423 lines
41 KiB
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423 lines
41 KiB
XML
<document ID-DOI="10.5252/geodiversitas2022v44a25" ID-GBIF-Dataset="d0184974-bea3-4f97-a2f8-98887ec4f87e" ID-ISSN="1638-9395" ID-Zenodo-Dep="6902033" ID-ZooBank="urn:lsid:zoobank.org:pub:EA12DCB7-A5BE-4763-B805-25087EBD726D" approvalRequired="9" approvalRequired_for_taxonomicNames="9" checkinTime="1658759598753" checkinUser="felipe" docAuthor="Allain, Ronan, Vullo, Romain, Rozada, Lee, Anquetin, Jérémy, Bourgeais, Renaud, Goedert, Jean, Lasseron, Maxime, Martin, Jeremy E., Pérez-García, Adán, Fabrègues, Claire Peyre De, Royo-Torres, Rafael, Augier, Dominique & Bailly, Gilles" docDate="2022" docId="03C887B9FFA1FFF27124A509FEA2F85A" docLanguage="en" docName="Geodiversitas.44.25.683-752.pdf.imf" docOrigin="Geodiversitas 44 (25)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.3:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="3" docTitle="Goniopholis undetermined" docType="treatment" docVersion="5" lastPageNumber="706" masterDocId="FFF1FFC1FFB6FFEB753AA238FFEBFFB1" masterDocTitle="Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J / K boundary" masterLastPageNumber="752" masterPageNumber="683" pageNumber="704" updateTime="1659037062892" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J / K boundary</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Allain, Ronan</mods:namePart>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vullo, Romain</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rozada, Lee</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Anquetin, Jérémy</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bourgeais, Renaud</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Goedert, Jean</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lasseron, Maxime</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Martin, Jeremy E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pérez-García, Adán</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fabrègues, Claire Peyre De</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Royo-Torres, Rafael</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Augier, Dominique</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bailly, Gilles</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Geodiversitas</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-07-21</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>44</mods:number>
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<mods:detail type="issue">
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<mods:number>25</mods:number>
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<mods:start>683</mods:start>
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<mods:end>752</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5252/geodiversitas2022v44a25</mods:identifier>
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<mods:identifier type="GBIF-Dataset">d0184974-bea3-4f97-a2f8-98887ec4f87e</mods:identifier>
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<mods:identifier type="ISSN">1638-9395</mods:identifier>
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<mods:identifier type="Zenodo-Dep">6902033</mods:identifier>
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<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:EA12DCB7-A5BE-4763-B805-25087EBD726D</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6929037" ID-Zenodo-Dep="6929037" LSID="urn:lsid:plazi:treatment:03C887B9FFA1FFF27124A509FEA2F85A" httpUri="http://treatment.plazi.org/id/03C887B9FFA1FFF27124A509FEA2F85A" lastPageId="25" lastPageNumber="706" pageId="23" pageNumber="704">
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<subSubSection box="[1054,1214,1841,1868]" pageId="23" pageNumber="704" type="nomenclature">
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<paragraph blockId="23.[1054,1214,1841,1868]" box="[1054,1214,1841,1868]" pageId="23" pageNumber="704">
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<heading box="[1054,1214,1841,1868]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="704" reason="2">
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<taxonomicName box="[1054,1214,1841,1868]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" kingdom="Animalia" order="Crocodylia" pageId="23" pageNumber="704" phylum="Chordata" rank="species" species="undetermined">
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<emphasis box="[1054,1177,1841,1867]" italics="true" pageId="23" pageNumber="704">Goniopholis</emphasis>
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sp.
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="25" lastPageNumber="706" pageId="23" pageNumber="704" type="description">
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<paragraph blockId="23.[1087,1181,1873,1900]" box="[1087,1181,1873,1900]" pageId="23" pageNumber="704">
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(
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<figureCitation box="[1096,1172,1873,1900]" captionStart="FIG" captionStartId="25.[132,143,1323,1340]" captionTargetBox="[132,1455,215,1279]" captionTargetId="figure-275@25.[132,1455,215,1279]" captionTargetPageId="25" captionText="FIG. 18. — Goniopholidid crocodyliform Angeac-Charente: A, D, left dentary of Goniopholis sp. (ANG18-5925) in dorsal (A) and ventral (D) views; B, C, skull of Goniopholis sp. (ANG18-5914, ANG18-5920, ANG18-5921) in dorsal (B) and ventral (C) views. Scale bar: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.6902077" httpUri="https://zenodo.org/record/6902077/files/figure.png" pageId="23" pageNumber="704">Fig. 18</figureCitation>
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)
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</paragraph>
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<paragraph blockId="23.[813,1456,1938,2028]" box="[813,957,1938,1964]" pageId="23" pageNumber="704">DESCRIPTION</paragraph>
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<paragraph blockId="23.[813,1456,1938,2028]" lastBlockId="24.[130,777,1140,2028]" lastPageId="24" lastPageNumber="705" pageId="23" pageNumber="704">
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In addition to fragmentary cranial remains and numerous isolated teeth, a nearly complete skull and mandibular elements (ANG1 8-5914, 5920, 5921 and 5925) of a single individual have been excavated and prepared in 2018 (
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<figureCitation box="[586,659,1172,1198]" captionStart="FIG" captionStartId="25.[132,143,1323,1340]" captionTargetBox="[132,1455,215,1279]" captionTargetId="figure-275@25.[132,1455,215,1279]" captionTargetPageId="25" captionText="FIG. 18. — Goniopholidid crocodyliform Angeac-Charente: A, D, left dentary of Goniopholis sp. (ANG18-5925) in dorsal (A) and ventral (D) views; B, C, skull of Goniopholis sp. (ANG18-5914, ANG18-5920, ANG18-5921) in dorsal (B) and ventral (C) views. Scale bar: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.6902077" httpUri="https://zenodo.org/record/6902077/files/figure.png" pageId="24" pageNumber="705">Fig. 18</figureCitation>
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). The skull is broken transversally in front of the orbits and the rostrum is split longitudinally along the nasals and premaxillae. The associated mandible preserves both dentaries. In comparison with other goniopholidids from Europe, this skull is notable for its exquisite preservation, being minimally compacted dorsoventrally.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.6902075" ID-Zenodo-Dep="6902075" httpUri="https://zenodo.org/record/6902075/files/figure.png" pageId="24" pageNumber="705" startId="24.[132,143,1035,1052]" targetBox="[132,1455,215,993]" targetPageId="24">
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<paragraph blockId="24.[132,1456,1035,1074]" pageId="24" pageNumber="705">
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FIG. 17. — Crocodyliform teeth from Angeac-Charente:
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<emphasis bold="true" box="[593,627,1035,1052]" pageId="24" pageNumber="705">A -D</emphasis>
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, broadened teeth of
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<taxonomicName box="[800,946,1035,1052]" class="Reptilia" family="Atoposauridae" genus="Theriosuchus" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="species" species="undetermined">
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<emphasis box="[800,911,1035,1052]" italics="true" pageId="24" pageNumber="705">Theriosuchus</emphasis>
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sp.
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</taxonomicName>
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, ANG M-13 (
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<emphasis bold="true" box="[1051,1064,1035,1052]" pageId="24" pageNumber="705">A</emphasis>
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), ANG M-07 (
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<emphasis bold="true" box="[1177,1190,1035,1052]" pageId="24" pageNumber="705">B</emphasis>
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), ANG M-19 (
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<emphasis bold="true" box="[1304,1318,1035,1052]" pageId="24" pageNumber="705">C</emphasis>
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), ANG M-12 (
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<emphasis bold="true" box="[1431,1445,1035,1052]" pageId="24" pageNumber="705">D</emphasis>
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);
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<emphasis bold="true" box="[132,144,1057,1074]" pageId="24" pageNumber="705">E</emphasis>
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, tooth of
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<taxonomicName authorityName="Dollo" authorityYear="1883" box="[225,349,1057,1074]" class="Reptilia" family="Bernissartiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="family">Bernissartiidae</taxonomicName>
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indet. (ANG M-11). Scale bar: 1 mm.
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</paragraph>
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</caption>
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<paragraph blockId="24.[130,777,1140,2028]" pageId="24" pageNumber="705">
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Based on recent works (
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<bibRefCitation author="SALISBURY S. W. & WILLIS P. M. A. & PEITZ S. & SANDER P. M." box="[400,609,1395,1421]" journalOrPublisher="Special Papers in Palaeontology" pageId="24" pageNumber="705" pagination="121 - 148" part="60" refId="ref48575" refString="SALISBURY S. W., WILLIS P. M. A., PEITZ S. & SANDER P. M. 1999. - The crocodilian Goniopholis simus from the lower Cretaceous of north-western Germany. Special Papers in Palaeontology 60: 121 - 148." title="The crocodilian Goniopholis simus from the lower Cretaceous of north-western Germany" type="journal article" year="1999">
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Salisbury
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<emphasis box="[501,549,1395,1421]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
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1999
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</bibRefCitation>
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;
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<bibRefCitation author="SCHWARZ D." box="[620,768,1395,1421]" journalOrPublisher="Palaeontology" pageId="24" pageNumber="705" pagination="185 - 208" part="45" refId="ref48771" refString="SCHWARZ D. 2002. - A New Species of Goniopholis from the Upper Jurassic of Portugal. Palaeontology 45: 185 - 208. https: // doi. org / 10.1111 / 1475 - 4983.00233" title="A New Species of Goniopholis from the Upper Jurassic of Portugal" type="journal article" year="2002">Schwarz 2002</bibRefCitation>
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;
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<bibRefCitation author="ANDRADE M. B. & HORNUNG J. J." box="[132,339,1427,1453]" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="24" pageNumber="705" pagination="352 - 368" part="31" refId="ref35562" refString="ANDRADE M. B. & HORNUNG J. J. 2011. - A new look into the periorbital morphology of Goniopholis (Mesoeucrocodylia: Neosuchia) and related forms. Journal of Vertebrate Paleontology 31 (2): 352 - 368. https: // doi. org / 10.1080 / 02724634.2011.550353" title="A new look into the periorbital morphology of Goniopholis (Mesoeucrocodylia: Neosuchia) and related forms" type="journal article" year="2011">
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Andrade
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<emphasis box="[229,279,1427,1453]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
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2011
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</bibRefCitation>
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;
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<bibRefCitation author="Buscalioni" box="[351,578,1427,1453]" etAl="et al." firstAuthor="Buscalioni" pageId="24" pageNumber="705" pagination="103 - 122" refId="ref37078" refString="BUSCALIONI A. D., ALCALA L., ESPILEZ E. & MAMPEL L. 2013. - European Goniopholididae from the Early Albian Escucha Formation in Arino (Teruel, Aragon, Spain). Spanish Journal of Palaeontology 28: 103 - 122. https: // doi. org / 10.7203 / sjp. 28.1.17835" type="journal article" year="2013">
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Buscalioni
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<emphasis box="[468,517,1427,1453]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
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2013
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</bibRefCitation>
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; Puértolas-Pascual
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<emphasis box="[132,182,1459,1485]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
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2015;
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<bibRefCitation author="Martin" box="[254,458,1459,1485]" etAl="et al." firstAuthor="Martin" pageId="24" pageNumber="705" pagination="1222534" refId="ref43294" refString="MARTIN J. E., DELFINO M. & SMITH T. 2016 a. - Osteology and affinities of Dollo's goniopholidid (Mesoeucrocodylia) from the Early Cretaceous of Bernissart, Belgium. Journal of Vertebrate Paleontology 36 (6): e 1222534. https: // doi. org / 10.1080 / 0272 4634.2016. 1222534" type="journal article" year="2016" yearSuffix="a">
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Martin
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<emphasis box="[336,386,1459,1485]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
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2016a
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</bibRefCitation>
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), several diagnostic characters used in combination permit assignment of this specimen to
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<taxonomicName authorityName="Cope" authorityYear="1875" box="[132,315,1523,1549]" class="Reptilia" family="Goniopholididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="family">Goniopholididae</taxonomicName>
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, including: 1) a mesorostrine condition with a rostrum to skull ratio
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<emphasis box="[387,404,1555,1580]" italics="true" pageId="24" pageNumber="705">c.</emphasis>
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0.65; 2) constricted outline of the upper jaw at the level of the premaxillary-maxillary suture; 3) double-caniniform maxillary teeth 4+5; 4) confluent dentary alveoli 3+4; 5) orbits smaller than supratemporal fenestrae and slightly smaller than frontal width at interorbital level; 6) skull table and dorsal surface of quadratojugal and jugal densely covered by large circular cupules; 7) internal choanae anteriorly bound by palatines; 8) semi-arched and widely expanded premaxillae with fifth alveolus more laterally placed than the rest of the premaxillary tooth row; and 9) absence of contact between ectopterygoid and posterior margin of maxillary tooth row.
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</paragraph>
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<paragraph blockId="24.[130,777,1140,2028]" lastBlockId="25.[131,775,1491,2027]" lastPageId="25" lastPageNumber="706" pageId="24" pageNumber="705">
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The Angeac-Charente specimen (ANG18-5914, 5920, 5921 and 5925) also exhibits several characters that permit provisional assignment to the genus
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<taxonomicName authorityName="Owen" authorityYear="1841" box="[500,619,2001,2027]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
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<emphasis box="[500,619,2001,2027]" italics="true" pageId="24" pageNumber="705">Goniopholis</emphasis>
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</taxonomicName>
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. Among these, antorbital and interorbital ornamentation is informative. The presence of periorbital crests on the prefrontal and lacrimal are present in
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<taxonomicName authority="(Andrade & Hornung 2011)" baseAuthorityName="Andrade & Hornung" baseAuthorityYear="2011" box="[967,1411,1204,1230]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
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<emphasis box="[967,1090,1204,1230]" italics="true" pageId="24" pageNumber="705">Goniopholis</emphasis>
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(
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<bibRefCitation author="Andrade & Hornung" box="[1106,1402,1204,1230]" firstAuthor="Andrade" pageId="24" pageNumber="705" pagination="352 - 368" refId="ref35562" refString="ANDRADE M. B. & HORNUNG J. J. 2011. - A new look into the periorbital morphology of Goniopholis (Mesoeucrocodylia: Neosuchia) and related forms. Journal of Vertebrate Paleontology 31 (2): 352 - 368. https: // doi. org / 10.1080 / 02724634.2011.550353" type="journal article" year="2011">Andrade & Hornung 2011</bibRefCitation>
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)
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</taxonomicName>
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but absent in
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<taxonomicName authorityName="Salisbury & Naish" authorityYear="2011" box="[914,1137,1236,1262]" class="Reptilia" family="Goniopholididae" genus="Anteophthalmosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
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<emphasis box="[914,1137,1236,1262]" italics="true" pageId="24" pageNumber="705">Anteophthalmosuchus</emphasis>
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</taxonomicName>
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and
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<taxonomicName box="[1189,1312,1236,1262]" class="Reptilia" family="Goniopholididae" genus="Hulkepholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
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<emphasis box="[1189,1312,1236,1262]" italics="true" pageId="24" pageNumber="705">Hulkepholis</emphasis>
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</taxonomicName>
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(
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<bibRefCitation author="SALISBURY S. W. & NAISH D." journalOrPublisher="The Palaeontological Association, London" pageId="24" pageNumber="705" pagination="305 - 369" refId="ref48547" refString="SALISBURY S. W. & NAISH D. 2011. - Crocodilians, in EnglishWealden Fossils. The Palaeontological Association, London: 305 - 369." title="Crocodilians, in EnglishWealden Fossils" type="book chapter" year="2011">Salisbury & Naish 2011</bibRefCitation>
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;
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<bibRefCitation author="Buscalioni" box="[949,1177,1267,1294]" etAl="et al." firstAuthor="Buscalioni" pageId="24" pageNumber="705" pagination="103 - 122" refId="ref37078" refString="BUSCALIONI A. D., ALCALA L., ESPILEZ E. & MAMPEL L. 2013. - European Goniopholididae from the Early Albian Escucha Formation in Arino (Teruel, Aragon, Spain). Spanish Journal of Palaeontology 28: 103 - 122. https: // doi. org / 10.7203 / sjp. 28.1.17835" type="journal article" year="2013">
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Buscalioni
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<emphasis box="[1067,1116,1267,1293]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
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|
2013
|
|
</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="Martin" box="[1189,1393,1267,1293]" etAl="et al." firstAuthor="Martin" pageId="24" pageNumber="705" pagination="1222534" refId="ref43294" refString="MARTIN J. E., DELFINO M. & SMITH T. 2016 a. - Osteology and affinities of Dollo's goniopholidid (Mesoeucrocodylia) from the Early Cretaceous of Bernissart, Belgium. Journal of Vertebrate Paleontology 36 (6): e 1222534. https: // doi. org / 10.1080 / 0272 4634.2016. 1222534" type="journal article" year="2016" yearSuffix="a">
|
|
Martin
|
|
<emphasis box="[1271,1321,1267,1293]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
|
|
2016a
|
|
</bibRefCitation>
|
|
). The interorbital ridge is restricted to the frontal in
|
|
<taxonomicName box="[1333,1457,1299,1325]" class="Reptilia" family="Goniopholididae" genus="Hulkepholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1333,1457,1299,1325]" italics="true" pageId="24" pageNumber="705">Hulkepholis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation author="SALISBURY S. W. & NAISH D." box="[822,1097,1331,1357]" journalOrPublisher="The Palaeontological Association, London" pageId="24" pageNumber="705" pagination="305 - 369" refId="ref48547" refString="SALISBURY S. W. & NAISH D. 2011. - Crocodilians, in EnglishWealden Fossils. The Palaeontological Association, London: 305 - 369." title="Crocodilians, in EnglishWealden Fossils" type="book chapter" year="2011">Salisbury & Naish 2011</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="Buscalioni" box="[1114,1360,1331,1357]" etAl="et al." firstAuthor="Buscalioni" pageId="24" pageNumber="705" pagination="103 - 122" refId="ref37078" refString="BUSCALIONI A. D., ALCALA L., ESPILEZ E. & MAMPEL L. 2013. - European Goniopholididae from the Early Albian Escucha Formation in Arino (Teruel, Aragon, Spain). Spanish Journal of Palaeontology 28: 103 - 122. https: // doi. org / 10.7203 / sjp. 28.1.17835" type="journal article" year="2013">
|
|
Buscalioni
|
|
<emphasis box="[1239,1294,1331,1357]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
|
|
2013
|
|
</bibRefCitation>
|
|
). In the Angeac-Charente specimen, the presence of both antorbital and interorbital crests is also a character shared with
|
|
<taxonomicName baseAuthorityName="Andrade & Hornung" baseAuthorityYear="2011" box="[1324,1447,1395,1421]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1324,1447,1395,1421]" italics="true" pageId="24" pageNumber="705">Goniopholis</emphasis>
|
|
</taxonomicName>
|
|
. A short postorbital spine is present in the Angeac-Charente specimen, a condition similar to other specimens of the genus
|
|
<taxonomicName baseAuthorityName="Andrade & Hornung" baseAuthorityYear="2011" box="[811,934,1491,1517]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
|
|
<emphasis box="[811,934,1491,1517]" italics="true" pageId="24" pageNumber="705">Goniopholis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation author="SALISBURY S. W. & WILLIS P. M. A. & PEITZ S. & SANDER P. M." box="[953,1178,1491,1517]" journalOrPublisher="Special Papers in Palaeontology" pageId="24" pageNumber="705" pagination="121 - 148" part="60" refId="ref48575" refString="SALISBURY S. W., WILLIS P. M. A., PEITZ S. & SANDER P. M. 1999. - The crocodilian Goniopholis simus from the lower Cretaceous of north-western Germany. Special Papers in Palaeontology 60: 121 - 148." title="The crocodilian Goniopholis simus from the lower Cretaceous of north-western Germany" type="journal article" year="1999">
|
|
Salisbury
|
|
<emphasis box="[1060,1113,1491,1517]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
|
|
1999
|
|
</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="SCHWARZ D." box="[1193,1347,1491,1517]" journalOrPublisher="Palaeontology" pageId="24" pageNumber="705" pagination="185 - 208" part="45" refId="ref48771" refString="SCHWARZ D. 2002. - A New Species of Goniopholis from the Upper Jurassic of Portugal. Palaeontology 45: 185 - 208. https: // doi. org / 10.1111 / 1475 - 4983.00233" title="A New Species of Goniopholis from the Upper Jurassic of Portugal" type="journal article" year="2002">Schwarz 2002</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="ANDRADE M. B. & HORNUNG J. J." journalOrPublisher="Journal of Vertebrate Paleontology" pageId="24" pageNumber="705" pagination="352 - 368" part="31" refId="ref35562" refString="ANDRADE M. B. & HORNUNG J. J. 2011. - A new look into the periorbital morphology of Goniopholis (Mesoeucrocodylia: Neosuchia) and related forms. Journal of Vertebrate Paleontology 31 (2): 352 - 368. https: // doi. org / 10.1080 / 02724634.2011.550353" title="A new look into the periorbital morphology of Goniopholis (Mesoeucrocodylia: Neosuchia) and related forms" type="journal article" year="2011">
|
|
Andrade
|
|
<emphasis box="[813,861,1523,1549]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
|
|
2011
|
|
</bibRefCitation>
|
|
) but unlike
|
|
<taxonomicName authorityName="Salisbury & Naish" authorityYear="2011" box="[1041,1256,1523,1549]" class="Reptilia" family="Goniopholididae" genus="Anteophthalmosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1041,1256,1523,1549]" italics="true" pageId="24" pageNumber="705">Anteophthalmosuchus</emphasis>
|
|
</taxonomicName>
|
|
in which this spine is longer, almost contacting the anterior margin of the orbits (
|
|
<bibRefCitation author="SALISBURY S. W. & NAISH D." box="[822,1080,1586,1613]" journalOrPublisher="The Palaeontological Association, London" pageId="24" pageNumber="705" pagination="305 - 369" refId="ref48547" refString="SALISBURY S. W. & NAISH D. 2011. - Crocodilians, in EnglishWealden Fossils. The Palaeontological Association, London: 305 - 369." title="Crocodilians, in EnglishWealden Fossils" type="book chapter" year="2011">Salisbury & Naish 2011</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="Martin" box="[1093,1299,1586,1612]" etAl="et al." firstAuthor="Martin" pageId="24" pageNumber="705" pagination="1222534" refId="ref43294" refString="MARTIN J. E., DELFINO M. & SMITH T. 2016 a. - Osteology and affinities of Dollo's goniopholidid (Mesoeucrocodylia) from the Early Cretaceous of Bernissart, Belgium. Journal of Vertebrate Paleontology 36 (6): e 1222534. https: // doi. org / 10.1080 / 0272 4634.2016. 1222534" type="journal article" year="2016" yearSuffix="a">
|
|
Martin
|
|
<emphasis box="[1176,1226,1586,1612]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
|
|
2016a
|
|
</bibRefCitation>
|
|
). The Angeac-Charente specimen is also similar in its maxillary tooth count (20) to
|
|
<taxonomicName authority="(Andrade et al. 2011)" baseAuthorityName="Andrade" baseAuthorityYear="2011" box="[898,1350,1650,1677]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="species" species="kiplingi">
|
|
<emphasis box="[898,1109,1650,1676]" italics="true" pageId="24" pageNumber="705">Goniopholis kiplingi</emphasis>
|
|
(
|
|
<bibRefCitation author="ANDRADE M. B. & HORNUNG J. J." box="[1125,1341,1650,1676]" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="24" pageNumber="705" pagination="352 - 368" part="31" refId="ref35562" refString="ANDRADE M. B. & HORNUNG J. J. 2011. - A new look into the periorbital morphology of Goniopholis (Mesoeucrocodylia: Neosuchia) and related forms. Journal of Vertebrate Paleontology 31 (2): 352 - 368. https: // doi. org / 10.1080 / 02724634.2011.550353" title="A new look into the periorbital morphology of Goniopholis (Mesoeucrocodylia: Neosuchia) and related forms" type="journal article" year="2011">
|
|
Andrade
|
|
<emphasis box="[1225,1278,1650,1676]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
|
|
2011
|
|
</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
and with
|
|
<taxonomicName authorityName="Salisbury & Naish" authorityYear="2011" box="[814,1032,1682,1708]" class="Reptilia" family="Goniopholididae" genus="Anteophthalmosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
|
|
<emphasis box="[814,1032,1682,1708]" italics="true" pageId="24" pageNumber="705">Anteophthalmosuchus</emphasis>
|
|
</taxonomicName>
|
|
, which possesses 19 alveoli (
|
|
<bibRefCitation author="Martin" etAl="et al." firstAuthor="Martin" pageId="24" pageNumber="705" pagination="1222534" refId="ref43294" refString="MARTIN J. E., DELFINO M. & SMITH T. 2016 a. - Osteology and affinities of Dollo's goniopholidid (Mesoeucrocodylia) from the Early Cretaceous of Bernissart, Belgium. Journal of Vertebrate Paleontology 36 (6): e 1222534. https: // doi. org / 10.1080 / 0272 4634.2016. 1222534" type="journal article" year="2016" yearSuffix="a">
|
|
Martin
|
|
<emphasis box="[1407,1456,1682,1708]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
|
|
2016a
|
|
</bibRefCitation>
|
|
). The maxillary alveolar count has not been established for
|
|
<taxonomicName authority="(Salisbury et al. 1999)" baseAuthorityName="Salisbury" baseAuthorityYear="1999" box="[852,1292,1746,1772]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="species" species="simus">
|
|
<emphasis box="[852,1040,1746,1772]" italics="true" pageId="24" pageNumber="705">Goniopholis simus</emphasis>
|
|
(
|
|
<bibRefCitation author="SALISBURY S. W. & WILLIS P. M. A. & PEITZ S. & SANDER P. M." box="[1057,1281,1746,1772]" journalOrPublisher="Special Papers in Palaeontology" pageId="24" pageNumber="705" pagination="121 - 148" part="60" refId="ref48575" refString="SALISBURY S. W., WILLIS P. M. A., PEITZ S. & SANDER P. M. 1999. - The crocodilian Goniopholis simus from the lower Cretaceous of north-western Germany. Special Papers in Palaeontology 60: 121 - 148." title="The crocodilian Goniopholis simus from the lower Cretaceous of north-western Germany" type="journal article" year="1999">
|
|
Salisbury
|
|
<emphasis box="[1164,1217,1746,1772]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
|
|
1999
|
|
</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
. However, this count is less than that of the goniopholidid
|
|
<taxonomicName baseAuthorityName="Salisbury & Naish" baseAuthorityYear="2011" box="[1252,1447,1778,1804]" class="Reptilia" family="Goniopholididae" genus="Hulkepholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="species" species="willetti">
|
|
<emphasis box="[1252,1447,1778,1804]" italics="true" pageId="24" pageNumber="705">Hulkepholis willetti</emphasis>
|
|
</taxonomicName>
|
|
, in which at least 24 maxillary alveoli are present (
|
|
<bibRefCitation author="SALISBURY S. W. & NAISH D." journalOrPublisher="The Palaeontological Association, London" pageId="24" pageNumber="705" pagination="305 - 369" refId="ref48547" refString="SALISBURY S. W. & NAISH D. 2011. - Crocodilians, in EnglishWealden Fossils. The Palaeontological Association, London: 305 - 369." title="Crocodilians, in EnglishWealden Fossils" type="book chapter" year="2011">Salisbury & Naish 2011</bibRefCitation>
|
|
). Obvious differences in rostrum proportions are observed with
|
|
<taxonomicName authorityName="Buffetaut & Hutt" authorityYear="1980" box="[1004,1119,1873,1899]" class="Reptilia" family="Goniopholididae" genus="Vectisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1004,1119,1873,1899]" italics="true" pageId="24" pageNumber="705">Vectisuchus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName box="[1173,1296,1873,1899]" class="Reptilia" family="Goniopholididae" genus="Hulkepholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1173,1296,1873,1899]" italics="true" pageId="24" pageNumber="705">Hulkepholis</emphasis>
|
|
</taxonomicName>
|
|
, which possess long and narrow rostra (
|
|
<bibRefCitation author="Buffetaut & Hutt" box="[1068,1316,1905,1932]" firstAuthor="Buffetaut" pageId="24" pageNumber="705" pagination="385 - 390" refId="ref36950" refString="BUFFETAUT E. & HUTT S. 1980. - Vectisuchus leptognathus, n. g. n. sp., a slender-snouted goniopholid crocodilian from the Wealden of the Isle of Wight. Neues Jahrbuch fur Geologie und Palaontologie Monatshefte 7: 385 - 390. https: // doi. org / 10.1127 / njgpm / 1980 / 1980 / 385" type="journal article" year="1980">Buffetaut & Hutt 1980</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="SALISBURY S. W. & NAISH D." journalOrPublisher="The Palaeontological Association, London" pageId="24" pageNumber="705" pagination="305 - 369" refId="ref48547" refString="SALISBURY S. W. & NAISH D. 2011. - Crocodilians, in EnglishWealden Fossils. The Palaeontological Association, London: 305 - 369." title="Crocodilians, in EnglishWealden Fossils" type="book chapter" year="2011">Salisbury & Naish 2011</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="Buscalioni" box="[957,1196,1937,1963]" etAl="et al." firstAuthor="Buscalioni" pageId="24" pageNumber="705" pagination="103 - 122" refId="ref37078" refString="BUSCALIONI A. D., ALCALA L., ESPILEZ E. & MAMPEL L. 2013. - European Goniopholididae from the Early Albian Escucha Formation in Arino (Teruel, Aragon, Spain). Spanish Journal of Palaeontology 28: 103 - 122. https: // doi. org / 10.7203 / sjp. 28.1.17835" type="journal article" year="2013">
|
|
Buscalioni
|
|
<emphasis box="[1079,1132,1937,1963]" italics="true" pageId="24" pageNumber="705">et al.</emphasis>
|
|
2013
|
|
</bibRefCitation>
|
|
). As in
|
|
<taxonomicName authorityName="Salisbury & Naish" authorityYear="2011" class="Reptilia" family="Goniopholididae" genus="Anteophthalmosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="24" pageNumber="705">Anteophthalmosuchus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName baseAuthorityName="Andrade & Hornung" baseAuthorityYear="2011" box="[935,1057,1969,1995]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="24" pageNumber="705" phylum="Chordata" rank="genus">
|
|
<emphasis box="[935,1057,1969,1995]" italics="true" pageId="24" pageNumber="705">Goniopholis</emphasis>
|
|
</taxonomicName>
|
|
, the palpebral in the specimen from Angeac-Charente is small and not large and triangular as in
|
|
<taxonomicName box="[132,255,1491,1517]" class="Reptilia" family="Goniopholididae" genus="Hulkepholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="25" pageNumber="706" phylum="Chordata" rank="genus">
|
|
<emphasis box="[132,255,1491,1517]" italics="true" pageId="25" pageNumber="706">Hulkepholis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation author="SALISBURY S. W. & NAISH D." box="[270,526,1491,1517]" journalOrPublisher="The Palaeontological Association, London" pageId="25" pageNumber="706" pagination="305 - 369" refId="ref48547" refString="SALISBURY S. W. & NAISH D. 2011. - Crocodilians, in EnglishWealden Fossils. The Palaeontological Association, London: 305 - 369." title="Crocodilians, in EnglishWealden Fossils" type="book chapter" year="2011">Salisbury & Naish 2011</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="Buscalioni" box="[539,768,1491,1517]" etAl="et al." firstAuthor="Buscalioni" pageId="25" pageNumber="706" pagination="103 - 122" refId="ref37078" refString="BUSCALIONI A. D., ALCALA L., ESPILEZ E. & MAMPEL L. 2013. - European Goniopholididae from the Early Albian Escucha Formation in Arino (Teruel, Aragon, Spain). Spanish Journal of Palaeontology 28: 103 - 122. https: // doi. org / 10.7203 / sjp. 28.1.17835" type="journal article" year="2013">
|
|
Buscalioni
|
|
<emphasis box="[657,707,1491,1517]" italics="true" pageId="25" pageNumber="706">et al.</emphasis>
|
|
2013
|
|
</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="Martin" box="[132,336,1523,1549]" etAl="et al." firstAuthor="Martin" pageId="25" pageNumber="706" pagination="1222534" refId="ref43294" refString="MARTIN J. E., DELFINO M. & SMITH T. 2016 a. - Osteology and affinities of Dollo's goniopholidid (Mesoeucrocodylia) from the Early Cretaceous of Bernissart, Belgium. Journal of Vertebrate Paleontology 36 (6): e 1222534. https: // doi. org / 10.1080 / 0272 4634.2016. 1222534" type="journal article" year="2016" yearSuffix="a">
|
|
Martin
|
|
<emphasis box="[214,264,1523,1549]" italics="true" pageId="25" pageNumber="706">et al.</emphasis>
|
|
2016a
|
|
</bibRefCitation>
|
|
). Comparison with
|
|
<taxonomicName authorityName="Owen" authorityYear="1879" box="[549,686,1523,1549]" class="Reptilia" family="Goniopholididae" genus="Nannosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="25" pageNumber="706" phylum="Chordata" rank="genus">
|
|
<emphasis box="[549,686,1523,1549]" italics="true" pageId="25" pageNumber="706">Nannosuchus</emphasis>
|
|
</taxonomicName>
|
|
will not be discussed here as this taxon has been viewed either as a juvenile of
|
|
<taxonomicName authority="(Salisbury 2002)" baseAuthorityName="Salisbury" baseAuthorityYear="2002" box="[256,567,1586,1613]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="25" pageNumber="706" phylum="Chordata" rank="genus">
|
|
<emphasis box="[256,379,1586,1612]" italics="true" pageId="25" pageNumber="706">Goniopholis</emphasis>
|
|
(
|
|
<bibRefCitation author="SALISBURY S. W." box="[395,558,1586,1613]" journalOrPublisher="Special Papers in Palaeontology" pageId="25" pageNumber="706" pagination="121 - 144" part="68" refId="ref48512" refString="SALISBURY S. W. 2002. - Crocodilians from the lower Cretaceous (Berriasian) Purbeck Limestone Group of Dorset, Southern England. Special Papers in Palaeontology 68: 121 - 144." title="Crocodilians from the lower Cretaceous (Berriasian) Purbeck Limestone Group of Dorset, Southern England" type="journal article" year="2002">Salisbury 2002</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
or as a valid taxon (
|
|
<bibRefCitation author="ANDRADE M. B. & HORNUNG J. J." box="[141,355,1618,1644]" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="25" pageNumber="706" pagination="352 - 368" part="31" refId="ref35562" refString="ANDRADE M. B. & HORNUNG J. J. 2011. - A new look into the periorbital morphology of Goniopholis (Mesoeucrocodylia: Neosuchia) and related forms. Journal of Vertebrate Paleontology 31 (2): 352 - 368. https: // doi. org / 10.1080 / 02724634.2011.550353" title="A new look into the periorbital morphology of Goniopholis (Mesoeucrocodylia: Neosuchia) and related forms" type="journal article" year="2011">
|
|
Andrade
|
|
<emphasis box="[241,292,1618,1644]" italics="true" pageId="25" pageNumber="706">et al.</emphasis>
|
|
2011
|
|
</bibRefCitation>
|
|
). Therefore, considering the characters mentioned above, we tentatively refer the Angeac-Charente specimens to
|
|
<taxonomicName box="[278,438,1682,1709]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" kingdom="Animalia" order="Crocodylia" pageId="25" pageNumber="706" phylum="Chordata" rank="species" species="undetermined">
|
|
<emphasis box="[278,401,1682,1708]" italics="true" pageId="25" pageNumber="706">Goniopholis</emphasis>
|
|
sp.
|
|
</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.6902077" ID-Zenodo-Dep="6902077" httpUri="https://zenodo.org/record/6902077/files/figure.png" pageId="25" pageNumber="706" startId="25.[132,143,1323,1340]" targetBox="[132,1455,215,1279]" targetPageId="25">
|
|
<paragraph blockId="25.[132,1455,1323,1362]" pageId="25" pageNumber="706">
|
|
FIG. 18. — Goniopholidid crocodyliform Angeac-Charente:
|
|
<emphasis bold="true" box="[624,637,1323,1340]" pageId="25" pageNumber="706">A</emphasis>
|
|
,
|
|
<emphasis bold="true" box="[648,662,1323,1340]" pageId="25" pageNumber="706">D</emphasis>
|
|
, left dentary of
|
|
<taxonomicName box="[794,927,1323,1340]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" kingdom="Animalia" order="Crocodylia" pageId="25" pageNumber="706" phylum="Chordata" rank="species" species="undetermined">
|
|
<emphasis box="[794,894,1323,1340]" italics="true" pageId="25" pageNumber="706">Goniopholis</emphasis>
|
|
sp.
|
|
</taxonomicName>
|
|
(ANG18-5925) in dorsal (
|
|
<emphasis bold="true" box="[1138,1151,1323,1340]" pageId="25" pageNumber="706">A</emphasis>
|
|
) and ventral (
|
|
<emphasis bold="true" box="[1265,1279,1323,1340]" pageId="25" pageNumber="706">D</emphasis>
|
|
) views;
|
|
<emphasis bold="true" box="[1347,1360,1323,1340]" pageId="25" pageNumber="706">B</emphasis>
|
|
,
|
|
<emphasis bold="true" box="[1371,1385,1323,1340]" pageId="25" pageNumber="706">C</emphasis>
|
|
, skull of
|
|
<taxonomicName box="[132,264,1345,1362]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" kingdom="Animalia" order="Crocodylia" pageId="25" pageNumber="706" phylum="Chordata" rank="species" species="undetermined">
|
|
<emphasis box="[132,232,1345,1362]" italics="true" pageId="25" pageNumber="706">Goniopholis</emphasis>
|
|
sp.
|
|
</taxonomicName>
|
|
(ANG18-5914, ANG18-5920, ANG18-5921) in dorsal (
|
|
<emphasis bold="true" box="[712,725,1345,1362]" pageId="25" pageNumber="706">B</emphasis>
|
|
) and ventral (
|
|
<emphasis bold="true" box="[838,852,1345,1362]" pageId="25" pageNumber="706">C</emphasis>
|
|
) views. Scale bar: 10 cm.
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection pageId="25" pageNumber="706" type="discussion">
|
|
<paragraph blockId="25.[131,775,1491,2027]" pageId="25" pageNumber="706">
|
|
A detailed description of the Angeac-Charente material will be provided elsewhere. In the meantime, a study of the numerous goniopholid specimens from Cherves-de-Cognac and assigned to
|
|
<taxonomicName authority="(Raslan-Loubatie 2007)" baseAuthorityName="Raslan-Loubatie" baseAuthorityYear="2007" box="[318,725,1810,1836]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="25" pageNumber="706" phylum="Chordata" rank="genus">
|
|
<emphasis box="[318,447,1810,1836]" italics="true" pageId="25" pageNumber="706">Goniopholis</emphasis>
|
|
(Raslan-Loubatié 2007)
|
|
</taxonomicName>
|
|
will provide a useful basis for discussing morphological variability within a goniopholidid population. Preliminary observations concerning the Angeac-Charente specimen do not reveal any obvious morphological differences between the goniopholidids at both localities but further study is required.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |