, separated from each other by distinct V-shaped notches; first tooth low, barely indicated; second, widest, low but more projecting than first, anterior margin much shorter than posterior; third, narrower at base but more projecting than preceding, anterior margin much shorter than posterior; fourth, similar to third but much smaller, posterior margin continuous with carapace posterolateral margin; maximum carapace width across tips of third teeth, short transverse groove between second, third teeth, continuing onto branchial region. Posterolateral margin longer than anterolateral margin, straight to slightly convex. Posterior margin with central region slightly concave.
</paragraph>
<caption id="DF1E6664FFD8C72FFF34427EDDCAF86D" httpUri="https://zenodo.org/record/223768/files/figure.png" pageId="2" pageNumber="376" targetBox="[151,1435,193,1873]" targetPageId="2">
<paragraph id="8BDE36ECFFD8C72FFF34427EDDCAF86D" blockId="2.[151,1436,1895,1948]" pageId="2" pageNumber="376">
<emphasis id="B915EAFEFFD8C72FFF34427ED941F88D" bold="true" box="[151,274,1895,1918]" pageId="2" pageNumber="376">FIGURE 1.</emphasis>
<taxonomicName id="4C614D6FFFD8C72FFEBB427ED986F88C" box="[280,469,1895,1917]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="376" phylum="Arthropoda" rank="species" species="joanneae" status="sp. nov.">
<emphasis id="B915EAFEFFD8C72FFEBB427ED986F88C" box="[280,469,1895,1917]" italics="true" pageId="2" pageNumber="376">Xanthias joanneae</emphasis>
</taxonomicName>
<emphasis id="B915EAFEFFD8C72FFE784271DA62F88C" bold="true" box="[475,561,1896,1917]" pageId="2" pageNumber="376">
<taxonomicNameLabel id="A2265785FFD8C72FFE784271DA7EF88C" box="[475,557,1896,1917]" pageId="2" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
live coloration. A, holotype, male, 7.9 × 5.1 mm (NMCR 39080); B, paratype, female, 9.4 × 6.0 mm (NMCR 39081). Photos by (A) Peter K.L. Ng/Tan Swee-Hee (NUS); (B) Chan Tin-Yam/Lin Chia-Wei (NTOU).
</paragraph>
</caption>
<caption id="DF1E6664FFD9C72EFF3442A9DA3FF814" httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="3" pageNumber="377" targetBox="[292,1282,193,1921]" targetPageId="3">
<paragraph id="8BDE36ECFFD9C72EFF3442A9DA3FF814" blockId="3.[151,1436,1968,2021]" pageId="3" pageNumber="377">
<emphasis id="B915EAFEFFD9C72EFF3442A9D941F837" bold="true" box="[151,274,1968,1991]" pageId="3" pageNumber="377">FIGURE 2.</emphasis>
<taxonomicName id="4C614D6FFFD9C72EFEBB42A8D986F836" box="[280,469,1969,1991]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="377" phylum="Arthropoda" rank="species" species="joanneae" status="sp. nov.">
<emphasis id="B915EAFEFFD9C72EFEBB42A8D986F836" box="[280,469,1969,1991]" italics="true" pageId="3" pageNumber="377">Xanthias joanneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2265785FFD9C72EFE7842A8DA78F837" box="[475,555,1969,1990]" pageId="3" pageNumber="377" rank="species">
<emphasis id="B915EAFEFFD9C72EFE7842A8DA78F837" bold="true" box="[475,555,1969,1990]" pageId="3" pageNumber="377">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view.
</paragraph>
</caption>
<caption id="DF1E6664FFDEC729FF34426AD9FCF836" httpUri="https://zenodo.org/record/223770/files/figure.png" pageId="4" pageNumber="378" targetBox="[151,1435,193,1886]" targetPageId="4">
<paragraph id="8BDE36ECFFDEC729FF34426AD9FCF836" blockId="4.[151,1436,1907,1991]" pageId="4" pageNumber="378">
<emphasis id="B915EAFEFFDEC729FF34426AD946F878" bold="true" box="[151,277,1907,1930]" pageId="4" pageNumber="378">FIGURE 3.</emphasis>
<taxonomicName id="4C614D6FFFDEC729FEBE426DD98FF87B" box="[285,476,1908,1930]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="378" phylum="Arthropoda" rank="species" species="joanneae" status="sp. nov.">
<emphasis id="B915EAFEFFDEC729FEBE426DD98FF87B" box="[285,476,1908,1930]" italics="true" pageId="4" pageNumber="378">Xanthias joanneae</emphasis>
</taxonomicName>
<emphasis id="B915EAFEFFDEC729FE47426DDA64F878" bold="true" box="[484,567,1908,1929]" pageId="4" pageNumber="378">
<taxonomicNameLabel id="A2265785FFDEC729FE47426DDA64F878" box="[484,567,1908,1929]" pageId="4" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, female, 9.4 × 6.0 mm (NMCR 39081), Pamilacan Is., Bohol Sea (A–C); holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea (D). A, habitus, dorsal view; B, habitus, anterior view; C, D, left chela, external view.
</paragraph>
</caption>
<caption id="DF1E6664FFDFC728FF344299DB86F803" httpUri="https://zenodo.org/record/223771/files/figure.png" pageId="5" pageNumber="379" targetBox="[179,1383,193,1862]" targetPageId="5">
<paragraph id="8BDE36ECFFDFC728FF344299DB86F803" blockId="5.[151,1436,1920,2034]" pageId="5" pageNumber="379">
<emphasis id="B915EAFEFFDFC728FF344299D941F864" bold="true" box="[151,274,1920,1943]" pageId="5" pageNumber="379">FIGURE 4.</emphasis>
<taxonomicName id="4C614D6FFFDFC728FEBB4299D986F867" box="[280,469,1920,1942]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="joanneae" status="sp. nov.">
<emphasis id="B915EAFEFFDFC728FEBB4299D986F867" box="[280,469,1920,1942]" italics="true" pageId="5" pageNumber="379">Xanthias joanneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2265785FFDFC728FE7F4298DA7FF867" box="[476,556,1921,1942]" pageId="5" pageNumber="379" rank="species">
<emphasis id="B915EAFEFFDFC728FE7F4298DA7FF867" bold="true" box="[476,556,1921,1942]" pageId="5" pageNumber="379">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, male, 7.9 × 5.1 mm (NMCR 39080). A, left third maxilliped, external view; B, anterior thoracic sternum, ventral view; C, carpus of right cheliped, dorso-external view; D, abdomen, ventral view; E, right P5, dorsal view; left G1 (F, G, I, J), F, external view; G, internal view; I, distal tip, external view; J, distal tip, internal view; H, left G2, external view. Scale bars: A–E = 1.0 mm; F–H = 0.5 mm; I, J = 0.25 mm.
</paragraph>
</caption>
<paragraph id="8BDE36ECFFDCC72BFF64458EDB2FFE76" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="380">
Orbits (
<figureCitation id="135A2A69FFDCC72BFE87458ED929FF5E" box="[292,378,151,175]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="6" pageNumber="380">Figs. 2</figureCitation>
C, 3B) suboval, margins subcristate; supraorbital margin with 2 small notches, smooth; infraorbital margin finely granular, infraorbital tooth bluntly triangular, apex rounded. Eyes well developed, eyestalks short, stout, corneas large. Antennular fossae transversely oval; antennules folding transversely. Basal article of antenna short, subrectangular; flagellum freely entering orbital hiatus, short, tip not reaching exorbital angle. Epistome broad, lateral regions more produced posteriorly than central region, regions separated by deep notches; posterior margin with broad median triangular prominence on central region, convex on lateral regions. Endostome smooth, with distinct, oblique ridges on posterior half.
</paragraph>
<paragraph id="8BDE36ECFFDCC72BFF64448DDBE0FDCD" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="380">
Third maxillipeds (
<figureCitation id="135A2A69FFDCC72BFE0C448DDA55FE5D" box="[431,518,404,428]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="6" pageNumber="380">Figs. 2</figureCitation>
B, 4A) finely granular; palp tapering distally, articles subcylindrical; merus subquadrate, anterolateral angle rounded, slightly projecting, anterior margin gently convex except for slight depression mesially; ischium about twice as long as merus, sulcate submedially, mesial margin cristate, lined with submarginal setae; exopod stout, margins straight, tapering slightly toward distal end, with subdistal triangular projection beneath mesial margin, flagellum as long as width of merus.
</paragraph>
<paragraph id="8BDE36ECFFDCC72BFF64475EDC32FCAD" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="380">
Male thoracic sternum (
<figureCitation id="135A2A69FFDCC72BFE70475EDA72FDAE" box="[467,545,583,607]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="6" pageNumber="380">Figs. 2</figureCitation>
B, 4B) finely granular; sternites 1, 2 fused, but boundary between them marked by row of setae; deep suture between sternites 2, 3; sternites 3, 4 mostly fused, with remnants of suture only at lateral edges continued mesially by deep transverse groove; sternite 4 with prominent median longitudinal line extending through most of its exposed length, lateral margins mostly straight; median longitudinal line interrupted on sternite 4 within anterior region of sternoabdominal cavity, continued at posterior limit of sternite 4, absent in sternites 5, 6, continued in sternites 7, 8 without interruption; sutures 4/5, 5/6 joining towards median longitudinal axis of thoracic sternum; small, central, triangular, non-calcified area present between sternites 6, 7 within sternoabdominal cavity; press-button on anterior half of sternite 5, closer to suture 4/5.
</paragraph>
<paragraph id="8BDE36ECFFDCC72BFF64467EDBE1FBA6" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="380">
Chelipeds (
<figureCitation id="135A2A69FFDCC72BFEEA467ED9C9FC8E" box="[329,410,871,895]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="6" pageNumber="380">Figs. 2</figureCitation>
A, B, 3D, 4C) symmetrical, moderately robust, external surfaces generally smooth. Meri short, distal end not extending beyond carapace anterolateral margin in dorsal view. Carpi with rugose external surface because of deep, curved groove; internal angle with blunt, triangular projection. Palm smooth, with shallow but distinct groove on dorso-external surface only. Fingers thick, each with longitudinal, submarginal groove, both pigmented throughout most of length except at hemi-cupuliform tips, pigment extending well into external, internal surfaces of palm, cutting margins with variably sized teeth; dactylus curved, distinctly longer than superior margin of palm, with ocellus near proximal end; fixed finger slightly deflexed.
</paragraph>
<paragraph id="8BDE36ECFFDCC72BFF64417DDA75FA89" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="380">
Ambulatory legs (
<figureCitation id="135A2A69FFDCC72BFE3A417DD9B9FB8D" box="[409,490,1124,1148]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="6" pageNumber="380">Figs. 2</figureCitation>
A, 3A, 4E) flattened, broad, moderate in length, P3, P4 longest, coxa-to-dactylus length about equal to carapace width, P5 shortest, coxa-to-dactylus length about 0.9 times carapace width; anterior margins of meri, carpi, propodi sharp but not distinctly cristate. Meri subrectangular, median length about 2.2–2.4 times greatest width; anterior margin gently convex throughout length, posterior margin convex proximally, concave distally. Carpi curved, distal end distinctly wider than proximal end. Propodi subrectangular, median length about 1.9–2.1 times greatest width; anterior, posterior margins convex, posterior margin denticulate, lined with stiff setae. Dactyli about as long as propodi, stout proximally, tapering distally, terminating in sharp chitinous claw; margins lined with stiff setae.
</paragraph>
<paragraph id="8BDE36ECFFDCC72BFF64409DDB3EF9DD" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="380">
Abdomen (
<figureCitation id="135A2A69FFDCC72BFEEA409DD9CFFA6D" box="[329,412,1412,1436]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="6" pageNumber="380">Figs. 2</figureCitation>
B, 4D) long relative to thoracic sternum, tip of telson reaching level anterior to sternal condyles of P1 coxae; somites 1, 2 trapezoidal, much wider than long; somites 3–5 fused, base of somite 3 widest, distal somites progressively narrower, combined lateral margin gently concave; somite 6 subquadrate, slightly wider than long, lateral margins straight; telson subtriangular, angles rounded, basal width about 1.3 times median length, base distinctly wider than adjacent distal end of somite 6.
</paragraph>
<paragraph id="8BDE36ECFFDCC72BFF644321DD39F969" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="380">
G1 (
<figureCitation id="135A2A69FFDCC72BFF594321D96DF9A1" box="[250,318,1592,1616]" captionStart="FIGURE 4" captionStartId="5.[151,250,1920,1943]" captionTargetBox="[179,1383,193,1862]" captionTargetId="figure@5.[171,1393,193,1873]" captionTargetPageId="5" captionText="FIGURE 4. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080). A, left third maxilliped, external view; B, anterior thoracic sternum, ventral view; C, carpus of right cheliped, dorso-external view; D, abdomen, ventral view; E, right P 5, dorsal view; left G 1 (F, G, I, J), F, external view; G, internal view; I, distal tip, external view; J, distal tip, internal view; H, left G 2, external view. Scale bars: A – E = 1.0 mm; F – H = 0.5 mm; I, J = 0.25 mm." httpUri="https://zenodo.org/record/223771/files/figure.png" pageId="6" pageNumber="380">Fig. 4</figureCitation>
F, G, I, J) moderate in length, stout, distal half curving laterally; distal tip with narrow, semi-fluted apical lobe, few spiniform granules immediately proximal to it, several (14) plumose subterminal setae. G2 (
<figureCitation id="135A2A69FFDCC72BFACC4345D8F6F969" captionStart="FIGURE 4" captionStartId="5.[151,250,1920,1943]" captionTargetBox="[179,1383,193,1862]" captionTargetId="figure@5.[171,1393,193,1873]" captionTargetPageId="5" captionText="FIGURE 4. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080). A, left third maxilliped, external view; B, anterior thoracic sternum, ventral view; C, carpus of right cheliped, dorso-external view; D, abdomen, ventral view; E, right P 5, dorsal view; left G 1 (F, G, I, J), F, external view; G, internal view; I, distal tip, external view; J, distal tip, internal view; H, left G 2, external view. Scale bars: A – E = 1.0 mm; F – H = 0.5 mm; I, J = 0.25 mm." httpUri="https://zenodo.org/record/223771/files/figure.png" pageId="6" pageNumber="380">Fig. 4</figureCitation>
H) sigmoidal in shape, about one-third length of G1. Penis emerging near base of sternal condyle of P5 coxa.
</paragraph>
<paragraph id="8BDE36ECFFDCC72BFF6443BDD885F89E" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="380">
<emphasis id="B915EAFEFFDCC72BFF6443BDD9EFF94D" bold="true" box="[199,444,1700,1724]" pageId="6" pageNumber="380">Female morphology.</emphasis>
The female (
<figureCitation id="135A2A69FFDCC72BFDF743BDDACCF94D" box="[596,671,1700,1724]" captionStart="FIGURE 1" captionStartId="2.[151,250,1895,1918]" captionTargetBox="[151,1435,193,1873]" captionTargetId="figure@2.[151,1436,193,1874]" captionTargetPageId="2" captionText="FIGURE 1. Xanthias joanneae sp. nov., live coloration. A, holotype, male, 7.9 × 5.1 mm (NMCR 39080); B, paratype, female, 9.4 × 6.0 mm (NMCR 39081). Photos by (A) Peter K. L. Ng / Tan Swee-Hee (NUS); (B) Chan Tin-Yam / Lin Chia-Wei (NTOU)." httpUri="https://zenodo.org/record/223768/files/figure.png" pageId="6" pageNumber="380">Figs. 1</figureCitation>
B, 3A–C) is similar in morphology to the male except in the sexual characters and chelipeds. The abdomen is generally wider and oval in shape, with all the somites freely articulated. The sternoabdominal cavity is also wider but shallower, and the median longitudinal line is of a similar pattern as in the male. The vulvae are oval, without an operculum but with a vulvar cover, and located on the mesial third of sternite 6. The chelipeds are less robust than in the male, and the pigment on the fingers does not extend into the palm.
</paragraph>
<paragraph id="8BDE36ECFFDCC72AFF644262DB1FFF25" blockId="6.[151,1437,151,2012]" lastBlockId="7.[151,1436,151,284]" lastPageId="7" lastPageNumber="381" pageId="6" pageNumber="380">
<emphasis id="B915EAFEFFDCC72BFF644262D9DBF865" bold="true" box="[199,392,1915,1940]" pageId="6" pageNumber="380">Live coloration.</emphasis>
Yellow-orange base (
<figureCitation id="135A2A69FFDCC72BFD2A4265DA9DF865" box="[649,718,1916,1940]" captionStart="FIGURE 1" captionStartId="2.[151,250,1895,1918]" captionTargetBox="[151,1435,193,1873]" captionTargetId="figure@2.[151,1436,193,1874]" captionTargetPageId="2" captionText="FIGURE 1. Xanthias joanneae sp. nov., live coloration. A, holotype, male, 7.9 × 5.1 mm (NMCR 39080); B, paratype, female, 9.4 × 6.0 mm (NMCR 39081). Photos by (A) Peter K. L. Ng / Tan Swee-Hee (NUS); (B) Chan Tin-Yam / Lin Chia-Wei (NTOU)." httpUri="https://zenodo.org/record/223768/files/figure.png" pageId="6" pageNumber="380">Fig. 1</figureCitation>
), with several ocelli, bluish-white spots ringed with maroon. About 40 or more ocelli on the dorsal surface of carapace and fewer on the external surfaces of the chelipeds. Ambulatory legs with 2–4 ocelli each on the dorsal surfaces of meri, carpi and propodi and white bands at the joints. Third maxillipeds, thoracic sternum and abdomen much lighter in color than the dorsal surfaces, but also decorated with ocelli. Eyes with one ocellus on each eyestalk.
</paragraph>
</subSubSection>
<subSubSection id="C37B6567FFDDC72AFF6445F9DA8DFEED" pageId="7" pageNumber="381" type="etymology">
<paragraph id="8BDE36ECFFDDC72AFF6445F9DA8DFEED" blockId="7.[151,1436,151,284]" pageId="7" pageNumber="381">
<emphasis id="B915EAFEFFDDC72AFF6445F9D91DFF09" bold="true" box="[199,334,224,248]" pageId="7" pageNumber="381">Etymology.</emphasis>
This strikingly colored species is named after the author’s wife, Joanne (née Uy), who has been constantly supportive of his carcinological research.
</paragraph>
</subSubSection>
<caption id="DF1E6664FFDDC72AFF34428EDB33F83C" httpUri="https://zenodo.org/record/223772/files/figure.png" pageId="7" pageNumber="381" targetBox="[151,1435,337,1921]" targetPageId="7">
<paragraph id="8BDE36ECFFDDC72AFF34428EDB33F83C" blockId="7.[151,1436,1943,1997]" pageId="7" pageNumber="381">
<emphasis id="B915EAFEFFDDC72AFF34428ED946F85C" bold="true" box="[151,277,1943,1966]" pageId="7" pageNumber="381">FIGURE 5.</emphasis>
<taxonomicName id="4C614D6FFFDDC72AFEBC4281DA22F85F" authority="Sakai, 1961" authorityName="Sakai" authorityYear="1961" box="[287,625,1944,1966]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="381" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFDDC72AFEBC4281D9B9F85F" box="[287,490,1944,1966]" italics="true" pageId="7" pageNumber="381">Xanthias maculatus</emphasis>
Sakai, 1961
</taxonomicName>
, holotype, male, 9.5 × 6.0 mm (NMST), Okinoyama, Sagami Bay, Japan. A, dorsal view; B, ventral view. Photos from Hironori Komatsu (NMST).
</paragraph>
</caption>
<subSubSection id="C37B6567FFD2C724FF644581DADEFB8D" lastPageId="9" lastPageNumber="383" pageId="8" pageNumber="382" type="discussion">
<paragraph id="8BDE36ECFFD2C725FF644581D943FDE6" blockId="8.[151,1437,151,2012]" pageId="8" pageNumber="382">
<emphasis id="B915EAFEFFD2C725FF644581D968FF41" bold="true" box="[199,315,152,176]" pageId="8" pageNumber="382">Remarks.</emphasis>
<taxonomicName id="4C614D6FFFD2C725FEE54581DAECFF5E" authority="Sakai, 1961" authorityName="Sakai" authorityYear="1961" box="[326,703,151,176]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FEE54581DA7BFF41" box="[326,552,152,176]" italics="true" pageId="8" pageNumber="382">Xanthias maculatus</emphasis>
Sakai, 1961
</taxonomicName>
, was originally described from two male specimens, with one being designated as the
<typeStatus id="54DA884EFFD2C725FE0E45A5DA41FF25" box="[429,530,188,212]" pageId="8" pageNumber="382" type="holotype">holotype</typeStatus>
, whereas the other was not explicitly called a
<typeStatus id="54DA884EFFD2C725FB8D45A4DCDCFF24" box="[1070,1167,189,213]" pageId="8" pageNumber="382" type="paratype">paratype</typeStatus>
though it was listed as material examined in the describing paper. The
<typeStatus id="54DA884EFFD2C725FD6245C6DB70FF06" box="[705,803,223,247]" pageId="8" pageNumber="382" type="holotype">holotype</typeStatus>
(9.5 × 6.0 mm) was collected by the then Emperor of
<collectingCountry id="F376767CFFD2C725FF34441CD884FEED" box="[151,215,261,284]" name="Japan" pageId="8" pageNumber="382">Japan</collectingCountry>
from Okinoyama, near Jogashima Island in Sagami Bay, Kanagawa Prefecture, at a depth of
<specimenCount id="9D67FD65FFD2C725FAB1441DDD06FEED" box="[1298,1365,260,284]" pageId="8" pageNumber="382" type="generic">
<quantity id="4C999B09FFD2C725FAB1441DDD03FEED" box="[1298,1360,260,284]" metricMagnitude="1" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="382" unit="m" value="85.0">85 m</quantity>
.
</specimenCount>
Sakai (1961) also mentioned that the other male was collected from Tosa Bay, in Shikoku farther to the southwest. This species’ most easily observed distinguishing character is the coloration of the live specimens, consisting of a yellow carapace and pereopods, decorated with several ocelli (purplish-blue spots ringed with maroon) (Sakai 1961: 141, fig. 1e–g; 1965: 142, pl. 71 fig. 1). It has since been reported from other parts of
<collectingCountry id="F376767CFFD2C725FB07448CDCBBFE5D" box="[1188,1256,405,428]" name="Japan" pageId="8" pageNumber="382">Japan</collectingCountry>
: Kii Nagashima and Ishigaki Island (Sakai 1976: 429, pl. 154 fig. 3); Nii-jima (Takeda 1978: 77, tab. 1); Shiono-misaki (Takeda 1979: 153, tab. 1); Nanki (Miyake 1983: 113, pl. 38 fig. 3); and Amami-Oshima (Takeda &amp; Komatsu 2005: 282, fig.5A, B).
</paragraph>
<paragraph id="8BDE36ECFFD2C725FF64473CDDD1FC1D" blockId="8.[151,1437,151,2012]" pageId="8" pageNumber="382">
It should be noted that
<taxonomicName id="4C614D6FFFD2C725FE6B473CDA09FDCD" box="[456,602,548,572]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FE6B473CDA09FDCD" box="[456,602,548,572]" italics="true" pageId="8" pageNumber="382">X. maculatus</emphasis>
</taxonomicName>
is not well known. The description by Sakai (1961) is not detailed enough for comparisons with current species, and there have not been any published photographs of the
<typeStatus id="54DA884EFFD2C725FB6B475EDD7DFDAE" box="[1224,1326,583,607]" pageId="8" pageNumber="382" type="holotype">holotype</typeStatus>
. The only figures available until recently were line drawings of the entire animal and its G1 (Sakai 1961: fig. 1e–g) as well as a color painting of the entire animal in dorsal view (presumably the
<typeStatus id="54DA884EFFD2C725FC394796DBADFD56" box="[922,1022,655,679]" pageId="8" pageNumber="382" type="holotype">holotype</typeStatus>
) (Sakai 1965: pl. 71 fig. 1; 1976: pl. 154 fig. 3). Examination of the photographs of the
<typeStatus id="54DA884EFFD2C725FD6E47ADDB7CFD3D" box="[717,815,692,716]" pageId="8" pageNumber="382" type="holotype">holotype</typeStatus>
(
<figureCitation id="135A2A69FFD2C725FC9E47ADDB2CFD3D" box="[829,895,692,716]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1966]" captionTargetBox="[151,1435,337,1921]" captionTargetId="figure@7.[151,1436,337,1922]" captionTargetPageId="7" captionText="FIGURE 5. Xanthias maculatus Sakai, 1961, holotype, male, 9.5 × 6.0 mm (NMST), Okinoyama, Sagami Bay, Japan. A, dorsal view; B, ventral view. Photos from Hironori Komatsu (NMST)." httpUri="https://zenodo.org/record/223772/files/figure.png" pageId="8" pageNumber="382">Fig. 5</figureCitation>
) shows that there are some inaccuracies in these early figures, particularly in the proportions of the ambulatory legs (i.e., the
<typeStatus id="54DA884EFFD2C725FBAD47CEDC23FD1E" box="[1038,1136,727,751]" pageId="8" pageNumber="382" type="holotype">holotype</typeStatus>
has stouter legs than was previously depicted). Miyake (1983) provided a short description and the first published color photograph for this species, with a specimen (12.6 ×
<quantity id="4C999B09FFD2C725FDB24606DA39FCC6" box="[529,618,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="8" pageNumber="382" unit="mm" value="8.3">8.3 mm</quantity>
) collected from a depth of
<quantity id="4C999B09FFD2C725FC3C4606DC50FCC9" box="[927,1027,799,824]" metricMagnitude="1" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="8" pageNumber="382" unit="m" value="65.0" valueMax="70.0" valueMin="60.0">60–70 m</quantity>
in Nanki (). This specimen (Miyake1983: pl. 38 fig. 3) compares favorably with the earlier figures by Sakai (1961, 1965, 1976), with the same number and location of the ocelli, and similar carapace and pereopod dimensions as the
<typeStatus id="54DA884EFFD2C725FB1E467EDD71FC8E" box="[1213,1314,871,895]" pageId="8" pageNumber="382" type="holotype">holotype</typeStatus>
. Although Miyake (1983) reports this specimen as a male, the characteristic pigmentation of the cheliped fingers, which extends into the external and internal surfaces of the palm, is not observed, indicating that the specimen may be a female instead. There is no doubt, however, that this specimen and the
<typeStatus id="54DA884EFFD2C725FC1246CDDC40FC1D" box="[945,1043,980,1004]" pageId="8" pageNumber="382" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C614D6FFFD2C725FB9B46CCDC99FC1D" box="[1080,1226,980,1004]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FB9B46CCDC99FC1D" box="[1080,1226,980,1004]" italics="true" pageId="8" pageNumber="382">X. maculatus</emphasis>
</taxonomicName>
are conspecific.
</paragraph>
<paragraph id="8BDE36ECFFD2C725FF6446EEDA92FA89" blockId="8.[151,1437,151,2012]" pageId="8" pageNumber="382">
The photographs published by Takeda &amp; Komatsu (2005: fig. 5A, B) of two specimens collected near Amami- Oshima, in the northern Ryukyu archipelago, one of an ovigerous female (10.7 ×
<quantity id="4C999B09FFD2C725FB954105DCC3FBC5" box="[1078,1168,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="8" pageNumber="382" unit="mm" value="6.9">6.9 mm</quantity>
; NMST-Cr 16194), and another, a smaller male (no data), exhibit considerable variation with each other and from the earlier published figures of this species. Takeda &amp; Komatsu (2005: 282, 283), however, gave no detailed description of these specimens, only remarking on the variable size and number of the “spots” (ocelli) on the carapace, pereopods and thoracic sternum, and raising the possibility that there were more than one species involved. It is noteworthy to mention that the female from Amami-Oshima (Takeda &amp; Komatsu 2005: fig. 5A) closely resembles the present new species, although it has not been examined for this study. It is highly likely, therefore, that there are at least two species in the territorial waters of
<collectingCountry id="F376767CFFD2C725FDB14000DA01FAC1" box="[530,594,1305,1328]" name="Japan" pageId="8" pageNumber="382">Japan</collectingCountry>
that are being referred to as “
<taxonomicName id="4C614D6FFFD2C725FC3E4001DC28FAC1" box="[925,1147,1304,1328]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FC3E4001DC28FAC1" box="[925,1147,1304,1328]" italics="true" pageId="8" pageNumber="382">Xanthias maculatus</emphasis>
</taxonomicName>
”. To avoid confusion, the following comparisons to
<taxonomicName id="4C614D6FFFD2C725FE1C4024DA01FAA5" box="[447,594,1340,1364]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FE1C4024DA01FAA5" box="[447,594,1340,1364]" italics="true" pageId="8" pageNumber="382">X. maculatus</emphasis>
</taxonomicName>
in this paper refer to the
<typeStatus id="54DA884EFFD2C725FCD04025DB86FAA5" box="[883,981,1340,1364]" pageId="8" pageNumber="382" type="holotype">holotype</typeStatus>
(
<figureCitation id="135A2A69FFD2C725FC454025DC7AFAA5" box="[998,1065,1340,1364]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1966]" captionTargetBox="[151,1435,337,1921]" captionTargetId="figure@7.[151,1436,337,1922]" captionTargetPageId="7" captionText="FIGURE 5. Xanthias maculatus Sakai, 1961, holotype, male, 9.5 × 6.0 mm (NMST), Okinoyama, Sagami Bay, Japan. A, dorsal view; B, ventral view. Photos from Hironori Komatsu (NMST)." httpUri="https://zenodo.org/record/223772/files/figure.png" pageId="8" pageNumber="382">Fig. 5</figureCitation>
) and also, where relevant, to the specimen figured by Miyake (1983: pl. 38 fig. 3).
</paragraph>
<paragraph id="8BDE36ECFFD2C724FF64409DDD4EFF06" blockId="8.[151,1437,151,2012]" lastBlockId="9.[151,1436,151,1149]" lastPageId="9" lastPageNumber="383" pageId="8" pageNumber="382">
The new species collected from the Bohol Sea is most similar to
<taxonomicName id="4C614D6FFFD2C725FC3C409DDD58FA6D" authority="Sakai, 1961" authorityName="Sakai" authorityYear="1961" box="[927,1291,1412,1436]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FC3C409DDC2EFA6D" box="[927,1149,1412,1436]" italics="true" pageId="8" pageNumber="382">Xanthias maculatus</emphasis>
Sakai, 1961
</taxonomicName>
, primarily in having a smooth carapace and pereopods, and having several ocelli decorating the carapace and pereopods (
<figureCitation id="135A2A69FFD2C725FAF140B1DDC5FA31" box="[1362,1430,1448,1472]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1966]" captionTargetBox="[151,1435,337,1921]" captionTargetId="figure@7.[151,1436,337,1922]" captionTargetPageId="7" captionText="FIGURE 5. Xanthias maculatus Sakai, 1961, holotype, male, 9.5 × 6.0 mm (NMST), Okinoyama, Sagami Bay, Japan. A, dorsal view; B, ventral view. Photos from Hironori Komatsu (NMST)." httpUri="https://zenodo.org/record/223772/files/figure.png" pageId="8" pageNumber="382">Fig. 5</figureCitation>
; also Sakai 1961: 141, fig. 1e–g; 1965: 142, pl. 71 fig. 1; 1976: 429, pl. 154 fig. 3).
<taxonomicName id="4C614D6FFFD2C725FBFE40D5DD62FA15" box="[1117,1329,1484,1508]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="joanneae" status="sp. nov.">
<emphasis id="B915EAFEFFD2C725FBFE40D5DD62FA15" box="[1117,1329,1484,1508]" italics="true" pageId="8" pageNumber="382">Xanthias joanneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2265785FFD2C725FA9A40D5DDC6FA15" box="[1337,1429,1484,1508]" pageId="8" pageNumber="382" rank="species">
<emphasis id="B915EAFEFFD2C725FA9A40D5DDC6FA15" bold="true" box="[1337,1429,1484,1508]" pageId="8" pageNumber="382">sp. nov.</emphasis>
</taxonomicNameLabel>
, however, differs from
<taxonomicName id="4C614D6FFFD2C725FE2C40E8DA72F9F9" box="[399,545,1520,1544]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FE2C40E8DA72F9F9" box="[399,545,1520,1544]" italics="true" pageId="8" pageNumber="382">X. maculatus</emphasis>
</taxonomicName>
in these aspects: 1) though roughly similar in size, there are more ocelli, at least 40, on the dorsal surface of the carapace (
<figureCitation id="135A2A69FFD2C725FDDB430DDAEBF9DD" box="[632,696,1556,1580]" captionStart="FIGURE 1" captionStartId="2.[151,250,1895,1918]" captionTargetBox="[151,1435,193,1873]" captionTargetId="figure@2.[151,1436,193,1874]" captionTargetPageId="2" captionText="FIGURE 1. Xanthias joanneae sp. nov., live coloration. A, holotype, male, 7.9 × 5.1 mm (NMCR 39080); B, paratype, female, 9.4 × 6.0 mm (NMCR 39081). Photos by (A) Peter K. L. Ng / Tan Swee-Hee (NUS); (B) Chan Tin-Yam / Lin Chia-Wei (NTOU)." httpUri="https://zenodo.org/record/223768/files/figure.png" pageId="8" pageNumber="382">Fig. 1</figureCitation>
A, B) (in contrast to fewer, about 14, in
<taxonomicName id="4C614D6FFFD2C725FB2F430CDD73F9DD" box="[1164,1312,1556,1580]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FB2F430CDD73F9DD" box="[1164,1312,1556,1580]" italics="true" pageId="8" pageNumber="382">X. maculatus</emphasis>
</taxonomicName>
; see Sakai 1961: fig. 1e; 1965: pl. 71 fig. 1; 1976: pl. 154 fig. 3; Miyake 1983: pl. 38 fig. 3); 2) the dorsal carapace regions (especially 2M, 3M) are not well demarcated (
<figureCitation id="135A2A69FFD2C725FD3D4345DAB8F985" box="[670,747,1628,1652]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="8" pageNumber="382">Figs. 2</figureCitation>
A, 3A) (regions easily discernible, particularly 2M and 3M in
<taxonomicName id="4C614D6FFFD2C725FF344398D97EF969" box="[151,301,1664,1688]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FF344398D97EF969" box="[151,301,1664,1688]" italics="true" pageId="8" pageNumber="382">X. maculatus</emphasis>
</taxonomicName>
;
<figureCitation id="135A2A69FFD2C725FE9C4399D9D6F969" box="[319,389,1664,1688]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1966]" captionTargetBox="[151,1435,337,1921]" captionTargetId="figure@7.[151,1436,337,1922]" captionTargetPageId="7" captionText="FIGURE 5. Xanthias maculatus Sakai, 1961, holotype, male, 9.5 × 6.0 mm (NMST), Okinoyama, Sagami Bay, Japan. A, dorsal view; B, ventral view. Photos from Hironori Komatsu (NMST)." httpUri="https://zenodo.org/record/223772/files/figure.png" pageId="8" pageNumber="382">Fig. 5</figureCitation>
A, also Sakai 1961: fig. 1e; Miyake 1983: pl. 38 fig. 3); 3) the teeth on the anterolateral margins of the carapace have more rounded apices, and are closer together, separated by narrow, V-shaped notches (
<figureCitation id="135A2A69FFD2C725FF3C43D1D8A0F911" box="[159,243,1736,1760]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="8" pageNumber="382">Figs. 2</figureCitation>
A, 3A) (teeth with more acute apices, and farther apart, as crests separated by wide troughs, in
<taxonomicName id="4C614D6FFFD2C725FA2743D0D959F8F5" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FA2743D0D959F8F5" italics="true" pageId="8" pageNumber="382">X. maculatus</emphasis>
</taxonomicName>
;
<figureCitation id="135A2A69FFD2C725FEB843F5D933F8F5" box="[283,352,1772,1796]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1966]" captionTargetBox="[151,1435,337,1921]" captionTargetId="figure@7.[151,1436,337,1922]" captionTargetPageId="7" captionText="FIGURE 5. Xanthias maculatus Sakai, 1961, holotype, male, 9.5 × 6.0 mm (NMST), Okinoyama, Sagami Bay, Japan. A, dorsal view; B, ventral view. Photos from Hironori Komatsu (NMST)." httpUri="https://zenodo.org/record/223772/files/figure.png" pageId="8" pageNumber="382">Fig. 5</figureCitation>
A); 4) sternite 4 is narrower and has mostly straight lateral margins (
<figureCitation id="135A2A69FFD2C725FB2243F5DC81F8F5" box="[1153,1234,1772,1796]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="8" pageNumber="382">Figs. 2</figureCitation>
B, 4B) (sternite 4 wider, with convex lateral margins in
<taxonomicName id="4C614D6FFFD2C725FDE54208DA8AF8D9" box="[582,729,1808,1832]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FDE54208DA8AF8D9" box="[582,729,1808,1832]" italics="true" pageId="8" pageNumber="382">X. maculatus</emphasis>
</taxonomicName>
;
<figureCitation id="135A2A69FFD2C725FD494209DB7EF8D9" box="[746,813,1808,1832]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1966]" captionTargetBox="[151,1435,337,1921]" captionTargetId="figure@7.[151,1436,337,1922]" captionTargetPageId="7" captionText="FIGURE 5. Xanthias maculatus Sakai, 1961, holotype, male, 9.5 × 6.0 mm (NMST), Okinoyama, Sagami Bay, Japan. A, dorsal view; B, ventral view. Photos from Hironori Komatsu (NMST)." httpUri="https://zenodo.org/record/223772/files/figure.png" pageId="8" pageNumber="382">Fig. 5</figureCitation>
B); 5) the external surface of the palm of the chelipeds does not have any longitudinal ridges (
<figureCitation id="135A2A69FFD2C725FDF1422DDAF1F8BD" box="[594,674,1844,1868]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="8" pageNumber="382">Figs. 2</figureCitation>
B, 3C, D) (palm external surface has two longitudinal ridges on its lower half in
<taxonomicName id="4C614D6FFFD2C725FE8F4240D9ECF881" box="[300,447,1880,1904]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FE8F4240D9ECF881" box="[300,447,1880,1904]" italics="true" pageId="8" pageNumber="382">X. maculatus</emphasis>
</taxonomicName>
;
<figureCitation id="135A2A69FFD2C725FE6E4241DA5CF881" box="[461,527,1880,1904]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1966]" captionTargetBox="[151,1435,337,1921]" captionTargetId="figure@7.[151,1436,337,1922]" captionTargetPageId="7" captionText="FIGURE 5. Xanthias maculatus Sakai, 1961, holotype, male, 9.5 × 6.0 mm (NMST), Okinoyama, Sagami Bay, Japan. A, dorsal view; B, ventral view. Photos from Hironori Komatsu (NMST)." httpUri="https://zenodo.org/record/223772/files/figure.png" pageId="8" pageNumber="382">Fig. 5</figureCitation>
B); and 6) the ambulatory legs have noticeably broader and stouter meri, with the ratio of the width of the P5 merus to carapace length = 0.22 (
<figureCitation id="135A2A69FFD2C725FCE64265DBC0F865" box="[837,915,1916,1940]" captionStart="FIGURE 2" captionStartId="3.[151,250,1968,1991]" captionTargetBox="[292,1282,193,1921]" captionTargetId="figure@3.[292,1283,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080), Panglao Is., Bohol Sea, Philippines. A, dorsal view; B, ventral view; C, anterior view." httpUri="https://zenodo.org/record/223769/files/figure.png" pageId="8" pageNumber="382">Figs. 2</figureCitation>
A, 3B, 4E) (ambulatory meri more slender, P5 merus width/CL ratio =
<quantity id="4C999B09FFD2C725FE1242B9DA51F849" box="[433,514,1952,1976]" metricMagnitude="-3" metricUnit="m" metricValue="4.318" pageId="8" pageNumber="382" unit="in" value="0.17">0.17 in</quantity>
<taxonomicName id="4C614D6FFFD2C725FDA842B8DAF3F849" box="[523,672,1952,1976]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD2C725FDA842B8DAF3F849" box="[523,672,1952,1976]" italics="true" pageId="8" pageNumber="382">X. maculatus</emphasis>
</taxonomicName>
;
<figureCitation id="135A2A69FFD2C725FD1242B9DAA5F849" box="[689,758,1952,1976]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1966]" captionTargetBox="[151,1435,337,1921]" captionTargetId="figure@7.[151,1436,337,1922]" captionTargetPageId="7" captionText="FIGURE 5. Xanthias maculatus Sakai, 1961, holotype, male, 9.5 × 6.0 mm (NMST), Okinoyama, Sagami Bay, Japan. A, dorsal view; B, ventral view. Photos from Hironori Komatsu (NMST)." httpUri="https://zenodo.org/record/223772/files/figure.png" pageId="8" pageNumber="382">Fig. 5</figureCitation>
A). Although the G1 of both species are similar in general form, some differences can still be noted: that of
<taxonomicName id="4C614D6FFFD2C725FD1942DCDB13F82D" box="[698,832,1989,2012]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="382" phylum="Arthropoda" rank="species" species="joanneae" status="sp. nov.">
<emphasis id="B915EAFEFFD2C725FD1942DCDB13F82D" box="[698,832,1989,2012]" italics="true" pageId="8" pageNumber="382">X. joanneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2265785FFD2C725FCE542DDDBCCF82D" box="[838,927,1988,2012]" pageId="8" pageNumber="382" rank="species">
<emphasis id="B915EAFEFFD2C725FCE542DDDBCCF82D" bold="true" box="[838,927,1988,2012]" pageId="8" pageNumber="382">sp. nov.</emphasis>
</taxonomicNameLabel>
appears to be stouter, with fewer subterminal setae and a narrower apical lobe (
<figureCitation id="135A2A69FFD3C724FDB5458EDA09FF5E" box="[534,602,151,175]" captionStart="FIGURE 4" captionStartId="5.[151,250,1920,1943]" captionTargetBox="[179,1383,193,1862]" captionTargetId="figure@5.[171,1393,193,1873]" captionTargetPageId="5" captionText="FIGURE 4. Xanthias joanneae sp. nov., holotype, male, 7.9 × 5.1 mm (NMCR 39080). A, left third maxilliped, external view; B, anterior thoracic sternum, ventral view; C, carpus of right cheliped, dorso-external view; D, abdomen, ventral view; E, right P 5, dorsal view; left G 1 (F, G, I, J), F, external view; G, internal view; I, distal tip, external view; J, distal tip, internal view; H, left G 2, external view. Scale bars: A – E = 1.0 mm; F – H = 0.5 mm; I, J = 0.25 mm." httpUri="https://zenodo.org/record/223771/files/figure.png" pageId="9" pageNumber="383">Fig. 4</figureCitation>
F, G, I, J), whereas that of
<taxonomicName id="4C614D6FFFD3C724FC2A4580DC4EFF41" box="[905,1053,152,176]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD3C724FC2A4580DC4EFF41" box="[905,1053,152,176]" italics="true" pageId="9" pageNumber="383">X. maculatus</emphasis>
</taxonomicName>
, as illustrated by Sakai (1961), is relatively more slender, with more subterminal setae and a wider apical lobe (see Sakai 1961: fig. 1f, g). Together, these differences are sufficient to consider
<taxonomicName id="4C614D6FFFD3C724FDD545F8DAAFFF09" box="[630,764,225,248]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="joanneae" status="sp. nov.">
<emphasis id="B915EAFEFFD3C724FDD545F8DAAFFF09" box="[630,764,225,248]" italics="true" pageId="9" pageNumber="383">X. joanneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2265785FFD3C724FCA045F9DB0EFF09" box="[771,861,224,248]" pageId="9" pageNumber="383" rank="species">
<emphasis id="B915EAFEFFD3C724FCA045F9DB0EFF09" bold="true" box="[771,861,224,248]" pageId="9" pageNumber="383">sp. nov.</emphasis>
</taxonomicNameLabel>
as a distinct species from
<taxonomicName id="4C614D6FFFD3C724FB2745F8DD45FF09" box="[1156,1302,224,248]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD3C724FB2745F8DD45FF09" box="[1156,1302,224,248]" italics="true" pageId="9" pageNumber="383">X. maculatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDE36ECFFD3C724FF64441DD9BBFDAE" blockId="9.[151,1436,151,1149]" pageId="9" pageNumber="383">
A male and three females collected from the intertidal zone in
<collectingCountry id="F376767CFFD3C724FC24441DDBABFEED" box="[903,1016,260,284]" name="Kenya" pageId="9" pageNumber="383">Mombasa</collectingCountry>
,
<collectingCountry id="F376767CFFD3C724FBA0441CDC1CFEEC" box="[1027,1103,261,285]" name="Kenya" pageId="9" pageNumber="383">Kenya</collectingCountry>
, in east Africa, were reported by Serène (1984) as “
<taxonomicName id="4C614D6FFFD3C724FE2C4431D9A0FEB1" box="[399,499,296,320]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="B915EAFEFFD3C724FE2C4431D9A0FEB1" box="[399,499,296,320]" italics="true" pageId="9" pageNumber="383">Xanthias</emphasis>
</taxonomicName>
aff.
<taxonomicName id="4C614D6FFFD3C724FD8A4431DACFFEB1" box="[553,668,296,320]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD3C724FD8A4431DACFFEB1" box="[553,668,296,320]" italics="true" pageId="9" pageNumber="383">maculatus</emphasis>
</taxonomicName>
”. He provided a black-and-white photograph of the male specimen and a line drawing of the distal tip of the G1 (Serène 1984: fig. 114, pl. 28 fig. A). Serène apparently had doubts on the conspecificity of the
<collectingCountry id="F376767CFFD3C724FE084469DA51FE79" box="[427,514,368,392]" name="Kenya" pageId="9" pageNumber="383">Kenyan</collectingCountry>
specimens with
<taxonomicName id="4C614D6FFFD3C724FD1F4468DB1DFE79" box="[700,846,368,392]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD3C724FD1F4468DB1DFE79" box="[700,846,368,392]" italics="true" pageId="9" pageNumber="383">X. maculatus</emphasis>
</taxonomicName>
, pointing out that they had a straighter, less undulate frontal margin, less prominent/distinct supraorbital margin and carapace regions, lower, less produced carapace anterolateral teeth, indistinct tuberosities on the cheliped carpus, and shorter, wider ambulatory legs. He further points out that specimens he collected from Nhatrang,
<collectingCountry id="F376767CFFD3C724FD5944C5DB0FFE05" box="[762,860,476,500]" name="Vietnam" pageId="9" pageNumber="383">Vietnam</collectingCountry>
, were identical to the
<collectingCountry id="F376767CFFD3C724FBF344C4DCF4FE04" box="[1104,1191,477,501]" name="Kenya" pageId="9" pageNumber="383">Kenyan</collectingCountry>
specimens, at least in the proportions of the ambulatory legs. But Serène stopped short of describing them as a new species, suggesting that these variations may be intraspecific and that more material from different localities still needed to be examined (Serène 1984: 198).
</paragraph>
<paragraph id="8BDE36ECFFD3C724FF644775DCD5FC55" blockId="9.[151,1436,151,1149]" pageId="9" pageNumber="383">
Compared with
<taxonomicName id="4C614D6FFFD3C724FE224774DA59FD75" box="[385,522,621,644]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="joanneae" status="sp. nov.">
<emphasis id="B915EAFEFFD3C724FE224774DA59FD75" box="[385,522,621,644]" italics="true" pageId="9" pageNumber="383">X. joanneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2265785FFD3C724FDB74775DA23FD75" box="[532,624,620,644]" pageId="9" pageNumber="383" rank="species">
<emphasis id="B915EAFEFFD3C724FDB74775DA23FD75" bold="true" box="[532,624,620,644]" pageId="9" pageNumber="383">sp. nov.</emphasis>
</taxonomicNameLabel>
, the carapace of the
<collectingCountry id="F376767CFFD3C724FCC54774DBEEFD74" box="[870,957,621,645]" name="Kenya" pageId="9" pageNumber="383">Kenyan</collectingCountry>
specimen appears more inflated dorsally, with the first two teeth on the carapace anterolateral margin virtually effaced, the last two teeth less pronounced, and external surface of the cheliped carpus smooth, without grooves or tuberosities. Furthermore, the G1 (Serène 1984: fig. 114) is also more slender, and, unlike in the new species and in
<taxonomicName id="4C614D6FFFD3C724FC7047C0DC36FD01" box="[979,1125,728,752]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD3C724FC7047C0DC36FD01" box="[979,1125,728,752]" italics="true" pageId="9" pageNumber="383">X. maculatus</emphasis>
</taxonomicName>
, appears to have two apical lobes instead of one. It is very likely that the
<collectingCountry id="F376767CFFD3C724FD6447E4DB4DFCE4" box="[711,798,765,789]" name="Kenya" pageId="9" pageNumber="383">Kenyan</collectingCountry>
specimens comprise a distinct species from either
<taxonomicName id="4C614D6FFFD3C724FA2747E4D959FCC9" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B915EAFEFFD3C724FA2747E4D959FCC9" italics="true" pageId="9" pageNumber="383">X. maculatus</emphasis>
</taxonomicName>
or
<taxonomicName id="4C614D6FFFD3C724FE8C4638D9E5FCC9" box="[303,438,801,824]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="joanneae" status="sp. nov.">
<emphasis id="B915EAFEFFD3C724FE8C4638D9E5FCC9" box="[303,438,801,824]" italics="true" pageId="9" pageNumber="383">X. joanneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2265785FFD3C724FE1F4639DA45FCC9" box="[444,534,800,824]" pageId="9" pageNumber="383" rank="species">
<emphasis id="B915EAFEFFD3C724FE1F4639DA45FCC9" bold="true" box="[444,534,800,824]" pageId="9" pageNumber="383">sp. nov.</emphasis>
</taxonomicNameLabel>
, but the specimens, presently deposited at MNHN, have to be re-examined for a proper description. There are no published figures of the specimen from Nhatrang which Serène (1984) referred to, and it could not be examined for this work, although it may still be in the Nhatrang Oceanography Institute, which currently maintains his collection of crustaceans from that region (P. Castro pers. comm.).
</paragraph>
<paragraph id="8BDE36ECFFD3C724FF6446A9DADEFB8D" blockId="9.[151,1436,151,1149]" pageId="9" pageNumber="383">
<emphasis id="B915EAFEFFD3C724FF6446A9DAC5FC39" bold="true" box="[199,662,943,968]" pageId="9" pageNumber="383">Ecology and geographical distribution.</emphasis>
The
<typeStatus id="54DA884EFFD3C724FD6D46A9DAADFC39" box="[718,766,944,968]" pageId="9" pageNumber="383">type</typeStatus>
material of
<taxonomicName id="4C614D6FFFD3C724FC2646A8DC58FC39" box="[901,1035,945,968]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="383" phylum="Arthropoda" rank="species" species="joanneae" status="sp. nov.">
<emphasis id="B915EAFEFFD3C724FC2646A8DC58FC39" box="[901,1035,945,968]" italics="true" pageId="9" pageNumber="383">X. joanneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2265785FFD3C724FBB246A9DC38FC39" box="[1041,1131,944,968]" pageId="9" pageNumber="383" rank="species">
<emphasis id="B915EAFEFFD3C724FBB246A9DC38FC39" bold="true" box="[1041,1131,944,968]" pageId="9" pageNumber="383">sp. nov.</emphasis>
</taxonomicNameLabel>
were all collected from the Bohol Sea, off the islands of Panglao, Balicasag and Pamilacan, at a depth of
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2–
<specimenCount id="9D67FD65FFD3C724FB9446CDDCD1FC1D" box="[1079,1154,980,1004]" pageId="9" pageNumber="383" type="generic">110 m</specimenCount>
</quantity>
. The microhabitats were described by the diver-collectors as reef platforms, slopes or walls, as well as the floors of submarine caves, with the substrate consisting mostly of coral rubble, but some also including sand and gravel. Most were collected by hand or by coral brushing at diving depth (&lt;
<quantity id="4C999B09FFD3C724FD374126DA82FBA6" box="[660,721,1087,1111]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="383" unit="m" value="40.0">40 m</quantity>
), but some specimens were collected at greater depths (
<quantity id="4C999B09FFD3C724FAEB4126D8FDFB8D" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="9" pageNumber="383" unit="m" value="85.0" valueMax="110.0" valueMin="60.0">60–110 m</quantity>
) utilizing tangle nets (see Ng
<emphasis id="B915EAFEFFD3C724FE5D417CDA7CFB8D" box="[510,559,1124,1148]" italics="true" pageId="9" pageNumber="383">et al</emphasis>
., 2009).
</paragraph>
</subSubSection>
</treatment>
</document>
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