589 lines
85 KiB
XML
589 lines
85 KiB
XML
<document id="480590F31ED6DA7B535D47D448C73AC6" ID-DOI="10.5281/zenodo.4525775" ID-ISSN="1638-9387" ID-RefBankID="FFB7FFA2FF95FFF4165CFF8EFFC6077C" ID-Zenodo-Dep="4525775" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612879640108" checkinUser="felipe" docAuthor="Salvini-Plawen, Luitfried von" docDate="2008" docId="0394ED6FFFB90105EF91FAB02163FB29" docLanguage="en" docName="Zoosystema.30.4.873-897.pdf" docOrigin="Zoosystema 30 (4)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Alexandromenia gulaglandulata Salvini-Plawen 2008, n. sp." docType="treatment" docVersion="3" lastPageNumber="896" masterDocId="FFAD9517FFA90112EF3CFF9C2008FFA3" masterDocTitle="Contributions to West European Cavibelonia (Mollusca, Solenogastres) with two new species" masterLastPageNumber="897" masterPageNumber="873" pageNumber="889" updateTime="1698869147205" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="AA06B350C313BEDB8EA104EBD0EF74AB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="498161323D19A526824F0C6FEA5F8A9F">
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<mods:title id="56C49D26503C8C18FEEB9DC7F3A7665E">Contributions to West European Cavibelonia (Mollusca, Solenogastres) with two new species</mods:title>
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</mods:titleInfo>
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<mods:name id="98ABBC37FA8D9A553C41B6F0FE73E77F" type="personal">
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<mods:role id="41E1436A9967C0B2DB3D3171FB616E0A">
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<mods:roleTerm id="D8D1A88D9FA112DBDEA4427BC62BF4EE">Author</mods:roleTerm>
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<mods:namePart id="1CE715E706B313AE3ACB41645D4A86C0">Salvini-Plawen, Luitfried von</mods:namePart>
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<mods:affiliation id="DB61D64ABF40EBAD8948F021351DD65A">Zoologie, Universität Wien, Althanstr. 14, A- 1090 Vienna (Austria) luitfried. salvini-plawen @ univie. ac. at</mods:affiliation>
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<mods:typeOfResource id="E5427D47D685380960B3939CDBC4C0EB">text</mods:typeOfResource>
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<mods:titleInfo id="32FB1DB4709FAF799FA1FF9183FDFA47">
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<mods:title id="232521DDAD0E66A657C53428FD82139F">Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part id="DD1E4F22E80A0570BA74A7107391BE59">
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<mods:date id="B1E2F3946248EB587867DB8F8B267D5E">2008</mods:date>
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<mods:detail id="C8D47C4008CF0B57E6C85BFB9342786B" type="volume">
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<mods:number id="F994A894E2DE8437794292DC391BD51E">30</mods:number>
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<mods:number id="C6C2D4FC451DB5349C798F405419D209">4</mods:number>
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<mods:end id="B9E654EBF2805DD606D1F4F2C58FC084">897</mods:end>
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<mods:classification id="F1C4B96573E4AFB5CF6A29517FF15DE4">journal article</mods:classification>
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<mods:identifier id="463C948C207E81C57393E30384601DA4" type="DOI">10.5281/zenodo.4525775</mods:identifier>
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<mods:identifier id="51D085CEDDFB9FF77B1CC43E06538D93" type="ISSN">1638-9387</mods:identifier>
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<treatment id="0394ED6FFFB90105EF91FAB02163FB29" LSID="urn:lsid:plazi:treatment:0394ED6FFFB90105EF91FAB02163FB29" httpUri="http://treatment.plazi.org/id/0394ED6FFFB90105EF91FAB02163FB29" lastPageId="23" lastPageNumber="896" pageId="16" pageNumber="889">
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<subSubSection id="C3270FF2FFB90102EF91FAB0220BFAC4" pageId="16" pageNumber="889" type="nomenclature">
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<paragraph id="8B825C79FFB90102EF91FAB0220BFAC4" blockId="16.[173,587,1324,1383]" pageId="16" pageNumber="889">
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<taxonomicName id="4C3D27FAFFB90102EF91FAB02201FAE4" ID-CoL="BRBP" authority="Salvini-Plawen, 2008" authorityName="Salvini-Plawen" authorityYear="2008" box="[173,521,1324,1351]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="16" pageNumber="889" phylum="Mollusca" rank="species" species="gulaglandulata" status="sp. nov.">
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<emphasis id="B949806BFFB90102EF91FAB02201FAE4" bold="true" box="[173,521,1324,1351]" italics="true" pageId="16" pageNumber="889">Alexandromenia gulaglandulata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A3D10FFB90102ED2CFAB12243FAEB" box="[528,587,1325,1352]" pageId="16" pageNumber="889" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="130640FCFFB90102EFC7FAD0217AFAC4" box="[251,370,1356,1383]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="17.[113,124,648,665]" captionStartId-1="17.[113,124,1519,1536]" captionStartId-2="17.[677,688,789,806]" captionStartId-3="18.[113,124,1155,1172]" captionTargetBox-0="[191,569,215,611]" captionTargetBox-1="[114,646,735,1477]" captionTargetBox-2="[678,1207,256,745]" captionTargetBox-3="[127,1182,219,1100]" captionTargetId-0="figure-13@17.[191,570,214,611]" captionTargetId-1="figure-35@17.[103,656,710,1496]" captionTargetId-2="figure-54@17.[678,1208,214,745]" captionTargetId-3="figure-14@18.[90,1231,177,1118]" captionTargetPageId-0="17" captionTargetPageId-1="17" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="FIG. 16. — Alexandromenia gulaglandulata n. sp., holotype in preserved state (anterior end upper left), 21 mm long." captionText-1="FIG. 17. — Alexandromenia gulaglandulata n.sp., mantle sclerites. Scale bar: 50 μm." captionText-2="FIG. 18. — Alexandromenia gulaglandulata n. sp., cross section through mantle. Abbreviations: ca, anterior caecum of midgut; cu, cuticle;gl, lateroventral foregut glands;lm, longitudinal musculature;pap, epidermal papillae;rm, matrix with transverse musculature of body wall; sp2, radial spicules. Scale bar: 150 μm." captionText-3="FIG. 19. — Alexandromenia gulaglandulata n. sp., organization of the anterior body viewed from left. Abbreviations: ao, atrial sense organ; es, oesophagus; gb, buccal ganglion; gc, cerebral ganglion; gl, lateroventral foregut glandular organ; go, gonad; ma, mantle; mg, midgut; pi, pedal pit; pgl, peripheral packages of pre-radular glands; ph, pharynx; vrs, ventral radula sack. Scale bar: 1000 μm." figureDoi-0="http://doi.org/10.5281/zenodo.4525809" figureDoi-1="http://doi.org/10.5281/zenodo.4525813" figureDoi-2="http://doi.org/10.5281/zenodo.4525811" figureDoi-3="http://doi.org/10.5281/zenodo.4525815" httpUri-0="https://zenodo.org/record/4525809/files/figure.png" httpUri-1="https://zenodo.org/record/4525813/files/figure.png" httpUri-2="https://zenodo.org/record/4525811/files/figure.png" httpUri-3="https://zenodo.org/record/4525815/files/figure.png" pageId="16" pageNumber="889">Figs 16-19</figureCitation>
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;
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<figureCitation id="130640FCFFB90102EE42FAD021A6FAC4" box="[382,430,1356,1383]" captionStart="FIG" captionStartId="19.[113,124,881,898]" captionTargetBox="[147,1129,248,821]" captionTargetId="figure-13@19.[113,1209,221,856]" captionTargetPageId="19" captionText="FIG. 20. — Two types of the outlet of the lateroventral foregut glandular organs with terminal cone (and opening papilla) within Amphimeniidae (schematised): A, simple cone as in Alexandromenia gulaglandulata n. sp.; B, cone with internal ductule such as in A. antarctica Salvini-Plawen, 1978. Abbreviations: con, cone; cp, cone pouch (sheath); dg, duct of glandular organ; du, ductule of cone; pa, opening papilla; ph, pharyngeal cavity; ri, median ridge of duct." figureDoi="http://doi.org/10.5281/zenodo.4525817" httpUri="https://zenodo.org/record/4525817/files/figure.png" pageId="16" pageNumber="889">20A</figureCitation>
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;
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<figureCitation id="130640FCFFB90102EE85FAD021F3FAC4" box="[441,507,1356,1383]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="20.[113,124,1075,1092]" captionStartId-1="20.[677,688,1436,1453]" captionStartId-2="21.[113,124,1032,1049]" captionStartId-3="22.[113,124,1073,1090]" captionTargetBox-0="[117,1206,214,1031]" captionTargetBox-1="[820,1062,1166,1394]" captionTargetBox-2="[203,1163,219,985]" captionTargetBox-3="[117,1206,215,1031]" captionTargetId-0="figure-14@20.[117,1206,214,1031]" captionTargetId-1="figure-69@20.[811,1075,1151,1405]" captionTargetId-2="figure-13@21.[200,1173,199,1000]" captionTargetId-3="figure-14@22.[117,1206,214,1031]" captionTargetPageId-0="20" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionText-0="FIG. 21. — Alexandromenia gulaglandulata n. sp., cross section through posterior pharynx, just anterior to radula. Abbreviations: ds, dorsal sinus; gl, lateroventral foregut glands; pgl, pre-radular packages of glands; ph, pharynx. Scale bar: 150 μm." captionText-1="FIG. 22. — Alexandromenia gulaglandulata n. sp.,one radula plate. Scale bar: 20 μm." captionText-2="FIG. 23.— Alexandromenia gulaglandulata n.sp.,organization of the posterior body viewed from left.Abbreviations:co, suprarectal commissure;div, diverticula;fo, pedal folds (foot);ma, mantle;mg, midgut;pc, pericardium;ro, respiratory organs;rs, receptaculum seminis; sd, spawning duct; sdg, spawning duct glands; so, dorsoterminal sense organ; spg, suprapallial glands. Scale bar: 500 μm." captionText-3="FIG. 24.— Alexandromenia gulaglandulata n. sp.,cross section through anterior mantle cavity with irregular,slender diverticula.Abbreviations: div, diverticle; gl, suprapallial glands; mc, mantle cavity; sd, spawning duct; sdg, spawning duct glands. Scale bar: 150 μm." figureDoi-0="http://doi.org/10.5281/zenodo.4525819" figureDoi-1="http://doi.org/10.5281/zenodo.4525821" figureDoi-2="http://doi.org/10.5281/zenodo.4525823" figureDoi-3="http://doi.org/10.5281/zenodo.4525825" httpUri-0="https://zenodo.org/record/4525819/files/figure.png" httpUri-1="https://zenodo.org/record/4525821/files/figure.png" httpUri-2="https://zenodo.org/record/4525823/files/figure.png" httpUri-3="https://zenodo.org/record/4525825/files/figure.png" pageId="16" pageNumber="889">21-24</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3270FF2FFB90102EF4DFA1424BDFCFA" pageId="16" pageNumber="889" type="materials_examined">
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<paragraph id="8B825C79FFB90102EF4DFA142172FA19" blockId="16.[112,647,1416,1466]" pageId="16" pageNumber="889">
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<materialsCitation id="3B555624FFB90102EF4DFA142172FA19" collectionCode="MNHN" pageId="16" pageNumber="889" specimenCode="MNHN 9835" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5486E2DBFFB90102EF4DFA1420D0FA3D" box="[113,216,1416,1439]" pageId="16" pageNumber="889" type="holotype">HOLOTYPE</typeStatus>
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. — Sclerites, section series on slides; midbody in alcohol (
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<specimenCode id="DB9BF402FFB90102EFE2FA3E2167FA19" box="[222,367,1442,1466]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="889">MNHN 9835</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8B825C79FFB90102EF4DFA492267FA4E" blockId="16.[113,623,1492,1517]" box="[113,623,1492,1517]" pageId="16" pageNumber="889">
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<collectionCode id="ED2CC4BCFFB90102EF4DFA49210CFA48" box="[113,260,1493,1516]" pageId="16" pageNumber="889">TYPE LOCALITY</collectionCode>
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. —
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<taxonomicName id="4C3D27FAFFB90102EE0CFA4821CDFA4E" authority="Cruise" authorityName="Cruise" box="[304,453,1492,1517]" class="Insecta" family="Coccinellidae" genus="Thalassa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="889" phylum="Arthropoda" rank="genus">
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<emphasis id="B949806BFFB90102EE0CFA482176FA4F" box="[304,382,1492,1516]" italics="true" pageId="16" pageNumber="889">Thalassa</emphasis>
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Cruise
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</taxonomicName>
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73 Z, stn Z 435.
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</paragraph>
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<paragraph id="8B825C79FFB90102EF4DF99B226BF900" blockId="16.[112,648,1542,1699]" pageId="16" pageNumber="889">
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||
<collectionCode id="ED2CC4BCFFB90102EF4DF99B2130F9BE" box="[113,312,1543,1566]" pageId="16" pageNumber="889">MATERIAL EXAMINED</collectionCode>
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. —
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<emphasis id="B949806BFFB90102EE5CF99B223DF9BD" bold="true" box="[352,565,1542,1567]" pageId="16" pageNumber="889">West European Basin.</emphasis>
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<taxonomicName id="4C3D27FAFFB90102ED05F99B20B8F99A" authority="Cruise" authorityName="Cruise" class="Insecta" family="Coccinellidae" genus="Thalassa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="889" phylum="Arthropoda" rank="genus">
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<emphasis id="B949806BFFB90102ED05F99B228EF9BC" box="[569,646,1543,1567]" italics="true" pageId="16" pageNumber="889">Thalassa</emphasis>
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Cruise
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</taxonomicName>
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73 Z (
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<collectionCode id="ED2CC4BCFFB90102EFD2F9BD2151F99A" box="[238,345,1569,1593]" pageId="16" pageNumber="889">CENTOB</collectionCode>
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), stn Z 435,
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<geoCoordinate id="EE093ABEFFB90102EEF2F9BD222CF99A" box="[462,548,1569,1593]" degrees="48" direction="north" minutes="40" orientation="latitude" pageId="16" pageNumber="889" precision="925" value="48.666668">48°40’N</geoCoordinate>
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,
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<geoCoordinate id="EE093ABEFFB90102ED10F9BD228AF99A" box="[556,642,1569,1593]" degrees="09" direction="west" minutes="53" orientation="longitude" pageId="16" pageNumber="889" precision="925" value="-9.883333">09°53’W</geoCoordinate>
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,
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<quantity id="4CC5F19CFFB90102EF4CF9A020B6F9F7" box="[112,190,1596,1620]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="16" pageNumber="889" unit="m" value="1050.0">1050 m</quantity>
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,
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<date id="FF837AB9FFB90102EFFBF9A0213AF9F7" box="[199,306,1596,1620]" pageId="16" pageNumber="889" value="1973-10-26">26.X.1973</date>
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,
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<specimenCount id="9D3B97F0FFB90102EE00F9A021A4F9F7" box="[316,428,1596,1620]" pageId="16" pageNumber="889" type="generic">1 specimen</specimenCount>
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||
,
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||
<quantity id="4CC5F19CFFB90102EE89F9A021F3F9F7" box="[437,507,1596,1620]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="16" pageNumber="889" unit="mm" value="21.0">21 mm</quantity>
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||
long, without keel formation (
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<figureCitation id="130640FCFFB90102EE2FF9CA2156F9CD" box="[275,350,1622,1646]" captionStart="FIG" captionStartId="17.[113,124,648,665]" captionTargetBox="[191,569,215,611]" captionTargetId="figure-13@17.[191,570,214,611]" captionTargetPageId="17" captionText="FIG. 16. — Alexandromenia gulaglandulata n. sp., holotype in preserved state (anterior end upper left), 21 mm long." figureDoi="http://doi.org/10.5281/zenodo.4525809" httpUri="https://zenodo.org/record/4525809/files/figure.png" pageId="16" pageNumber="889">Fig. 16</figureCitation>
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||
). Th e preserved animal has a rounded, coiled body, somewhat higher (
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<quantity id="4CC5F19CFFB90102ED34F9ED2289F92A" box="[520,641,1649,1673]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="16" pageNumber="889" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.8-3.5 mm</quantity>
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||
) than broad (
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||
<quantity id="4CC5F19CFFB90102EFD6F917216CF900" box="[234,356,1675,1699]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="16" pageNumber="889" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4-2.8 mm</quantity>
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), and is thicker anteriorly.
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</paragraph>
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<paragraph id="8B825C79FFB90102ED87FCBB24BDFCFA" blockId="16.[678,1209,807,857]" pageId="16" pageNumber="889">
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Both body ends of the specimen were serially cross sectioned in paraffi n of
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||
<quantity id="4CC5F19CFFB90102ECB1FCDD23C6FCFA" box="[909,974,833,857]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="889" unit="mm" value="10.0">10 µm</quantity>
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and stained with Azan.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C3270FF2FFB90102ED99FCED242BFC1D" pageId="16" pageNumber="889" type="etymology">
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<paragraph id="8B825C79FFB90102ED99FCED242BFC1D" blockId="16.[677,1210,881,958]" pageId="16" pageNumber="889">
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||
ETYMOLOGY. — Latin:
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<emphasis id="B949806BFFB90102ECAAFCED23B7FC2A" box="[918,959,881,905]" italics="true" pageId="16" pageNumber="889">gula</emphasis>
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||
, gullet, throat, pharynx;
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<emphasis id="B949806BFFB90102ED9AFC1722F1FC00" box="[678,761,907,931]" italics="true" pageId="16" pageNumber="889">glandula</emphasis>
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||
, small gland;
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||
<emphasis id="B949806BFFB90102ECBAFC1023BCFC00" box="[902,948,908,931]" italics="true" pageId="16" pageNumber="889">-atus</emphasis>
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||
, provided with. Referring to the different
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||
<typeStatus id="5486E2DBFFB90102EC03FC3A2379FC1D" box="[831,881,934,958]" pageId="16" pageNumber="889">types</typeStatus>
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||
of foregut glands.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C3270FF2FFB90102ED99FC4A24ADFA6A" pageId="16" pageNumber="889" type="diagnosis">
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||
<paragraph id="8B825C79FFB90102ED99FC4A24ADFA6A" blockId="16.[675,1211,981,1481]" pageId="16" pageNumber="889">
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DIAGNOSIS. — 21 × max.
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||
<quantity id="4CC5F19CFFB90102EC9CFC4923E7FC4E" box="[928,1007,981,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="16" pageNumber="889" unit="mm" value="3.5">3.5 mm</quantity>
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||
, without keel formation. Mantle papillae long-stalked with most distal swellings at the same level. With 18-11 (terminally 4) pedal folds that do not enter the mantle cavity; mantle cavity posteriorly with 16 respiratory folds, anteriorly with irregular, slender sacculations; with suprapallial glands. Transverse body wall musculature embedded in matrix. Atrial papillae slender, single. With four sets of foregut glands: pharyngeal glands, pre-radular packages of glands, paired lateroventral foregut glandular organs with duct leading into pouch through ductule-less cone and not opening on a papilla, and oesophageal glands. Radula plates
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||
<quantity id="4CC5F19CFFB90102EC17FA8F2384FA88" box="[811,908,1299,1323]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="16" pageNumber="889" unit="mm" value="60.0" valueMax="65.0" valueMin="55.0">55-65 µm</quantity>
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wide, with one denticle at each side; radula support (bolster, odontophore) symmetrically arranged turgescent cells; midgut caecum short, unpaired throughout. Fused section of spawning ducts anteriorly still with glands, muscular opening ventral at anterior border of mantle cavity opening. Suprarectal commissure long; dorsoterminal sense organ above mantle cavity.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3270FF2FFB90105ED99FA772163FB29" lastPageId="23" lastPageNumber="896" pageId="16" pageNumber="889" type="description">
|
||
<paragraph id="8B825C79FFB90102ED99FA77233DF9A0" blockId="16.[674,1210,1515,1700]" box="[677,821,1515,1541]" pageId="16" pageNumber="889">DESCRIPTION</paragraph>
|
||
<paragraph id="8B825C79FFB90102ED99F9962381F987" blockId="16.[674,1210,1515,1700]" box="[677,905,1546,1572]" pageId="16" pageNumber="889">
|
||
<emphasis id="B949806BFFB90102ED99F9962381F987" box="[677,905,1546,1572]" italics="true" pageId="16" pageNumber="889">Mantle and body wall</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFB90103ED99F9B6228DF907" blockId="16.[674,1210,1515,1700]" lastBlockId="17.[111,647,1610,1700]" lastPageId="17" lastPageNumber="890" pageId="16" pageNumber="889">
|
||
The
|
||
<quantity id="4CC5F19CFFB90102EDEDF9B6235AF9E6" box="[721,850,1578,1605]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="16" pageNumber="889" unit="mm" value="200.0" valueMax="250.0" valueMin="150.0">150-250 µm</quantity>
|
||
thick cuticle with densely arranged, long-stalked epidermis papillae, whose distal swellings are nearly all at the same level (
|
||
<figureCitation id="130640FCFFB90102ECE7F9F5242FF927" box="[987,1063,1641,1668]" captionStart="FIG" captionStartId="17.[677,688,789,806]" captionTargetBox="[678,1207,256,745]" captionTargetId="figure-54@17.[678,1208,214,745]" captionTargetPageId="17" captionText="FIG. 18. — Alexandromenia gulaglandulata n. sp., cross section through mantle. Abbreviations: ca, anterior caecum of midgut; cu, cuticle;gl, lateroventral foregut glands;lm, longitudinal musculature;pap, epidermal papillae;rm, matrix with transverse musculature of body wall; sp2, radial spicules. Scale bar: 150 μm." figureDoi="http://doi.org/10.5281/zenodo.4525811" httpUri="https://zenodo.org/record/4525811/files/figure.png" pageId="16" pageNumber="889">Fig. 18</figureCitation>
|
||
). The sclerites are mostly eroded, but fragments and negatives in the cuticle show two kinds of acicular, slightly curved, hollow spicules (
|
||
<figureCitation id="130640FCFFB80103EE29F9F52156F927" box="[277,350,1641,1668]" captionStart="FIG" captionStartId="17.[113,124,1519,1536]" captionTargetBox="[114,646,735,1477]" captionTargetId="figure-35@17.[103,656,710,1496]" captionTargetPageId="17" captionText="FIG. 17. — Alexandromenia gulaglandulata n.sp., mantle sclerites. Scale bar: 50 μm." figureDoi="http://doi.org/10.5281/zenodo.4525813" httpUri="https://zenodo.org/record/4525813/files/figure.png" pageId="17" pageNumber="890">Fig.17</figureCitation>
|
||
): small, 60-90 ×
|
||
<quantity id="4CC5F19CFFB80103ED3FF9F52246F927" box="[515,590,1641,1668]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="17" pageNumber="890" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4-8 µm</quantity>
|
||
long, tangential needles, densely arranged in several layers,
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFB80103EC7DFF732480FEA5" blockId="17.[833,1160,233,265]" box="[833,1160,233,265]" pageId="17" pageNumber="890">cu pap lm sp</paragraph>
|
||
<caption id="DF420CF1FFB80103EF4DFD142238FD13" ID-DOI="http://doi.org/10.5281/zenodo.4525809" ID-Zenodo-Dep="4525809" httpUri="https://zenodo.org/record/4525809/files/figure.png" pageId="17" pageNumber="890" startId="17.[113,124,648,665]" targetBox="[191,569,215,611]" targetPageId="17">
|
||
<paragraph id="8B825C79FFB80103EF4DFD142238FD13" blockId="17.[113,646,647,688]" pageId="17" pageNumber="890">
|
||
FIG. 16. —
|
||
<taxonomicName id="4C3D27FAFFB80103EFEFFD1B21E9FD3B" authority="Salvini-Plawen, 2008" authorityName="Salvini-Plawen" authorityYear="2008" box="[211,481,647,664]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="17" pageNumber="890" phylum="Mollusca" rank="species" species="gulaglandulata" status="sp. nov.">
|
||
<emphasis id="B949806BFFB80103EFEFFD1B21E9FD3B" box="[211,481,647,664]" italics="true" pageId="17" pageNumber="890">Alexandromenia gulaglandulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A3D10FFB80103EED4FD142215FD3A" box="[488,541,648,665]" pageId="17" pageNumber="890" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype in preserved state (anterior end upper left), 21 mm long.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF420CF1FFB80103ED99FC892453FC23" ID-DOI="http://doi.org/10.5281/zenodo.4525811" ID-Zenodo-Dep="4525811" httpUri="https://zenodo.org/record/4525811/files/figure.png" pageId="17" pageNumber="890" startId="17.[677,688,789,806]" targetBox="[678,1207,256,745]" targetPageId="17">
|
||
<paragraph id="8B825C79FFB80103ED99FC892453FC23" blockId="17.[675,1211,788,899]" pageId="17" pageNumber="890">
|
||
FIG. 18. —
|
||
<taxonomicName id="4C3D27FAFFB80103EC3FFC882400FC86" authority="Salvini-Plawen, 2008" authorityName="Salvini-Plawen" authorityYear="2008" box="[771,1032,788,805]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="17" pageNumber="890" phylum="Mollusca" rank="species" species="gulaglandulata" status="sp. nov.">
|
||
<emphasis id="B949806BFFB80103EC3FFC882400FC86" box="[771,1032,788,805]" italics="true" pageId="17" pageNumber="890">Alexandromenia gulaglandulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A3D10FFB80103EB32FC892449FC85" box="[1038,1089,789,806]" pageId="17" pageNumber="890" rank="species">n. sp.</taxonomicNameLabel>
|
||
, cross section through mantle. Abbreviations:
|
||
<emphasis id="B949806BFFB80103EC8CFCB723CEFC9F" bold="true" box="[944,966,811,828]" pageId="17" pageNumber="890">ca</emphasis>
|
||
, anterior caecum of midgut;
|
||
<emphasis id="B949806BFFB80103ED9AFCDE22B3FCF0" bold="true" box="[678,699,834,851]" pageId="17" pageNumber="890">cu</emphasis>
|
||
, cuticle;
|
||
<emphasis id="B949806BFFB80103EDC3FCDE2306FCF0" bold="true" box="[767,782,834,851]" pageId="17" pageNumber="890">gl</emphasis>
|
||
, lateroventral foregut glands;
|
||
<emphasis id="B949806BFFB80103ECC9FCDE2402FCF0" bold="true" box="[1013,1034,834,851]" pageId="17" pageNumber="890">lm</emphasis>
|
||
, longitudinal musculature;
|
||
<emphasis id="B949806BFFB80103EDF7FCC422E3FCCA" bold="true" box="[715,747,856,873]" pageId="17" pageNumber="890">pap</emphasis>
|
||
, epidermal papillae;
|
||
<emphasis id="B949806BFFB80103ECB6FCC423A9FCCA" bold="true" box="[906,929,856,873]" pageId="17" pageNumber="890">rm</emphasis>
|
||
, matrix with transverse musculature of body wall;
|
||
<emphasis id="B949806BFFB80103EC2FFCF32339FC20" bold="true" box="[787,817,879,899]" pageId="17" pageNumber="890">
|
||
sp
|
||
<subScript id="17B95E3CFFB80103EC16FCEB2339FC20" attach="right" box="[810,817,887,899]" fontSize="5" pageId="17" pageNumber="890">2</subScript>
|
||
</emphasis>
|
||
, radial spicules. Scale bar: 150 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B825C79FFB80103ED9EFC4C2394FA8B" blockId="17.[674,1209,976,1320]" pageId="17" pageNumber="890">
|
||
and strong radial spicules (
|
||
<figureCitation id="130640FCFFB80103EC82FC4C2405FC48" box="[958,1037,976,1003]" captionStart="FIG" captionStartId="17.[677,688,789,806]" captionTargetBox="[678,1207,256,745]" captionTargetId="figure-54@17.[678,1208,214,745]" captionTargetPageId="17" captionText="FIG. 18. — Alexandromenia gulaglandulata n. sp., cross section through mantle. Abbreviations: ca, anterior caecum of midgut; cu, cuticle;gl, lateroventral foregut glands;lm, longitudinal musculature;pap, epidermal papillae;rm, matrix with transverse musculature of body wall; sp2, radial spicules. Scale bar: 150 μm." figureDoi="http://doi.org/10.5281/zenodo.4525811" httpUri="https://zenodo.org/record/4525811/files/figure.png" pageId="17" pageNumber="890">Fig. 18</figureCitation>
|
||
, sp
|
||
<subScript id="17B95E3CFFB80103EB0DFC412433FC4D" attach="right" box="[1073,1083,989,1006]" fontSize="7" pageId="17" pageNumber="890">2</subScript>
|
||
) measuring 120-200 ×
|
||
<quantity id="4CC5F19CFFB80103EC24FC6C2389FBA8" box="[792,897,1008,1035]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="17" pageNumber="890" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10-15 µm</quantity>
|
||
(the shorter and more slender ones towards ventral), some up to 225 ×
|
||
<quantity id="4CC5F19CFFB80103EB54FB8C24BCFB88" box="[1128,1204,1040,1067]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="890" unit="mm" value="25.0">25 µm</quantity>
|
||
. The low epidermis (only
|
||
<quantity id="4CC5F19CFFB80103EC89FBB3240FFBE9" box="[949,1031,1071,1098]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="17" pageNumber="890" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4-8 µm</quantity>
|
||
) is underlain by a
|
||
<quantity id="4CC5F19CFFB80103ED8BFBD322F5FBC9" box="[695,765,1103,1130]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="890" unit="mm" value="70.0">70 µm</quantity>
|
||
thick matrix. This contains the transverse (“circular”) musculature (
|
||
<figureCitation id="130640FCFFB80103EC88FBF3240CFB29" box="[948,1028,1135,1162]" captionStart="FIG" captionStartId="17.[677,688,789,806]" captionTargetBox="[678,1207,256,745]" captionTargetId="figure-54@17.[678,1208,214,745]" captionTargetPageId="17" captionText="FIG. 18. — Alexandromenia gulaglandulata n. sp., cross section through mantle. Abbreviations: ca, anterior caecum of midgut; cu, cuticle;gl, lateroventral foregut glands;lm, longitudinal musculature;pap, epidermal papillae;rm, matrix with transverse musculature of body wall; sp2, radial spicules. Scale bar: 150 μm." figureDoi="http://doi.org/10.5281/zenodo.4525811" httpUri="https://zenodo.org/record/4525811/files/figure.png" pageId="17" pageNumber="890">Fig. 18</figureCitation>
|
||
), which is rather scattered in the anterior body. The longitudinal musculature of the body wall is indistinctly grouped into bundles; bundles somewhat enlarged ventrally without,however,forming a distinct,paired musculus longitudinalis ventralis.
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFB80103ED99FAD12399FAC5" blockId="17.[674,1210,1356,1700]" box="[677,913,1356,1382]" pageId="17" pageNumber="890">
|
||
<emphasis id="B949806BFFB80103ED99FAD12399FAC5" box="[677,913,1356,1382]" italics="true" pageId="17" pageNumber="890">Foot and mantle cavity</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFB80103ED99FAF024B2F907" blockId="17.[674,1210,1356,1700]" pageId="17" pageNumber="890">
|
||
The posterior half of the pedal pit shows 18 slender folds which enter the pedal groove and extend along the oesophageal region; in the posterior body, 11 of these folds are reduced to eight below the anterior mantle cavity,ultimately to four folds directly in front of the opening of the mantle cavity.Th e mantle cavity forms 16 symmetrically arranged respiratory folds. Anterior to the ventral opening, these are replaced by peripheral, irregular, slender diverticula (up to
|
||
<quantity id="4CC5F19CFFB80103ED99F91522F1F907" box="[677,761,1673,1700]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="890" unit="mm" value="200.0">200 µm</quantity>
|
||
long) or sacculations of the roof (
|
||
<figureCitation id="130640FCFFB80103EB62F91524A5F907" box="[1118,1197,1673,1700]" captionStart="FIG" captionStartId="22.[113,124,1073,1090]" captionTargetBox="[117,1206,215,1031]" captionTargetId="figure-14@22.[117,1206,214,1031]" captionTargetPageId="22" captionText="FIG. 24.— Alexandromenia gulaglandulata n. sp.,cross section through anterior mantle cavity with irregular,slender diverticula.Abbreviations: div, diverticle; gl, suprapallial glands; mc, mantle cavity; sd, spawning duct; sdg, spawning duct glands. Scale bar: 150 μm." figureDoi="http://doi.org/10.5281/zenodo.4525825" httpUri="https://zenodo.org/record/4525825/files/figure.png" pageId="17" pageNumber="890">Fig. 24</figureCitation>
|
||
);
|
||
</paragraph>
|
||
<caption id="DF420CF1FFB80103EF4DFA73210CF9B4" ID-DOI="http://doi.org/10.5281/zenodo.4525813" ID-Zenodo-Dep="4525813" httpUri="https://zenodo.org/record/4525813/files/figure.png" pageId="17" pageNumber="890" startId="17.[113,124,1519,1536]" targetBox="[114,646,735,1477]" targetPageId="17">
|
||
<paragraph id="8B825C79FFB80103EF4DFA73210CF9B4" blockId="17.[112,648,1518,1559]" pageId="17" pageNumber="890">
|
||
FIG. 17. —
|
||
<taxonomicName id="4C3D27FAFFB80103EFFBFA7221CFFA5C" authority="Salvini-Plawen, 2008" authorityName="Salvini-Plawen" authorityYear="2008" box="[199,455,1518,1535]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="17" pageNumber="890" phylum="Mollusca" rank="species" species="gulaglandulata" status="sp. nov.">
|
||
<emphasis id="B949806BFFB80103EFFBFA7221CFFA5C" box="[199,455,1518,1535]" italics="true" pageId="17" pageNumber="890">Alexandromenia gulaglandulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A3D10FFB80103EEF7FA7321F4F9A3" box="[459,508,1519,1536]" pageId="17" pageNumber="890" rank="species">n.sp.</taxonomicNameLabel>
|
||
, mantle sclerites. Scale bar: 50 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF420CF1FFBB0100EF4DFB1F24B3FB62" ID-DOI="http://doi.org/10.5281/zenodo.4525815" ID-Zenodo-Dep="4525815" httpUri="https://zenodo.org/record/4525815/files/figure.png" pageId="18" pageNumber="891" startId="18.[113,124,1155,1172]" targetBox="[127,1182,219,1100]" targetPageId="18">
|
||
<paragraph id="8B825C79FFBB0100EF4DFB1F24B3FB62" blockId="18.[113,1211,1154,1217]" pageId="18" pageNumber="891">
|
||
FIG. 19. —
|
||
<taxonomicName id="4C3D27FAFFBB0100EFF1FB1E21DAFB30" authority="Salvini-Plawen, 2008" authorityName="Salvini-Plawen" authorityYear="2008" box="[205,466,1154,1171]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="18" pageNumber="891" phylum="Mollusca" rank="species" species="gulaglandulata" status="sp. nov.">
|
||
<emphasis id="B949806BFFBB0100EFF1FB1E21DAFB30" box="[205,466,1154,1171]" italics="true" pageId="18" pageNumber="891">Alexandromenia gulaglandulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A3D10FFBB0100EEE4FB1F2203FB37" box="[472,523,1155,1172]" pageId="18" pageNumber="891" rank="species">n. sp.</taxonomicNameLabel>
|
||
, organization of the anterior body viewed from left. Abbreviations:
|
||
<emphasis id="B949806BFFBB0100EB05FB1E2447FB30" bold="true" box="[1081,1103,1154,1171]" pageId="18" pageNumber="891">ao</emphasis>
|
||
, atrial sense organ;
|
||
<emphasis id="B949806BFFBB0100EF96FB0520B7FB09" bold="true" box="[170,191,1177,1194]" pageId="18" pageNumber="891">es</emphasis>
|
||
, oesophagus;
|
||
<emphasis id="B949806BFFBB0100EE06FB052159FB09" bold="true" box="[314,337,1177,1194]" pageId="18" pageNumber="891">gb</emphasis>
|
||
, buccal ganglion;
|
||
<emphasis id="B949806BFFBB0100EED6FB052208FB09" bold="true" box="[490,512,1177,1194]" pageId="18" pageNumber="891">gc</emphasis>
|
||
, cerebral ganglion;
|
||
<emphasis id="B949806BFFBB0100ED98FB0522BCFB09" bold="true" box="[676,692,1177,1194]" pageId="18" pageNumber="891">gl</emphasis>
|
||
, lateroventral foregut glandular organ;
|
||
<emphasis id="B949806BFFBB0100ECCAFB052405FB09" bold="true" box="[1014,1037,1177,1194]" pageId="18" pageNumber="891">go</emphasis>
|
||
, gonad;
|
||
<emphasis id="B949806BFFBB0100EB6AFB05247AFB09" bold="true" box="[1110,1138,1177,1194]" pageId="18" pageNumber="891">ma</emphasis>
|
||
, mantle;
|
||
<emphasis id="B949806BFFBB0100EF4DFB2C2085FB62" bold="true" box="[113,141,1200,1217]" pageId="18" pageNumber="891">mg</emphasis>
|
||
, midgut;
|
||
<emphasis id="B949806BFFBB0100EFE0FB2C20E4FB62" bold="true" box="[220,236,1200,1217]" pageId="18" pageNumber="891">pi</emphasis>
|
||
, pedal pit;
|
||
<emphasis id="B949806BFFBB0100EE75FB2C216DFB62" bold="true" box="[329,357,1200,1217]" pageId="18" pageNumber="891">pgl</emphasis>
|
||
, peripheral packages of pre-radular glands;
|
||
<emphasis id="B949806BFFBB0100EDEBFB2C22E5FB62" bold="true" box="[727,749,1200,1217]" pageId="18" pageNumber="891">ph</emphasis>
|
||
, pharynx;
|
||
<emphasis id="B949806BFFBB0100EC79FB2C2368FB62" bold="true" box="[837,864,1200,1217]" pageId="18" pageNumber="891">vrs</emphasis>
|
||
, ventral radula sack. Scale bar: 1000 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B825C79FFBB0100EF4DFA91222EF986" blockId="18.[111,647,1293,1573]" pageId="18" pageNumber="891">
|
||
they delimit the mantle cavity against the circular hindgut; the
|
||
<quantity id="4CC5F19CFFBB0100EFC0FAB12159FAEB" box="[252,337,1325,1352]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="891" unit="mm" value="800.0">800 µm</quantity>
|
||
long suprarectal commissure is located near the end of the pericardium. The muscular terminal portion of the unpaired spawning duct opens at the anterior border of the mantle cavity opening.From the suprarectal commissure backwards, the whole mantle cavity is coated by a dense mass of suprapallial glands. The terminal sense organ lies above the central mantle cavity (
|
||
<figureCitation id="130640FCFFBB0100EEF6F9962211F986" box="[458,537,1546,1573]" captionStart="FIG" captionStartId="21.[113,124,1032,1049]" captionTargetBox="[203,1163,219,985]" captionTargetId="figure-13@21.[200,1173,199,1000]" captionTargetPageId="21" captionText="FIG. 23.— Alexandromenia gulaglandulata n.sp.,organization of the posterior body viewed from left.Abbreviations:co, suprarectal commissure;div, diverticula;fo, pedal folds (foot);ma, mantle;mg, midgut;pc, pericardium;ro, respiratory organs;rs, receptaculum seminis; sd, spawning duct; sdg, spawning duct glands; so, dorsoterminal sense organ; spg, suprapallial glands. Scale bar: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.4525823" httpUri="https://zenodo.org/record/4525823/files/figure.png" pageId="18" pageNumber="891">Fig. 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFBB0100EF4DF9D62113F9C7" blockId="18.[113,649,1610,1700]" box="[113,283,1610,1636]" pageId="18" pageNumber="891">
|
||
<emphasis id="B949806BFFBB0100EF4DF9D62113F9C7" box="[113,283,1610,1636]" italics="true" pageId="18" pageNumber="891">Alimentary tract</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFBB0100EF4DF9F524B4F907" blockId="18.[113,649,1610,1700]" lastBlockId="18.[675,1212,1293,1700]" pageId="18" pageNumber="891">
|
||
The anterior sense organ is located within the common atriobuccal cavity and is delimited by the horseshoe-shaped ciliary tract; the atrial papillae are single and slender. Th e cerebral ganglion (
|
||
<quantity id="4CC5F19CFFBB0100EB5DFAB124BFFAEB" box="[1121,1207,1325,1352]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="891" unit="mm" value="400.0">400 µm</quantity>
|
||
broad,
|
||
<quantity id="4CC5F19CFFBB0100EDD1FAD02348FAC4" box="[749,832,1356,1383]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="891" unit="mm" value="200.0">200 µm</quantity>
|
||
long,
|
||
<quantity id="4CC5F19CFFBB0100ECBCFAD023DBFAC4" box="[896,979,1356,1383]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="891" unit="mm" value="200.0">200 µm</quantity>
|
||
high) is located above the buccal cavity between atrium and mouth opening; it has three pairs of cerebral nerves, each with a small basal swelling and a separate lateral origin of the connectives. Th e wide anterior pharynx shows several irregular foldings and is surrounded by distinct, but loosely arranged musculature as well as by indistinct pharyngeal glands. More posteriorly, the lumen becomes X-shaped and the musculature more distinctly circular; ventro-laterally, the paired foregut glandular organ opens pre-radularly
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFBA0101EF83FF7D246FFF59" blockId="19.[191,208,225,250]" box="[191,1127,225,250]" lastBlockId="19.[1109,1127,225,250]" pageId="19" pageNumber="892">A B</paragraph>
|
||
<caption id="DF420CF1FFBA0101EF4DFCED22DFFC65" ID-DOI="http://doi.org/10.5281/zenodo.4525817" ID-Zenodo-Dep="4525817" httpUri="https://zenodo.org/record/4525817/files/figure.png" pageId="19" pageNumber="892" startId="19.[113,124,881,898]" targetBox="[147,1129,248,821]" targetPageId="19">
|
||
<paragraph id="8B825C79FFBA0101EF4DFCED22DFFC65" blockId="19.[113,1211,881,966]" pageId="19" pageNumber="892">
|
||
FIG. 20. — Two types of the outlet of the lateroventral foregut glandular organs with terminal cone (and opening papilla) within
|
||
<taxonomicName id="4C3D27FAFFBA0101EF4FFC1420FCFC3A" authorityName="Salvini-Plawen" authorityYear="1972" box="[115,244,904,921]" class="Solenogastres" family="Amphimeniidae" kingdom="Animalia" order="Cavibelonia" pageId="19" pageNumber="892" phylum="Mollusca" rank="family">Amphimeniidae</taxonomicName>
|
||
(schematised):
|
||
<emphasis id="B949806BFFBA0101EE45FC1B218EFC3B" bold="true" box="[377,390,903,920]" pageId="19" pageNumber="892">A</emphasis>
|
||
, simple cone as in
|
||
<taxonomicName id="4C3D27FAFFBA0101ED16FC1B2326FC3B" authority="Salvini-Plawen, 2008" authorityName="Salvini-Plawen" authorityYear="2008" box="[554,814,903,920]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="19" pageNumber="892" phylum="Mollusca" rank="species" species="gulaglandulata" status="sp. nov.">
|
||
<emphasis id="B949806BFFBA0101ED16FC1B2326FC3B" box="[554,814,903,920]" italics="true" pageId="19" pageNumber="892">Alexandromenia gulaglandulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A3D10FFBA0101EC08FC14236FFC3A" box="[820,871,904,921]" pageId="19" pageNumber="892" rank="species">n. sp.</taxonomicNameLabel>
|
||
;
|
||
<emphasis id="B949806BFFBA0101EC52FC1B2373FC3B" bold="true" box="[878,891,903,920]" pageId="19" pageNumber="892">B</emphasis>
|
||
, cone with internal ductule such as in
|
||
<taxonomicName id="4C3D27FAFFBA0101EF4DFC02219BFC0C" authority="Salvini-Plawen, 1978" authorityName="Salvini-Plawen" authorityYear="1978" box="[113,403,926,943]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="19" pageNumber="892" phylum="Mollusca" rank="species" species="antarctica">
|
||
<emphasis id="B949806BFFBA0101EF4DFC0220D2FC0C" box="[113,218,926,943]" italics="true" pageId="19" pageNumber="892">A. antarctica</emphasis>
|
||
Salvini-Plawen, 1978
|
||
</taxonomicName>
|
||
. Abbreviations:
|
||
<emphasis id="B949806BFFBA0101ED24FC022231FC0C" bold="true" box="[536,569,926,943]" pageId="19" pageNumber="892">con</emphasis>
|
||
, cone;
|
||
<emphasis id="B949806BFFBA0101ED45FC022287FC0C" bold="true" box="[633,655,926,943]" pageId="19" pageNumber="892">cp</emphasis>
|
||
, cone pouch (sheath);
|
||
<emphasis id="B949806BFFBA0101EC6EFC022361FC0C" bold="true" box="[850,873,926,943]" pageId="19" pageNumber="892">dg</emphasis>
|
||
, duct of glandular organ;
|
||
<emphasis id="B949806BFFBA0101EB7FFC022451FC0C" bold="true" box="[1091,1113,926,943]" pageId="19" pageNumber="892">du</emphasis>
|
||
, ductule of cone;
|
||
<emphasis id="B949806BFFBA0101EF99FC2920B3FC65" bold="true" box="[165,187,949,966]" pageId="19" pageNumber="892">pa</emphasis>
|
||
, opening papilla;
|
||
<emphasis id="B949806BFFBA0101EE73FC29216DFC65" bold="true" box="[335,357,949,966]" pageId="19" pageNumber="892">ph</emphasis>
|
||
, pharyngeal cavity;
|
||
<emphasis id="B949806BFFBA0101ED30FC292210FC65" bold="true" box="[524,536,949,966]" pageId="19" pageNumber="892">ri</emphasis>
|
||
, median ridge of duct.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B825C79FFBA0101EF4DFB8C2373FBE9" blockId="19.[111,650,1040,1700]" lastBlockId="19.[674,1211,1040,1700]" pageId="19" pageNumber="892">
|
||
(
|
||
<figureCitation id="130640FCFFBA0101EF45FB8C20CAFB88" box="[121,194,1040,1067]" captionStart="FIG" captionStartId="18.[113,124,1155,1172]" captionTargetBox="[127,1182,219,1100]" captionTargetId="figure-14@18.[90,1231,177,1118]" captionTargetPageId="18" captionText="FIG. 19. — Alexandromenia gulaglandulata n. sp., organization of the anterior body viewed from left. Abbreviations: ao, atrial sense organ; es, oesophagus; gb, buccal ganglion; gc, cerebral ganglion; gl, lateroventral foregut glandular organ; go, gonad; ma, mantle; mg, midgut; pi, pedal pit; pgl, peripheral packages of pre-radular glands; ph, pharynx; vrs, ventral radula sack. Scale bar: 1000 μm." figureDoi="http://doi.org/10.5281/zenodo.4525815" httpUri="https://zenodo.org/record/4525815/files/figure.png" pageId="19" pageNumber="892">Fig.19</figureCitation>
|
||
). Th e region between these openings and the radula is widened and characterised by peripheral packages of an additional
|
||
<typeStatus id="5486E2DBFFBA0101EEB6FBD321B0FBC9" box="[394,440,1103,1130]" pageId="19" pageNumber="892">type</typeStatus>
|
||
of strongly stained glands; the cells of each package empty together via slender stalks into the pharynx (
|
||
<figureCitation id="130640FCFFBA0101EE81FB122205FB0A" box="[445,525,1166,1193]" captionStart="FIG" captionStartId="18.[113,124,1155,1172]" captionTargetBox="[127,1182,219,1100]" captionTargetId="figure-14@18.[90,1231,177,1118]" captionTargetPageId="18" captionText="FIG. 19. — Alexandromenia gulaglandulata n. sp., organization of the anterior body viewed from left. Abbreviations: ao, atrial sense organ; es, oesophagus; gb, buccal ganglion; gc, cerebral ganglion; gl, lateroventral foregut glandular organ; go, gonad; ma, mantle; mg, midgut; pi, pedal pit; pgl, peripheral packages of pre-radular glands; ph, pharynx; vrs, ventral radula sack. Scale bar: 1000 μm." figureDoi="http://doi.org/10.5281/zenodo.4525815" httpUri="https://zenodo.org/record/4525815/files/figure.png" pageId="19" pageNumber="892">Figs 19</figureCitation>
|
||
;
|
||
<figureCitation id="130640FCFFBA0101ED2BFB12223EFB0A" box="[535,566,1166,1193]" captionStart="FIG" captionStartId="20.[113,124,1075,1092]" captionTargetBox="[117,1206,214,1031]" captionTargetId="figure-14@20.[117,1206,214,1031]" captionTargetPageId="20" captionText="FIG. 21. — Alexandromenia gulaglandulata n. sp., cross section through posterior pharynx, just anterior to radula. Abbreviations: ds, dorsal sinus; gl, lateroventral foregut glands; pgl, pre-radular packages of glands; ph, pharynx. Scale bar: 150 μm." figureDoi="http://doi.org/10.5281/zenodo.4525819" httpUri="https://zenodo.org/record/4525819/files/figure.png" pageId="19" pageNumber="892">21</figureCitation>
|
||
). These packages differ distinctly from those of the paired lateroventral foregut glandular organs; both
|
||
<typeStatus id="5486E2DBFFBA0101ED73FB52228EFB4A" box="[591,646,1230,1257]" pageId="19" pageNumber="892">types</typeStatus>
|
||
partially overlap in the radula region (
|
||
<figureCitation id="130640FCFFBA0101EEC2FB712244FAAB" box="[510,588,1261,1288]" captionStart="FIG" captionStartId="20.[113,124,1075,1092]" captionTargetBox="[117,1206,214,1031]" captionTargetId="figure-14@20.[117,1206,214,1031]" captionTargetPageId="20" captionText="FIG. 21. — Alexandromenia gulaglandulata n. sp., cross section through posterior pharynx, just anterior to radula. Abbreviations: ds, dorsal sinus; gl, lateroventral foregut glands; pgl, pre-radular packages of glands; ph, pharynx. Scale bar: 150 μm." figureDoi="http://doi.org/10.5281/zenodo.4525819" httpUri="https://zenodo.org/record/4525819/files/figure.png" pageId="19" pageNumber="892">Fig. 21</figureCitation>
|
||
). The ducts of the paired foregut glandular organs show a median ridge and are encircled by musculature. The anterior-most portion of each duct curves medially and widens to a pouch. Th e median ridge of the pouch is enlarged to form a cone (
|
||
<figureCitation id="130640FCFFBA0101EEE9FA10223BFA04" box="[469,563,1420,1447]" captionStart="FIG" captionStartId="19.[113,124,881,898]" captionTargetBox="[147,1129,248,821]" captionTargetId="figure-13@19.[113,1209,221,856]" captionTargetPageId="19" captionText="FIG. 20. — Two types of the outlet of the lateroventral foregut glandular organs with terminal cone (and opening papilla) within Amphimeniidae (schematised): A, simple cone as in Alexandromenia gulaglandulata n. sp.; B, cone with internal ductule such as in A. antarctica Salvini-Plawen, 1978. Abbreviations: con, cone; cp, cone pouch (sheath); dg, duct of glandular organ; du, ductule of cone; pa, opening papilla; ph, pharyngeal cavity; ri, median ridge of duct." figureDoi="http://doi.org/10.5281/zenodo.4525817" httpUri="https://zenodo.org/record/4525817/files/figure.png" pageId="19" pageNumber="892">Fig. 20A</figureCitation>
|
||
) which, in this species, has no ductule. Both pouches open directly into a deep, slender cleft (“duct”) of the pharynx; no protruding opening papilla is developed. Each of the paired ducts of the glandular organs is, in the radula region, continuous with the dorsal and ventral limb of a collecting duct. Th is duct, in turn, receives the often ramified outleading ductules of the gland clusters; these clusters are thus more or less semicircularly arranged at both sides of the postradular foregut.
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFBA0106ED87FBD32195F986" blockId="19.[674,1211,1040,1700]" lastBlockId="20.[111,647,1166,1573]" lastPageId="20" lastPageNumber="893" pageId="19" pageNumber="892">
|
||
The radula bends into an unpaired anterio-ventral sack containing the radula plates. Th e plates are basally
|
||
<quantity id="4CC5F19CFFBA0101EDEBFB122317FB0A" box="[727,799,1166,1193]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="892" unit="mm" value="65.0">65 µm</quantity>
|
||
, distally
|
||
<quantity id="4CC5F19CFFBA0101EC44FB1223B5FB0A" box="[888,957,1166,1193]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="892" unit="mm" value="50.0">50 µm</quantity>
|
||
broad and about
|
||
<quantity id="4CC5F19CFFBA0101EB48FB1224B1FB0A" box="[1140,1209,1166,1193]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="892" unit="mm" value="30.0">30 µm</quantity>
|
||
high with a
|
||
<quantity id="4CC5F19CFFBA0101EC18FB322387FB6A" box="[804,911,1198,1225]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="19" pageNumber="892" unit="mm" value="28.5" valueMax="30.0" valueMin="27.0">27-30 µm</quantity>
|
||
long denticle at both lateral borders (
|
||
<figureCitation id="130640FCFFBA0101EDC3FB522342FB4A" box="[767,842,1230,1257]" captionStart="FIG" captionStartId="20.[677,688,1436,1453]" captionTargetBox="[820,1062,1166,1394]" captionTargetId="figure-69@20.[811,1075,1151,1405]" captionTargetPageId="20" captionText="FIG. 22. — Alexandromenia gulaglandulata n. sp.,one radula plate. Scale bar: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.4525821" httpUri="https://zenodo.org/record/4525821/files/figure.png" pageId="19" pageNumber="892">Fig. 22</figureCitation>
|
||
); no central denticles were detected. The radula support consists of a paired bolster of at least five turgescent cells each. The buccal ganglia (diam.150 ×
|
||
<quantity id="4CC5F19CFFBA0101EC16FAB12372FAEB" box="[810,890,1325,1352]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="892" unit="mm" value="100.0">100 µm</quantity>
|
||
) closely adjoin (distance
|
||
<quantity id="4CC5F19CFFBA0101EB53FAB124BBFAEB" box="[1135,1203,1325,1352]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="892" unit="mm" value="50.0">50 µm</quantity>
|
||
) and are located ventro-posterior to the radula sheath. The postradular foregut (oesophagus) initially narrows without structural change and is provided with circular musculature as well as several strong radial muscle bundles. It is surrounded by densely arranged, small groups of long-necked oesophageal glands. More posteriorly, this portion telescopically intrudes into the wide second oesophageal region, which has a weak circular musculature and groups of gland cells. The oesophagus narrows again and axially intrudes into the midgut. Th e anterior caecum of the midgut is short, extending above the oesophagus only. The midgut enlarges laterally to flank both sides of most of the oesophagus.Both the midgut and caecum have a middorsal ciliary tract. The dorsoventral muscle bundles, about
|
||
<quantity id="4CC5F19CFFBD0106EE2FFA91216EFA8B" box="[275,358,1293,1320]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="893" unit="mm" value="300.0">300 µm</quantity>
|
||
apart, cause regular ventrolateral midgut pouches. Below the pericardium and between the spawning ducts, the midgut continues into the hindgut, which is ciliated throughout; the anterior hindgut is surrounded by loose longitudinal muscles,posteriorly by both longitudinal and circular musculature.Th e
|
||
<quantity id="4CC5F19CFFBD0106EE1AFA57217FFA45" box="[294,375,1483,1510]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="893" unit="mm" value="800.0">800 µm</quantity>
|
||
long, medullar suprarectal commissure (diam.
|
||
<quantity id="4CC5F19CFFBD0106EE00FA77218AF9A5" box="[316,386,1515,1542]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="893" unit="mm" value="90.0">90 µm</quantity>
|
||
) is located below the end of the pericardium (
|
||
<figureCitation id="130640FCFFBD0106EE7EF9962199F986" box="[322,401,1546,1573]" captionStart="FIG" captionStartId="21.[113,124,1032,1049]" captionTargetBox="[203,1163,219,985]" captionTargetId="figure-13@21.[200,1173,199,1000]" captionTargetPageId="21" captionText="FIG. 23.— Alexandromenia gulaglandulata n.sp.,organization of the posterior body viewed from left.Abbreviations:co, suprarectal commissure;div, diverticula;fo, pedal folds (foot);ma, mantle;mg, midgut;pc, pericardium;ro, respiratory organs;rs, receptaculum seminis; sd, spawning duct; sdg, spawning duct glands; so, dorsoterminal sense organ; spg, suprapallial glands. Scale bar: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.4525823" httpUri="https://zenodo.org/record/4525823/files/figure.png" pageId="20" pageNumber="893">Fig. 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF420CF1FFBD0106EF4DFBAF2444FBF8" ID-DOI="http://doi.org/10.5281/zenodo.4525819" ID-Zenodo-Dep="4525819" httpUri="https://zenodo.org/record/4525819/files/figure.png" pageId="20" pageNumber="893" startId="20.[113,124,1075,1092]" targetBox="[117,1206,214,1031]" targetPageId="20">
|
||
<paragraph id="8B825C79FFBD0106EF4DFBAF2444FBF8" blockId="20.[113,1211,1075,1115]" pageId="20" pageNumber="893">
|
||
FIG. 21. —
|
||
<taxonomicName id="4C3D27FAFFBD0106EFEDFBAF21D0FBE7" authority="Salvini-Plawen, 2008" authorityName="Salvini-Plawen" authorityYear="2008" box="[209,472,1075,1092]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="20" pageNumber="893" phylum="Mollusca" rank="species" species="gulaglandulata" status="sp. nov.">
|
||
<emphasis id="B949806BFFBD0106EFEDFBAF21D0FBE7" box="[209,472,1075,1092]" italics="true" pageId="20" pageNumber="893">Alexandromenia gulaglandulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A3D10FFBD0106EEE3FBAF221BFBE7" box="[479,531,1075,1092]" pageId="20" pageNumber="893" rank="species">n. sp.</taxonomicNameLabel>
|
||
, cross section through posterior pharynx, just anterior to radula. Abbreviations:
|
||
<emphasis id="B949806BFFBD0106EF4FFBD52080FBF9" bold="true" box="[115,136,1097,1114]" pageId="20" pageNumber="893">ds</emphasis>
|
||
, dorsal sinus;
|
||
<emphasis id="B949806BFFBD0106EE3DFBD52119FBF9" bold="true" box="[257,273,1097,1114]" pageId="20" pageNumber="893">gl</emphasis>
|
||
, lateroventral foregut glands;
|
||
<emphasis id="B949806BFFBD0106ED36FBD5222EFBF9" bold="true" box="[522,550,1097,1114]" pageId="20" pageNumber="893">pgl</emphasis>
|
||
, pre-radular packages of glands;
|
||
<emphasis id="B949806BFFBD0106EC03FBD5235DFBF9" bold="true" box="[831,853,1097,1114]" pageId="20" pageNumber="893">ph</emphasis>
|
||
, pharynx. Scale bar: 150 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF420CF1FFBD0106ED99FA00233FFA60" ID-DOI="http://doi.org/10.5281/zenodo.4525821" ID-Zenodo-Dep="4525821" httpUri="https://zenodo.org/record/4525821/files/figure.png" pageId="20" pageNumber="893" startId="20.[677,688,1436,1453]" targetBox="[820,1062,1166,1394]" targetPageId="20">
|
||
<paragraph id="8B825C79FFBD0106ED99FA00233FFA60" blockId="20.[675,1211,1435,1475]" pageId="20" pageNumber="893">
|
||
FIG. 22. —
|
||
<taxonomicName id="4C3D27FAFFBD0106EDC6FA0723F1FA0F" authority="Salvini-Plawen, 2008" authorityName="Salvini-Plawen" authorityYear="2008" box="[762,1017,1435,1452]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="20" pageNumber="893" phylum="Mollusca" rank="species" species="gulaglandulata" status="sp. nov.">
|
||
<emphasis id="B949806BFFBD0106EDC6FA0723F1FA0F" box="[762,1017,1435,1452]" italics="true" pageId="20" pageNumber="893">Alexandromenia gulaglandulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A3D10FFBD0106ECC1FA002425FA0E" box="[1021,1069,1436,1453]" pageId="20" pageNumber="893" rank="species">n. sp.</taxonomicNameLabel>
|
||
,one radula plate. Scale bar: 20 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B825C79FFBD0106EF4DF9D62156F9C7" blockId="20.[113,646,1610,1700]" box="[113,350,1610,1636]" pageId="20" pageNumber="893">
|
||
<emphasis id="B949806BFFBD0106EF4DF9D62156F9C7" box="[113,350,1610,1636]" italics="true" pageId="20" pageNumber="893">Gonopericardial system</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFBD0106EF4DF9F524B4F907" blockId="20.[113,646,1610,1700]" lastBlockId="20.[675,1212,1546,1700]" pageId="20" pageNumber="893">
|
||
The animal is mature, with the paired gonad extending to the second oesophageal region; the eggs are up to
|
||
<quantity id="4CC5F19CFFBD0106EDE3F9962339F986" box="[735,817,1546,1573]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="893" unit="mm" value="150.0">150 µm</quantity>
|
||
in diameter. The gonopericardioducts are continuous with the dorso-frontal pericardium, which shows a paired anterior pouch. Both the ventricle and the single auricle of the heart are a free tube within the pericardium; the connection
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFBC0107EF8EFD8320CCFD90" blockId="21.[178,196,543,563]" box="[178,196,543,563]" pageId="21" pageNumber="894">rs</paragraph>
|
||
<paragraph id="8B825C79FFBC0107EBB2FD0A24A9FD09" blockId="21.[1166,1185,662,682]" box="[1166,1185,662,682]" pageId="21" pageNumber="894">ro</paragraph>
|
||
<caption id="DF420CF1FFBC0107EF4DFB942471FBE5" ID-DOI="http://doi.org/10.5281/zenodo.4525823" ID-Zenodo-Dep="4525823" httpUri="https://zenodo.org/record/4525823/files/figure.png" pageId="21" pageNumber="894" startId="21.[113,124,1032,1049]" targetBox="[203,1163,219,985]" targetPageId="21">
|
||
<paragraph id="8B825C79FFBC0107EF4DFB942471FBE5" blockId="21.[112,1211,942,1094]" pageId="21" pageNumber="894">
|
||
FIG. 23.—
|
||
<taxonomicName id="4C3D27FAFFBC0107EFFAFB9B21CCFBBB" authority="Salvini-Plawen, 2008" authorityName="Salvini-Plawen" authorityYear="2008" box="[198,452,1031,1048]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="21" pageNumber="894" phylum="Mollusca" rank="species" species="gulaglandulata" status="sp. nov.">
|
||
<emphasis id="B949806BFFBC0107EFFAFB9B21CCFBBB" box="[198,452,1031,1048]" italics="true" pageId="21" pageNumber="894">Alexandromenia gulaglandulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A3D10FFBC0107EEFBFB9421FFFBBA" box="[455,503,1032,1049]" pageId="21" pageNumber="894" rank="species">n.sp.</taxonomicNameLabel>
|
||
,organization of the posterior body viewed from left.Abbreviations:
|
||
<emphasis id="B949806BFFBC0107EB2CFB9B242EFBBB" bold="true" box="[1040,1062,1031,1048]" pageId="21" pageNumber="894">co</emphasis>
|
||
, suprarectal commissure;
|
||
<emphasis id="B949806BFFBC0107EF86FB8220DBFB8C" bold="true" box="[186,211,1054,1071]" pageId="21" pageNumber="894">div</emphasis>
|
||
, diverticula;
|
||
<emphasis id="B949806BFFBC0107EE04FB822141FB8C" bold="true" box="[312,329,1054,1071]" pageId="21" pageNumber="894">fo</emphasis>
|
||
, pedal folds (foot);
|
||
<emphasis id="B949806BFFBC0107EEDDFB8221F4FB8C" bold="true" box="[481,508,1054,1071]" pageId="21" pageNumber="894">ma</emphasis>
|
||
, mantle;
|
||
<emphasis id="B949806BFFBC0107ED79FB822269FB8C" bold="true" box="[581,609,1054,1071]" pageId="21" pageNumber="894">mg</emphasis>
|
||
, midgut;
|
||
<emphasis id="B949806BFFBC0107ED97FB8222C9FB8C" bold="true" box="[683,705,1054,1071]" pageId="21" pageNumber="894">pc</emphasis>
|
||
, pericardium;
|
||
<emphasis id="B949806BFFBC0107EC08FB82234EFB8C" bold="true" box="[820,838,1054,1071]" pageId="21" pageNumber="894">ro</emphasis>
|
||
, respiratory organs;
|
||
<emphasis id="B949806BFFBC0107ECD5FB8223F2FB8C" bold="true" box="[1001,1018,1054,1071]" pageId="21" pageNumber="894">rs</emphasis>
|
||
, receptaculum seminis;
|
||
<emphasis id="B949806BFFBC0107EF4CFBA8208DFBE6" bold="true" box="[112,133,1076,1093]" pageId="21" pageNumber="894">sd</emphasis>
|
||
, spawning duct;
|
||
<emphasis id="B949806BFFBC0107EE29FBA8213EFBE6" bold="true" box="[277,310,1076,1093]" pageId="21" pageNumber="894">sdg</emphasis>
|
||
, spawning duct glands;
|
||
<emphasis id="B949806BFFBC0107ED3EFBA8221FFBE6" bold="true" box="[514,535,1076,1093]" pageId="21" pageNumber="894">so</emphasis>
|
||
, dorsoterminal sense organ;
|
||
<emphasis id="B949806BFFBC0107EC36FBA82323FBE6" bold="true" box="[778,811,1076,1093]" pageId="21" pageNumber="894">spg</emphasis>
|
||
, suprapallial glands. Scale bar: 500 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B825C79FFBC0107EF4DFB122392FB6A" blockId="21.[111,647,1166,1700]" lastBlockId="21.[675,1208,1166,1225]" pageId="21" pageNumber="894">
|
||
of both portions is strongly narrowed. The two pericardioducts emerge lateroterminally. There are no vesiculae seminales. A small, posteriorly directed receptaculum seminis is laterally differentiated at the opening of each pericardioduct into the anteriomost, lateral spawning duct (
|
||
<figureCitation id="130640FCFFBC0107EE66FAB121A0FAEB" box="[346,424,1325,1352]" captionStart="FIG" captionStartId="21.[113,124,1032,1049]" captionTargetBox="[203,1163,219,985]" captionTargetId="figure-13@21.[200,1173,199,1000]" captionTargetPageId="21" captionText="FIG. 23.— Alexandromenia gulaglandulata n.sp.,organization of the posterior body viewed from left.Abbreviations:co, suprarectal commissure;div, diverticula;fo, pedal folds (foot);ma, mantle;mg, midgut;pc, pericardium;ro, respiratory organs;rs, receptaculum seminis; sd, spawning duct; sdg, spawning duct glands; so, dorsoterminal sense organ; spg, suprapallial glands. Scale bar: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.4525823" httpUri="https://zenodo.org/record/4525823/files/figure.png" pageId="21" pageNumber="894">Fig. 23</figureCitation>
|
||
). The ciliated, simple spawning ducts are paired along most of their length and fuse near the end region of the pericardium. The subepithelially arranged, elongate and slender spawning glands are aggregated; they surround the ducts as a dense mass and open intercellularly. The unpaired spawning duct is provided with circular musculature and anteriorly (region of rectum) still shows openings of glands. Th e posterior portion of the unpaired spawning duct (below anterior mantle cavity) has a sphincter and radial muscles within a bulge at the bottom of the mantle cavity (
|
||
<figureCitation id="130640FCFFBC0107ED16F9152271F907" box="[554,633,1673,1700]" captionStart="FIG" captionStartId="22.[113,124,1073,1090]" captionTargetBox="[117,1206,215,1031]" captionTargetId="figure-14@22.[117,1206,214,1031]" captionTargetPageId="22" captionText="FIG. 24.— Alexandromenia gulaglandulata n. sp.,cross section through anterior mantle cavity with irregular,slender diverticula.Abbreviations: div, diverticle; gl, suprapallial glands; mc, mantle cavity; sd, spawning duct; sdg, spawning duct glands. Scale bar: 150 μm." figureDoi="http://doi.org/10.5281/zenodo.4525825" httpUri="https://zenodo.org/record/4525825/files/figure.png" pageId="21" pageNumber="894">Fig. 24</figureCitation>
|
||
); the ventro-terminal outlet opens into the beginning mantle cavity opening.
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFBC0107ED99FB732336FAA5" blockId="21.[674,1212,1263,1700]" box="[677,830,1263,1289]" pageId="21" pageNumber="894">COMPARISONS</paragraph>
|
||
<paragraph id="8B825C79FFBC0104ED99FA91220AF986" blockId="21.[674,1212,1263,1700]" lastBlockId="22.[111,648,1166,1700]" lastPageId="22" lastPageNumber="895" pageId="21" pageNumber="894">
|
||
All organizational characters of the single specimen classify it as a member of the
|
||
<taxonomicName id="4C3D27FAFFBC0107ECC7FAB124BCFAEB" authorityName="Salvini-Plawen" authorityYear="1972" box="[1019,1204,1325,1352]" class="Solenogastres" family="Amphimeniidae" kingdom="Animalia" order="Cavibelonia" pageId="21" pageNumber="894" phylum="Mollusca" rank="family">Amphimeniidae</taxonomicName>
|
||
. The key features here include the configuration of the lateral foregut glandular organs, the monoserial radula and the organization of the spawning ducts (see Salvini-Plawen 1972, 1978). Except in
|
||
<taxonomicName id="4C3D27FAFFBC0107ED99FA5723CBFA45" authority="Schwabl, 1961" authorityName="Schwabl" authorityYear="1961" box="[677,963,1483,1510]" class="Solenogastres" family="Amphimeniidae" genus="Plathymenia" kingdom="Animalia" order="Cavibelonia" pageId="21" pageNumber="894" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949806BFFBC0107ED99FA57232BFA46" box="[677,803,1483,1509]" italics="true" pageId="21" pageNumber="894">Plathymenia</emphasis>
|
||
Schwabl, 1961
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3D27FAFFBC0107ECC4FA572494FA46" authorityName="Heath" authorityYear="1911" box="[1016,1180,1483,1509]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="21" pageNumber="894" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949806BFFBC0107ECC4FA572494FA46" box="[1016,1180,1483,1509]" italics="true" pageId="21" pageNumber="894">Alexandromenia</emphasis>
|
||
</taxonomicName>
|
||
(?)
|
||
<taxonomicName id="4C3D27FAFFBC0107ED98FA77237DF9A5" authority="Odhner, 1921" authorityName="Odhner" authorityYear="1921" box="[676,885,1515,1542]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="21" pageNumber="894" phylum="Mollusca" rank="species" species="crassa">
|
||
<emphasis id="B949806BFFBC0107ED98FA7722D3F9A7" box="[676,731,1515,1540]" italics="true" pageId="21" pageNumber="894">crassa</emphasis>
|
||
<bibRefCitation id="EFAC2188FFBC0107EDDCFA77237DF9A5" author="ODHNER N." box="[736,885,1515,1542]" pageId="21" pageNumber="894" pagination="1 - 86" refId="ref10832" refString="ODHNER N. 1921. - Norwegian Solenogastres. Bergens Museums Aarbok 1918 - 1919 3: 1 - 86." type="journal article" year="1921">Odhner, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
, all members of the family show a pre-radular outlet of the ventro-lateral foregut glandular organs.In most species this organ possesses a terminal cone within a pouch or sheath (
|
||
<figureCitation id="130640FCFFBC0107EB5FF9D624A7F9C6" box="[1123,1199,1610,1637]" captionStart="FIG" captionStartId="19.[113,124,881,898]" captionTargetBox="[147,1129,248,821]" captionTargetId="figure-13@19.[113,1209,221,856]" captionTargetPageId="19" captionText="FIG. 20. — Two types of the outlet of the lateroventral foregut glandular organs with terminal cone (and opening papilla) within Amphimeniidae (schematised): A, simple cone as in Alexandromenia gulaglandulata n. sp.; B, cone with internal ductule such as in A. antarctica Salvini-Plawen, 1978. Abbreviations: con, cone; cp, cone pouch (sheath); dg, duct of glandular organ; du, ductule of cone; pa, opening papilla; ph, pharyngeal cavity; ri, median ridge of duct." figureDoi="http://doi.org/10.5281/zenodo.4525817" httpUri="https://zenodo.org/record/4525817/files/figure.png" pageId="21" pageNumber="894">Fig. 20</figureCitation>
|
||
) that opens by formation of a papilla intruding into the pharynx (
|
||
<figureCitation id="130640FCFFBC0107EC16F915238FF907" box="[810,903,1673,1700]" captionStart="FIG" captionStartId="19.[113,124,881,898]" captionTargetBox="[147,1129,248,821]" captionTargetId="figure-13@19.[113,1209,221,856]" captionTargetPageId="19" captionText="FIG. 20. — Two types of the outlet of the lateroventral foregut glandular organs with terminal cone (and opening papilla) within Amphimeniidae (schematised): A, simple cone as in Alexandromenia gulaglandulata n. sp.; B, cone with internal ductule such as in A. antarctica Salvini-Plawen, 1978. Abbreviations: con, cone; cp, cone pouch (sheath); dg, duct of glandular organ; du, ductule of cone; pa, opening papilla; ph, pharyngeal cavity; ri, median ridge of duct." figureDoi="http://doi.org/10.5281/zenodo.4525817" httpUri="https://zenodo.org/record/4525817/files/figure.png" pageId="21" pageNumber="894">Fig.20B</figureCitation>
|
||
). Th e details of the outleading duct, however, differ:this may open directly, without cone formation, into the pharynx (with or without a papilla), such as in
|
||
<taxonomicName id="4C3D27FAFFBF0104EE72FB512171FAAB" authority="Handl & Salvini-Plawen, 2002" authorityName="Handl & Salvini-Plawen" authorityYear="2002" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="species" species="pilosa">
|
||
<emphasis id="B949806BFFBF0104EE72FB512230FB44" box="[334,568,1229,1255]" italics="true" pageId="22" pageNumber="895">Alexandromenia pilosa</emphasis>
|
||
Handl & Salvini-Plawen, 2002
|
||
</taxonomicName>
|
||
; the duct may open into the pouch (sheath) of the cone (
|
||
<figureCitation id="130640FCFFBF0104EEFAFA912220FA8B" box="[454,552,1293,1320]" captionStart="FIG" captionStartId="19.[113,124,881,898]" captionTargetBox="[147,1129,248,821]" captionTargetId="figure-13@19.[113,1209,221,856]" captionTargetPageId="19" captionText="FIG. 20. — Two types of the outlet of the lateroventral foregut glandular organs with terminal cone (and opening papilla) within Amphimeniidae (schematised): A, simple cone as in Alexandromenia gulaglandulata n. sp.; B, cone with internal ductule such as in A. antarctica Salvini-Plawen, 1978. Abbreviations: con, cone; cp, cone pouch (sheath); dg, duct of glandular organ; du, ductule of cone; pa, opening papilla; ph, pharyngeal cavity; ri, median ridge of duct." figureDoi="http://doi.org/10.5281/zenodo.4525817" httpUri="https://zenodo.org/record/4525817/files/figure.png" pageId="22" pageNumber="895">Fig. 20A</figureCitation>
|
||
), such as in the present species (no papilla); the outleading duct may be directly continuous with a (somewhat eccentric) internal ductule of the cone (
|
||
<figureCitation id="130640FCFFBF0104ED2BFAF02270FA24" box="[535,632,1388,1415]" captionStart="FIG" captionStartId="19.[113,124,881,898]" captionTargetBox="[147,1129,248,821]" captionTargetId="figure-13@19.[113,1209,221,856]" captionTargetPageId="19" captionText="FIG. 20. — Two types of the outlet of the lateroventral foregut glandular organs with terminal cone (and opening papilla) within Amphimeniidae (schematised): A, simple cone as in Alexandromenia gulaglandulata n. sp.; B, cone with internal ductule such as in A. antarctica Salvini-Plawen, 1978. Abbreviations: con, cone; cp, cone pouch (sheath); dg, duct of glandular organ; du, ductule of cone; pa, opening papilla; ph, pharyngeal cavity; ri, median ridge of duct." figureDoi="http://doi.org/10.5281/zenodo.4525817" httpUri="https://zenodo.org/record/4525817/files/figure.png" pageId="22" pageNumber="895">Fig. 20B</figureCitation>
|
||
), such as in
|
||
<taxonomicName id="4C3D27FAFFBF0104EFEBFA10223EFA04" authority="Salvini-Plawen, 1978" authorityName="Salvini-Plawen" authorityYear="1978" box="[215,566,1420,1447]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="species" species="antarctica">
|
||
<emphasis id="B949806BFFBF0104EFEBFA10215DFA06" box="[215,341,1420,1446]" italics="true" pageId="22" pageNumber="895">A. antarctica</emphasis>
|
||
Salvini-Plawen, 1978
|
||
</taxonomicName>
|
||
. Several species exhibit both a connection with the pouch and a ductule within the cone (Salvini-Plawen 1978); in some species, several slender ductules pass through the cone (
|
||
<bibRefCitation id="EFAC2188FFBF0104EE52F99621FEF986" author="HEATH H." box="[366,502,1546,1573]" pageId="22" pageNumber="895" pagination="1 - 179" refId="ref10542" refString="HEATH H. 1911. - Reports on the scientific results of the expedition to the tropical Pacific, XIV. The Solenogastres. Memoirs of the Museum of Comparative Zoology at Harvard College 45 (1): 1 - 179." type="journal article" year="1911">Heath 1911</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF420CF1FFBF0104EF4DFBAD24A6FBFA" ID-DOI="http://doi.org/10.5281/zenodo.4525825" ID-Zenodo-Dep="4525825" httpUri="https://zenodo.org/record/4525825/files/figure.png" pageId="22" pageNumber="895" startId="22.[113,124,1073,1090]" targetBox="[117,1206,215,1031]" targetPageId="22">
|
||
<paragraph id="8B825C79FFBF0104EF4DFBAD24A6FBFA" blockId="22.[113,1210,1072,1113]" pageId="22" pageNumber="895">
|
||
FIG. 24.—
|
||
<taxonomicName id="4C3D27FAFFBF0104EFFAFBAC21CCFBE2" authority="Salvini-Plawen, 2008" authorityName="Salvini-Plawen" authorityYear="2008" box="[198,452,1072,1089]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="species" species="gulaglandulata" status="sp. nov.">
|
||
<emphasis id="B949806BFFBF0104EFFAFBAC21CCFBE2" box="[198,452,1072,1089]" italics="true" pageId="22" pageNumber="895">Alexandromenia gulaglandulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A3D10FFBF0104EEFBFBAD21FFFBE1" box="[455,503,1073,1090]" pageId="22" pageNumber="895" rank="species">n. sp.</taxonomicNameLabel>
|
||
,cross section through anterior mantle cavity with irregular,slender diverticula.Abbreviations:
|
||
<emphasis id="B949806BFFBF0104EF98FBDB20B6FBFB" bold="true" box="[164,190,1095,1112]" pageId="22" pageNumber="895">div</emphasis>
|
||
, diverticle;
|
||
<emphasis id="B949806BFFBF0104EE22FBDB2126FBFB" bold="true" box="[286,302,1095,1112]" pageId="22" pageNumber="895">gl</emphasis>
|
||
, suprapallial glands;
|
||
<emphasis id="B949806BFFBF0104EEE2FBDB21F2FBFB" bold="true" box="[478,506,1095,1112]" pageId="22" pageNumber="895">mc</emphasis>
|
||
, mantle cavity;
|
||
<emphasis id="B949806BFFBF0104ED41FBDB229AFBFB" bold="true" box="[637,658,1095,1112]" pageId="22" pageNumber="895">sd</emphasis>
|
||
, spawning duct;
|
||
<emphasis id="B949806BFFBF0104EC1EFBDB234BFBFB" bold="true" box="[802,835,1095,1112]" pageId="22" pageNumber="895">sdg</emphasis>
|
||
, spawning duct glands. Scale bar: 150 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B825C79FFBF0104EFB4F9B623B8F9C6" blockId="22.[111,648,1166,1700]" lastBlockId="22.[675,1211,1166,1700]" pageId="22" pageNumber="895">
|
||
Within the
|
||
<taxonomicName id="4C3D27FAFFBF0104EE36F9B621B3F9E6" authorityName="Salvini-Plawen" authorityYear="1972" box="[266,443,1578,1605]" class="Solenogastres" family="Amphimeniidae" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="family">Amphimeniidae</taxonomicName>
|
||
, the present specimen clearly belongs to the genus
|
||
<taxonomicName id="4C3D27FAFFBF0104EEE1F9D6228EF9C7" authorityName="Heath" authorityYear="1911" box="[477,646,1610,1636]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949806BFFBF0104EEE1F9D6228EF9C7" box="[477,646,1610,1636]" italics="true" pageId="22" pageNumber="895">Alexandromenia</emphasis>
|
||
</taxonomicName>
|
||
(see diagnosis above and the comparative tables in Salvini-Plawen 1978 as well as García-Álvarez
|
||
<emphasis id="B949806BFFBF0104ED6AF916228FF900" box="[598,647,1673,1699]" italics="true" pageId="22" pageNumber="895">et al.</emphasis>
|
||
2000). An identity with the incompletely known
|
||
<taxonomicName id="4C3D27FAFFBF0104ED99FB332314FB6B" authorityName="Leloup" authorityYear="1949" box="[677,796,1199,1224]" class="Solenogastres" family="Amphimeniidae" genus="Meromenia" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949806BFFBF0104ED99FB332314FB6B" box="[677,796,1199,1224]" italics="true" pageId="22" pageNumber="895">Meromenia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C3D27FAFFBF0104EC15FB33245CFB6A" authority="Leloup, 1949" authorityName="Leloup" authorityYear="1949" box="[809,1108,1198,1225]" class="Solenogastres" family="Amphimeniidae" genus="Meromenia" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="species" species="hirondellei">
|
||
<emphasis id="B949806BFFBF0104EC15FB3323B4FB6B" box="[809,956,1198,1224]" italics="true" pageId="22" pageNumber="895">M. hirondellei</emphasis>
|
||
<bibRefCitation id="EFAC2188FFBF0104ECFEFB32245CFB6A" author="LELOUP E." box="[962,1108,1198,1225]" pageId="22" pageNumber="895" pagination="1 - 6" refId="ref10765" refString="LELOUP E. 1949. - Meromenia hirondellei g. nov., sp. nov., Solenogastre du Golfe de Gascogne. Bulletin de l'Institut royal des Sciences naturelles de Belgique 25 (1): 1 - 6." type="journal article" year="1949">Leloup, 1949</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the Bay of Biscay) can be excluded based on the lack of respiratory organs and midgut pouches. Eight
|
||
<taxonomicName id="4C3D27FAFFBF0104ED99FA912346FA84" authorityName="Heath" authorityYear="1911" box="[677,846,1293,1319]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949806BFFBF0104ED99FA912346FA84" box="[677,846,1293,1319]" italics="true" pageId="22" pageNumber="895">Alexandromenia</emphasis>
|
||
</taxonomicName>
|
||
species are currently known. The present specimen differs from these by the packages of additional foregut glands in the region between the outlets of the lateral glandular organs and the radula apparatus. Th is also includes the three known Atlantic representatives:
|
||
<taxonomicName id="4C3D27FAFFBF0104ECC3FA3022D7FA45" authority="Odhner, 1921" authorityName="Odhner" authorityYear="1921" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="species" species="crassa">
|
||
<emphasis id="B949806BFFBF0104ECC3FA30245FFA66" box="[1023,1111,1452,1478]" italics="true" pageId="22" pageNumber="895">A. crassa</emphasis>
|
||
<bibRefCitation id="EFAC2188FFBF0104EB62FA3722D7FA45" author="ODHNER N." pageId="22" pageNumber="895" pagination="1 - 86" refId="ref10832" refString="ODHNER N. 1921. - Norwegian Solenogastres. Bergens Museums Aarbok 1918 - 1919 3: 1 - 86." type="journal article" year="1921">Odhner, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the Hjelteŋord (Bergen area,
|
||
<collectingCountry id="F32A1CE9FFBF0104EB66FA5724BAFA45" box="[1114,1202,1483,1510]" name="Norway" pageId="22" pageNumber="895">Norway</collectingCountry>
|
||
) at
|
||
<quantity id="4CC5F19CFFBF0104EDFDFA77234AF9A5" box="[705,834,1515,1542]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="22" pageNumber="895" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100-200 m</quantity>
|
||
,
|
||
<taxonomicName id="4C3D27FAFFBF0104EC73FA77247DF9A5" authority="Leloup, 1946" authorityName="Leloup" authorityYear="1946" box="[847,1141,1515,1542]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="species" species="grimaldii">
|
||
<emphasis id="B949806BFFBF0104EC73FA7723DBF9A6" box="[847,979,1515,1541]" italics="true" pageId="22" pageNumber="895">A. grimaldii</emphasis>
|
||
<bibRefCitation id="EFAC2188FFBF0104ECE0FA77247DF9A5" author="LELOUP E." box="[988,1141,1515,1542]" pageId="22" pageNumber="895" pagination="1 - 12" refId="ref10732" refString="LELOUP E. 1946. - Alexandromenia grimaldii sp. nov., Solenogastre bathypelagique. Bulletin du Musee royal d'Histoire naturelle de Belgique 22 (16): 1 - 12." type="journal article" year="1946">Leloup, 1946</bibRefCitation>
|
||
</taxonomicName>
|
||
, from off the Azores at
|
||
<quantity id="4CC5F19CFFBF0104EC62F99623BFF986" box="[862,951,1546,1573]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="22" pageNumber="895" unit="m" value="1250.0">1250 m</quantity>
|
||
, and
|
||
<taxonomicName id="4C3D27FAFFBF0104ECCFF9972398F9E6" authority="Handl & Salvini-Plawen, 2002" authorityName="Handl & Salvini-Plawen" authorityYear="2002" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="species" species="pilosa">
|
||
<emphasis id="B949806BFFBF0104ECCFF9972445F987" box="[1011,1101,1546,1573]" italics="true" pageId="22" pageNumber="895">A. pilosa</emphasis>
|
||
Handl & Salvini-Plawen, 2002
|
||
</taxonomicName>
|
||
, from the Trondheimsŋord (
|
||
<collectingCountry id="F32A1CE9FFBF0104ED90F9D6230EF9C6" box="[684,774,1610,1637]" name="Norway" pageId="22" pageNumber="895">Norway</collectingCountry>
|
||
) at
|
||
<quantity id="4CC5F19CFFBF0104EC13F9D623A4F9C6" box="[815,940,1610,1637]" metricMagnitude="2" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="22" pageNumber="895" unit="m" value="210.0" valueMax="240.0" valueMin="180.0">
|
||
180-
|
||
<specimenCount id="9D3B97F0FFBF0104EC5CF9D623A4F9C6" box="[864,940,1610,1637]" pageId="22" pageNumber="895" type="generic">240 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFBF0105ED87F9F521B9FE72" blockId="22.[675,1211,1166,1700]" lastBlockId="23.[111,649,216,1162]" lastPageId="23" lastPageNumber="896" pageId="22" pageNumber="895">
|
||
The poorly described
|
||
<taxonomicName id="4C3D27FAFFBF0104ECA4F9F623E4F920" authorityName="Odhner" authorityYear="1921" box="[920,1004,1642,1668]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="22" pageNumber="895" phylum="Mollusca" rank="species" species="crassa">
|
||
<emphasis id="B949806BFFBF0104ECA4F9F623E4F920" box="[920,1004,1642,1668]" italics="true" pageId="22" pageNumber="895">A. crassa</emphasis>
|
||
</taxonomicName>
|
||
, apart from its “pair of small salivary glands opening into the radula sac” (
|
||
<bibRefCitation id="EFAC2188FFBE0105EF45FF44214BFF50" author="ODHNER N." box="[121,323,216,243]" pageId="23" pageNumber="896" pagination="1 - 86" refId="ref10832" refString="ODHNER N. 1921. - Norwegian Solenogastres. Bergens Museums Aarbok 1918 - 1919 3: 1 - 86." type="journal article" year="1921">Odhner 1921: 22</bibRefCitation>
|
||
) – which casts doubt on the generic classification – also differs from the animal at hand by the dorso-median keel of spicules, by the fairly stout body (10 ×
|
||
<quantity id="4CC5F19CFFBE0105EEB6FEAB21FEFEF1" box="[394,502,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="23" pageNumber="896" unit="mm" value="3.15" valueMax="3.3" valueMin="3.0">3-3.3 mm</quantity>
|
||
) with 9 pedal folds, by the
|
||
<quantity id="4CC5F19CFFBE0105EFC7FECB2158FED1" box="[251,336,343,370]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="23" pageNumber="896" unit="mm" value="160.0">160 µm</quantity>
|
||
broad radula plates of different outline, by the long midgut caecum extending beyond the cerebral ganglion, and by the dorsally extended receptacula seminis.
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFBE0105EFB4FE492255FCAE" blockId="23.[111,649,216,1162]" pageId="23" pageNumber="896">
|
||
<taxonomicName id="4C3D27FAFFBE0105EFB4FE49219DFE4C" authorityName="Leloup" authorityYear="1946" box="[136,405,469,495]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="23" pageNumber="896" phylum="Mollusca" rank="species" species="grimaldii">
|
||
<emphasis id="B949806BFFBE0105EFB4FE49219DFE4C" box="[136,405,469,495]" italics="true" pageId="23" pageNumber="896">Alexandromenia grimaldii</emphasis>
|
||
</taxonomicName>
|
||
measures 20 ×
|
||
<quantity id="4CC5F19CFFBE0105ED0FFE49228EFE53" box="[563,646,469,496]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="23" pageNumber="896" unit="mm" value="2.3">2.3 mm</quantity>
|
||
with an about
|
||
<quantity id="4CC5F19CFFBE0105EE35FE692154FDB3" box="[265,348,501,528]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="23" pageNumber="896" unit="mm" value="140.0">140 µm</quantity>
|
||
thick cuticle; it differs from the present specimen in the number (7-3) of pedal folds, whereby the median fold connects with the mantle cavity, in the voluminous midgut caecum, in the hindgut being continuous with the ventral midgut only, as well as in the receptacula seminis being enlargements of the distal pericardioducts (
|
||
<bibRefCitation id="EFAC2188FFBE0105EF46FD4F210EFD4D" author="LELOUP E." box="[122,262,723,750]" pageId="23" pageNumber="896" pagination="1 - 12" refId="ref10732" refString="LELOUP E. 1946. - Alexandromenia grimaldii sp. nov., Solenogastre bathypelagique. Bulletin du Musee royal d'Histoire naturelle de Belgique 22 (16): 1 - 12." type="journal article" year="1946">Leloup 1946</bibRefCitation>
|
||
). It is particularly characterised by the U-shaped radula teeth (Salvini-Plawen 1972).
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFBE0105EFB4FC8E227EFB88" blockId="23.[111,649,216,1162]" pageId="23" pageNumber="896">
|
||
<taxonomicName id="4C3D27FAFFBE0105EFB4FC8E216DFC8F" authorityName="Handl & Salvini-Plawen" authorityYear="2002" box="[136,357,786,812]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="23" pageNumber="896" phylum="Mollusca" rank="species" species="pilosa">
|
||
<emphasis id="B949806BFFBE0105EFB4FC8E216DFC8F" box="[136,357,786,812]" italics="true" pageId="23" pageNumber="896">Alexandromenia pilosa</emphasis>
|
||
</taxonomicName>
|
||
, measuring 9 ×
|
||
<quantity id="4CC5F19CFFBE0105ED39FC8E228BFC8E" box="[517,643,786,813]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="23" pageNumber="896" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2-1.4 mm</quantity>
|
||
, has only 5-3 pedal folds, club-shaped epidermis papillae, sclerites of one
|
||
<typeStatus id="5486E2DBFFBE0105EE40FCCD21A2FCCF" box="[380,426,849,876]" pageId="23" pageNumber="896">type</typeStatus>
|
||
only, a subepithelial matrix outside the transverse musculature, an anteriorly bilobed midgut caecum, a short suprarectal commissure (about
|
||
<quantity id="4CC5F19CFFBE0105EE05FC2C2184FC68" box="[313,396,944,971]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="896" unit="mm" value="100.0">100 µm</quantity>
|
||
), and anteriorly directed receptacula seminis; it lacks suprapallial glands and no turgescent cells of the radula support or bolster are mentioned (Handl & Salvini-Plawen 2002).
|
||
</paragraph>
|
||
<paragraph id="8B825C79FFBE0105EFB4FBB32163FB29" blockId="23.[111,649,216,1162]" pageId="23" pageNumber="896">
|
||
The present specimen is thus organizationally well separated and represents a proper, new species:
|
||
<taxonomicName id="4C3D27FAFFBE0105EF4CFBF32121FB2B" authority="Salvini-Plawen, 2008" authorityName="Salvini-Plawen" authorityYear="2008" box="[112,297,1134,1161]" class="Solenogastres" family="Amphimeniidae" genus="Alexandromenia" kingdom="Animalia" order="Cavibelonia" pageId="23" pageNumber="896" phylum="Mollusca" rank="species" species="gulaglandulata" status="sp. nov.">
|
||
<emphasis id="B949806BFFBE0105EF4CFBF32121FB2B" box="[112,297,1134,1161]" italics="true" pageId="23" pageNumber="896">A. gulaglandulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A3D10FFBE0105EE0CFBF32163FB29" box="[304,363,1135,1162]" pageId="23" pageNumber="896" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |