487 lines
53 KiB
XML
487 lines
53 KiB
XML
<document id="F3630529A848487CB2589B0A2A3496F1" ID-DOI="10.11646/zootaxa.5486.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="13209805" ID-ZooBank="0B69BD10-AB31-456F-98E0-3073787C5DB6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1722787646408" checkinUser="plazi" docAuthor="Tchesunov, Alexei V. & Fedyaeva, Maria A." docDate="2024" docId="03C887B3FFFC43622ADE951AFA7DE491" docLanguage="en" docName="zootaxa.5486.1.5.pdf" docOrigin="Zootaxa 5486 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5486.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Angursa olenevskii Tchesunov & Fedyaeva, 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="127" masterDocId="FFF1FFCBFFFE43642A499451FFF6E614" masterDocTitle="A new species of Angursa (Tardigrada, Heterotardigrada) from the White Sea, North Russia" masterLastPageNumber="128" masterPageNumber="121" pageNumber="123" updateTime="1722950837948" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:titleInfo id="C7DD2BA753D54C4480D6915E8AD5B06B">
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<mods:title id="A166F5B02E4686069CBAE4F8E9FC4C90">A new species of Angursa (Tardigrada, Heterotardigrada) from the White Sea, North Russia</mods:title>
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<mods:name id="EE84879C9332AFCC93112BDA390B0F43" type="personal">
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<mods:roleTerm id="9E80D97043C0B1E9C58A4DC9BCA46FDA">Author</mods:roleTerm>
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<mods:namePart id="AC2FE5032367A1425E5F6084433F1029">Tchesunov, Alexei V.</mods:namePart>
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<mods:nameIdentifier id="77E556CE8922849C422E3AB83E303A98" type="ORCID">0000-0003-2365-910X</mods:nameIdentifier>
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<mods:affiliation id="903308EE6A2A5BA5CD028976177AF502">Department of Invertebrate Zoology, Faculty of Biology, M. V. Lomonosov’s Moscow University, 119234 Moscow, Russia</mods:affiliation>
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<mods:nameIdentifier id="0FB0BB6A2A4ACE4C03B0EE1BD18D0B1A" type="email">chesunov@yandex.ru</mods:nameIdentifier>
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<mods:roleTerm id="7C43F500FD20AC3EC92E43077D6117A0">Author</mods:roleTerm>
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<mods:namePart id="4149B503EA1CA8B3141EAB7384667698">Fedyaeva, Maria A.</mods:namePart>
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<mods:affiliation id="F070734F758748AE5193671EF4F1ACF1">N. A. Pertsov White Sea Biological Station, Faculty of Biology, M. V. Lomonosov’s Moscow University, 119234 Moscow, Russia</mods:affiliation>
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<mods:nameIdentifier id="A2475CBDAFF0C16BC694CC9965EDFCD2" type="email">mariaf92@mail.ru</mods:nameIdentifier>
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<mods:title id="3CE1638DC6F2248AEAF99E9506446C51">Zootaxa</mods:title>
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<mods:part id="E9D56BAC92A9897010D380337DC71A0E">
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<mods:date id="90EF5B06081D942CFC81FBEF49CD0B17">2024</mods:date>
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<mods:number id="E40FED982F04DBDD155DB4246A6843BC">2024-07-24</mods:number>
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<mods:number id="6032FB035323B1036A9793F1A0A02291">5486</mods:number>
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<mods:url id="9AED1DE3DCBDA5DE0FDA855AD342F05C">http://dx.doi.org/10.11646/zootaxa.5486.1.5</mods:url>
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<mods:classification id="E7739770E8DCEC8669F85B775939E360">journal article</mods:classification>
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<treatment id="03C887B3FFFC43622ADE951AFA7DE491" LSID="urn:lsid:plazi:treatment:03C887B3FFFC43622ADE951AFA7DE491" httpUri="http://treatment.plazi.org/id/03C887B3FFFC43622ADE951AFA7DE491" lastPageId="6" lastPageNumber="127" pageId="2" pageNumber="123">
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<subSubSection id="C37B652EFFFC43662ADE951AFE12E771" box="[151,484,330,357]" pageId="2" pageNumber="123" type="nomenclature">
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<paragraph id="8BDE36A5FFFC43662ADE951AFE12E771" blockId="2.[151,484,330,393]" box="[151,484,330,357]" pageId="2" pageNumber="123">
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<heading id="D09681C9FFFC43662ADE951AFE12E771" bold="true" box="[151,484,330,357]" fontSize="11" level="1" pageId="2" pageNumber="123" reason="1">
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<emphasis id="B915EAB7FFFC43662ADE951AFE12E771" bold="true" box="[151,484,330,357]" pageId="2" pageNumber="123">
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<taxonomicName id="4C614D26FFFC43662ADE951AFE76E771" authorityName="Tchesunov & Fedyaeva" authorityYear="2024" box="[151,384,330,357]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="123" phylum="Tardigrada" rank="species" species="olenevskii" status="sp. nov.">
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<emphasis id="B915EAB7FFFC43662ADE951AFE76E771" bold="true" box="[151,384,330,357]" italics="true" pageId="2" pageNumber="123">Angursa olenevskii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22657CCFFFC43662BCE951AFE12E771" box="[391,484,331,357]" pageId="2" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B652EFFFC43662ADE953EFEAFE79D" box="[151,345,367,393]" pageId="2" pageNumber="123" type="description">
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<paragraph id="8BDE36A5FFFC43662ADE953EFEAFE79D" blockId="2.[151,484,330,393]" box="[151,345,367,393]" pageId="2" pageNumber="123">
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<figureCitation id="135A2A20FFFC43662ADE953EFF0DE79D" box="[151,251,367,393]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[152,255,1971,1995]" captionStartId-1="3.[152,255,1424,1448]" captionStartId-2="5.[152,255,1414,1438]" captionTargetBox-0="[380,1210,677,1937]" captionTargetBox-1="[383,1209,194,1386]" captionTargetBox-2="[340,1247,181,1389]" captionTargetId-0="figure-127@2.[367,1219,669,1948]" captionTargetId-1="figure-17@3.[375,1213,181,1400]" captionTargetId-2="figure-17@5.[340,1247,181,1389]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURE 1. Angursa olenevskii sp. n., heads.A holotype female, dorsal view. B paratype 1, female, ventral view.The continuous punctations of the cuticle is shown just as a band. Scale bars 10 μm." captionText-1="FIGURE 2. Angursa olenevskii sp. n., details. A paratype 2, female, entire. B paratype 1, female 1, hind body ventrally. C holotype female, left leg IV, dorsal view. The continuous punctations of the cuticle is shown just as a band. Scale bars: A 20 μm, B and C 10 μm." captionText-2="FIGURE 3. Angursa olenevskii sp. n., SEM. A specimen 1 male, entire, ventral view. B specimen 2 female, entire, lateroventral view. C specimen 3, entire, lateral view. D specimen 1, left foot I, ventral view. E specimen I, right foot II, toes with claws withdrawn in hoods. F specimen 4, left foot IV, toes with claws protruded. G specimen 3, primary clava and cirrus A. H specimen 1, male gonopore and anus. Scale bars: A 10 μm, B and C 20 μm, D–F, H 2 μm, G 5 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13209807" figureDoi-1="http://doi.org/10.5281/zenodo.13209809" figureDoi-2="http://doi.org/10.5281/zenodo.13209811" httpUri-0="https://zenodo.org/record/13209807/files/figure.png" httpUri-1="https://zenodo.org/record/13209809/files/figure.png" httpUri-2="https://zenodo.org/record/13209811/files/figure.png" pageId="2" pageNumber="123">Figs 1–3</figureCitation>
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,
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<tableCitation id="C6E3031EFFFC43662B4F953EFEAFE79D" box="[262,345,367,393]" captionStart="TABLE 1" captionStartId="4.[151,239,909,933]" captionTargetPageId="4" captionText="TABLE 1. Morphometrics of Angursa olenevskii type specimens (μm)" pageId="2" pageNumber="123">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B652EFFFC43662ADE95E7FF24E491" pageId="2" pageNumber="123" type="diagnosis">
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<paragraph id="8BDE36A5FFFC43662ADE95E7FEE6E7C4" blockId="2.[151,272,438,464]" box="[151,272,438,464]" pageId="2" pageNumber="123">
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<emphasis id="B915EAB7FFFC43662ADE95E7FEE6E7C4" bold="true" box="[151,272,438,464]" pageId="2" pageNumber="123">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BDE36A5FFFC43662ADE95AEFF24E491" blockId="2.[151,1437,511,645]" pageId="2" pageNumber="123">
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<taxonomicName id="4C614D26FFFC43662ADE95AEFF00E40D" authorityName="Pollock" authorityYear="1979" box="[151,246,511,537]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="123" phylum="Tardigrada" rank="genus">
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<emphasis id="B915EAB7FFFC43662ADE95AEFF00E40D" box="[151,246,511,537]" italics="true" pageId="2" pageNumber="123">Angursa</emphasis>
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</taxonomicName>
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with ballon-shaped primary clava inserted on pedestal together with cirrus A; cirrus A two to three times longer than primary clava; secondary and tertiary clavae not evident; cirrus E simple, tapered to the end; legs I–III with equal setose sensory organs on coxae; external digits with fine peduncle along their length and attached to the claw.
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</paragraph>
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</subSubSection>
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||
<caption id="DF1E662DFFFC43662AD193E2FCBEE1E4" ID-DOI="http://doi.org/10.5281/zenodo.13209807" ID-Zenodo-Dep="13209807" httpUri="https://zenodo.org/record/13209807/files/figure.png" pageId="2" pageNumber="123" startId="2.[152,255,1971,1995]" targetBox="[380,1210,677,1937]" targetPageId="2" targetType="figure">
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<paragraph id="8BDE36A5FFFC43662AD193E2FCBEE1E4" blockId="2.[151,1436,1971,2032]" pageId="2" pageNumber="123">
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<emphasis id="B915EAB7FFFC43662AD193E2FEE1E1DF" bold="true" box="[152,279,1971,1995]" pageId="2" pageNumber="123">FIGURE 1.</emphasis>
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<taxonomicName id="4C614D26FFFC43662B5293E2FE2CE1DF" authorityName="Tchesunov & Fedyaeva" authorityYear="2024" box="[283,474,1971,1995]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="123" phylum="Tardigrada" rank="species" species="olenevskii" status="sp. nov.">
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<emphasis id="B915EAB7FFFC43662B5293E2FE2CE1DF" box="[283,474,1971,1995]" italics="true" pageId="2" pageNumber="123">Angursa olenevskii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22657CCFFFC43662B9693E5FDE3E1DF" box="[479,533,1972,1995]" pageId="2" pageNumber="123" rank="species">sp. n.</taxonomicNameLabel>
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, heads.
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<emphasis id="B915EAB7FFFC4366282A93E2FD83E1DF" bold="true" box="[611,629,1971,1995]" pageId="2" pageNumber="123">A</emphasis>
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holotype female, dorsal view.
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<emphasis id="B915EAB7FFFC436629ED93E2FC43E1DF" bold="true" box="[932,949,1971,1995]" pageId="2" pageNumber="123">B</emphasis>
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paratype 1, female, ventral view. The continuous punctations of the cuticle is shown just as a band. Scale bars 10 μm.
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</paragraph>
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</caption>
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<caption id="DF1E662DFFFD43672AD191C1FECFE3E5" ID-DOI="http://doi.org/10.5281/zenodo.13209809" ID-Zenodo-Dep="13209809" httpUri="https://zenodo.org/record/13209809/files/figure.png" pageId="3" pageNumber="124" startId="3.[152,255,1424,1448]" targetBox="[383,1209,194,1386]" targetPageId="3" targetType="figure">
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<paragraph id="8BDE36A5FFFD43672AD191C1FECFE3E5" blockId="3.[151,1436,1424,1521]" pageId="3" pageNumber="124">
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<emphasis id="B915EAB7FFFD43672AD191C1FEECE3BC" bold="true" box="[152,282,1424,1448]" pageId="3" pageNumber="124">FIGURE 2.</emphasis>
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<taxonomicName id="4C614D26FFFD43672B6A91C0FE10E3BC" authorityName="Tchesunov & Fedyaeva" authorityYear="2024" box="[291,486,1424,1448]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="124" phylum="Tardigrada" rank="species" species="olenevskii" status="sp. nov.">
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<emphasis id="B915EAB7FFFD43672B6A91C0FE10E3BC" box="[291,486,1424,1448]" italics="true" pageId="3" pageNumber="124">Angursa olenevskii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22657CCFFFD43672BA791C0FDDEE3BC" box="[494,552,1425,1448]" pageId="3" pageNumber="124" rank="species">sp. n.</taxonomicNameLabel>
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, details.
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<emphasis id="B915EAB7FFFD436728CF91C1FD6EE3BC" bold="true" box="[646,664,1424,1448]" pageId="3" pageNumber="124">A</emphasis>
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paratype 2, female, entire.
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<emphasis id="B915EAB7FFFD436729FE91C1FC3EE3BC" bold="true" box="[951,968,1424,1448]" pageId="3" pageNumber="124">B</emphasis>
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paratype 1, female 1, hind body ventrally.
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<emphasis id="B915EAB7FFFD43672FC391C1FA6AE3BC" bold="true" box="[1418,1436,1424,1448]" pageId="3" pageNumber="124">C</emphasis>
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holotype female, left leg IV, dorsal view. The continuous punctations of the cuticle is shown just as a band. Scale bars: A 20 μm, B and C 10 μm.
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</paragraph>
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</caption>
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<subSubSection id="C37B652EFFFD43672ADE9212FBA2E0B0" box="[151,1108,1603,1701]" pageId="3" pageNumber="124" type="etymology">
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<paragraph id="8BDE36A5FFFD43672ADE9212FEE8E049" blockId="3.[151,286,1603,1629]" box="[151,286,1603,1629]" pageId="3" pageNumber="124">
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<emphasis id="B915EAB7FFFD43672ADE9212FEE8E049" bold="true" box="[151,286,1603,1629]" pageId="3" pageNumber="124">Etymology</emphasis>
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</paragraph>
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<paragraph id="8BDE36A5FFFD43672ADE92DBFBA2E0B0" blockId="3.[151,1108,1674,1701]" box="[151,1108,1674,1701]" pageId="3" pageNumber="124">The species name refers to the Olenevski Island where the new tardigrade was found.</paragraph>
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</subSubSection>
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<subSubSection id="C37B652EFFFD43602ADE92A6FCF6E709" lastPageId="4" lastPageNumber="125" pageId="3" pageNumber="124" type="materials_examined">
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<paragraph id="8BDE36A5FFFD43672ADE92A6FEB0E104" blockId="3.[151,326,1782,1809]" box="[151,326,1782,1809]" pageId="3" pageNumber="124">
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<emphasis id="B915EAB7FFFD43672ADE92A6FEB0E104" bold="true" box="[151,326,1782,1809]" pageId="3" pageNumber="124">Type material</emphasis>
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</paragraph>
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<paragraph id="8BDE36A5FFFD43672ADE936FFE52E1D1" blockId="3.[151,1437,1854,1989]" pageId="3" pageNumber="124">
|
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<materialsCitation id="3B093CF8FFFD43672ADE936FFB5AE168" collectionCode="V" country="Russia" location="Lomonosov" municipality="University Zoological Museum" pageId="3" pageNumber="124" specimenCount="3" stateProvince="Moscow" typeStatus="holotype">
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||
<typeStatus id="54DA8807FFFD43672ADE936FFF09E14C" box="[151,255,1854,1880]" pageId="3" pageNumber="124" type="holotype">Holotype</typeStatus>
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||
(female) and
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<specimenCount id="9D67FD2CFFFD43672BDC936FFDCBE14D" box="[405,573,1854,1881]" count="3" pageId="3" pageNumber="124" type="generic" typeStatus="paratype">three paratypes</specimenCount>
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in permanent glycerine slide each are deposited in collection M.
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<collectionCode id="ED70AE60FFFD43672EB0936EFAF9E14D" box="[1273,1295,1855,1881]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="124" type="Museum">V</collectionCode>
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.
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<location id="8EBE607EFFFD43672F5F936EFA6AE14D" LSID="urn:lsid:plazi:treatment:03C887B3FFFC43622ADE951AFA7DE491:8EBE607EFFFD43672F5F936EFA6AE14D" box="[1302,1436,1855,1881]" country="Russia" municipality="University Zoological Museum" name="Lomonosov" pageId="3" pageNumber="124" stateProvince="Moscow">Lomonosov</location>
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||
Moscow State
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||
<collectingMunicipality id="6BBAACDFFFFD43672B0F9333FD46E169" box="[326,688,1890,1917]" pageId="3" pageNumber="124">University Zoological Museum</collectingMunicipality>
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,
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<collectingRegion id="49A5F847FFFD436728F69332FCD7E169" box="[703,801,1891,1917]" country="Russia" name="Moscow" pageId="3" pageNumber="124">Moscow</collectingRegion>
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,
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||
<collectingCountry id="F3767635FFFD436729799333FC8DE168" box="[816,891,1890,1916]" name="Russia" pageId="3" pageNumber="124">Russia</collectingCountry>
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under inventory numbers
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||
</materialsCitation>
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||
:
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||
<materialsCitation id="3B093CF8FFFD43672EF39333FEC9E1B5" collectionCode="ZMMU" country="Russia" pageId="3" pageNumber="124" specimenCount="1" stateProvince="Moscow" typeStatus="holotype">
|
||
<typeStatus id="54DA8807FFFD43672EF39333FAD3E168" box="[1210,1317,1890,1916]" pageId="3" pageNumber="124" type="holotype">Holotype</typeStatus>
|
||
, female— 1
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||
<collectionCode id="ED70AE60FFFD43672AF993D6FEF1E1B5" box="[176,263,1927,1953]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="124" type="Museum">ZMMU</collectionCode>
|
||
R-2
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||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B093CF8FFFD43672B0793D6FD1DE1B5" box="[334,747,1926,1953]" collectionCode="ZMMU" country="Russia" pageId="3" pageNumber="124" specimenCount="1" stateProvince="Moscow" typeStatus="paratype">
|
||
<typeStatus id="54DA8807FFFD43672B0793D6FE46E1B5" box="[334,432,1927,1953]" pageId="3" pageNumber="124" type="paratype">Paratype</typeStatus>
|
||
1, female—2
|
||
<collectionCode id="ED70AE60FFFD4367281493D6FD42E1B5" box="[605,692,1927,1953]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="124" type="Museum">ZMMU</collectionCode>
|
||
R-3
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B093CF8FFFD436728B293D6FAFDE1B5" box="[763,1291,1926,1953]" collectionCode="ZMMU" country="Russia" pageId="3" pageNumber="124" specimenCount="1" stateProvince="Moscow" typeStatus="paratype">
|
||
<typeStatus id="54DA8807FFFD436728B293D6FCABE1B5" box="[763,861,1927,1953]" pageId="3" pageNumber="124" type="paratype">Paratype</typeStatus>
|
||
2, gender unknown—2
|
||
<collectionCode id="ED70AE60FFFD43672E3593D6FB25E1B5" box="[1148,1235,1927,1953]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="124" type="Museum">ZMMU</collectionCode>
|
||
R-4
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B093CF8FFFD43672F5393D6FE56E1D1" collectionCode="ZMMU" country="Russia" pageId="3" pageNumber="124" specimenCount="1" stateProvince="Moscow" typeStatus="paratype">
|
||
<typeStatus id="54DA8807FFFD43672F5393D6FA8AE1B5" box="[1306,1404,1927,1953]" pageId="3" pageNumber="124" type="paratype">Paratype</typeStatus>
|
||
3, female—4
|
||
<collectionCode id="ED70AE60FFFD43672B5D93FAFE9DE1D1" box="[276,363,1963,1989]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="124" type="Museum">ZMMU</collectionCode>
|
||
R-5
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDE36A5FFFA43602ADE94C6FEC1E6A5" blockId="4.[151,311,151,177]" box="[151,311,151,177]" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602ADE94C6FEC1E6A5" bold="true" box="[151,311,151,177]" pageId="4" pageNumber="125">Type locality</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDE36A5FFFA43602ADE948EFCF6E709" blockId="4.[151,1436,223,286]" pageId="4" pageNumber="125">
|
||
White Sea, Kandalaksha Bay, Karelian coast, Olenevski Island,
|
||
<geoCoordinate id="EE555062FFFA436029CB948EFC14E6EE" box="[898,994,223,250]" degrees="66" direction="north" minutes="52" orientation="latitude" pageId="4" pageNumber="125" precision="925" value="66.86667">66°52’N</geoCoordinate>
|
||
and
|
||
<geoCoordinate id="EE555062FFFA43602E69948EFB89E6ED" box="[1056,1151,223,249]" degrees="33" direction="east" minutes="11" orientation="longitude" pageId="4" pageNumber="125" precision="925" value="33.183334">33°11’E</geoCoordinate>
|
||
, intertidal sandy littoral, medium sand,
|
||
<quantity id="4C999B40FFFA43602B729552FE5CE70A" box="[315,426,259,286]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="4" pageNumber="125" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 cm</quantity>
|
||
sediment depth.
|
||
<date id="FFDF1065FFFA436028219552FD0DE709" box="[616,763,259,285]" pageId="4" pageNumber="125" value="2021-08">August 2021</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B652EFFFA43622ADE953EFCE5E6C1" lastPageId="6" lastPageNumber="127" pageId="4" pageNumber="125" type="description">
|
||
<paragraph id="8BDE36A5FFFA43602ADE953EFEDEE79D" blockId="4.[151,296,367,393]" box="[151,296,367,393]" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602ADE953EFEDEE79D" bold="true" box="[151,296,367,393]" pageId="4" pageNumber="125">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDE36A5FFFA43602ADE95E6FB89E7E1" blockId="4.[151,1437,439,861]" pageId="4" pageNumber="125">Body elongate, cylindrical, its length to width ratio 3.6–4.5. Body cuticle smooth, without sculpture and surface differentiations, but looking indistinctly punctated because of dense intracuticular pillars.</paragraph>
|
||
<paragraph id="8BDE36A5FFFA43602A8E95AEFCC9E549" blockId="4.[151,1437,439,861]" pageId="4" pageNumber="125">
|
||
Cephalic region provided with complete set of appendages. All the appendages, except cA and primary clavae consist of two parts with indistinct borders, a high slim basal truncate cone (scapus) and a distal acute cirrus. The basal cones and pedestals rise up smoothly from the body surface. Internal cirri situated a bit dorsally and a bit more close to one another than the external cirri. Cirrus A and primary clava inserted together on the stout truncate conical pedestal, the clava just posterior and ventral to the cirrus A. The primary clava generally balloon-shaped. Cirrus A exceeds the clava two to three times in length (fig. 1 A, B; 3 G). Secondary and tertiary clavae not visible in light nor in scanning electron microscope. There is a pair of small, slightly pigmented bodies (ocelli) situated interiorly just posterior to the cirrus A+clava pedestals (fig. 1 A); the ocelli distinctly visible but in only
|
||
<typeStatus id="54DA8807FFFA43602EC296AAFB1BE501" box="[1163,1261,763,789]" pageId="4" pageNumber="125" type="holotype">holotype</typeStatus>
|
||
female. Cirri E situated dorso-laterally close to posterior margin of the body and consist also of basal truncate conical pedestal and long pointed flagellar cirrus without annulation (fig. 2 A, C).
|
||
</paragraph>
|
||
<caption id="DF1E662DFFFA43602ADE97DCFC7DE5B1" box="[151,907,909,933]" pageId="4" pageNumber="125" startId="4.[151,239,909,933]" targetBox="[163,1423,948,1796]" targetIsTable="true" targetPageId="4" targetType="table">
|
||
<paragraph id="8BDE36A5FFFA43602ADE97DCFC7DE5B1" blockId="4.[151,907,909,933]" box="[151,907,909,933]" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602ADE97DCFEFCE5B1" bold="true" box="[151,266,909,933]" pageId="4" pageNumber="125">TABLE 1.</emphasis>
|
||
Morphometrics of
|
||
<taxonomicName id="4C614D26FFFA43602B9097DCFD53E5B1" authorityName="Tchesunov & Fedyaeva" authorityYear="2024" box="[473,677,909,933]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="4" pageNumber="125" phylum="Tardigrada" rank="species" species="olenevskii">
|
||
<emphasis id="B915EAB7FFFA43602B9097DCFD53E5B1" box="[473,677,909,933]" italics="true" pageId="4" pageNumber="125">Angursa olenevskii</emphasis>
|
||
</taxonomicName>
|
||
type specimens (μm)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDE36A5FFFA43602AEA97E5FB2BE117" pageId="4" pageNumber="125">
|
||
<table id="F961C405FFFABC9B2AEA97E5FA79E110" box="[163,1423,948,1796]" gridcols="5" gridrows="20" pageId="4" pageNumber="125">
|
||
<tr id="355134E7FFFABC9B2AEA97E5FA79E5FE" box="[163,1423,948,1002]" gridrow="0" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA97E5FE31E5FE" box="[163,455,948,1002]" gridcol="0" gridrow="0" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA97E5FEE4E5D8" bold="true" box="[163,274,948,972]" pageId="4" pageNumber="125">Character</emphasis>
|
||
</th>
|
||
<th id="76805D9BFFFABC9B2BAA97E5FD6DE5FE" box="[483,667,948,1002]" gridcol="1" gridrow="0" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602BAA97E4FD6DE5D8" bold="true" box="[483,667,948,973]" pageId="4" pageNumber="125">Holotype, female</emphasis>
|
||
</th>
|
||
<th id="76805D9BFFFABC9B288F97E5FCB4E5FE" box="[710,834,948,1002]" gridcol="2" gridrow="0" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA4360288F97E4FCF8E5FE" bold="true" pageId="4" pageNumber="125">Paratype 1, female</emphasis>
|
||
</th>
|
||
<th id="76805D9BFFFABC9B29DF97E5FB94E5FE" box="[918,1122,948,1002]" gridcol="3" gridrow="0" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA436029DF97E4FC0DE5FE" bold="true" pageId="4" pageNumber="125">Paratype 2, gender unknown</emphasis>
|
||
</th>
|
||
<th id="76805D9BFFFABC9B2E8D97E5FA79E5FE" box="[1220,1423,948,1002]" gridcol="4" gridrow="0" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602E8D97E4FA79E5D8" bold="true" box="[1220,1423,948,973]" pageId="4" pageNumber="125">Paratype 3, female</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA97ABFA79E206" box="[163,1423,1018,1042]" gridrow="1" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA97ABFE31E206" box="[163,455,1018,1042]" gridcol="0" gridrow="1" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA97ABFE8CE206" bold="true" box="[163,378,1018,1042]" pageId="4" pageNumber="125">Position to observer</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA97ABFD6DE206" box="[483,667,1018,1042]" gridcol="1" gridrow="1" pageId="4" pageNumber="125">dorsal</td>
|
||
<td id="76805D9BFFFABC9B288F97ABFCB4E206" box="[710,834,1018,1042]" gridcol="2" gridrow="1" pageId="4" pageNumber="125">ventral</td>
|
||
<td id="76805D9BFFFABC9B29DF97ABFB94E206" box="[918,1122,1018,1042]" gridcol="3" gridrow="1" pageId="4" pageNumber="125">lateral</td>
|
||
<td id="76805D9BFFFABC9B2E8D97ABFA79E206" box="[1220,1423,1018,1042]" gridcol="4" gridrow="1" pageId="4" pageNumber="125">ventral</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA9073FA79E22F" box="[163,1423,1058,1083]" gridrow="2" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA9073FE31E22F" box="[163,455,1058,1083]" gridcol="0" gridrow="2" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA9073FED3E22E" bold="true" box="[163,293,1058,1082]" pageId="4" pageNumber="125">Body length</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA9073FD6DE22F" box="[483,667,1058,1083]" gridcol="1" gridrow="2" pageId="4" pageNumber="125">158</td>
|
||
<td id="76805D9BFFFABC9B288F9073FCB4E22F" box="[710,834,1058,1083]" gridcol="2" gridrow="2" pageId="4" pageNumber="125">131</td>
|
||
<td id="76805D9BFFFABC9B29DF9073FB94E22F" box="[918,1122,1058,1083]" gridcol="3" gridrow="2" pageId="4" pageNumber="125">107</td>
|
||
<td id="76805D9BFFFABC9B2E8D9073FA79E22F" box="[1220,1423,1058,1083]" gridcol="4" gridrow="2" pageId="4" pageNumber="125">157</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA901BFA79E294" box="[163,1423,1098,1152]" gridrow="3" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA901BFE31E294" box="[163,455,1098,1152]" gridcol="0" gridrow="3" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA901BFED5E26B" bold="true" pageId="4" pageNumber="125">Body width between II and III leg pairs</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA901BFD6DE294" box="[483,667,1098,1152]" gridcol="1" gridrow="3" pageId="4" pageNumber="125">38</td>
|
||
<td id="76805D9BFFFABC9B288F901BFCB4E294" box="[710,834,1098,1152]" gridcol="2" gridrow="3" pageId="4" pageNumber="125">31</td>
|
||
<td id="76805D9BFFFABC9B29DF901BFB94E294" box="[918,1122,1098,1152]" gridcol="3" gridrow="3" pageId="4" pageNumber="125">30</td>
|
||
<td id="76805D9BFFFABC9B2E8D901BFA79E294" box="[1220,1423,1098,1152]" gridcol="4" gridrow="3" pageId="4" pageNumber="125">35</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA90DEFA79E2BC" box="[163,1423,1167,1192]" gridrow="4" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA90DEFE31E2BC" box="[163,455,1167,1192]" gridcol="0" gridrow="4" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA90DEFECBE2B3" bold="true" box="[163,317,1167,1191]" pageId="4" pageNumber="125">Median cirrus</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA90DEFD6DE2BC" box="[483,667,1167,1192]" gridcol="1" gridrow="4" pageId="4" pageNumber="125">?</td>
|
||
<td id="76805D9BFFFABC9B288F90DEFCB4E2BC" box="[710,834,1167,1192]" gridcol="2" gridrow="4" pageId="4" pageNumber="125">5</td>
|
||
<td id="76805D9BFFFABC9B29DF90DEFB94E2BC" box="[918,1122,1167,1192]" gridcol="3" gridrow="4" pageId="4" pageNumber="125">8.1</td>
|
||
<td id="76805D9BFFFABC9B2E8D90DEFA79E2BC" box="[1220,1423,1167,1192]" gridcol="4" gridrow="4" pageId="4" pageNumber="125">?</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA90E9FA79E2C4" box="[163,1423,1208,1232]" gridrow="5" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA90E9FE31E2C4" box="[163,455,1208,1232]" gridcol="0" gridrow="5" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA90E9FEB5E2C4" bold="true" box="[163,323,1208,1232]" pageId="4" pageNumber="125">Internal cirrus</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA90E9FD6DE2C4" box="[483,667,1208,1232]" gridcol="1" gridrow="5" pageId="4" pageNumber="125">11</td>
|
||
<td id="76805D9BFFFABC9B288F90E9FCB4E2C4" box="[710,834,1208,1232]" gridcol="2" gridrow="5" pageId="4" pageNumber="125">7.8</td>
|
||
<td id="76805D9BFFFABC9B29DF90E9FB94E2C4" box="[918,1122,1208,1232]" gridcol="3" gridrow="5" pageId="4" pageNumber="125">9.8</td>
|
||
<td id="76805D9BFFFABC9B2E8D90E9FA79E2C4" box="[1220,1423,1208,1232]" gridcol="4" gridrow="5" pageId="4" pageNumber="125">11</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA90B1FA79E2ED" box="[163,1423,1248,1273]" gridrow="6" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA90B1FE31E2ED" box="[163,455,1248,1273]" gridcol="0" gridrow="6" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA90B1FEBEE2EC" bold="true" box="[163,328,1248,1272]" pageId="4" pageNumber="125">External cirrus</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA90B1FD6DE2ED" box="[483,667,1248,1273]" gridcol="1" gridrow="6" pageId="4" pageNumber="125">8</td>
|
||
<td id="76805D9BFFFABC9B288F90B1FCB4E2ED" box="[710,834,1248,1273]" gridcol="2" gridrow="6" pageId="4" pageNumber="125">9.8</td>
|
||
<td id="76805D9BFFFABC9B29DF90B1FB94E2ED" box="[918,1122,1248,1273]" gridcol="3" gridrow="6" pageId="4" pageNumber="125">8.1</td>
|
||
<td id="76805D9BFFFABC9B2E8D90B1FA79E2ED" box="[1220,1423,1248,1273]" gridcol="4" gridrow="6" pageId="4" pageNumber="125">8</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA9159FA79E335" box="[163,1423,1288,1313]" gridrow="7" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA9159FE31E335" box="[163,455,1288,1313]" gridcol="0" gridrow="7" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA9159FEF7E334" bold="true" box="[163,257,1288,1312]" pageId="4" pageNumber="125">Cirrus A</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA9159FD6DE335" box="[483,667,1288,1313]" gridcol="1" gridrow="7" pageId="4" pageNumber="125">21</td>
|
||
<td id="76805D9BFFFABC9B288F9159FCB4E335" box="[710,834,1288,1313]" gridcol="2" gridrow="7" pageId="4" pageNumber="125">13.4</td>
|
||
<td id="76805D9BFFFABC9B29DF9159FB94E335" box="[918,1122,1288,1313]" gridcol="3" gridrow="7" pageId="4" pageNumber="125">21</td>
|
||
<td id="76805D9BFFFABC9B2E8D9159FA79E335" box="[1220,1423,1288,1313]" gridcol="4" gridrow="7" pageId="4" pageNumber="125">21</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA9161FA79E35D" box="[163,1423,1328,1353]" gridrow="8" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA9161FE31E35D" box="[163,455,1328,1353]" gridcol="0" gridrow="8" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA9161FECAE35D" bold="true" box="[163,316,1328,1353]" pageId="4" pageNumber="125">Primary clava</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA9161FD6DE35D" box="[483,667,1328,1353]" gridcol="1" gridrow="8" pageId="4" pageNumber="125">9.5</td>
|
||
<td id="76805D9BFFFABC9B288F9161FCB4E35D" box="[710,834,1328,1353]" gridcol="2" gridrow="8" pageId="4" pageNumber="125">6.8</td>
|
||
<td id="76805D9BFFFABC9B29DF9161FB94E35D" box="[918,1122,1328,1353]" gridcol="3" gridrow="8" pageId="4" pageNumber="125">7</td>
|
||
<td id="76805D9BFFFABC9B2E8D9161FA79E35D" box="[1220,1423,1328,1353]" gridcol="4" gridrow="8" pageId="4" pageNumber="125">9.5</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA9109FA79E365" box="[163,1423,1368,1393]" gridrow="9" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA9109FE31E365" box="[163,455,1368,1393]" gridcol="0" gridrow="9" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA9109FEF4E365" bold="true" box="[163,258,1368,1393]" pageId="4" pageNumber="125">Cirrus E</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA9109FD6DE365" box="[483,667,1368,1393]" gridcol="1" gridrow="9" pageId="4" pageNumber="125">31</td>
|
||
<td id="76805D9BFFFABC9B288F9109FCB4E365" box="[710,834,1368,1393]" gridcol="2" gridrow="9" pageId="4" pageNumber="125">21</td>
|
||
<td id="76805D9BFFFABC9B29DF9109FB94E365" box="[918,1122,1368,1393]" gridcol="3" gridrow="9" pageId="4" pageNumber="125">27.2</td>
|
||
<td id="76805D9BFFFABC9B2E8D9109FA79E365" box="[1220,1423,1368,1393]" gridcol="4" gridrow="9" pageId="4" pageNumber="125">31</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA91D0FA79E38E" box="[163,1423,1409,1434]" gridrow="10" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA91D0FE31E38E" box="[163,455,1409,1434]" gridcol="0" gridrow="10" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA91D0FE80E38D" bold="true" box="[163,374,1409,1433]" pageId="4" pageNumber="125">Sense organ of leg I</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA91D0FD6DE38E" box="[483,667,1409,1434]" gridcol="1" gridrow="10" pageId="4" pageNumber="125">7.8</td>
|
||
<td id="76805D9BFFFABC9B288F91D0FCB4E38E" box="[710,834,1409,1434]" gridcol="2" gridrow="10" pageId="4" pageNumber="125">6</td>
|
||
<td id="76805D9BFFFABC9B29DF91D0FB94E38E" box="[918,1122,1409,1434]" gridcol="3" gridrow="10" pageId="4" pageNumber="125">?</td>
|
||
<td id="76805D9BFFFABC9B2E8D91D0FA79E38E" box="[1220,1423,1409,1434]" gridcol="4" gridrow="10" pageId="4" pageNumber="125">7.8</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA91F8FA79E3D6" box="[163,1423,1449,1474]" gridrow="11" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA91F8FE31E3D6" box="[163,455,1449,1474]" gridcol="0" gridrow="11" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA91F8FE76E3D5" bold="true" box="[163,384,1449,1473]" pageId="4" pageNumber="125">Sense organ of leg II</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA91F8FD6DE3D6" box="[483,667,1449,1474]" gridcol="1" gridrow="11" pageId="4" pageNumber="125">7.2</td>
|
||
<td id="76805D9BFFFABC9B288F91F8FCB4E3D6" box="[710,834,1449,1474]" gridcol="2" gridrow="11" pageId="4" pageNumber="125">7.9</td>
|
||
<td id="76805D9BFFFABC9B29DF91F8FB94E3D6" box="[918,1122,1449,1474]" gridcol="3" gridrow="11" pageId="4" pageNumber="125">7.9</td>
|
||
<td id="76805D9BFFFABC9B2E8D91F8FA79E3D6" box="[1220,1423,1449,1474]" gridcol="4" gridrow="11" pageId="4" pageNumber="125">7.2</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA9180FA79E3FE" box="[163,1423,1489,1514]" gridrow="12" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA9180FE31E3FE" box="[163,455,1489,1514]" gridcol="0" gridrow="12" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA9180FE7CE3FE" bold="true" box="[163,394,1489,1514]" pageId="4" pageNumber="125">Sense organ of leg III</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA9180FD6DE3FE" box="[483,667,1489,1514]" gridcol="1" gridrow="12" pageId="4" pageNumber="125">11.3</td>
|
||
<td id="76805D9BFFFABC9B288F9180FCB4E3FE" box="[710,834,1489,1514]" gridcol="2" gridrow="12" pageId="4" pageNumber="125">9.3</td>
|
||
<td id="76805D9BFFFABC9B29DF9180FB94E3FE" box="[918,1122,1489,1514]" gridcol="3" gridrow="12" pageId="4" pageNumber="125">9.4</td>
|
||
<td id="76805D9BFFFABC9B2E8D9180FA79E3FE" box="[1220,1423,1489,1514]" gridcol="4" gridrow="12" pageId="4" pageNumber="125">11.3</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA91A8FA79E006" box="[163,1423,1529,1554]" gridrow="13" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA91A8FE31E006" box="[163,455,1529,1554]" gridcol="0" gridrow="13" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA91A8FE7EE006" bold="true" box="[163,392,1529,1554]" pageId="4" pageNumber="125">Sense organ of leg IV</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA91A8FD6DE006" box="[483,667,1529,1554]" gridcol="1" gridrow="13" pageId="4" pageNumber="125">6.5</td>
|
||
<td id="76805D9BFFFABC9B288F91A8FCB4E006" box="[710,834,1529,1554]" gridcol="2" gridrow="13" pageId="4" pageNumber="125">5.6</td>
|
||
<td id="76805D9BFFFABC9B29DF91A8FB94E006" box="[918,1122,1529,1554]" gridcol="3" gridrow="13" pageId="4" pageNumber="125">5.5</td>
|
||
<td id="76805D9BFFFABC9B2E8D91A8FA79E006" box="[1220,1423,1529,1554]" gridcol="4" gridrow="13" pageId="4" pageNumber="125">6.5</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA9273FA79E02E" box="[163,1423,1570,1594]" gridrow="14" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA9273FE31E02E" box="[163,455,1570,1594]" gridcol="0" gridrow="14" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA9273FEDAE02E" bold="true" box="[163,300,1570,1594]" pageId="4" pageNumber="125">Claw of leg I</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA9273FD6DE02E" box="[483,667,1570,1594]" gridcol="1" gridrow="14" pageId="4" pageNumber="125">2</td>
|
||
<td id="76805D9BFFFABC9B288F9273FCB4E02E" box="[710,834,1570,1594]" gridcol="2" gridrow="14" pageId="4" pageNumber="125">2.4</td>
|
||
<td id="76805D9BFFFABC9B29DF9273FB94E02E" box="[918,1122,1570,1594]" gridcol="3" gridrow="14" pageId="4" pageNumber="125">2.7</td>
|
||
<td id="76805D9BFFFABC9B2E8D9273FA79E02E" box="[1220,1423,1570,1594]" gridcol="4" gridrow="14" pageId="4" pageNumber="125">2</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA921BFA79E077" box="[163,1423,1610,1635]" gridrow="15" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA921BFE31E077" box="[163,455,1610,1635]" gridcol="0" gridrow="15" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA921BFEC0E076" bold="true" box="[163,310,1610,1634]" pageId="4" pageNumber="125">Claw of leg II</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA921BFD6DE077" box="[483,667,1610,1635]" gridcol="1" gridrow="15" pageId="4" pageNumber="125">2.6</td>
|
||
<td id="76805D9BFFFABC9B288F921BFCB4E077" box="[710,834,1610,1635]" gridcol="2" gridrow="15" pageId="4" pageNumber="125">2.4</td>
|
||
<td id="76805D9BFFFABC9B29DF921BFB94E077" box="[918,1122,1610,1635]" gridcol="3" gridrow="15" pageId="4" pageNumber="125">2.7</td>
|
||
<td id="76805D9BFFFABC9B2E8D921BFA79E077" box="[1220,1423,1610,1635]" gridcol="4" gridrow="15" pageId="4" pageNumber="125">2.6</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA9223FA79E09F" box="[163,1423,1650,1675]" gridrow="16" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA9223FE31E09F" box="[163,455,1650,1675]" gridcol="0" gridrow="16" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA9223FEDAE09F" bold="true" box="[163,300,1650,1675]" pageId="4" pageNumber="125">Claw of leg I</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA9223FD6DE09F" box="[483,667,1650,1675]" gridcol="1" gridrow="16" pageId="4" pageNumber="125">2.6</td>
|
||
<td id="76805D9BFFFABC9B288F9223FCB4E09F" box="[710,834,1650,1675]" gridcol="2" gridrow="16" pageId="4" pageNumber="125">2.4</td>
|
||
<td id="76805D9BFFFABC9B29DF9223FB94E09F" box="[918,1122,1650,1675]" gridcol="3" gridrow="16" pageId="4" pageNumber="125">3.7</td>
|
||
<td id="76805D9BFFFABC9B2E8D9223FA79E09F" box="[1220,1423,1650,1675]" gridcol="4" gridrow="16" pageId="4" pageNumber="125">2.6</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA92CBFA79E0A7" box="[163,1423,1690,1715]" gridrow="17" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA92CBFE31E0A7" box="[163,455,1690,1715]" gridcol="0" gridrow="17" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA92CBFEDAE0A7" bold="true" box="[163,300,1690,1715]" pageId="4" pageNumber="125">Claw of leg I</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA92CBFD6DE0A7" box="[483,667,1690,1715]" gridcol="1" gridrow="17" pageId="4" pageNumber="125">2.9</td>
|
||
<td id="76805D9BFFFABC9B288F92CBFCB4E0A7" box="[710,834,1690,1715]" gridcol="2" gridrow="17" pageId="4" pageNumber="125">2.8</td>
|
||
<td id="76805D9BFFFABC9B29DF92CBFB94E0A7" box="[918,1122,1690,1715]" gridcol="3" gridrow="17" pageId="4" pageNumber="125">3.8</td>
|
||
<td id="76805D9BFFFABC9B2E8D92CBFA79E0A7" box="[1220,1423,1690,1715]" gridcol="4" gridrow="17" pageId="4" pageNumber="125">2.9</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA9292FA79E0CF" box="[163,1423,1731,1755]" gridrow="18" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA9292FE31E0CF" box="[163,455,1731,1755]" gridcol="0" gridrow="18" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA9292FED7E0CF" bold="true" box="[163,289,1731,1755]" pageId="4" pageNumber="125">Bulb length</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA9292FD6DE0CF" box="[483,667,1731,1755]" gridcol="1" gridrow="18" pageId="4" pageNumber="125">16</td>
|
||
<td id="76805D9BFFFABC9B288F9292FCB4E0CF" box="[710,834,1731,1755]" gridcol="2" gridrow="18" pageId="4" pageNumber="125">?</td>
|
||
<td id="76805D9BFFFABC9B29DF9292FB94E0CF" box="[918,1122,1731,1755]" gridcol="3" gridrow="18" pageId="4" pageNumber="125">14</td>
|
||
<td id="76805D9BFFFABC9B2E8D9292FA79E0CF" box="[1220,1423,1731,1755]" gridcol="4" gridrow="18" pageId="4" pageNumber="125">19</td>
|
||
</tr>
|
||
<tr id="355134E7FFFABC9B2AEA92BAFA79E110" box="[163,1423,1771,1796]" gridrow="19" pageId="4" pageNumber="125">
|
||
<th id="76805D9BFFFABC9B2AEA92BAFE31E110" box="[163,455,1771,1796]" gridcol="0" gridrow="19" pageId="4" pageNumber="125">
|
||
<emphasis id="B915EAB7FFFA43602AEA92BAFEEDE117" bold="true" box="[163,283,1771,1795]" pageId="4" pageNumber="125">Bulb width</emphasis>
|
||
</th>
|
||
<td id="76805D9BFFFABC9B2BAA92BAFD6DE110" box="[483,667,1771,1796]" gridcol="1" gridrow="19" pageId="4" pageNumber="125">14</td>
|
||
<td id="76805D9BFFFABC9B288F92BAFCB4E110" box="[710,834,1771,1796]" gridcol="2" gridrow="19" pageId="4" pageNumber="125">13</td>
|
||
<td id="76805D9BFFFABC9B29DF92BAFB94E110" box="[918,1122,1771,1796]" gridcol="3" gridrow="19" pageId="4" pageNumber="125">11</td>
|
||
<td id="76805D9BFFFABC9B2E8D92BAFA79E110" box="[1220,1423,1771,1796]" gridcol="4" gridrow="19" pageId="4" pageNumber="125">15</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BDE36A5FFFA43602A8E9306FB77E1AD" blockId="4.[151,1437,1879,1978]" pageId="4" pageNumber="125">Each limb bears a sensory organ located dorsally on coxa (basal part) of the leg. Sensory organs of the legs I–III appear as simple spines equal in shape and size (fig. 2 A, 3 C). Leg IV sensory organ shaped as short ovoid to hemispherical papilla with a minute apical spine and without a cuticular sheath (fig. 2 A).</paragraph>
|
||
<caption id="DF1E662DFFFB43612AD191D7FB89E01E" ID-DOI="http://doi.org/10.5281/zenodo.13209811" ID-Zenodo-Dep="13209811" httpUri="https://zenodo.org/record/13209811/files/figure.png" pageId="5" pageNumber="126" startId="5.[152,255,1414,1438]" targetBox="[340,1247,181,1389]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BDE36A5FFFB43612AD191D7FB89E01E" blockId="5.[151,1437,1414,1546]" pageId="5" pageNumber="126">
|
||
<emphasis id="B915EAB7FFFB43612AD191D7FEECE38A" bold="true" box="[152,282,1414,1438]" pageId="5" pageNumber="126">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C614D26FFFB43612B6A91D7FE10E38A" authorityName="Tchesunov & Fedyaeva" authorityYear="2024" box="[291,486,1414,1438]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="126" phylum="Tardigrada" rank="species" species="olenevskii" status="sp. nov.">
|
||
<emphasis id="B915EAB7FFFB43612B6A91D7FE10E38A" box="[291,486,1414,1438]" italics="true" pageId="5" pageNumber="126">Angursa olenevskii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22657CCFFFB43612BA691D7FDDFE389" box="[495,553,1414,1437]" pageId="5" pageNumber="126" rank="species">sp. n.</taxonomicNameLabel>
|
||
, SEM.
|
||
<emphasis id="B915EAB7FFFB4361283091D7FD7DE38A" bold="true" box="[633,651,1414,1438]" pageId="5" pageNumber="126">A</emphasis>
|
||
specimen 1 male, entire, ventral view.
|
||
<emphasis id="B915EAB7FFFB43612E6E91D7FBCEE38A" bold="true" box="[1063,1080,1414,1438]" pageId="5" pageNumber="126">B</emphasis>
|
||
specimen 2 female, entire, lateroventral view.
|
||
<emphasis id="B915EAB7FFFB43612B6A91FBFEC3E3D6" bold="true" box="[291,309,1450,1474]" pageId="5" pageNumber="126">C</emphasis>
|
||
specimen 3, entire, lateral view.
|
||
<emphasis id="B915EAB7FFFB436128C391FBFD6AE3D6" bold="true" box="[650,668,1450,1474]" pageId="5" pageNumber="126">D</emphasis>
|
||
specimen 1, left foot I, ventral view.
|
||
<emphasis id="B915EAB7FFFB43612E6B91FBFBC5E3D6" bold="true" box="[1058,1075,1450,1474]" pageId="5" pageNumber="126">E</emphasis>
|
||
specimen I, right foot II, toes with claws withdrawn in hoods.
|
||
<emphasis id="B915EAB7FFFB43612BE7919FFE4BE3F2" bold="true" box="[430,445,1486,1510]" pageId="5" pageNumber="126">F</emphasis>
|
||
specimen 4, left foot IV, toes with claws protruded.
|
||
<emphasis id="B915EAB7FFFB4361299A919FFC11E3F2" bold="true" box="[979,999,1486,1510]" pageId="5" pageNumber="126">G</emphasis>
|
||
specimen 3, primary clava and cirrus A.
|
||
<emphasis id="B915EAB7FFFB43612FC0919FFA6BE3F2" bold="true" box="[1417,1437,1486,1510]" pageId="5" pageNumber="126">H</emphasis>
|
||
specimen 1, male gonopore and anus. Scale bars: A 10 μm, B and C 20 μm, D–F, H 2 μm, G 5 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDE36A5FFFB43612A8E920DFA7CE166" blockId="5.[151,1437,1628,2015]" pageId="5" pageNumber="126">Limbs conical, with barely discernible coxa and femur, and retractable distal part, tibia and tarsus with four digits terminating with claws. Digits unequal in length, laterally flattened and transparent. Internal digits slightly to significantly longer (1.3–1.5 times) than the external digits. External digits have a long peduncle extended along the ventral edge to the claw while internal digits do not have peduncle. Internal digits proximally differentiated in proximal pads while the external digits are not (fig. 2 C, 3 D). Claws minute, equal in shape and size, crescent-like in overall appearance. Primary point and secondary point broadly collocated; accessory point present on all claws but clearly discernible in only scanning electron and not in optical microscope (fig. 3 F). Claws may be projected or withdrawn into a fine transparent hood-like claw sheath. External and internal claws of all leg pairs equal in size.</paragraph>
|
||
<paragraph id="8BDE36A5FFFB43612A8E932DFBCEE1AE" blockId="5.[151,1437,1628,2015]" pageId="5" pageNumber="126">Mouth opening subterminal. Stylets short and straight. Muscular bulb ovoid, its length to width ratio 1.14–1.25. Stylets short and straight, 10–16 μm long. Midgut pale brownish. No anal papillae.</paragraph>
|
||
<paragraph id="8BDE36A5FFFB43622A8E9395FCE5E6C1" blockId="5.[151,1437,1628,2015]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="127" pageId="5" pageNumber="126">Female gonopore a small six-lobed rosette at a short distance anterior to the longitudinal anal slit (fig. 2 B). Spermathecas and ducts poorly discernible. Male gonopore an arcuate transversal slit at anterior edge of a round areole just anterior to the longitudinal anal slit (fig. 3 H).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B652EFFF843622ADE9577FA7DE491" pageId="6" pageNumber="127" type="diagnosis">
|
||
<paragraph id="8BDE36A5FFF843622ADE9577FE55E754" blockId="6.[151,419,294,320]" box="[151,419,294,320]" pageId="6" pageNumber="127">
|
||
<heading id="D09681C9FFF843622ADE9577FE55E754" bold="true" box="[151,419,294,320]" fontSize="11" level="1" pageId="6" pageNumber="127" reason="1">
|
||
<emphasis id="B915EAB7FFF843622ADE9577FE55E754" bold="true" box="[151,419,294,320]" pageId="6" pageNumber="127">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDE36A5FFF843622ADE953EFA7DE491" blockId="6.[151,1437,367,645]" pageId="6" pageNumber="127">
|
||
Comparison of
|
||
<taxonomicName id="4C614D26FFF843622B0C953EFE0AE79D" box="[325,508,367,393]" pageId="6" pageNumber="127">
|
||
<emphasis id="B915EAB7FFF843622B0C953EFE52E79D" box="[325,420,367,393]" italics="true" pageId="6" pageNumber="127">Angursa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
is given on the
|
||
<tableCitation id="C6E3031EFFF8436228E7953EFCD5E79D" box="[686,803,367,393]" captionStart="TABLE 2" captionStartId="6.[151,239,692,716]" captionTargetPageId="6" captionText="TABLE 2. Comparison of Angursa olenevskyi sp. n. with other Angursa species" pageId="6" pageNumber="127">
|
||
Table 2.
|
||
<taxonomicName id="4C614D26FFF843622946953EFCD5E79D" authorityName="Tchesunov & Fedyaeva" authorityYear="2024" box="[783,803,367,393]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="127" phylum="Tardigrada" rank="species" species="olenevskii">
|
||
<emphasis id="B915EAB7FFF843622946953EFCD5E79D" box="[783,803,367,393]" italics="true" pageId="6" pageNumber="127">A</emphasis>
|
||
</taxonomicName>
|
||
</tableCitation>
|
||
. olenevskii is easily distinguished from other congeneric species by having well-developed sensory organs on all the legs. Most
|
||
<taxonomicName id="4C614D26FFF8436229FD95C2FB9DE7B9" box="[948,1131,403,429]" pageId="6" pageNumber="127">
|
||
<emphasis id="B915EAB7FFF8436229FD95C2FBE5E7B9" box="[948,1043,403,429]" italics="true" pageId="6" pageNumber="127">Angursa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
have no sensory organs on legs II and III except
|
||
<taxonomicName id="4C614D26FFF843622BC095E6FDFBE7C5" authorityName=", Noda" authorityYear="1985" box="[393,525,439,465]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="127" phylum="Tardigrada" rank="species" species="clavifera">
|
||
<emphasis id="B915EAB7FFF843622BC095E6FDFBE7C5" box="[393,525,439,465]" italics="true" pageId="6" pageNumber="127">A. clavifera</emphasis>
|
||
</taxonomicName>
|
||
where those organs are quite tiny and barely visible. Another feature differing
|
||
<taxonomicName id="4C614D26FFF843622FCD95E6FEF1E7E1" authorityName="Tchesunov & Fedyaeva" authorityYear="2024" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="127" phylum="Tardigrada" rank="species" species="olenevskii">
|
||
<emphasis id="B915EAB7FFF843622FCD95E6FEF1E7E1" italics="true" pageId="6" pageNumber="127">A. olenevskii</emphasis>
|
||
</taxonomicName>
|
||
from all other
|
||
<taxonomicName id="4C614D26FFF843622BE5958AFD94E7E1" box="[428,610,475,501]" pageId="6" pageNumber="127">
|
||
<emphasis id="B915EAB7FFF843622BE5958AFDFDE7E1" box="[428,523,475,501]" italics="true" pageId="6" pageNumber="127">Angursa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
is evident claw and toe structures. Peduncles of
|
||
<taxonomicName id="4C614D26FFF843622E33958AFAFFE7E1" authorityName="Tchesunov & Fedyaeva" authorityYear="2024" box="[1146,1289,475,501]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="127" phylum="Tardigrada" rank="species" species="olenevskii">
|
||
<emphasis id="B915EAB7FFF843622E33958AFAFFE7E1" box="[1146,1289,475,501]" italics="true" pageId="6" pageNumber="127">A. olenevskii</emphasis>
|
||
</taxonomicName>
|
||
are extended along the ventral edge of digits from the base to the claw while the peduncles of all other congeneric species present a short curved bar at the base of the external digit stretched or even not stretched to basal third of the digits. Finally,
|
||
<taxonomicName id="4C614D26FFF843622ADE9616FED0E475" authorityName="Tchesunov & Fedyaeva" authorityYear="2024" box="[151,294,583,609]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="127" phylum="Tardigrada" rank="species" species="olenevskii">
|
||
<emphasis id="B915EAB7FFF843622ADE9616FED0E475" box="[151,294,583,609]" italics="true" pageId="6" pageNumber="127">A. olenevskii</emphasis>
|
||
</taxonomicName>
|
||
differs from almost all species of
|
||
<taxonomicName id="4C614D26FFF8436228E99616FD09E475" authorityName="Pollock" authorityYear="1979" box="[672,767,583,609]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="127" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B915EAB7FFF8436228E99616FD09E475" box="[672,767,583,609]" italics="true" pageId="6" pageNumber="127">Angursa</emphasis>
|
||
</taxonomicName>
|
||
(with possible exception of
|
||
<taxonomicName id="4C614D26FFF843622E739616FB42E475" authorityName="Pollock" authorityYear="1979" box="[1082,1204,583,609]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="127" phylum="Tardigrada" rank="species" species="bicuspis">
|
||
<emphasis id="B915EAB7FFF843622E739616FB42E475" box="[1082,1204,583,609]" italics="true" pageId="6" pageNumber="127">A. bicuspis</emphasis>
|
||
</taxonomicName>
|
||
) by lack of even any traces of secondary and tertiary clavae which could not be observed neither with light microscope nor with SEM.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |