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<document id="2D182950608599DD8A209338B39BE0CD" ID-CLB-Dataset="32718" ID-DOI="10.11646/zootaxa.4290.2.1" ID-GBIF-Dataset="43ebda6c-201a-46af-8b54-e7980b8d79e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="828965" ID-ZooBank="52D6E3C3-BA0A-49C7-865C-00F3F5E5C5D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110315309" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2017" docId="03C887A8FFD4FFD8FF737FAD5B14B597" docLanguage="en" docName="zootaxa.4290.2.1.pdf" docOrigin="Zootaxa 4290 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monteithocanthon koombooloomba Gunter &amp; Weir, new species" docType="treatment" docVersion="7" lastPageNumber="232" masterDocId="FFF1FFD0FFCBFFC7FFE47F3A5F6FB348" masterDocTitle="Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species" masterLastPageNumber="243" masterPageNumber="201" pageNumber="232" updateTime="1698465402479" updateUser="ExternalLinkService">
<mods:mods id="FBF93829E3BB6E38DBCF43CEC3D2D89F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6F2B6693B36052F9830CBB6F1347E0F6">
<mods:title id="2A2F37047773310B3EDDCFAD6E26B5EE">Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species</mods:title>
</mods:titleInfo>
<mods:name id="F44F9B452CF380CD0E72796D4B9CC2BA" type="personal">
<mods:role id="ACCD1EECD406480DE6D544045E7B9152">
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</mods:role>
<mods:namePart id="6D53457F446D7CAD3A63415079EA69B7">Gunter, Nicole L.</mods:namePart>
</mods:name>
<mods:name id="71699F299C574E1CC4E14AB0CAF0A1EC" type="personal">
<mods:role id="F5A878316739C18C3E7A858E83D1A47B">
<mods:roleTerm id="189895185F7824717343F23F4E48EC4A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="405F82FA7C5E68B977ABC9C4C256330F">Weir, Thomas A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CC1E7A96F3C948F8ABB94474E43AB0A1">text</mods:typeOfResource>
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<mods:title id="CC035EF2964DE23EEEF226BBF142119C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1996D8FFA40E493DF14D2E51B8C0D86C">
<mods:date id="CB4F6BD2C02184D5DED166D2ADA28B91">2017</mods:date>
<mods:detail id="B2FD53FB878260D8B71453E8EC5F2D8E" type="volume">
<mods:number id="E1ECAAFBA8F85806BF5EF0C75A46029A">4290</mods:number>
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<mods:detail id="1A98B2F1401B78C923462BF27507F728" type="issue">
<mods:number id="6E31DE961FA51D74F7D56E9B6A260924">2</mods:number>
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<mods:start id="0FB317C4A41A67A93F139F06461E2593">201</mods:start>
<mods:end id="FABDA266DC99C9DD6EFACBFEE67A20F5">243</mods:end>
</mods:extent>
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<mods:classification id="1B280EBFE833EDA03E3F2F2CD82D1DED">journal article</mods:classification>
<mods:identifier id="5A285C9679AC723CD4C7A27F2AA19191" type="CLB-Dataset">32718</mods:identifier>
<mods:identifier id="12D1FF2752A31FAD7C44A7C5970BC61A" type="DOI">10.11646/zootaxa.4290.2.1</mods:identifier>
<mods:identifier id="90A12406585FA27172B8D4018A076733" type="GBIF-Dataset">43ebda6c-201a-46af-8b54-e7980b8d79e1</mods:identifier>
<mods:identifier id="4C687D8E5503FAAD4B0132BAF3301DB8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="31044271F8E3D43E0A02A160A896F032" type="Zenodo-Dep">828965</mods:identifier>
<mods:identifier id="2D91FB2EA6045699AF473E314634CA83" type="ZooBank">52D6E3C3-BA0A-49C7-865C-00F3F5E5C5D4</mods:identifier>
</mods:mods>
<treatment id="03C887A8FFD4FFD8FF737FAD5B14B597" ID-DOI="http://doi.org/10.5281/zenodo.6033018" ID-GBIF-Taxon="132549210" ID-Zenodo-Dep="6033018" LSID="urn:lsid:plazi:treatment:03C887A8FFD4FFD8FF737FAD5B14B597" httpUri="http://treatment.plazi.org/id/03C887A8FFD4FFD8FF737FAD5B14B597" lastPageNumber="232" pageId="31" pageNumber="232">
<subSubSection id="C37B6535FFD4FFD8FF737FAD5EA3B39B" pageId="31" pageNumber="232" type="nomenclature">
<paragraph id="8BDE36BEFFD4FFD8FF737FAD5CCAB3FA" blockId="31.[151,933,151,211]" box="[151,933,151,178]" pageId="31" pageNumber="232">
<heading id="D09681D2FFD4FFD8FF737FAD5CCAB3FA" bold="true" box="[151,933,151,178]" fontSize="11" level="1" pageId="31" pageNumber="232" reason="1">
<emphasis id="B915EAACFFD4FFD8FF737FAD5CCAB3FA" bold="true" box="[151,933,151,178]" pageId="31" pageNumber="232">
<taxonomicName id="4C614D3DFFD4FFD8FF737FAD5C60B3FA" ID-CoL="73Q87" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" box="[151,783,151,178]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="koombooloomba" status="sp. nov.">
<emphasis id="B915EAACFFD4FFD8FF737FAD5D28B3F9" bold="true" box="[151,583,151,177]" italics="true" pageId="31" pageNumber="232">Monteithocanthon koombooloomba</emphasis>
Gunter &amp; Weir
</taxonomicName>
<taxonomicNameLabel id="A22657D7FFD4FFD8FCF27FA25CCAB3FA" box="[790,933,152,178]" pageId="31" pageNumber="232" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDE36BEFFD4FFD8FF737F805EA3B39B" blockId="31.[151,933,151,211]" box="[151,460,186,211]" pageId="31" pageNumber="232">
<figureCitation id="135A2A3BFFD4FFD8FF737F805F8CB39B" box="[151,227,186,211]" captionStart="FIGURE 5" captionStartId="13.[151,250,1554,1576]" captionTargetBox="[170,1385,589,1501]" captionTargetId="figure@13.[151,1436,589,1532]" captionTargetPageId="13" captionText="FIGURE 5. Distributional maps of species belonging to the new genera Matthewsius and Monteithocanthon including close up insets of A, Pilbara region, Western Australia; B, northeastern Queensland; C, southeastern Queensland; D, southwestern Western Australia; E, southeastern South Australia; F, central to southeastern New South Wales." httpUri="https://zenodo.org/record/828975/files/figure.png" pageId="31" pageNumber="232">Figs. 5</figureCitation>
B, 7K, 8D, 9D, 10D.
</paragraph>
</subSubSection>
<subSubSection id="C37B6535FFD4FFD8FF737E385B85B15F" pageId="31" pageNumber="232" type="materials_examined">
<paragraph id="8BDE36BEFFD4FFD8FF737E385B85B15F" blockId="31.[151,1437,258,1759]" pageId="31" pageNumber="232">
<materialsCitation id="3B093CE3FFD4FFD8FF737E385B66B277" ID-GBIF-Occurrence="1572325628" collectingDate="1989-12-08" county="Dam" latitude="-17.904167" location="Thompson" longLatPrecision="21" longitude="145.62167" municipality="Monteith" pageId="31" pageNumber="232" specimenCount="1" specimenCount-male="1" stateProvince="Queensland" typeStatus="holotype">
<emphasis id="B915EAACFFD4FFD8FF737E385EE8B253" bold="true" box="[151,391,258,283]" pageId="31" pageNumber="232">Material examined.</emphasis>
<collectingRegion id="49A5F85CFFD4FFD8FE717E385D71B253" box="[405,542,258,283]" country="Australia" name="Queensland" pageId="31" pageNumber="232">Queensland</collectingRegion>
:
<typeStatus id="54DA881CFFD4FFD8FDCB7E385DFAB253" box="[559,661,258,283]" pageId="31" pageNumber="232" type="holotype">holotype</typeStatus>
:
<specimenCount id="9D67FD37FFD4FFD8FD437E395DD3B252" box="[679,700,259,282]" pageId="31" pageNumber="232" type="male">♂</specimenCount>
<geoCoordinate id="EE555079FFD4FFD8FD2D7E385C25B253" box="[713,842,258,283]" direction="south" orientation="latitude" pageId="31" pageNumber="232" precision="15" value="-17.904167">17°54'15''S</geoCoordinate>
,
<geoCoordinate id="EE555079FFD4FFD8FCB87E385C81B253" box="[860,1006,258,283]" direction="east" orientation="longitude" pageId="31" pageNumber="232" precision="15" value="145.62167">145°37'18&quot;E</geoCoordinate>
,
<locationDeviation id="9A6C2F9CFFD4FFD8FBE47E385A63B253" box="[1024,1292,258,283]" pageId="31" pageNumber="232">
<quantity id="4C999B5BFFD4FFD8FBE47E385B3AB253" box="[1024,1109,258,283]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="232" unit="km" value="1.5">1.5 km</quantity>
N Tully River
</locationDeviation>
crossing, S
<taxonomicName id="4C614D3DFFD4FFD8FF737E1C5E34B277" box="[151,347,294,319]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="koombooloomba">Koombooloomba</taxonomicName>
<collectingCounty id="62BF4E32FFD4FFD8FE817E1D5EF1B276" box="[357,414,295,318]" pageId="31" pageNumber="232">Dam</collectingCounty>
,
<collectingMunicipality id="6BBAACC4FFD4FFD8FE497E1C5D78B277" box="[429,535,294,319]" pageId="31" pageNumber="232">Monteith</collectingMunicipality>
,
<location id="8EBE6065FFD4FFD8FDC17E1C5DCFB277" LSID="urn:lsid:plazi:treatment:03C887A8FFD4FFD8FF737FAD5B14B597:8EBE6065FFD4FFD8FDC17E1C5DCFB277" box="[549,672,294,319]" county="Dam" latitude="-17.904167" longLatPrecision="21" longitude="145.62167" municipality="Monteith" name="Thompson" pageId="31" pageNumber="232" stateProvince="Queensland">Thompson</location>
, &amp;
<location id="8EBE6065FFD4FFD8FD2A7E1C5C42B277" LSID="urn:lsid:plazi:treatment:03C887A8FFD4FFD8FF737FAD5B14B597:8EBE6065FFD4FFD8FD2A7E1C5C42B277" box="[718,813,294,319]" county="Dam" latitude="-17.904167" longLatPrecision="21" longitude="145.62167" municipality="Monteith" name="Janetzki" pageId="31" pageNumber="232" stateProvince="Queensland">Janetzki</location>
,
<collectingDate id="EF9BE996FFD4FFD8FCDF7E1D5B66B277" box="[827,1033,294,319]" pageId="31" pageNumber="232" value="1989-12-08">
<date id="FFDF107EFFD4FFD8FCDF7E1D5B66B277" box="[827,1033,294,319]" pageId="31" pageNumber="232" value="1989-12-08">8 December 1989</date>
</collectingDate>
</materialsCitation>
–
<materialsCitation id="3B093CE3FFD4FFD8FBFD7E1C5D07B2CF" ID-GBIF-Occurrence="1572325627" collectingDate="1989-12-08" collectingDateMax="1990-01-05" collectingDateMin="1989-12-08" collectorName="Dam &amp; Monteith &amp; Thompson &amp; Janetzki" latitude="-17.904167" location="Tully River" longLatPrecision="21" longitude="145.62167" pageId="31" pageNumber="232" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectingDate id="EF9BE996FFD4FFD8FBFD7E1C5BA6B277" box="[1049,1225,294,319]" pageId="31" pageNumber="232" value="1990-01-05">
<date id="FFDF107EFFD4FFD8FBFD7E1C5BA6B277" box="[1049,1225,294,319]" pageId="31" pageNumber="232" value="1990-01-05">5 January 1990</date>
</collectingDate>
(in QM T54139);
<typeStatus id="54DA881CFFD4FFD8FF737E715E68B22B" box="[151,263,331,355]" pageId="31" pageNumber="232" type="paratype">paratypes</typeStatus>
:
<geoCoordinate id="EE555079FFD4FFD8FEFE7E705EF5B22B" box="[282,410,330,355]" direction="south" orientation="latitude" pageId="31" pageNumber="232" precision="15" value="-17.904167">17°54'15''S</geoCoordinate>
,
<geoCoordinate id="EE555079FFD4FFD8FE4B7E705D2FB22B" box="[431,576,330,355]" direction="east" orientation="longitude" pageId="31" pageNumber="232" precision="15" value="145.62167">145°37'18&quot;E</geoCoordinate>
,
<quantity id="4C999B5BFFD4FFD8FDB17E705DC4B22B" box="[597,683,330,355]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="232" unit="km" value="1.5">1.5 km</quantity>
N
<location id="8EBE6065FFD4FFD8FD3B7E705C06B22B" LSID="urn:lsid:plazi:treatment:03C887A8FFD4FFD8FF737FAD5B14B597:8EBE6065FFD4FFD8FD3B7E705C06B22B" box="[735,873,330,355]" latitude="-17.904167" longLatPrecision="21" longitude="145.62167" name="Tully River" pageId="31" pageNumber="232" stateProvince="Queensland">Tully River</location>
crossing, S
<taxonomicName id="4C614D3DFFD4FFD8FBE97E705BBEB22B" box="[1037,1233,330,355]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="koombooloomba">Koombooloomba</taxonomicName>
<collectorName id="26945368FFD4FFD8FB057E715A75B22A" box="[1249,1306,331,354]" pageId="31" pageNumber="232">Dam</collectorName>
,
<collectorName id="26945368FFD4FFD8FACA7E705AF7B22B" box="[1326,1432,330,355]" pageId="31" pageNumber="232">Monteith</collectorName>
,
<collectorName id="26945368FFD4FFD8FF737E545E7DB2CF" box="[151,274,366,391]" pageId="31" pageNumber="232">Thompson</collectorName>
, &amp;
<collectorName id="26945368FFD4FFD8FEDC7E545EF8B2CF" box="[312,407,366,391]" pageId="31" pageNumber="232">Janetzki</collectorName>
,
<collectingDate id="EF9BE996FFD4FFD8FE457E555D07B2CF" box="[417,616,366,391]" pageId="31" pageNumber="232" value="1989-12-08">
<date id="FFDF107EFFD4FFD8FE457E555D07B2CF" box="[417,616,366,391]" pageId="31" pageNumber="232" value="1989-12-08">8 December 1989</date>
</collectingDate>
</materialsCitation>
–
<date id="FFDF107EFFD4FFD8FD937E545C4FB2CF" box="[631,800,366,391]" pageId="31" pageNumber="232" value="1990-01-05">5 January 1990</date>
(in QM T54140 ♀, T54142 ♂, in ANIC 25-055641 (QM T54141 ♂, T54144 ♀, and T54143 ♀); [17°50'00''S, 145°36'00&quot;E],
<taxonomicName id="4C614D3DFFD4FFD8FC717EA85B33B2E3" box="[917,1116,402,427]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="koombooloomba">Koombooloomba</taxonomicName>
, R.W. Taylor &amp; J. Feehan,
<date id="FFDF107EFFD4FFD8FA6A7EA95E64B287" pageId="31" pageNumber="232" value="1971-07-04">4 July 1971</date>
,
<materialsCitation id="3B093CE3FFD4FFD8FEF87E8D5BB8B2BB" ID-GBIF-Occurrence="1572325725" collectingDate="1989-12-08" collectionCode="ANIC" county="Tully River" latitude="-17.916666" location="Thompson" longLatPrecision="21" longitude="145.63333" municipality="Monteith" pageId="31" pageNumber="232" specimenCode="ANIC 25-055631" specimenCount="1" specimenCount-male="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="ED70AE7BFFD4FFD8FEF87E8D5E0FB287" box="[284,352,439,463]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="31" pageNumber="232">ANIC</collectionCode>
berlesate #356, rainforest (in
<specimenCode id="DBC79EC5FFD4FFD8FD2D7E8D5CFCB287" box="[713,915,438,463]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="31" pageNumber="232">
<collectionCode id="ED70AE7BFFD4FFD8FD2D7E8D5C61B287" box="[713,782,439,463]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="31" pageNumber="232">ANIC</collectionCode>
25-055631
</specimenCode>
<specimenCount id="9D67FD37FFD4FFD8FC7B7E8D5CD9B287" box="[927,950,439,463]" pageId="31" pageNumber="232" type="male">♂</specimenCount>
); [
<geoCoordinate id="EE555079FFD4FFD8FC3C7E8C5B35B287" box="[984,1114,438,463]" direction="south" orientation="latitude" pageId="31" pageNumber="232" precision="15" value="-17.916666">17°55'00''S</geoCoordinate>
,
<geoCoordinate id="EE555079FFD4FFD8FB8F7E8C5B91B287" box="[1131,1278,438,463]" direction="east" orientation="longitude" pageId="31" pageNumber="232" precision="15" value="145.63333">145°38'00&quot;E</geoCoordinate>
],
<collectingCounty id="62BF4E32FFD4FFD8FAF27E8C5AF3B287" box="[1302,1436,438,463]" pageId="31" pageNumber="232">Tully River</collectingCounty>
crossing,
<locationDeviation id="9A6C2F9CFFD4FFD8FEE07EE15D1AB2BA" box="[260,629,474,499]" pageId="31" pageNumber="232">
<quantity id="4C999B5BFFD4FFD8FEE07EE15E22B2BB" box="[260,333,474,499]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="232" unit="km" value="10.0">10 km</quantity>
S
<taxonomicName id="4C614D3DFFD4FFD8FE8B7EE05D5CB2BB" box="[367,563,474,499]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="koombooloomba">Koombooloomba</taxonomicName>
Dam
</locationDeviation>
,
<collectingMunicipality id="6BBAACC4FFD4FFD8FD677EE05D82B2BB" box="[643,749,474,499]" pageId="31" pageNumber="232">Monteith</collectingMunicipality>
,
<location id="8EBE6065FFD4FFD8FD1E7EE05C1AB2BB" LSID="urn:lsid:plazi:treatment:03C887A8FFD4FFD8FF737FAD5B14B597:8EBE6065FFD4FFD8FD1E7EE05C1AB2BB" box="[762,885,474,499]" county="Tully River" latitude="-17.916666" longLatPrecision="21" longitude="145.63333" municipality="Monteith" name="Thompson" pageId="31" pageNumber="232" stateProvince="Queensland">Thompson</location>
, &amp;
<location id="8EBE6065FFD4FFD8FC457EE05B6FB2BB" LSID="urn:lsid:plazi:treatment:03C887A8FFD4FFD8FF737FAD5B14B597:8EBE6065FFD4FFD8FC457EE05B6FB2BB" box="[929,1024,474,499]" county="Tully River" latitude="-17.916666" longLatPrecision="21" longitude="145.63333" municipality="Monteith" name="Janetzki" pageId="31" pageNumber="232" stateProvince="Queensland">Janetzki</location>
,
<collectingDate id="EF9BE996FFD4FFD8FBE87EE15BB8B2BB" box="[1036,1239,474,499]" pageId="31" pageNumber="232" value="1989-12-08">
<date id="FFDF107EFFD4FFD8FBE87EE15BB8B2BB" box="[1036,1239,474,499]" pageId="31" pageNumber="232" value="1989-12-08">8 December 1989</date>
</collectingDate>
</materialsCitation>
–
<date id="FFDF107EFFD4FFD8FB027EE15AF7B2BB" box="[1254,1432,475,499]" pageId="31" pageNumber="232" value="1990-01-04">
4 January
<quantity id="4C999B5BFFD4FFD8FAB97EE15AF7B2BB" box="[1373,1432,475,499]" metricMagnitude="6" metricUnit="m" metricValue="1.9907499999999998" pageId="31" pageNumber="232" unit="m" value="1990750.0">1990</quantity>
</date>
, 750 m,
<collectingMethod id="52204EA9FFD4FFD8FF087EC45E95B15E" box="[236,506,510,535]" pageId="31" pageNumber="232">pitfall and intercept trap</collectingMethod>
(in QM T54135–137
<date id="FFDF107EFFD4FFD8FD157EC55C63B15F" box="[753,780,511,535]" pageId="31" pageNumber="232">3x</date>
♀, in ANIC 25-065499 ♀ (QM T54138)).
</paragraph>
</subSubSection>
<subSubSection id="C37B6535FFD4FFD8FF237D185D08B117" pageId="31" pageNumber="232" type="etymology">
<paragraph id="8BDE36BEFFD4FFD8FF237D185D08B117" blockId="31.[151,1437,258,1759]" pageId="31" pageNumber="232">
<emphasis id="B915EAACFFD4FFD8FF237D185E21B173" bold="true" box="[199,334,546,571]" pageId="31" pageNumber="232">Etymology.</emphasis>
Named for
<taxonomicName id="4C614D3DFFD4FFD8FE3E7D185DCEB173" box="[474,673,546,571]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="koombooloomba">Koombooloomba</taxonomicName>
, the locality where the entire
<typeStatus id="54DA881CFFD4FFD8FC1C7D195B47B173" box="[1016,1064,547,571]" pageId="31" pageNumber="232">type</typeStatus>
series was collected. This name should be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C37B6535FFD4FFD8FF237D505ED8B53B" pageId="31" pageNumber="232" type="description">
<paragraph id="8BDE36BEFFD4FFD8FF237D505C78B1CB" blockId="31.[151,1437,258,1759]" box="[199,791,618,643]" pageId="31" pageNumber="232">
<emphasis id="B915EAACFFD4FFD8FF237D505E36B1CB" bold="true" box="[199,345,618,643]" pageId="31" pageNumber="232">Description.</emphasis>
Colour: Black. Antennae white to grey.
</paragraph>
<paragraph id="8BDE36BEFFD4FFD8FF237DB45B3DB1EF" blockId="31.[151,1437,258,1759]" box="[199,1106,654,679]" pageId="31" pageNumber="232">
<typeStatus id="54DA881CFFD4FFD8FF237DB45E40B1EF" box="[199,303,654,679]" pageId="31" pageNumber="232" type="holotype">Holotype</typeStatus>
measurements:
<quantity id="4C999B5BFFD4FFD8FE017DB55D54B1EE" box="[485,571,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="232" unit="mm" value="2.4">2.4 mm</quantity>
in length,
<quantity id="4C999B5BFFD4FFD8FD567DB45C67B1EE" box="[690,776,654,679]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="31" pageNumber="232" unit="mm" value="1.6">1.6 mm</quantity>
across widest point of elytra.
</paragraph>
<paragraph id="8BDE36BEFFD4FFD8FF237D885EDDB182" blockId="31.[151,1437,258,1759]" box="[199,434,690,715]" pageId="31" pageNumber="232">
Total length:
<quantity id="4C999B5BFFD4FFD8FEB87D895EDDB182" box="[348,434,691,715]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="232" unit="mm" value="2.4">2.4 mm</quantity>
</paragraph>
<paragraph id="8BDE36BEFFD4FFD8FF237DEC5DD0B037" blockId="31.[151,1437,258,1759]" pageId="31" pageNumber="232">Male: Head: Broad U-shape between clypeal teeth, which are slightly upturned. Rest of margin of head feebly convex to eye. Mentum almost square, not bordered, with striations and setation sparse on posterior corners. Edge of head completely bordered, genal angle distinct. Dorsal parts of eyes narrow, separated by an interocular space of about 16 times eye width; eye canthus nearly dividing eye. Surface smooth, nitid; with sparse, round punctures; setation absent. Head width to length ratio 40:25.</paragraph>
<paragraph id="8BDE36BEFFD4FFD8FF237CB15C23B747" blockId="31.[151,1437,258,1759]" pageId="31" pageNumber="232">Pronotum: Anterior angles almost 90°. Basal angles distinct, basal margin rounded. Hypomeron surface finely reticulate with sparse punctation. Elongate punctures along basal margin almost extend to basal edge but diverging medially to leave an impunctate area. All punctures without setae. Lateral edge margined. Hypomeral stria 1/3 to 1/ 2 length of hypomeron. Pronotum width to length ratio 63:37.</paragraph>
<paragraph id="8BDE36BEFFD4FFD8FF237B205FB1B733" blockId="31.[151,1437,258,1759]" pageId="31" pageNumber="232">Elytra: Elytra with small flange at most, surface smooth. Intervals finely punctate and not reticulate. Striae 1–5 superficial but clearly visible, striae 6–8 obsolete. Ratio of length of elytra along suture to maximum elytral width 72:77.</paragraph>
<paragraph id="8BDE36BEFFD4FFD8FF237BBD5CC4B667" blockId="31.[151,1437,258,1759]" pageId="31" pageNumber="232">
Legs: Protibiae with 3 teeth on outer edge, which is serrate between teeth and proximal to them; each tooth with 1 group of setae dorsally, front edge straight between apical tooth and small inconspicuous spur. The tooth on underside near base of tarsus is a blunt projection. Protibiae inner apex expanded, with a comb and a brush of setae. Claws small and toothed. Mesotibiae slightly expanded, densely setose on inner side. Metatibiae bent with an indentation, with small tooth at inner apex and not crenulate (
<figureCitation id="135A2A3BFFD4FFD8FCAC7A2C5CE5B667" box="[840,906,1302,1327]" captionStart="FIGURE 8" captionStartId="26.[151,250,955,977]" captionTargetBox="[192,1404,193,933]" captionTargetId="figure@26.[182,1404,193,933]" captionTargetPageId="26" captionText="FIGURE 8. Male metatibia of species of Monteithocanthon new genus, all scale bars 0.25 mm. A, Monteithocanthon arator new combination; B, M. elliotensis new species; C, M. glaber new combination; D, M. koombooloomba new species; E, M. paraarator new species; F, M. peckorum new species; G, female metathoracic leg of M. arator." httpUri="https://zenodo.org/record/828981/files/figure.png" pageId="31" pageNumber="232">Fig. 8</figureCitation>
E).
</paragraph>
<paragraph id="8BDE36BEFFD4FFD8FF237A005B90B6D3" blockId="31.[151,1437,258,1759]" pageId="31" pageNumber="232">Abdomen: Pygidium with deep, curved transverse groove at the base about 3/4 the width of the pygidium and not reaching basal angle. Ventrites with fine reticulation, absent from middle 1/3, with a row of punctures along the anterior edge of ventrites 2–5, ventrite 6 finely punctate. Sutures between abdominal ventrites slight.</paragraph>
<paragraph id="8BDE36BEFFD4FFD8FF237A9C5B9CB54F" blockId="31.[151,1437,258,1759]" pageId="31" pageNumber="232">Pterothorax: Medial lobe of metaventrite finely punctate and broadly margined between mesocoxae. Mesometaventral suture arched. Lateral lobe of the metaventrite broadly margined anteriorly, with round punctures. Mesoventrite smooth, not notched anteriorly. Mesepimera smooth. Metanepisterna finely reticulate.</paragraph>
<paragraph id="8BDE36BEFFD4FFD8FF2379285D7BB563" blockId="31.[151,1437,258,1759]" box="[199,532,1554,1579]" pageId="31" pageNumber="232">Hind wings: Fully developed.</paragraph>
<paragraph id="8BDE36BEFFD4FFD8FF23790C5ED8B53B" blockId="31.[151,1437,258,1759]" pageId="31" pageNumber="232">Female: Protibiae not expanded at the inner apical angle with longer apical spur. Mesotibiae lack inner brush of setae. Metatibiae straight.</paragraph>
</subSubSection>
<subSubSection id="C37B6535FFD4FFD8FF2379445C7CB5F2" pageId="31" pageNumber="232" type="distribution">
<paragraph id="8BDE36BEFFD4FFD8FF2379445C7CB5F2" blockId="31.[151,1437,258,1759]" pageId="31" pageNumber="232">
<emphasis id="B915EAACFFD4FFD8FF2379445E0FB5DF" bold="true" box="[199,352,1662,1687]" pageId="31" pageNumber="232">Distribution.</emphasis>
See
<figureCitation id="135A2A3BFFD4FFD8FE7379445EB5B5DF" box="[407,474,1662,1687]" captionStart="FIGURE 5" captionStartId="13.[151,250,1554,1576]" captionTargetBox="[170,1385,589,1501]" captionTargetId="figure@13.[151,1436,589,1532]" captionTargetPageId="13" captionText="FIGURE 5. Distributional maps of species belonging to the new genera Matthewsius and Monteithocanthon including close up insets of A, Pilbara region, Western Australia; B, northeastern Queensland; C, southeastern Queensland; D, southwestern Western Australia; E, southeastern South Australia; F, central to southeastern New South Wales." httpUri="https://zenodo.org/record/828975/files/figure.png" pageId="31" pageNumber="232">Fig. 5</figureCitation>
B. Known only from the
<taxonomicName id="4C614D3DFFD4FFD8FD1779445CD8B5DF" box="[755,951,1662,1687]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="koombooloomba">Koombooloomba</taxonomicName>
area, northern
<collectingRegion id="49A5F85CFFD4FFD8FB8679445B89B5DF" box="[1122,1254,1662,1687]" country="Australia" name="Queensland" pageId="31" pageNumber="232">Queensland</collectingRegion>
at about
<specimenCount id="9D67FD37FFD4FFD8FAAA79445AF2B5DE" box="[1358,1437,1662,1687]" pageId="31" pageNumber="232" type="generic">
<quantity id="4C999B5BFFD4FFD8FAAA79445AF7B5DE" box="[1358,1432,1662,1687]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="31" pageNumber="232" unit="m" value="750.0">750 m</quantity>
.
</specimenCount>
Collected in rainforest in pitfall traps and intercept traps.
</paragraph>
</subSubSection>
<subSubSection id="C37B6535FFD4FFD8FF2379FC5B14B597" box="[199,1147,1734,1759]" pageId="31" pageNumber="232" type="discussion">
<paragraph id="8BDE36BEFFD4FFD8FF2379FC5B14B597" blockId="31.[151,1437,258,1759]" box="[199,1147,1734,1759]" pageId="31" pageNumber="232">
<emphasis id="B915EAACFFD4FFD8FF2379FC5E20B597" bold="true" box="[199,335,1734,1759]" pageId="31" pageNumber="232">Comments.</emphasis>
This species was referred to as “
<taxonomicName id="4C614D3DFFD4FFD8FD2479FD5C70B596" box="[704,799,1735,1758]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis id="B915EAACFFD4FFD8FD2479FD5C70B596" box="[704,799,1735,1758]" italics="true" pageId="31" pageNumber="232">Lepanus</emphasis>
</taxonomicName>
NQ22” in
<bibRefCitation id="EFF04B4FFFD4FFD8FC7F79FD5B18B597" box="[923,1143,1734,1759]" pageId="31" pageNumber="232" refString="Yeates, D. K., Gunter, N. L., Weir, T. &amp; Cameron, S. L. (2011) Developing a benchmark invertebrate group for biodiversity modelling. In: West, J. (Ed.), 21 st Century Taxonomy: Accelerating Research and Discovery of Australia's Biodiversity. Taxonomic Research and Identification Network Final Report- March 2011, pp. 93 - 99. Available from: www. environment. gov. au / system / files / pages / 56 c 08 f 6 f- 7 bf 7 - 4594 - 8175 - b 4 c 143110156 / files / hub-taxonomy. pdf (accessed 20 April 2017)" type="journal article">
Yeates
<emphasis id="B915EAACFFD4FFD8FC0F79FD5B73B597" box="[1003,1052,1734,1759]" italics="true" pageId="31" pageNumber="232">et al</emphasis>
. (2011)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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