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<document id="D94E2BB30DE6B1EEF673FE96ACCFD399" ID-DOI="10.11646/zootaxa.3981.3.7" ID-GBIF-Dataset="bfd83ee0-4e51-48ec-ba61-1bd5fcd80462" ID-ISSN="1175-5326" ID-Zenodo-Dep="241823" ID-ZooBank="E9FAE6B4-7430-45D9-8C61-CC7E8CCB2BA0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461248323986" checkinUser="plazi" docAuthor="Bidzilya, Oleksiy V." docDate="2015" docId="038DE56F513D8A54FF2E05B3FA6965FB" docLanguage="en" docName="zt03981p429.pdf" docOrigin="Zootaxa 3981 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Armatophallus Bidzilya, 2015, gen. n." docType="treatment" docVersion="8" lastPageNumber="415" masterDocId="FFB49D17513C8A56FFB9056EFFF8615A" masterDocTitle="Armatophallus gen. n., a new genus of gelechiid moths (Lepidoptera, Gelechiidae) from the Afrotropical and Oriental regions" masterLastPageNumber="429" masterPageNumber="413" pageNumber="414" updateTime="1698606970243" updateUser="plazi">
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<mods:title id="70EB45408E97C60DC54A5581F2024288">Armatophallus gen. n., a new genus of gelechiid moths (Lepidoptera, Gelechiidae) from the Afrotropical and Oriental regions</mods:title>
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<mods:namePart id="5E8DA980C3C1BE9BCC40DF52C6BF9A8E">Bidzilya, Oleksiy V.</mods:namePart>
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<mods:date id="249216A45CF844C37056BEB88E9934D8">2015</mods:date>
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<treatment id="038DE56F513D8A54FF2E05B3FA6965FB" ID-DOI="http://doi.org/10.5281/zenodo.6118563" ID-GBIF-Taxon="119635460" ID-Zenodo-Dep="6118563" LSID="urn:lsid:plazi:treatment:038DE56F513D8A54FF2E05B3FA6965FB" httpUri="http://treatment.plazi.org/id/038DE56F513D8A54FF2E05B3FA6965FB" lastPageId="2" lastPageNumber="415" pageId="1" pageNumber="414">
<subSubSection id="C33E07F2513D8A57FF2E05B3FD786060" pageId="1" pageNumber="414" type="nomenclature">
<paragraph id="8B9B5479513D8A57FF2E05B3FE5D61AD" blockId="1.[151,421,221,247]" box="[151,421,221,247]" pageId="1" pageNumber="414">
<heading id="D0D3E315513D8A57FF2E05B3FE5D61AD" bold="true" box="[151,421,221,247]" fontSize="11" level="1" pageId="1" pageNumber="414" reason="1">
<emphasis id="B950886B513D8A57FF2E05B3FE5D61AD" bold="true" box="[151,421,221,247]" pageId="1" pageNumber="414">
<taxonomicName id="4C242FFA513D8A57FF2E05B3FEB461AD" ID-CoL="93D6T" box="[151,332,221,247]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B950886B513D8A57FF2E05B3FEB461AD" bold="true" box="[151,332,221,247]" italics="true" pageId="1" pageNumber="414">Armatophallus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2633510513D8A57FEEA05B3FE5D61AD" box="[339,421,221,247]" pageId="1" pageNumber="414" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B5479513D8A57FF2E044AFD786060" blockId="1.[151,640,292,314]" box="[151,640,292,314]" pageId="1" pageNumber="414">
Type-species:
<taxonomicName id="4C242FFA513D8A57FE91044AFD786060" authority="Meyrick, 1918" authorityName="Meyrick" authorityYear="1918" box="[296,640,292,314]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="exoenota">
<emphasis id="B950886B513D8A57FE91044AFE1E6060" box="[296,486,292,314]" italics="true" pageId="1" pageNumber="414">Gelechia exoenota</emphasis>
<bibRefCitation id="EFB52988513D8A57FE52044AFD786060" author="Meyrick" box="[491,640,292,314]" pageId="1" pageNumber="414" refString="Meyrick, E. (1918) Descriptions of South African Micro-Lepidoptera. Annals of the Transvaal Museum, 6, 7 - 59." type="journal article" year="1918">Meyrick, 1918</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C33E07F2513D8A57FF2E0408FC6D6779" pageId="1" pageNumber="414" type="description">
<paragraph id="8B9B5479513D8A57FF2E0408FC6D6779" blockId="1.[151,1437,358,2003]" pageId="1" pageNumber="414">
<emphasis id="B950886B513D8A57FF2E0408FED96025" bold="true" box="[151,289,358,383]" pageId="1" pageNumber="414">Description</emphasis>
.
<emphasis id="B950886B513D8A57FE880408FE826025" bold="true" box="[305,378,358,383]" pageId="1" pageNumber="414">Head:</emphasis>
Smooth-scaled, yellow to light brown; labial palpus strongly up-curved, protruding far over head, segment 2 off-white to grey, usually with brown scales at base and before apex, slender, only slightly broader and approximately as long as segment 3, underside without modified scales, segment 3 slender, acute; ocelli absent; antenna in male slightly thicker than antenna in female, nearly 2/3 length of forewing, scape brown, shaft brown with narrow whitish rings at the base of all segments.
<emphasis id="B950886B513D8A57FCEB0498FC536355" bold="true" box="[850,939,502,527]" pageId="1" pageNumber="414">Thorax</emphasis>
: Brown, often mottled with ochre or black, tegulae usually darker, black; forewing elongate-ovate, wingspan 12.520.1 mm, brown or light brown; 3 black dots in cell, 2-3 black diffuse spots at base; costal margin usually distinctly mottled with black, big spots at 2/3 of costal margin, light brown or whitish diffuse spots at both margins at 2/33/4 length; hindwing trapezoidal, broad, tornus (anal angle) broadly rounded, terminal excavation indistinct, apex short, weakly pointed.
<emphasis id="B950886B513D8A57FAA607E8FA6C63C5" bold="true" box="[1311,1428,646,671]" pageId="1" pageNumber="414">Abdomen</emphasis>
: Segment VIII in male merged in basal half, sternum VIII broader than long, rounded or sub-rectangular, anterior margin with medial emargination, tergum VIII narrow, distinctly broader than long, posterior margin broadly rounded; female segment VII trapezoidal, about twice length of other abdominal segments; sternum II of both sexes with pair of venulae, medially with group of sensory setae (sensilla trichodea), apodemes well developed. Male genitalia with uncus a short rounded flap covered with long strong setae, sometimes (
<taxonomicName id="4C242FFA513D8A57FB750655FABE6209" box="[1228,1350,827,851]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="kuehnei">
<emphasis id="B950886B513D8A57FB750655FABE6209" box="[1228,1350,827,851]" italics="true" pageId="1" pageNumber="414">A. kuehnei</emphasis>
</taxonomicName>
) uncus deeply divided into two lobes (
<figureCitation id="131F48FC513D8A57FE4C0630FDB1622D" box="[501,585,862,887]" captionStart="FIGURES 20 22" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[305,1278,212,1914]" captionTargetId="figure@9.[273,1308,193,1924]" captionTargetPageId="9" captionText="FIGURES 20 22. Male genitalia of Armatophallus spp. 20. A. crudescens, syntype, Kenya, gen. slide 408 / 14, O. Bidzilya. 21. A. crudescens, Yemen, gen .. slide 552 / 14, O. Bidzilya. 22. A. kuehnei, gen. slide 182 / 12, O. Bidzilya." httpUri="https://zenodo.org/record/241827/files/figure.png" pageId="1" pageNumber="414">Fig. 22</figureCitation>
); paired rounded flaps (socii) arise from the base of uncus are developed in
<taxonomicName id="4C242FFA513D8A57FF2E06EDFEE362C0" box="[151,283,899,922]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="exoenota">
<emphasis id="B950886B513D8A57FF2E06EDFEE362C0" box="[151,283,899,922]" italics="true" pageId="1" pageNumber="414">A. exoenota</emphasis>
</taxonomicName>
(
<figureCitation id="131F48FC513D8A57FE9206ECFE4D62C1" box="[299,437,898,923]" captionStart="FIGURES 17 19" captionStartId="6.[151,264,1963,1986]" captionTargetBox="[318,1271,207,1925]" captionTargetId="figure@6.[281,1292,193,1930]" captionTargetPageId="6" captionText="FIGURES 17 19. Male genitalia of A. exoenota (Meyr). 17. Namibia, gen. slide 116 / 05, O. Bidzilya. 18. Tanzania, gen. slide 2310. 19. Cameroon, gen. slide 516 / 14, O. Bidzilya; 19 a. Phallus, Cameroon, gen. slide 394 / 14, O. Bidzilya." httpUri="https://zenodo.org/record/241826/files/figure.png" pageId="1" pageNumber="414">Figs. 1719</figureCitation>
); gnathos strong, long, evenly curved hook with pointed tip; tegumen weakly prolonged, trapezoidal, with deep, broad anterior emargination, lateral folds narrow, well developed; valva digitate, ventral margin usually with triangular tooth at about middle, distal portion moderately broad, densely covered with hairs, apex rounded, slightly exceeding the top of uncus; sacculus with broadly rounded base that is merged with valva, distal portion usually narrow extending to ½3/4 length of valva; posterior margin of vinculum divided by deep anteromedial emargination into big lobes covered with fine microtrichia and terminated into one or two (
<taxonomicName id="4C242FFA513D8A57FA3D0159FF136529" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="hackeri">
<emphasis id="B950886B513D8A57FA3D0159FF136529" italics="true" pageId="1" pageNumber="414">A. hackeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C242FFA513D8A57FF410135FE906529" box="[248,360,1115,1139]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B950886B513D8A57FF410135FE906529" box="[248,360,1115,1139]" italics="true" pageId="1" pageNumber="414">A. indicus</emphasis>
</taxonomicName>
) projections (
<figureCitation id="131F48FC513D8A57FDBD0134FD736529" box="[516,651,1114,1139]" captionStart="FIGURES 23 25" captionStartId="10.[151,264,1940,1963]" captionTargetBox="[325,1257,236,1910]" captionTargetId="figure@10.[293,1293,193,1920]" captionTargetPageId="10" captionText="FIGURES 23 25. Male genitalia of Armatophallus spp. 23. A. akagericus sp. n., holotype. 23 a. Phallus, gen. slide 426 / 14, O. Bidzilya. 24. A. hackeri sp. n., holotype. 25. A. indicus sp. n., holotype." httpUri="https://zenodo.org/record/241828/files/figure.png" pageId="1" pageNumber="414">Figs. 24, 25</figureCitation>
), except in
<taxonomicName id="4C242FFA513D8A57FCB20135FC796529" box="[779,897,1115,1139]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="kuehnei">
<emphasis id="B950886B513D8A57FCB20135FC796529" box="[779,897,1115,1139]" italics="true" pageId="1" pageNumber="414">A. kuehnei</emphasis>
</taxonomicName>
(
<figureCitation id="131F48FC513D8A57FC290134FC1C6529" box="[912,996,1114,1139]" captionStart="FIGURES 20 22" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[305,1278,212,1914]" captionTargetId="figure@9.[273,1308,193,1924]" captionTargetPageId="9" captionText="FIGURES 20 22. Male genitalia of Armatophallus spp. 20. A. crudescens, syntype, Kenya, gen. slide 408 / 14, O. Bidzilya. 21. A. crudescens, Yemen, gen .. slide 552 / 14, O. Bidzilya. 22. A. kuehnei, gen. slide 182 / 12, O. Bidzilya." httpUri="https://zenodo.org/record/241827/files/figure.png" pageId="1" pageNumber="414">Fig. 22</figureCitation>
); saccus short, broadly rounded with xshaped sclerotized support; phallus basally bulbous, distal portion narrow, straight or weakly curved, apex with 1 3 lateral tooth-shaped processes and with 13 horn-shaped arms; ductus ejaculatorius with very long, coiled, strongly sclerotized lamina. Female genitalia with papilla analis sub-ovate, covered with setae; apophyses posteriores straight, long and narrow; segment VIII sub-rectangular, sternum VIII evenly sclerotized, densely covered antero-medially with microtrichia, anterior margin usually strongly sclerotized with two sclerites curved inwards and connected medially near ostium (
<figureCitation id="131F48FC513D8A57FD0A005CFCB96411" box="[691,833,1330,1355]" captionStart="FIGURES 26 28" captionStartId="11.[151,264,1960,1983]" captionTargetBox="[295,1289,213,1915]" captionTargetId="figure@11.[269,1314,193,1933]" captionTargetPageId="11" captionText="FIGURES 26 28. Female genitalia of Armatophallus exoenota Meyr. 26. Namibia, gen. slide 126 / 05, O. Bidzilya. 27. Cameroon, gen. slide 395 / 14, O. Bidzilya. 28. Holotype of A. xylophaea." httpUri="https://zenodo.org/record/241829/files/figure.png" pageId="1" pageNumber="414">Figs. 2628</figureCitation>
), narrow postero-medial sclerite developed in some species (
<taxonomicName id="4C242FFA513D8A57FF450039FE7C6434" box="[252,388,1367,1390]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="exoenota">
<emphasis id="B950886B513D8A57FF450039FE7C6434" box="[252,388,1367,1390]" italics="true" pageId="1" pageNumber="414">A. exoenota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C242FFA513D8A57FE2E0039FDCE6434" box="[407,566,1366,1390]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="crudescens">
<emphasis id="B950886B513D8A57FE2E0039FDCE6434" box="[407,566,1366,1390]" italics="true" pageId="1" pageNumber="414">A. crudescens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C242FFA513D8A57FDF00039FD3B6434" box="[585,707,1366,1390]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="kuehnei">
<emphasis id="B950886B513D8A57FDF00039FD3B6434" box="[585,707,1366,1390]" italics="true" pageId="1" pageNumber="414">A. kuehnei</emphasis>
</taxonomicName>
); antrum broad, rounded, usually with sclerotized plate inside (
<figureCitation id="131F48FC513D8A57FF260014FED164C9" box="[159,297,1402,1427]" captionStart-0="FIGURES 26 28" captionStart-1="FIGURES 29 30" captionStart-2="FIGURES 31 32" captionStartId-0="11.[151,264,1960,1983]" captionStartId-1="12.[151,264,1923,1946]" captionStartId-2="13.[151,264,1898,1921]" captionTargetBox-0="[295,1289,213,1915]" captionTargetBox-1="[273,1286,219,1862]" captionTargetBox-2="[308,1266,206,1851]" captionTargetId-0="figure@11.[269,1314,193,1933]" captionTargetId-1="figure@12.[262,1324,193,1902]" captionTargetId-2="figure@13.[262,1324,193,1877]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="FIGURES 26 28. Female genitalia of Armatophallus exoenota Meyr. 26. Namibia, gen. slide 126 / 05, O. Bidzilya. 27. Cameroon, gen. slide 395 / 14, O. Bidzilya. 28. Holotype of A. xylophaea." captionText-1="FIGURES 29 30. Female genitalia of Armatophallus spp. 29. A. crudescens, Yemen, gen. slide 551 / 14, O. Bidzilya. 30. A. kuehnei, gen. slide 183 / 12, O. Bidzilya." captionText-2="FIGURES 31 32. Female genitalia of Armatophallus spp. 31. A. hackeri, Yemen, gen. slide 172 / 12, O. Bidzilya. 32. A. indicus, gen. slide 545 / 14, O. Bidzilya." httpUri-0="https://zenodo.org/record/241829/files/figure.png" httpUri-1="https://zenodo.org/record/241830/files/figure.png" httpUri-2="https://zenodo.org/record/241831/files/figure.png" pageId="1" pageNumber="414">Figs. 2631</figureCitation>
); apophyses anteriores broad at base, distal portion about as long or longer than segment VIII; ductus bursae moderately broad, distal portion usually with minute spikes, with abrupt transition to globular corpus bursae (
<taxonomicName id="4C242FFA513D8A57FF1900ADFEDD6480" box="[160,293,1475,1498]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="exoenota">
<emphasis id="B950886B513D8A57FF1900ADFEDD6480" box="[160,293,1475,1498]" italics="true" pageId="1" pageNumber="414">A. exoenota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C242FFA513D8A57FE8D00ADFE286481" box="[308,464,1475,1499]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="crudescens">
<emphasis id="B950886B513D8A57FE8D00ADFE286481" box="[308,464,1475,1499]" italics="true" pageId="1" pageNumber="414">A. crudescens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C242FFA513D8A57FE5900ADFDAF6481" box="[480,599,1475,1499]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="kuehnei">
<emphasis id="B950886B513D8A57FE5900ADFDAF6481" box="[480,599,1475,1499]" italics="true" pageId="1" pageNumber="414">A. kuehnei</emphasis>
</taxonomicName>
), signum at bottom of corpus bursae, strongly sclerotized prolonged plate with densely serrated edges and medial ridge; in some cases (
<taxonomicName id="4C242FFA513D8A57FCF00089FC4464A4" box="[841,956,1510,1534]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="hackeri">
<emphasis id="B950886B513D8A57FCF00089FC4464A4" box="[841,956,1510,1534]" italics="true" pageId="1" pageNumber="414">A. hackeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C242FFA513D8A57FC730089FBC364A4" box="[970,1083,1510,1534]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B950886B513D8A57FC730089FBC364A4" box="[970,1083,1510,1534]" italics="true" pageId="1" pageNumber="414">A. indicus</emphasis>
</taxonomicName>
) corpus bursae without signum (
<figureCitation id="131F48FC513D8A57FF260364FEDF6779" box="[159,295,1546,1571]" captionStart="FIGURES 31 32" captionStartId="13.[151,264,1898,1921]" captionTargetBox="[308,1266,206,1851]" captionTargetId="figure@13.[262,1324,193,1877]" captionTargetPageId="13" captionText="FIGURES 31 32. Female genitalia of Armatophallus spp. 31. A. hackeri, Yemen, gen. slide 172 / 12, O. Bidzilya. 32. A. indicus, gen. slide 545 / 14, O. Bidzilya." httpUri="https://zenodo.org/record/241831/files/figure.png" pageId="1" pageNumber="414">Figs. 31, 32</figureCitation>
), very narrow, gradually merges into the ductus bursae.
</paragraph>
</subSubSection>
<subSubSection id="C33E07F2513D8A57FF7E0340FE006731" pageId="1" pageNumber="414" type="distribution">
<paragraph id="8B9B5479513D8A57FF7E0340FE006731" blockId="1.[151,1437,358,2003]" pageId="1" pageNumber="414">
<emphasis id="B950886B513D8A57FF7E0340FE98671D" bold="true" box="[199,352,1582,1607]" pageId="1" pageNumber="414">Distribution.</emphasis>
<collectingCountry id="F33314E9513D8A57FED60340FDF1671D" box="[367,521,1582,1607]" name="South Africa" pageId="1" pageNumber="414">South Africa</collectingCountry>
,
<collectingCountry id="F33314E9513D8A57FDA50340FD87671D" box="[540,639,1582,1607]" name="Namibia" pageId="1" pageNumber="414">Namibia</collectingCountry>
,
<collectingCountry id="F33314E9513D8A57FD2B0340FCF5671D" box="[658,781,1582,1607]" name="Zimbabwe" pageId="1" pageNumber="414">Zimbabwe</collectingCountry>
,
<collectingCountry id="F33314E9513D8A57FCA60340FC85671D" box="[799,893,1582,1607]" name="Rwanda" pageId="1" pageNumber="414">Rwanda</collectingCountry>
,
<collectingCountry id="F33314E9513D8A57FC280340FBDF671D" box="[913,1063,1582,1607]" name="Gambia" pageId="1" pageNumber="414">The Gambia</collectingCountry>
,
<collectingCountry id="F33314E9513D8A57FB800341FB48671D" box="[1081,1200,1583,1607]" name="Cameroon" pageId="1" pageNumber="414">Cameroon</collectingCountry>
,
<collectingCountry id="F33314E9513D8A57FB7D0340FAD4671D" box="[1220,1324,1582,1607]" name="Tanzania" pageId="1" pageNumber="414">Tanzania</collectingCountry>
,
<collectingCountry id="F33314E9513D8A57FA870340FA60671D" box="[1342,1432,1582,1607]" name="Uganda" pageId="1" pageNumber="414">Uganda</collectingCountry>
,
<collectingCountry id="F33314E9513D8A57FF2E033DFF1B6731" box="[151,227,1619,1643]" name="Kenya" pageId="1" pageNumber="414">Kenya</collectingCountry>
,
<collectingCountry id="F33314E9513D8A57FF57033CFEA86731" box="[238,336,1618,1643]" name="Ethiopia" pageId="1" pageNumber="414">Ethiopia</collectingCountry>
,
<collectingCountry id="F33314E9513D8A57FEE2033DFE546730" box="[347,428,1619,1642]" name="Yemen" pageId="1" pageNumber="414">Yemen</collectingCountry>
,
<collectingCountry id="F33314E9513D8A57FE0E033CFE0C6731" box="[439,500,1618,1643]" name="India" pageId="1" pageNumber="414">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E07F2513D8A57FF7E0318FCD467D5" box="[199,812,1654,1679]" pageId="1" pageNumber="414" type="biology_ecology">
<paragraph id="8B9B5479513D8A57FF7E0318FCD467D5" blockId="1.[151,1437,358,2003]" box="[199,812,1654,1679]" pageId="1" pageNumber="414">
<emphasis id="B950886B513D8A57FF7E0318FEDF67D5" bold="true" box="[199,295,1654,1679]" pageId="1" pageNumber="414">Biology.</emphasis>
The early stages and hostplants are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33E07F2513D8A57FF7E03F4FE05678D" pageId="1" pageNumber="414" type="etymology">
<paragraph id="8B9B5479513D8A57FF7E03F4FE05678D" blockId="1.[151,1437,358,2003]" pageId="1" pageNumber="414">
<emphasis id="B950886B513D8A57FF7E03F4FEB667E9" bold="true" box="[199,334,1690,1715]" pageId="1" pageNumber="414">Etymology.</emphasis>
The generic name is derived from the Latin
<emphasis id="B950886B513D8A57FCF303F5FC5D67E8" box="[842,933,1691,1714]" italics="true" pageId="1" pageNumber="414">armatus</emphasis>
meaning armed or warrior, and refers to the arms on the apex of the phallus.
</paragraph>
</subSubSection>
<subSubSection id="C33E07F2513D8A54FF7E038CFA6965FB" lastPageId="2" lastPageNumber="415" pageId="1" pageNumber="414" type="discussion">
<paragraph id="8B9B5479513D8A54FF7E038CFC07601A" blockId="1.[151,1437,358,2003]" lastBlockId="2.[151,1437,151,1185]" lastPageId="2" lastPageNumber="415" pageId="1" pageNumber="414">
<emphasis id="B950886B513D8A57FF7E038CFEC367A1" bold="true" box="[199,315,1762,1787]" pageId="1" pageNumber="414">Remarks.</emphasis>
The new genus is recognizable by the following combination of characters: the phallus bearing apical arms, the posterior margin of the vinculum with well developed projections, the x-shaped support system of the saccus, the uncus densely covered with strong setae, a very long gnathos, and well developed lateral folds of the tegumen in the male genitalia. The female genitalia are characterized by sternum VIII covered with microtrichia, usually widened antrum, and the anterior margin of the sternite VIII with tendency to strong sclerotization. The presence of the long apical arms on the phallus seems to be unique within
<taxonomicName id="4C242FFA513D8A57FC4902F8FB8E66F5" box="[1008,1142,1942,1967]" class="Insecta" family="Gelechiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="family">Gelechiidae</taxonomicName>
and can be considered as autapomorphy of
<taxonomicName id="4C242FFA513D8A57FEDF02D5FDF46689" box="[358,524,1979,2003]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B950886B513D8A57FEDF02D5FDF46689" box="[358,524,1979,2003]" italics="true" pageId="1" pageNumber="414">Armatophallus</emphasis>
</taxonomicName>
. The well developed socii are rather rare in
<taxonomicName id="4C242FFA513D8A57FBA702D4FB5F6689" box="[1054,1191,1978,2003]" class="Insecta" family="Gelechiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="family">Gelechiidae</taxonomicName>
, but this character is known only in one species of the genus (
<taxonomicName id="4C242FFA513E8A54FDDB05F6FD1E61EA" box="[610,742,152,176]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="exoenota">
<emphasis id="B950886B513E8A54FDDB05F6FD1E61EA" box="[610,742,152,176]" italics="true" pageId="2" pageNumber="415">A. exoenota</emphasis>
</taxonomicName>
) and therefore cannot be considered an autapomorphy for the genus. The difference in the width of the antenna between male and female is an additional feature that may be used to define
<taxonomicName id="4C242FFA513E8A54FE87058EFE1C61A2" box="[318,484,224,248]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="414" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B950886B513E8A54FE87058EFE1C61A2" box="[318,484,224,248]" italics="true" pageId="2" pageNumber="415">Armatophallus</emphasis>
</taxonomicName>
species, but its importance for the classification remains unclear. It should be noted that sexual dimorphism in the width of the antenna is common in
<taxonomicName id="4C242FFA513E8A54FC7B046AFAF26046" authority="Meyrick, 1931" authorityName="Meyrick" authorityYear="1931" box="[962,1290,260,285]" class="Insecta" family="Gelechiidae" genus="Aphanostola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FC7B046AFBA86046" box="[962,1104,260,284]" italics="true" pageId="2" pageNumber="415">Aphanostola</emphasis>
Meyrick, 1931
</taxonomicName>
,
<taxonomicName id="4C242FFA513E8A54FAA5046BFEC7601A" authority="Meyrick, 1918" authorityName="Meyrick" authorityYear="1918" class="Insecta" family="Gelechiidae" genus="Leuronoma" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FAA5046BFA646046" box="[1308,1436,261,284]" italics="true" pageId="2" pageNumber="415">Leuronoma</emphasis>
<bibRefCitation id="EFB52988513E8A54FF2E0449FEC7601A" author="Meyrick" box="[151,319,295,320]" pageId="2" pageNumber="415" refString="Meyrick, E. (1918) Descriptions of South African Micro-Lepidoptera. Annals of the Transvaal Museum, 6, 7 - 59." type="journal article" year="1918">Meyrick, 1918</bibRefCitation>
</taxonomicName>
, and related unrevised genera of the subfamily Anomologinae.
</paragraph>
<paragraph id="8B9B5479513E8A54FF7E0423FE14624F" blockId="2.[151,1437,151,1185]" pageId="2" pageNumber="415">
Most of the diagnostic characters of
<taxonomicName id="4C242FFA513E8A54FDDF0422FCF4603E" box="[614,780,332,356]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="414" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B950886B513E8A54FDDF0422FCF4603E" box="[614,780,332,356]" italics="true" pageId="2" pageNumber="415">Armatophallus</emphasis>
</taxonomicName>
are found scattered across the subfamily
<taxonomicName id="4C242FFA513E8A54FB5D0422FA96603F" box="[1252,1390,332,357]" class="Insecta" family="Gelechiidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="subFamily" subFamily="Gelechiinae">Gelechiinae</taxonomicName>
: the x-shaped support system of the saccus, the uncus densely covered with strong setae and the well developed lateral folds of the tegumen are shared with
<taxonomicName id="4C242FFA513E8A54FDFC04FAFCA160F7" authority="Janse, 1951" authorityName="Janse" authorityYear="1951" box="[581,857,404,429]" class="Insecta" family="Gelechiidae" genus="Schizovalva" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FDFC04FAFD3360F6" box="[581,715,404,428]" italics="true" pageId="2" pageNumber="415">Schizovalva</emphasis>
Janse, 1951
</taxonomicName>
,
<emphasis id="B950886B513E8A54FCD004FBFC1560F6" box="[873,1005,405,428]" italics="true" pageId="2" pageNumber="415">Parapsectis</emphasis>
Meyrick, 1911, and
<taxonomicName id="4C242FFA513E8A54FB6704FAFF2B608A" authority="Billberg, 1820" authorityName="Billberg" authorityYear="1820" class="Insecta" family="Gelechiidae" genus="Athrips" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FB6704FAFAC960F6" box="[1246,1329,404,428]" italics="true" pageId="2" pageNumber="415">Athrips</emphasis>
Billberg, 1820
</taxonomicName>
. The microtrichia in sternum VIII in the female genitalia are widely distributed in the tribe
<taxonomicName id="4C242FFA513E8A54FB6B04D9FA64608A" box="[1234,1436,439,464]" class="Insecta" family="Gelechiidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="tribe" tribe="Gnorimoschemini">Gnorimoschemini</taxonomicName>
(
<bibRefCitation id="EFB52988513E8A54FF2704B2FE3960AE" author="Huemer" box="[158,449,476,501]" pageId="2" pageNumber="415" refString="Huemer, P. &amp; Karsholt, O. (2010) Gelechiidae II (Gelechiinae: Gnorimoschemini). In: Huemer, P., Karsholt, O. &amp; Nuss, M. (Eds.), Microlepidoptera of Europe. Vol. 6. Apollo Books, Stenstrup, pp. 1 - 586." type="book" year="2010">Huemer &amp; Karsholt 2010</bibRefCitation>
) but occur also in
<taxonomicName id="4C242FFA513E8A54FD2A04B2FCAE60AF" authority="Hubner" authorityName="Hubner" box="[659,854,476,501]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FD2A04B2FD0060AE" box="[659,760,476,500]" italics="true" pageId="2" pageNumber="415">Gelechia</emphasis>
Hübner
</taxonomicName>
, [1825],
<taxonomicName id="4C242FFA513E8A54FC0204B2FAF860AF" authority="Gozmany, 1955" authorityName="Gozmany" authorityYear="1955" box="[955,1280,476,501]" class="Insecta" family="Gelechiidae" genus="Mirificarma" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FC0204B2FBBC60AE" box="[955,1092,476,500]" italics="true" pageId="2" pageNumber="415">Mirificarma</emphasis>
Gozmány, 1955
</taxonomicName>
, and
<taxonomicName id="4C242FFA513E8A54FA8504B2FEDF6342" authority="Busck, 1939" authorityName="Busck" authorityYear="1939" class="Insecta" family="Gelechiidae" genus="Filatima" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FA8504B2FA6460AE" box="[1340,1436,476,500]" italics="true" pageId="2" pageNumber="415">Filatima</emphasis>
Busck, 1939
</taxonomicName>
. The narrow serrated signum of most
<taxonomicName id="4C242FFA513E8A54FD6C076EFC836342" box="[725,891,512,536]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="414" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B950886B513E8A54FD6C076EFC836342" box="[725,891,512,536]" italics="true" pageId="2" pageNumber="415">Armatophallus</emphasis>
</taxonomicName>
species is common in
<taxonomicName id="4C242FFA513E8A54FBC7076EFA606342" authority="Sattler, 1960" authorityName="Sattler" authorityYear="1960" box="[1150,1432,511,536]" class="Insecta" family="Gelechiidae" genus="Neofriseria" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FBC7076EFB056342" box="[1150,1277,512,536]" italics="true" pageId="2" pageNumber="415">Neofriseria</emphasis>
Sattler, 1960
</taxonomicName>
, and
<taxonomicName id="4C242FFA513E8A54FF71074AFE676367" authority="Hubner" authorityName="Hubner" box="[200,415,548,573]" class="Insecta" family="Gelechiidae" genus="Chionodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FF71074AFEB86366" box="[200,320,548,572]" italics="true" pageId="2" pageNumber="415">Chionodes</emphasis>
Hübner
</taxonomicName>
, [1825]. The evenly sclerotized sternum VIII with strongly sclerotized anterior margin in combination with a widened antrum indicates affinity of the new genus to
<taxonomicName id="4C242FFA513E8A54FC610726FBA0633A" box="[984,1112,584,608]" class="Insecta" family="Gelechiidae" genus="Neofriseria" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FC610726FBA0633A" box="[984,1112,584,608]" italics="true" pageId="2" pageNumber="415">Neofriseria</emphasis>
</taxonomicName>
. Most genera of
<taxonomicName id="4C242FFA513E8A54FAAF0729FA64633A" box="[1302,1436,583,608]" class="Insecta" family="Gelechiidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="subFamily" subFamily="Gelechiinae">Gelechiinae</taxonomicName>
have sternum and tergum VIII separate (not fused), and that is considered as a synapomorphy of the subfamily (
<bibRefCitation id="EFB52988513E8A54FF2707FEFE3B63F2" author="Huemer" box="[158,451,655,680]" pageId="2" pageNumber="415" refString="Huemer, P. &amp; Sattler, K. (1995) A taxonomic revision of Palaearctic Chionodes (Lepidoptera, Gelechiidae). Beitrage zur Entomolgie, 45, 3 - 108." type="journal article" year="1995">Huemer &amp; Sattler 1995</bibRefCitation>
,
<bibRefCitation id="EFB52988513E8A54FE6E07E1FD2B63F2" author="Karsholt" box="[471,723,655,680]" pageId="2" pageNumber="415" refString="Karsholt, O., Mutanen, M., Lee, S. &amp; Kaila, L. (2013) A molecular analysis of the Gelechiidae (Lepidoptera, Gelechioidea) with an interpretative grouping of its taxa. Systematic Entomology, 38, 334 - 348. http: // dx. doi. org / 10.1111 / syen. 12006" type="journal article" year="2013">Karsholt et al. 2013</bibRefCitation>
).
<taxonomicName id="4C242FFA513E8A54FD5407FEFC6B63F2" box="[749,915,656,680]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="414" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B950886B513E8A54FD5407FEFC6B63F2" box="[749,915,656,680]" italics="true" pageId="2" pageNumber="415">Armatophallus</emphasis>
</taxonomicName>
, along with
<taxonomicName id="4C242FFA513E8A54FB8307FEFB7463F2" box="[1082,1164,656,680]" class="Insecta" family="Gelechiidae" genus="Athrips" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FB8307FEFB7463F2" box="[1082,1164,656,680]" italics="true" pageId="2" pageNumber="415">Athrips</emphasis>
</taxonomicName>
and
<taxonomicName id="4C242FFA513E8A54FB6D07FEFA9B63FD" box="[1236,1379,656,679]" class="Insecta" family="Gelechiidae" genus="Parapsectris" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B950886B513E8A54FB6D07FEFA9B63FD" box="[1236,1379,656,679]" italics="true" pageId="2" pageNumber="415">Parapsectris</emphasis>
</taxonomicName>
, are characterized by the sternum and tergum VIII also separated laterally but fused in the basal portion (
<bibRefCitation id="EFB52988513E8A54FA8207DAFF2C63AA" author="Bidzilya" pageId="2" pageNumber="415" refString="Bidzilya, O. (2005) A review of the genus Athrips (Lepidoptera, Gelechiidae) in the Palaearctic region. Deutsche Entomologische Zeitschrift, 52 (1), 3 - 72. http: // dx. doi. org / 10.1002 / mmnd. 200310001" type="journal article" year="2005">Bidzilya 2005</bibRefCitation>
). This character indicates rather isolated position of these genera within
<taxonomicName id="4C242FFA513E8A54FBBF07B9FB7763AA" box="[1030,1167,727,752]" class="Insecta" family="Gelechiidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="subFamily" subFamily="Gelechiinae">Gelechiinae</taxonomicName>
, but their assignment to this subfamily seems justified.
</paragraph>
<paragraph id="8B9B5479513E8A54FF7E064EFA6965FB" blockId="2.[151,1437,151,1185]" pageId="2" pageNumber="415">
<taxonomicName id="4C242FFA513E8A54FF7E064EFE956262" box="[199,365,800,824]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="414" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B950886B513E8A54FF7E064EFE956262" box="[199,365,800,824]" italics="true" pageId="2" pageNumber="415">Armatophallus</emphasis>
</taxonomicName>
is rather homogeneous externally. All species have a similar wing pattern that consists of black dots at the base of the forewing, black spots at 2/3 of the costa, and a usually mottled black costal margin. Sexual dimorphism in the width of the antenna is observed in those
<taxonomicName id="4C242FFA513E8A54FCF10606FC1662DA" box="[840,1006,872,896]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="414" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B950886B513E8A54FCF10606FC1662DA" box="[840,1006,872,896]" italics="true" pageId="2" pageNumber="415">Armatophallus</emphasis>
</taxonomicName>
species that are known in both sexes. The male genitalia of all species are quite similar to each other too, except for
<taxonomicName id="4C242FFA513E8A54FB9D06E2FB5262FE" box="[1060,1194,908,932]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="exoenota">
<emphasis id="B950886B513E8A54FB9D06E2FB5262FE" box="[1060,1194,908,932]" italics="true" pageId="2" pageNumber="415">A. exoenota</emphasis>
</taxonomicName>
that has socii and
<taxonomicName id="4C242FFA513E8A54FA3D06E2FF166292" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="kuehnei">
<emphasis id="B950886B513E8A54FA3D06E2FF166292" italics="true" pageId="2" pageNumber="415">A. kuehnei</emphasis>
</taxonomicName>
that has a medially divided uncus and reduced projections of the posterior margin of the vinculum.
<taxonomicName id="4C242FFA513E8A54FA3D06DEFF1362B6" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="hackeri">
<emphasis id="B950886B513E8A54FA3D06DEFF1362B6" italics="true" pageId="2" pageNumber="415">A. hackeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C242FFA513E8A54FE9F06BAFE5862B6" box="[294,416,980,1004]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="indicius">
<emphasis id="B950886B513E8A54FE9F06BAFE5862B6" box="[294,416,980,1004]" italics="true" pageId="2" pageNumber="415">A. indicius</emphasis>
</taxonomicName>
can be separated from the rest of
<taxonomicName id="4C242FFA513E8A54FC9406BAFC2B62B6" box="[813,979,980,1004]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="414" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B950886B513E8A54FC9406BAFC2B62B6" box="[813,979,980,1004]" italics="true" pageId="2" pageNumber="415">Armatophallus</emphasis>
</taxonomicName>
species by a very long gnathos and the presence of lateral projection on the posterior margin of the vinculum. Contrary to the male genitalia, the female genitalia are more variable. They clearly divide into two groups. The first group comprises three species (
<emphasis id="B950886B513E8A54FA3D0172FDC86502" italics="true" pageId="2" pageNumber="415">
<taxonomicName id="4C242FFA513E8A54FA3D0172FF076502" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="exoenota">A. exoenota</taxonomicName>
,
<taxonomicName id="4C242FFA513E8A54FEB5012EFE536502" box="[268,427,1088,1112]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="crudescens">A. crudescens</taxonomicName>
,
<taxonomicName id="4C242FFA513E8A54FE01012EFDC86502" box="[440,560,1088,1112]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="kuehnei">A. kuehnei</taxonomicName>
</emphasis>
) with a corpus bursae that is clearly differentiated from the ductus bursae in combination with a well developed signum. Two species (
<emphasis id="B950886B513E8A54FCEA010AFBAE6526" box="[851,1110,1124,1148]" italics="true" pageId="2" pageNumber="415">
<taxonomicName id="4C242FFA513E8A54FCEA010AFC376526" box="[851,975,1124,1148]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="hackeri">A. hackeri</taxonomicName>
,
<taxonomicName id="4C242FFA513E8A54FC59010AFBAE6526" box="[992,1110,1124,1148]" class="Insecta" family="Gelechiidae" genus="Armatophallus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="indicus">A. indicus</taxonomicName>
</emphasis>
) with a ductus bursae that gradually merges into the very narrow corpus bursae in combination with reduced signum form the second group.
</paragraph>
</subSubSection>
</treatment>
</document>