173 lines
20 KiB
XML
173 lines
20 KiB
XML
<document id="449AE53BE0B9789E80232A96827E08D1" ID-DOI="10.11646/zootaxa.4755.1.6" ID-GBIF-Dataset="9f6752d6-7c9a-4756-acfe-badb2a0301d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3724291" ID-ZooBank="D0B3FFA8-C262-4CD6-98AB-B46C55FCEF6D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584968592865" checkinUser="plazi" docAuthor="Busanello, Dilson G. C., Irmler, Ulrich & Caron, Edilson" docDate="2020" docId="03C887D5FF84FF8483C6F9D5C485FD61" docLanguage="en" docName="zootaxa.4755.1.6.pdf" docOrigin="Zootaxa 4755 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudastenus latus Busanello & Irmler & Caron 2020, sp. nov." docType="treatment" docVersion="5" lastPageNumber="149" masterDocId="FFF1FFADFF8DFF8E8351FF82C634FFE6" masterDocTitle="Revision of Pseudastenus Bernhauer, synonym with Xenasterides Newton and six new species (Coleoptera, Staphylinidae, Paederinae)" masterLastPageNumber="154" masterPageNumber="139" pageNumber="148" updateTime="1698730789478" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="050CF6AADC7658A9E368DC4C093081D9">Revision of Pseudastenus Bernhauer, synonym with Xenasterides Newton and six new species (Coleoptera, Staphylinidae, Paederinae)</mods:title>
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<mods:name id="92611CDE3D139DBCBFF1EA5338AE07D0" type="personal">
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<mods:namePart id="DDB720F33A054A2763E4A1D3C8B2E3F8">Busanello, Dilson G. C.</mods:namePart>
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<mods:namePart id="F94DFEE0C935DC4669A871CEAE15226E">Irmler, Ulrich</mods:namePart>
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<mods:namePart id="068DDAEF86F544050CE8F7CD7E423D11">Caron, Edilson</mods:namePart>
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<mods:title id="CE8A65E3306AE690D0FCE166FBD4B370">Zootaxa</mods:title>
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<mods:part id="0D4F51FF115A8EA81F1373BE7CCE6825">
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<mods:date id="13195D9A7FC5F4A9B2393CE444D3FB56">2020</mods:date>
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<mods:number id="68DDA8A74C2662F0B131CC3709FC379F">2020-03-23</mods:number>
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<mods:identifier id="9908A78A358860CE016EF0E574CC7B8C" type="DOI">10.11646/zootaxa.4755.1.6</mods:identifier>
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<treatment id="03C887D5FF84FF8483C6F9D5C485FD61" ID-DOI="http://doi.org/10.5281/zenodo.3730231" ID-GBIF-Taxon="162696940" ID-Zenodo-Dep="3730231" LSID="urn:lsid:plazi:treatment:03C887D5FF84FF8483C6F9D5C485FD61" httpUri="http://treatment.plazi.org/id/03C887D5FF84FF8483C6F9D5C485FD61" lastPageId="10" lastPageNumber="149" pageId="9" pageNumber="148">
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<subSubSection id="C37B6548FF84FF8783C6F9D5C4EFF994" box="[151,731,1623,1650]" pageId="9" pageNumber="148" type="nomenclature">
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<paragraph id="8BDE36C3FF84FF8783C6F9D5C4EFF994" blockId="9.[151,731,1623,1686]" box="[151,731,1623,1650]" pageId="9" pageNumber="148">
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<heading id="D09681AFFF84FF8783C6F9D5C4EFF994" bold="true" box="[151,731,1623,1650]" fontSize="11" level="1" pageId="9" pageNumber="148" reason="1">
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<emphasis id="B915EAD1FF84FF8783C6F9D5C4EFF994" bold="true" box="[151,731,1623,1650]" pageId="9" pageNumber="148">
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<taxonomicName id="4C614D40FF84FF8783C6F9D5C444F997" ID-CoL="9V8WT" authority="Busanello& Caron" authorityName="Busanello & Irmler & Caron" authorityYear="2020" box="[151,624,1623,1650]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="148" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
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<emphasis id="B915EAD1FF84FF8783C6F9D5C74AF994" bold="true" box="[151,382,1623,1650]" italics="true" pageId="9" pageNumber="148">Pseudastenus latus</emphasis>
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Busanello& Caron
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22657AAFF84FF87812CF9DAC4EFF994" box="[637,731,1624,1650]" pageId="9" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6548FF84FF8783C6F9F9C72FF970" box="[151,283,1659,1686]" pageId="9" pageNumber="148" type="description">
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<paragraph id="8BDE36C3FF84FF8783C6F9F9C72FF970" blockId="9.[151,731,1623,1686]" box="[151,283,1659,1686]" pageId="9" pageNumber="148">
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<figureCitation id="135A2A46FF84FF8783C6F9F9C6C0F970" box="[151,244,1659,1686]" captionStart="FIGURES 40–45" captionStartId="11.[151,264,1932,1957]" captionTargetBox="[197,1381,213,1868]" captionTargetId="figure@11.[161,1427,181,1906]" captionTargetPageId="11" captionText="FIGURES 40–45. Dorsal view. 40. Pseudastenus ribeirocostae sp. nov.; 41. P. ferrugineus sp. nov.; 42. P. latus sp. nov.; 43. P. oculatus sp. nov.; 44. P. amazonicus sp. nov.; 45. P. schubarti sp. nov. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3724303" httpUri="https://zenodo.org/record/3724303/files/figure.png" pageId="9" pageNumber="148">Figs. 42</figureCitation>
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,
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<figureCitation id="135A2A46FF84FF8783AEF9F9C72FF970" box="[255,283,1659,1686]" captionStart="FIGURES 48–54" captionStartId="13.[151,264,1895,1920]" captionTargetBox="[232,1346,220,1805]" captionTargetId="figure@13.[164,1424,181,1869]" captionTargetPageId="13" captionText="FIGURES 48–54. Aedeagus of Pseudastenus, dorsal and lateral view, respectively, figures 51–54 including microscopic detail of basal slerites. 48. P. ribeirocostae sp. nov.; 49. P. ferrugineus sp. nov.; 50. P. latus sp. nov.; 51. P. barretoi; 52. P. oculatus sp. nov.; 53. P. amazonicus sp. nov.; 54. P. schubarti sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3724309" httpUri="https://zenodo.org/record/3724309/files/figure.png" pageId="9" pageNumber="148">50</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C37B6548FF84FF8783C6F93FC429F8A2" pageId="9" pageNumber="148" type="materials_examined">
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<paragraph id="8BDE36C3FF84FF8783C6F93FC429F8A2" blockId="9.[151,1437,1725,2041]" pageId="9" pageNumber="148">
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<emphasis id="B915EAD1FF84FF8783C6F93FC7B4F93E" bold="true" box="[151,384,1725,1752]" pageId="9" pageNumber="148">Material examined.</emphasis>
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<materialsCitation id="3B093C9EFF84FF8782D6F93CC429F8A2" ID-GBIF-Occurrence="2579576920" collectingDate="1978-02-03" collectingDateMax="1978-02-05" collectingDateMin="1978-02-03" collectionCode="FMNH" collectorName="B. Heppner" determinerName="E. Caron" location="Rio" municipality="Canton" pageId="9" pageNumber="148" specimenCount="2" stateProvince="Barinas" typeStatus="holotype">
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<typeStatus id="54DA8861FF84FF8782D6F93CC7C7F93E" box="[391,499,1726,1752]" pageId="9" pageNumber="148" type="holotype">Holotype</typeStatus>
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||
. Male, dissected, deposited in
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<collectionCode id="ED70AE06FF84FF878017F93DC5AEF93F" box="[838,922,1727,1753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="148" type="Museum">FMNH</collectionCode>
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, labels: (1) “VEN.:
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<collectingRegion id="49A5F821FF84FF878726F93CC2FBF93E" box="[1143,1231,1726,1752]" country="Venezuela" name="Barinas" pageId="9" pageNumber="148">Barinas</collectingRegion>
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,
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||
<quantity id="4C999B26FF84FF87878BF93CC315F93E" box="[1242,1313,1726,1752]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="148" unit="km" value="32.0">32 km</quantity>
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/E E1
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<collectingMunicipality id="6BBAACB9FF84FF878637F93CC689F91B" pageId="9" pageNumber="148">Canton</collectingMunicipality>
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,
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||
<location id="8EBE6018FF84FF87839AF960C6C7F91A" LSID="urn:lsid:plazi:treatment:03C887D5FF84FF8483C6F9D5C485FD61:8EBE6018FF84FF87839AF960C6C7F91A" box="[203,243,1762,1788]" municipality="Canton" name="Rio" pageId="9" pageNumber="148" stateProvince="Barinas">Rio</location>
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/Caparo Res. Sta. S.,/
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<date id="FFDF1003FF84FF8782B0F963C45DF91A" box="[481,617,1761,1788]" pageId="9" pageNumber="148" value="1978-02-03" valueMax="1978-02-05" valueMin="1978-02-03">
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<collectingDate id="EF9BE9EBFF84FF8782B0F963C45DF91A" box="[481,617,1761,1788]" pageId="9" pageNumber="148" value="1978-02-03" valueMax="1978-02-05" valueMin="1978-02-03">3–5-II-1978</collectingDate>
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</date>
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,/seasonal forest, J./
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<collectorName id="26945315FF84FF878001F960C5E0F91A" box="[848,980,1762,1788]" pageId="9" pageNumber="148">B. Heppner</collectorName>
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” [white label, printed in black]; (2) “At black/light” [white label, printed in black]; (3) “
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<typeStatus id="54DA8861FF84FF8781FDF884C576F8C6" box="[684,834,1798,1824]" pageId="9" pageNumber="148" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="4C614D40FF84FF87801AF884C2C8F8C6" authority="Busanello & Caron" authorityName="Busanello & Irmler & Caron" authorityYear="2020" box="[843,1276,1798,1824]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="148" phylum="Arthropoda" rank="species" species="latus">Pseudastenus latus Busanello & Caron</taxonomicName>
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/det.
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<determinerName id="7E02DA07FF84FF878663F885C3ACF8C6" box="[1330,1432,1798,1825]" pageId="9" pageNumber="148">E. Caron</determinerName>
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, 2019” [red label/ printed in black].
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37B6548FF84FF878396F8CFC307F86A" pageId="9" pageNumber="148" type="diagnosis">
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<paragraph id="8BDE36C3FF84FF878396F8CFC307F86A" blockId="9.[151,1437,1725,2041]" pageId="9" pageNumber="148">
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<emphasis id="B915EAD1FF84FF878396F8CFC776F88E" bold="true" box="[199,322,1869,1896]" pageId="9" pageNumber="148">Diagnosis.</emphasis>
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<taxonomicName id="4C614D40FF84FF87821AF8CCC410F88E" authorityName="Busanello & Irmler & Caron" authorityYear="2020" box="[331,548,1870,1896]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="148" phylum="Arthropoda" rank="species" species="latus">
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<emphasis id="B915EAD1FF84FF87821AF8CCC410F88E" box="[331,548,1870,1896]" italics="true" pageId="9" pageNumber="148">Pseudastenus latus</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C614D40FF84FF878193F8CDC56CF88E" baseAuthorityName="Bierig" baseAuthorityYear="1939" box="[706,856,1870,1896]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="148" phylum="Arthropoda" rank="species" species="plaumanni">
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<emphasis id="B915EAD1FF84FF878193F8CDC56CF88E" box="[706,856,1870,1896]" italics="true" pageId="9" pageNumber="148">P. plaumanni</emphasis>
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</taxonomicName>
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by the dark brown coloration of the head (lighter brown to yellowish in
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<taxonomicName id="4C614D40FF84FF8782C2F8F1C413F86A" baseAuthorityName="Bierig" baseAuthorityYear="1939" box="[403,551,1906,1932]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="148" phylum="Arthropoda" rank="species" species="plaumanni">
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<emphasis id="B915EAD1FF84FF8782C2F8F1C413F86A" box="[403,551,1906,1932]" italics="true" pageId="9" pageNumber="148">P. plaumanni</emphasis>
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</taxonomicName>
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) and mainly by the apical sclerites broader in the basal half (
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<figureCitation id="135A2A46FF84FF878783F8F0C312F86A" box="[1234,1318,1905,1932]" captionStart="FIGURES 48–54" captionStartId="13.[151,264,1895,1920]" captionTargetBox="[232,1346,220,1805]" captionTargetId="figure@13.[164,1424,181,1869]" captionTargetPageId="13" captionText="FIGURES 48–54. Aedeagus of Pseudastenus, dorsal and lateral view, respectively, figures 51–54 including microscopic detail of basal slerites. 48. P. ribeirocostae sp. nov.; 49. P. ferrugineus sp. nov.; 50. P. latus sp. nov.; 51. P. barretoi; 52. P. oculatus sp. nov.; 53. P. amazonicus sp. nov.; 54. P. schubarti sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3724309" httpUri="https://zenodo.org/record/3724309/files/figure.png" pageId="9" pageNumber="148">Fig. 50</figureCitation>
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).
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C37B6548FF84FF848396F817C338FE11" lastPageId="10" lastPageNumber="149" pageId="9" pageNumber="148" type="description">
|
||
<paragraph id="8BDE36C3FF84FF848396F817C338FE11" blockId="9.[151,1437,1725,2041]" lastBlockId="10.[151,1437,153,647]" lastPageId="10" lastPageNumber="149" pageId="9" pageNumber="148">
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||
<emphasis id="B915EAD1FF84FF878396F817C76DF856" bold="true" box="[199,345,1941,1968]" pageId="9" pageNumber="148">Description.</emphasis>
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Body length:
|
||
<quantity id="4C999B26FF84FF8782A7F814C468F857" box="[502,604,1942,1969]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="148" unit="mm" value="3.2">3.20 mm</quantity>
|
||
. Coloration: head, pronotum and appendices yellowish brown, head slightly darker; coloration of basal elytra similar to head, apical half similar to pronotum; abdomen dark brown. Head
|
||
<quantity id="4C999B26FF84FF8783C6F85FC6CEF81F" box="[151,250,2013,2041]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="148" unit="mm" value="0.45">0.45 mm</quantity>
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long,
|
||
<quantity id="4C999B26FF84FF87826EF85FC797F81F" box="[319,419,2013,2041]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="148" unit="mm" value="0.52">0.52 mm</quantity>
|
||
wide; eyes slightly prominent; 0.30 times as long as total length of head; 0.86 times as long as postocular length; approximately parallel; posterior margin straight; depressed on central vertex in front of neck. Pronotum
|
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<quantity id="4C999B26FF87FF84825CFF3EC745FF31" box="[269,369,188,215]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="149" unit="mm" value="0.45">0.45 mm</quantity>
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long,
|
||
<quantity id="4C999B26FF87FF8482E8FF3EC429FF31" box="[441,541,188,215]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="10" pageNumber="149" unit="mm" value="0.52">0.52 mm</quantity>
|
||
wide; widest close to anterior angles; anterior margin slightly curved; posterior margin straight; sides convergingposteriad. Elytra
|
||
<quantity id="4C999B26FF87FF84819DFF63C504FF1D" box="[716,816,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="10" pageNumber="149" unit="mm" value="0.64">0.64 mm</quantity>
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||
long,
|
||
<quantity id="4C999B26FF87FF848027FF62C5EEFF1D" box="[886,986,224,251]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="10" pageNumber="149" unit="mm" value="0.53">0.53 mm</quantity>
|
||
wide; conspicuously longer than pronotum; sides slightly convexly curved; widest in the middle; humeral angles rounded and obtuse, but conspicuously developed; hind wings developed; posterior angles nearly rectangular; posterior margin emarginate; suture 0.82 times as long as lateral side; microtuberculate punctation larger than on head and pronotum; with yellow pubescence pointing posteriad, longer than head and pronotum; posterior edge with fringe of sparse robust setae. Abdomen with dense punctation; with yellow pubescence pointing posteriad; pubescence conspicuously longer than on the rest of body. Aedeagus elongate;
|
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<quantity id="4C999B26FF87FF84828FFE3BC477FE35" box="[478,579,441,467]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="10" pageNumber="149" unit="mm" value="0.61">0.61 mm</quantity>
|
||
long; 4.18 times as long as wide; sides parallel for almost the entire length, without grooves on internal sac and apical sclerites enlarged in the basal half,
|
||
<quantity id="4C999B26FF87FF848750FE5FC250FE11" box="[1025,1124,477,503]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="149" unit="mm" value="0.11">0.11 mm</quantity>
|
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long (
|
||
<figureCitation id="135A2A46FF87FF8487FAFE5FC334FE11" box="[1195,1280,476,503]" captionStart="FIGURES 48–54" captionStartId="13.[151,264,1895,1920]" captionTargetBox="[232,1346,220,1805]" captionTargetId="figure@13.[164,1424,181,1869]" captionTargetPageId="13" captionText="FIGURES 48–54. Aedeagus of Pseudastenus, dorsal and lateral view, respectively, figures 51–54 including microscopic detail of basal slerites. 48. P. ribeirocostae sp. nov.; 49. P. ferrugineus sp. nov.; 50. P. latus sp. nov.; 51. P. barretoi; 52. P. oculatus sp. nov.; 53. P. amazonicus sp. nov.; 54. P. schubarti sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3724309" httpUri="https://zenodo.org/record/3724309/files/figure.png" pageId="10" pageNumber="149">Fig. 50</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B6548FF87FF848396FD83C485FD61" pageId="10" pageNumber="149" type="etymology">
|
||
<paragraph id="8BDE36C3FF87FF848396FD83C7CEFDD9" blockId="10.[151,1437,153,647]" pageId="10" pageNumber="149">
|
||
<emphasis id="B915EAD1FF87FF848396FD83C77AFDFD" bold="true" box="[199,334,513,539]" pageId="10" pageNumber="149">Etymology.</emphasis>
|
||
The specific name derives from the Latin word “latus” meaning broad and refers to the shape of apical sclerites of the aedeagus.
|
||
</paragraph>
|
||
<paragraph id="8BDE36C3FF87FF848396FDCAC3B5FD82" blockId="10.[151,1437,153,647]" box="[199,1409,584,612]" pageId="10" pageNumber="149">
|
||
<emphasis id="B915EAD1FF87FF848396FDCAC7E5FD85" bold="true" box="[199,465,584,611]" pageId="10" pageNumber="149">Geographical records.</emphasis>
|
||
<taxonomicName id="4C614D40FF87FF848289FDCBC499FD85" authorityName="Busanello & Irmler & Caron" authorityYear="2020" box="[472,685,585,611]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="149" phylum="Arthropoda" rank="species" species="latus">
|
||
<emphasis id="B915EAD1FF87FF848289FDCBC499FD85" box="[472,685,585,611]" italics="true" pageId="10" pageNumber="149">Pseudastenus latus</emphasis>
|
||
</taxonomicName>
|
||
is known from the southeast of the state of
|
||
<collectingRegion id="49A5F821FF87FF8487C6FDCBC2D9FD85" box="[1175,1261,585,611]" country="Venezuela" name="Barinas" pageId="10" pageNumber="149">Barinas</collectingRegion>
|
||
(
|
||
<collectingCountry id="F3767653FF87FF8487ADFDCBC342FD82" box="[1276,1398,585,612]" name="Venezuela" pageId="10" pageNumber="149">Venezuela</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDE36C3FF87FF848396FDEEC485FD61" blockId="10.[151,1437,153,647]" box="[199,689,620,647]" pageId="10" pageNumber="149">
|
||
<emphasis id="B915EAD1FF87FF848396FDEEC7BEFD61" bold="true" box="[199,394,620,647]" pageId="10" pageNumber="149">Biological notes.</emphasis>
|
||
No information on labels.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |