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<mods:title id="9CACC270C89703EA74987586D62CABBB">A review of the subfamily Methochinae from Taiwan (Hymenoptera: Tiphiidae) with description of a new species and redescription of the known species</mods:title>
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<mods:namePart id="C92ED66A3604F89443C8520302F79C7D">Narita, Keisuke</mods:namePart>
<mods:affiliation id="F6C550B1ADF008A9E32EBA9E42E1950D">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Motooka 744, Nishiku, Fukuoka, 819 - 0385, Japan.</mods:affiliation>
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<mods:namePart id="A069B5360F8E22A3FFABA365B0EFBAA9">Mita, Toshiharu</mods:namePart>
<mods:affiliation id="7F8354BB67D778959A0AD039CE7222A3">Entomological Laboratory, Faculty of Agriculture, Kyushu University, Motooka 744, Nishi-ku, Fukuoka, 819 - 0385, Japan. t 3 mita @ agr. kyushu-u. ac. jp</mods:affiliation>
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<treatment id="03F687D2FFB9E01B4C93FA5BFC52F810" ID-DOI="http://doi.org/10.5281/zenodo.4709455" ID-GBIF-Taxon="181875176" ID-Zenodo-Dep="4709455" LSID="urn:lsid:plazi:treatment:03F687D2FFB9E01B4C93FA5BFC52F810" httpUri="http://treatment.plazi.org/id/03F687D2FFB9E01B4C93FA5BFC52F810" lastPageNumber="327" pageId="24" pageNumber="327">
<subSubSection id="C345654FFFB9E01B4C93FA5BFD53FA5A" box="[151,665,1482,1509]" pageId="24" pageNumber="327" type="nomenclature">
<paragraph id="8BE036C4FFB9E01B4C93FA5BFD53FA5A" blockId="24.[151,665,1482,1545]" box="[151,665,1482,1509]" pageId="24" pageNumber="327">
<heading id="D0A881A8FFB9E01B4C93FA5BFD53FA5A" bold="true" box="[151,665,1482,1509]" fontSize="11" level="1" pageId="24" pageNumber="327" reason="1">
<emphasis id="B92BEAD6FFB9E01B4C93FA5BFD53FA5A" bold="true" box="[151,665,1482,1509]" pageId="24" pageNumber="327">
<taxonomicName id="4C5F4D47FFB9E01B4C93FA5BFDCFFA5B" authority="(Strand, 1913)" authorityName="Narita &amp; Mita" authorityYear="2021" baseAuthorityName="Strand" baseAuthorityYear="1913" box="[151,517,1482,1509]" class="Insecta" family="Tiphiidae" genus="Karlissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="327" phylum="Arthropoda" rank="species" species="tricha" status="comb. nov.">
<emphasis id="B92BEAD6FFB9E01B4C93FA5BFE80FA5A" bold="true" box="[151,330,1482,1509]" italics="true" pageId="24" pageNumber="327">Karlissa tricha</emphasis>
(
<bibRefCitation id="EFCE4B35FFB9E01B4D5FFA5BFE37FA5B" author="Strand, E." box="[347,509,1482,1508]" pageId="24" pageNumber="327" pagination="209 - 215" refId="ref15800" refString="Strand, E. (1913) H. Sauter's Formosa-Ausbeute. Entomologische Mitteilungen, 2, 209 - 215. https: // doi. org / 10.5962 / bhl. part. 14993" type="journal article" year="1913">Strand, 1913</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A21857ADFFB9E01B4E17FA5AFD53FA5A" box="[531,665,1483,1509]" pageId="24" pageNumber="327" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C345654FFFB9E01B4C93FA7EFEE4F9B6" box="[151,302,1519,1545]" pageId="24" pageNumber="327" type="description">
<paragraph id="8BE036C4FFB9E01B4C93FA7EFEE4F9B6" blockId="24.[151,665,1482,1545]" box="[151,302,1519,1545]" pageId="24" pageNumber="327">
<figureCitation id="13642A41FFB9E01B4C93FA7EFEE4F9B6" box="[151,302,1519,1545]" captionStart="FIGURES 117118" captionStartId="25.[151,264,794,818]" captionTargetBox="[154,1435,181,768]" captionTargetId="figure-17@25.[151,1436,181,770]" captionTargetPageId="25" captionText="FIGURES 117118. Karlissa tricha (Strand, 1913) comb. nov., holotype, male. 117 Dorsal habitus, 118 lateral habitus. The photos of K. tricha are courtesy of National Museum of Natural Science (Chan et al. 2018)." figureDoi="http://doi.org/10.5281/zenodo.4709487" httpUri="https://zenodo.org/record/4709487/files/figure.png" pageId="24" pageNumber="327">Figs 117118</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C345654FFFB9E01B4C93F9A2FDD6F9D5" pageId="24" pageNumber="327" type="reference_group">
<paragraph id="8BE036C4FFB9E01B4C93F9A2FBD1F9F4" blockId="24.[151,1051,1587,1642]" box="[151,1051,1587,1612]" pageId="24" pageNumber="327">
<treatmentCitationGroup id="AB4F11EAFFB9E01B4C93F9A2FBD1F9F4" box="[151,1051,1587,1612]" pageId="24" pageNumber="327">
<treatmentCitation id="0AFE10D5FFB9E01B4C93F9A2FDE7F9F4" author="Strand, E." box="[151,557,1587,1611]" page="214" pageId="24" pageNumber="327" year="1913">
<taxonomicName id="4C5F4D47FFB9E01B4C93F9A2FDE7F9F4" authority="Strand, 1913: 214" authorityName="Strand" authorityPageNumber="214" authorityYear="1913" box="[151,557,1587,1611]" class="Insecta" family="Tiphiidae" genus="Methoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="327" phylum="Arthropoda" rank="species" species="tricha">
<emphasis id="B92BEAD6FFB9E01B4C93F9A2FF3AF9F4" box="[151,240,1587,1611]" italics="true" pageId="24" pageNumber="327">Methoca</emphasis>
(sic!)
<materialsCitation id="3B373C99FFB9E01B4D36F9A2FDE7F9F4" ID-GBIF-Occurrence="3107345343" box="[306,557,1587,1611]" location="Liukwei" pageId="24" pageNumber="327" specimenCount="1" stateProvince="Kaohsiung">
<emphasis id="B92BEAD6FFB9E01B4D36F9A2FEA5F9F4" box="[306,367,1587,1611]" italics="true" pageId="24" pageNumber="327">tricha</emphasis>
<bibRefCitation id="EFCE4B35FFB9E01B4D71F9A2FDE7F9F4" author="Strand, E." box="[373,557,1587,1611]" pageId="24" pageNumber="327" pagination="209 - 215" refId="ref15800" refString="Strand, E. (1913) H. Sauter's Formosa-Ausbeute. Entomologische Mitteilungen, 2, 209 - 215. https: // doi. org / 10.5962 / bhl. part. 14993" type="journal article" year="1913">Strand, 1913: 214</bibRefCitation>
</materialsCitation>
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. Orig. desig.
<typeStatus id="54E48866FFB9E01B4EB7F9A5FD2FF9F3" box="[691,741,1588,1612]" pageId="24" pageNumber="327">Type</typeStatus>
locality:
<location id="8E80601FFFB9E01B4F41F9A2FC56F9F4" LSID="urn:lsid:plazi:treatment:03F687D2FFB9E01B4C93FA5BFC52F810:8E80601FFFB9E01B4F41F9A2FC56F9F4" box="[837,924,1587,1611]" name="Liukwei" pageId="24" pageNumber="327" stateProvince="Kaohsiung">Liukwei</location>
,
<collectingRegion id="499BF826FFB9E01B4FA2F9A2FBDDF9F4" box="[934,1047,1587,1611]" country="Taiwan" name="Kaohsiung" pageId="24" pageNumber="327">Kaohsiung</collectingRegion>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE036C4FFB9E01B4C93F9C3FDD6F9D5" blockId="24.[151,1051,1587,1642]" box="[151,540,1618,1642]" pageId="24" pageNumber="327">
<treatmentCitationGroup id="AB4F11EAFFB9E01B4C93F9C3FDD6F9D5" box="[151,540,1618,1642]" pageId="24" pageNumber="327">
<treatmentCitation id="0AFE10D5FFB9E01B4C93F9C3FE2FF9D5" author="Lin, K. S." box="[151,485,1618,1642]" page="183" pageId="24" pageNumber="327" year="1966">
<taxonomicName id="4C5F4D47FFB9E01B4C93F9C3FE2FF9D5" authority=": Lin, 1966: 183" authorityName=": Lin" authorityPageNumber="183" authorityYear="1966" box="[151,485,1618,1642]" class="Insecta" family="Tiphiidae" genus="Methocha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="327" phylum="Arthropoda" rank="species" species="tricha">
<emphasis id="B92BEAD6FFB9E01B4C93F9C3FE8AF9D5" box="[151,320,1618,1642]" italics="true" pageId="24" pageNumber="327">Methocha tricha</emphasis>
:
<bibRefCitation id="EFCE4B35FFB9E01B4D49F9C3FE2FF9D5" author="Lin, K. S." box="[333,485,1618,1642]" pageId="24" pageNumber="327" pagination="181 - 202" refId="ref15592" refString="Lin, K. S. (1966) The Methochidae of Taiwan (Hymenoptera: Scolioidea). Quarterly Journal of Taiwan Museum, 19, 181 - 202." type="journal article" year="1966">Lin, 1966: 183</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 199.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C345654FFFB9E01B4C93F909FD65F945" pageId="24" pageNumber="327" type="materials_examined">
<paragraph id="8BE036C4FFB9E01B4C93F909FD65F945" blockId="24.[151,1436,1687,1967]" pageId="24" pageNumber="327">
<emphasis id="B92BEAD6FFB9E01B4C93F909FE8CF90D" bold="true" box="[151,326,1688,1714]" pageId="24" pageNumber="327">Type material.</emphasis>
<materialsCitation id="3B373C99FFB9E01B4D49F909FD65F945" ID-GBIF-Occurrence="3107345330" collectingDate="1968-07-07" collectionCode="SDEI" pageId="24" pageNumber="327" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54E48866FFB9E01B4D49F909FE72F90D" box="[333,440,1688,1714]" pageId="24" pageNumber="327" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D59FD4DFFB9E01B4DC0F908FE11F90C" box="[452,475,1689,1715]" pageId="24" pageNumber="327" type="male"></specimenCount>
, Sokutsu,
<date id="FFE11004FFB9E01B4E54F909FD0EF90D" box="[592,708,1688,1714]" pageId="24" pageNumber="327" value="1968-07-07">
<collectingDate id="EFA5E9ECFFB9E01B4E54F909FD0EF90D" box="[592,708,1688,1714]" pageId="24" pageNumber="327" value="1968-07-07">7.vii.1968</collectingDate>
</date>
, collector unknown. [
<collectionCode id="ED4EAE01FFB9E01B4FB3F906FC37F90D" box="[951,1021,1687,1714]" pageId="24" pageNumber="327">SDEI</collectionCode>
]. The male
<typeStatus id="54E48866FFB9E01B4884F909FB28F90D" box="[1152,1250,1688,1714]" pageId="24" pageNumber="327" type="holotype">holotype</typeStatus>
deposited in the Senckenberg Deutsches Entomologisches Institut (SDEI, German Entomological Institute). The
<typeStatus id="54E48866FFB9E01B48EBF92DFA9BF969" box="[1263,1361,1724,1750]" pageId="24" pageNumber="327" type="holotype">holotype</typeStatus>
is not examined directly, but the photo was examined.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C345654FFFB9E01B4CC3F895FDE7F8A1" box="[199,557,1796,1822]" pageId="24" pageNumber="327" type="distribution">
<paragraph id="8BE036C4FFB9E01B4CC3F895FDE7F8A1" blockId="24.[151,1436,1687,1967]" box="[199,557,1796,1822]" pageId="24" pageNumber="327">
<emphasis id="B92BEAD6FFB9E01B4CC3F895FEABF8A1" bold="true" box="[199,353,1796,1822]" pageId="24" pageNumber="327">Distribution.</emphasis>
Southern
<collectingCountry id="F3487654FFB9E01B4DD7F895FDE3F8A1" box="[467,553,1796,1822]" name="Taiwan" pageId="24" pageNumber="327">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C345654FFFB9E01B4CC3F8B9FC52F810" pageId="24" pageNumber="327" type="discussion">
<paragraph id="8BE036C4FFB9E01B4CC3F8B9FC52F810" blockId="24.[151,1436,1687,1967]" pageId="24" pageNumber="327">
<emphasis id="B92BEAD6FFB9E01B4CC3F8B9FEF1F8FD" bold="true" box="[199,315,1832,1858]" pageId="24" pageNumber="327">Remarks.</emphasis>
This species shows an elongate pronotum and strongly flattened flagellum. These are diagnostic characters that define the genus
<taxonomicName id="4C5F4D47FFB9E01B4DFAF8DDFD97F8D9" authorityName="Krombein. A" authorityYear="1979" box="[510,605,1868,1894]" class="Insecta" family="Tiphiidae" genus="Karlissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAD6FFB9E01B4DFAF8DDFD97F8D9" box="[510,605,1868,1894]" italics="true" pageId="24" pageNumber="327">Karlissa</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCE4B35FFB9E01B4E6FF8DDFCEDF8D9" author="Krombein, K. V." box="[619,807,1868,1894]" pageId="24" pageNumber="327" pagination="424 - 434" refId="ref15490" refString="Krombein, K. V. (1979) Studies in the Tiphiidae, XII. A new genus of Methochinae with notes on the subgenera of Methocha Latreille (Hymenoptera Aculeata). Proceedings of the Entomological Society of Washington, 81 (3), 424 - 434." type="journal article" year="1979">Krombein, 1979</bibRefCitation>
;
<bibRefCitation id="EFCE4B35FFB9E01B4F37F8DDFCA5F8D9" author="Krombein, K. V." box="[819,879,1868,1894]" pageId="24" pageNumber="327" pagination="1 - 121" refId="ref15541" refString="Krombein, K. V. (1982) Biosystematic studies of Ceylonese wasps, IX: A monograph of the Tiphiidae (Hymenoptera: Vespoidea). Smithsonian Contributions to Zoology, 374, 1 - 121. https: // doi. org / 10.5479 / si. 00810282.374" type="journal article" year="1982">1982</bibRefCitation>
), therefore a new combination proposed.
<taxonomicName id="4C5F4D47FFB9E01B4939F8DDFA56F8D9" authorityName="Krombein. A" authorityYear="1979" box="[1341,1436,1868,1894]" class="Insecta" family="Tiphiidae" genus="Karlissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAD6FFB9E01B4939F8DDFA56F8D9" box="[1341,1436,1868,1894]" italics="true" pageId="24" pageNumber="327">Karlissa</emphasis>
</taxonomicName>
was a monotypic genus established by
<bibRefCitation id="EFCE4B35FFB9E01B4E57F8E1FCDDF835" author="Krombein, K. V." box="[595,791,1904,1930]" pageId="24" pageNumber="327" pagination="424 - 434" refId="ref15490" refString="Krombein, K. V. (1979) Studies in the Tiphiidae, XII. A new genus of Methochinae with notes on the subgenera of Methocha Latreille (Hymenoptera Aculeata). Proceedings of the Entomological Society of Washington, 81 (3), 424 - 434." type="journal article" year="1979">Krombein (1979)</bibRefCitation>
for the Ceylonese species,
<taxonomicName id="4C5F4D47FFB9E01B4855F8E1FED1F811" authority="(Cameron, 1897)" baseAuthorityName="Cameron" baseAuthorityYear="1897" class="Insecta" family="Tiphiidae" genus="Methoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="327" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B92BEAD6FFB9E01B4855F8E1FB79F835" box="[1105,1203,1904,1930]" italics="true" pageId="24" pageNumber="327">Methoca</emphasis>
(sic!)
<emphasis id="B92BEAD6FFB9E01B48FBF8E0FA87F835" box="[1279,1357,1905,1930]" italics="true" pageId="24" pageNumber="327">rugosa</emphasis>
(
<bibRefCitation id="EFCE4B35FFB9E01B4958F8E1FED9F811" author="Cameron, P." pageId="24" pageNumber="327" refId="ref15216" refString="Cameron, P. (1897) Hymenoptera Orientalis, or Contributions to a Knowledge of the Hymenoptera of the Oriental Zoological Region, Part VII. Memoirs and Proceedings of the Manchester Literary and Philosophical Society, 42 (11), 84 pp., pl. 4." type="book" year="1897">Cameron, 1897</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C5F4D47FFB9E01B4D22F805FE05F811" authorityName="Narita &amp; Mita" authorityYear="2021" baseAuthorityName="Strand" baseAuthorityYear="1913" box="[294,463,1940,1966]" class="Insecta" family="Tiphiidae" genus="Karlissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="327" phylum="Arthropoda" rank="species" species="tricha">
<emphasis id="B92BEAD6FFB9E01B4D22F805FE05F811" box="[294,463,1940,1966]" italics="true" pageId="24" pageNumber="327">Karlissa tricha</emphasis>
</taxonomicName>
is the second species placed in
<taxonomicName id="4C5F4D47FFB9E01B4F36F805FC5BF811" authorityName="Krombein. A" authorityYear="1979" box="[818,913,1940,1966]" class="Insecta" family="Tiphiidae" genus="Karlissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAD6FFB9E01B4F36F805FC5BF811" box="[818,913,1940,1966]" italics="true" pageId="24" pageNumber="327">Karlissa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>