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<document id="16C3ED136CE98BB69A82152C0F58C630" ID-CLB-Dataset="272041" ID-DOI="10.1093/zoolinnean/zlad013" ID-GBIF-Dataset="effa6181-8326-4c7d-8c91-809f7fabb862" ID-ISSN="0024-4082" ID-Zenodo-Dep="8432389" ID-ZooBank="C3E2DFF9-A0A6-41EA-A149-0F73A2BEE5E6" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1696939870042" checkinUser="plazi" docAuthor="Silva, Felipe De Vasconcelos, Pola, Marta &amp; Cervera, Juan Lucas" docDate="2023" docId="03A6878CFF856502FD43FD0CED08FAE5" docLanguage="en" docName="zlad013.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 199 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad013" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Myrella Odhner 1963" docType="treatment" docVersion="5" lastPageNumber="466" masterDocId="FF9FFFF4FF906517FC52FF84EE67FF8B" masterDocTitle="A stomach plate to divide them all: a phylogenetic reassessment of the family Tritoniidae (Nudibranchia: Cladobranchia)" masterLastPageNumber="476" masterPageNumber="445" pageNumber="466" updateTime="1699694776904" updateUser="plazi">
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<mods:title id="8DEEB2617D14361156659896E80EE5C4">A stomach plate to divide them all: a phylogenetic reassessment of the family Tritoniidae (Nudibranchia: Cladobranchia)</mods:title>
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<mods:namePart id="DF98B2DE428B78F63A7811C6B924A2D6">Silva, Felipe De Vasconcelos</mods:namePart>
<mods:affiliation id="9BB41D0B4560D39F4726461BC88EECAF">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales; Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain</mods:affiliation>
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<mods:namePart id="8E0873D9D7DA2C3435FA786508CFAFE6">Pola, Marta</mods:namePart>
<mods:affiliation id="EADD7133FE3B7D262C4E816CDDE2B29C">Departamento de Biología, Facultad de Ciencias, Universidad Autónoma de Madrid; Campus de Excelencia Internacional UAM + CSIC, C / Darwin, 2, 28049 Madrid, Spain</mods:affiliation>
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<mods:namePart id="AE3590B6BCC7F500F2709716E9098642">Cervera, Juan Lucas</mods:namePart>
<mods:affiliation id="746534F0985B482B72B56AE91F3BC17A">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales; Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain &amp; Instituto Universitario de Investigación Marina (INMAR), Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Universidad de Cádiz, Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain</mods:affiliation>
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<mods:title id="D8ABCB0AD658E47B2F19670E3EF36ADF">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="22033B6C40188958B18D0DD55B792945">2023</mods:date>
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<mods:title id="DE2AF425DB20D66E4902572CEABE5653">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="A65B7943665BB307CF1C4C72D37C2D68">2023-07-11</mods:number>
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<mods:number id="E6C55F20BBBBCEA051C51D5DE7B39A80">199</mods:number>
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<treatment id="03A6878CFF856502FD43FD0CED08FAE5" ID-DOI="http://doi.org/10.5281/zenodo.8432417" ID-GBIF-Taxon="213768873" ID-Zenodo-Dep="8432417" LSID="urn:lsid:plazi:treatment:03A6878CFF856502FD43FD0CED08FAE5" httpUri="http://treatment.plazi.org/id/03A6878CFF856502FD43FD0CED08FAE5" lastPageNumber="466" pageId="21" pageNumber="466">
<subSubSection id="C3156511FF856502FD43FD0CEC3CFD29" box="[273,603,648,675]" pageId="21" pageNumber="466" type="nomenclature">
<paragraph id="8BB0369AFF856502FD43FD0CEC3CFD29" blockId="21.[273,603,648,675]" box="[273,603,648,675]" pageId="21" pageNumber="466">
<heading id="D0F881F6FF856502FD43FD0CEC3CFD29" bold="true" box="[273,603,648,675]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="466" reason="2">
<emphasis id="B97BEA88FF856502FD43FD0CEC3CFD29" bold="true" box="[273,603,648,675]" pageId="21" pageNumber="466">
<taxonomicName id="4C0F4D19FF856502FD43FD0CEF93FD28" ID-CoL="BGKM5" authority="Odhner 1963" authorityName="Odhner" authorityYear="1963" box="[273,500,648,675]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502FD43FD0CEF38FD28" bold="true" box="[273,351,648,675]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
<bibRefCitation id="EF9E4B6BFF856502FD36FD0DEF93FD28" author="Odhner NH" box="[356,500,649,675]" pageId="21" pageNumber="466" pagination="48 - 62" refId="ref26607" refString="Odhner NH. On the taxonomy of the family Tritoniidae (Mollusca: Opisthobranchia). The Veliger 1963; 6: 48 - 62." type="journal article" year="1963">Odhner1963</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13342A1FFF856502FE56FD0DEC37FD28" box="[516,592,649,675]" captionStart="Figure 3" captionStartId="12.[129,194,1533,1557]" captionTargetBox="[321,1281,144,1505]" captionTargetId="figure-116@12.[321,1281,144,1505]" captionTargetPageId="12" captionText="Figure 3. Representative species of genera of Tritoniidae (with exception of Paratritonia). A, Tritonia hombergii (MNCN15.05/93379) (Portaferry, Ireland); B, Candiella plebeia (MNCN15.05/93393) (Portaferry, Ireland); C, Tritoniopsis elegans (MB28-0045061) (Zavora, Mozambique); D, Tritonidoxa capensis (MNCN15.05/93399) (Scoưburg, South Africa); E, Tritoniella belli (Antarctica); F, Marianina rosea (French Polynesia); G, Myrella challengeriana comb.nov.(Bahía Camarones, Argentina); H, Tochuina gigantea (Agnew Pass, USA); I, Tritonicula khaleesi (MZUSP113165) (Fortaleza, Brazil); J, Marionia cucullata (Golfo Nuevo, Argentina); (K) Marioniopsis levis ‘1’ (MNHNC/ MB28-004895) (Nacala, Mozambique). Photo A and B, courtesy of B. Picton; photo C and K courtesy of Y.Tibiriça; photo D courtesy of V. Fraser; photo E courtesy of M. Caballer; photo F courtesy of G. Rouse; photo G and J, courtesy of N. Baưini; photo H courtesy of S. Thiebaud; photo I by F. Silva." figureDoi="http://doi.org/10.5281/zenodo.8432399" httpUri="https://zenodo.org/record/8432399/files/figure.png" pageId="21" pageNumber="466">Fig. 3G</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3156511FF856502FC23FD39EF1EFD7F" pageId="21" pageNumber="466" type="reference_group">
<paragraph id="8BB0369AFF856502FC23FD39EF1EFD7F" blockId="21.[113,762,701,756]" pageId="21" pageNumber="466">
<taxonomicName id="4C0F4D19FF856502FC23FD39EC25FD5E" ID-CoL="17c37d57-d160-41c5-8a27-d4b075a6a8bc" authority="Mabille and Rochebrune 1889" authorityName="Mabille and Rochebrune" authorityYear="1889" box="[113,578,701,725]" class="Gastropoda" family="Tritoniidae" genus="Microlophus" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502FC23FD39EE8DFD5E" box="[113,234,701,725]" italics="true" pageId="21" pageNumber="466">Microlophus</emphasis>
Mabille and Rochebrune 1889
</taxonomicName>
, non-
<taxonomicName id="4C0F4D19FF856502FED3FD39EF1EFD7F" ID-CoL="17c37d57-d160-41c5-8a27-d4b075a6a8bc" authority="Dumeril and Bibron 1837" authorityName="Dumeril and Bibron" authorityYear="1837" class="Gastropoda" family="Tritoniidae" genus="Microlophus" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502FED3FD39EC9DFD5E" box="[641,762,701,725]" italics="true" pageId="21" pageNumber="466">Microlophus</emphasis>
Duméril and Bibron 1837
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3156511FF856502FC23FC97EE88FCE2" pageId="21" pageNumber="466" type="type_taxon">
<paragraph id="8BB0369AFF856502FC23FC97EE88FCE2" blockId="21.[113,763,786,873]" pageId="21" pageNumber="466">
<emphasis id="B97BEA88FF856502FC23FC97EE89FCA1" box="[113,238,787,810]" italics="true" pageId="21" pageNumber="466">
<typeStatus id="54B48838FF856502FC23FC97EEC6FCA1" box="[113,161,787,810]" pageId="21" pageNumber="466">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C0F4D19FF856502FCA8FC96EC19FCA0" ID-CoL="1f3e416a-c316-40b6-9643-fff0110e3043" authority="(Bergh 1884)" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Bergh" baseAuthorityYear="1884" box="[250,638,786,811]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="challengeriana" status="comb. nov.">
<emphasis id="B97BEA88FF856502FCA8FC96EF8DFCA1" bold="true" box="[250,490,786,810]" italics="true" pageId="21" pageNumber="466">Myrella challengeriana</emphasis>
(Bergh 1884)
</taxonomicName>
<emphasis id="B97BEA88FF856502FED1FC97EC9CFCA0" bold="true" box="[643,763,787,811]" pageId="21" pageNumber="466">
<taxonomicNameLabel id="A24857F3FF856502FED1FC97EC9CFCA0" box="[643,763,787,811]" pageId="21" pageNumber="466" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(=
<taxonomicName id="4C0F4D19FF856502FCC1FCB6EC1FFCC1" ID-CoL="b36e5309-f6b9-4e75-a24c-af04082ae7b2" authority="Mabille and Rochebrune 1889" authorityName="Mabille and Rochebrune" authorityYear="1889" box="[147,632,818,842]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="poirieri">
<emphasis id="B97BEA88FF856502FCC1FCB6EF4AFCC1" box="[147,301,818,842]" italics="true" pageId="21" pageNumber="466">Myrella poirieri</emphasis>
Mabille and Rochebrune 1889
</taxonomicName>
), by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C3156511FF856502FC23FC0DEE98FC75" pageId="21" pageNumber="466" type="diagnosis">
<paragraph id="8BB0369AFF856502FC23FC0DEE98FC75" blockId="21.[113,763,904,1022]" pageId="21" pageNumber="466">
<emphasis id="B97BEA88FF856502FC23FC0DEEB0FC2B" box="[113,215,905,928]" italics="true" pageId="21" pageNumber="466">Diagnosis:</emphasis>
Tricuspid rachidian tooth with minor folds. Masticatory borders denticulate with conical denticles. Penis thin, oħen conical at the distal end and flagelliform towards proximal end.
</paragraph>
</subSubSection>
<subSubSection id="C3156511FF856502FC23FB99EC43F9EA" pageId="21" pageNumber="466" type="description">
<paragraph id="8BB0369AFF856502FC23FB99EF3EFA27" blockId="21.[113,763,1053,1452]" pageId="21" pageNumber="466">
<emphasis id="B97BEA88FF856502FC23FB99EE97FBBE" box="[113,240,1053,1077]" italics="true" pageId="21" pageNumber="466">Morphology:</emphasis>
Body broad, length up to
<quantity id="4CF79B7FFF856502FE59FB99EC32FBBF" box="[523,597,1053,1077]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="21" pageNumber="466" unit="mm" value="80.0">80 mm</quantity>
(
<figureCitation id="13342A1FFF856502FE35FB99ECD3FBBE" box="[615,692,1053,1077]" captionStart="Figure 3" captionStartId="12.[129,194,1533,1557]" captionTargetBox="[321,1281,144,1505]" captionTargetId="figure-116@12.[321,1281,144,1505]" captionTargetPageId="12" captionText="Figure 3. Representative species of genera of Tritoniidae (with exception of Paratritonia). A, Tritonia hombergii (MNCN15.05/93379) (Portaferry, Ireland); B, Candiella plebeia (MNCN15.05/93393) (Portaferry, Ireland); C, Tritoniopsis elegans (MB28-0045061) (Zavora, Mozambique); D, Tritonidoxa capensis (MNCN15.05/93399) (Scoưburg, South Africa); E, Tritoniella belli (Antarctica); F, Marianina rosea (French Polynesia); G, Myrella challengeriana comb.nov.(Bahía Camarones, Argentina); H, Tochuina gigantea (Agnew Pass, USA); I, Tritonicula khaleesi (MZUSP113165) (Fortaleza, Brazil); J, Marionia cucullata (Golfo Nuevo, Argentina); (K) Marioniopsis levis ‘1’ (MNHNC/ MB28-004895) (Nacala, Mozambique). Photo A and B, courtesy of B. Picton; photo C and K courtesy of Y.Tibiriça; photo D courtesy of V. Fraser; photo E courtesy of M. Caballer; photo F courtesy of G. Rouse; photo G and J, courtesy of N. Baưini; photo H courtesy of S. Thiebaud; photo I by F. Silva." figureDoi="http://doi.org/10.5281/zenodo.8432399" httpUri="https://zenodo.org/record/8432399/files/figure.png" pageId="21" pageNumber="466">Fig. 3G</figureCitation>
). Oral veil broad, bilobed or entire. Five to 20 simple or branched velar processes. Notal border distinct. Six to 40 pairs of fully arborescent gills. Rachidian tooth tricuspid, denticulate (minor folds) with rectangular base. First lateral tooth differentiated. Masticatory borders of jaws denticulate with medium-sized (up to 50 µm) denticles with conical bases and sharp cusps. Ratio jaw/body length: 0.21–0.4. Cuticular folds absent. Gonopore at end of one-third of body length, furthest from rhinophores. Anus at middle of body length; nephroproct above anus. Ampulla cylindrical and coiled. Penis thin and flagelliform or conical at the distal end and flagelliform towards proximal end. Capsule and mucous gland distinct.
</paragraph>
<paragraph id="8BB0369AFF856502FC23FA48EC43F9EA" blockId="21.[113,762,1483,1633]" pageId="21" pageNumber="466">
<emphasis id="B97BEA88FF856502FC23FA48EC7FFA68" box="[113,536,1483,1507]" italics="true" pageId="21" pageNumber="466">
Species composition:
<taxonomicName id="4C0F4D19FF856502FD1CFA4FEC7FFA68" box="[334,536,1483,1507]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="antarctica">
<emphasis id="B97BEA88FF856502FD1CFA4FEC7FFA68" bold="true" box="[334,536,1483,1507]" italics="true" pageId="21" pageNumber="466">Myrella antarctica</emphasis>
</taxonomicName>
</emphasis>
(Pfeffer in Martens and Pfeffer 1886)
<emphasis id="B97BEA88FF856502FD7DFA6FEFC0F988" bold="true" box="[303,423,1515,1539]" pageId="21" pageNumber="466">comb. nov.</emphasis>
,
<taxonomicName id="4C0F4D19FF856502FDE1FA6EEC9DF988" authority="(Bergh 1898)" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Bergh" baseAuthorityYear="1898" box="[435,762,1514,1539]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="australis" status="comb. nov.">
<emphasis id="B97BEA88FF856502FDE1FA6EEC01F989" bold="true" box="[435,614,1514,1538]" italics="true" pageId="21" pageNumber="466">Myrella australis</emphasis>
(Bergh 1898)
</taxonomicName>
<emphasis id="B97BEA88FF856502FC23F98EEE8FF9A9" bold="true" box="[113,232,1546,1570]" pageId="21" pageNumber="466">
<taxonomicNameLabel id="A24857F3FF856502FC23F98EEE8FF9A9" box="[113,232,1546,1570]" pageId="21" pageNumber="466" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C0F4D19FF856502FCA6F98DEC1FF9A9" authority="(Bergh 1884)" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Bergh" baseAuthorityYear="1884" box="[244,632,1545,1570]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="challengeriana" status="comb. nov.">
<emphasis id="B97BEA88FF856502FCA6F98DEF83F9AA" bold="true" box="[244,484,1545,1569]" italics="true" pageId="21" pageNumber="466">Myrella challengeriana</emphasis>
(Bergh 1884)
</taxonomicName>
<emphasis id="B97BEA88FF856502FE2CF98EEC92F9A9" bold="true" box="[638,757,1546,1570]" pageId="21" pageNumber="466">
<taxonomicNameLabel id="A24857F3FF856502FE2CF98EEC92F9A9" box="[638,757,1546,1570]" pageId="21" pageNumber="466" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C0F4D19FF856502FC23F9ADEC2AF9CA" authority="(Ballesteros and Avila 2006)" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Ballesteros and Avila" baseAuthorityYear="2006" box="[113,589,1577,1601]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="dantarti" status="comb. nov.">
<emphasis id="B97BEA88FF856502FC23F9ADEF46F9CA" bold="true" box="[113,289,1577,1601]" italics="true" pageId="21" pageNumber="466">Myrella dantarti</emphasis>
(Ballesteros and Avila 2006)
</taxonomicName>
<emphasis id="B97BEA88FF856502FE07F9ADECAAF9CA" bold="true" box="[597,717,1577,1601]" pageId="21" pageNumber="466">
<taxonomicNameLabel id="A24857F3FF856502FE07F9ADECAAF9CA" box="[597,717,1577,1601]" pageId="21" pageNumber="466" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C0F4D19FF856502FC23F9CCEFCEF9EB" authority="(Odhner 1926)" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Odhner" baseAuthorityYear="1926" box="[113,425,1608,1632]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="vorax" status="comb. nov.">
<emphasis id="B97BEA88FF856502FC23F9CCEF64F9EB" bold="true" box="[113,259,1608,1632]" italics="true" pageId="21" pageNumber="466">Myrella vorax</emphasis>
(Odhner 1926)
</taxonomicName>
<emphasis id="B97BEA88FF856502FDFCF9CDEC43F9EA" bold="true" box="[430,548,1609,1633]" pageId="21" pageNumber="466">
<taxonomicNameLabel id="A24857F3FF856502FDFCF9CDEC43F9EA" box="[430,548,1609,1633]" pageId="21" pageNumber="466" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3156511FF856502FC23F9FBED08FAE5" pageId="21" pageNumber="466" type="discussion">
<paragraph id="8BB0369AFF856502FC23F9FBEA4EFEEF" blockId="21.[113,762,1663,1969]" lastBlockId="21.[810,1459,144,1390]" pageId="21" pageNumber="466">
<emphasis id="B97BEA88FF856502FC23F9FBEEB5F91C" box="[113,210,1663,1687]" italics="true" pageId="21" pageNumber="466">Remarks:</emphasis>
Based on our phylogenetic analyses, the clade formed by the Antarctic ‘
<taxonomicName id="4C0F4D19FF856502FD2CF91BEFB6F93D" authorityName="Cuvier" authorityYear="1798" box="[382,465,1695,1718]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502FD2CF91BEFB6F93D" box="[382,465,1695,1718]" italics="true" pageId="21" pageNumber="466">Tritonia</emphasis>
</taxonomicName>
’
<taxonomicName id="4C0F4D19FF856502FD8CF91AEC17F93D" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Bergh" baseAuthorityYear="1884" box="[478,624,1694,1718]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="challengeriana">
<emphasis id="B97BEA88FF856502FD8CF91AEC17F93D" box="[478,624,1694,1718]" italics="true" pageId="21" pageNumber="466">challengeriana</emphasis>
</taxonomicName>
and the sub-Antarctic ‘
<taxonomicName id="4C0F4D19FF856502FCB0F93AEF52F95E" authorityName="Cuvier" authorityYear="1798" box="[226,309,1726,1749]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502FCB0F93AEF52F95E" box="[226,309,1726,1749]" italics="true" pageId="21" pageNumber="466">Tritonia</emphasis>
</taxonomicName>
’
<taxonomicName id="4C0F4D19FF856502FD16F93AEFF0F95D" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Ballesteros and Avila" baseAuthorityYear="2006" box="[324,407,1726,1750]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="dantarti">
<emphasis id="B97BEA88FF856502FD16F93AEFF0F95D" box="[324,407,1726,1750]" italics="true" pageId="21" pageNumber="466">dantarti</emphasis>
</taxonomicName>
corresponds to a distinct genus, molecularly and morphologically different from
<taxonomicName id="4C0F4D19FF856502FEF5F95AEC9DF97E" authorityName="Cuvier" authorityYear="1798" box="[679,762,1758,1781]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus" sensu="stricto">
<emphasis id="B97BEA88FF856502FEF5F95AEC9DF97E" box="[679,762,1758,1781]" italics="true" pageId="21" pageNumber="466">Tritonia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24857F3FF856502FC23F979EEE8F89F" box="[113,143,1789,1812]" pageId="21" pageNumber="466" sensu="stricto">
<emphasis id="B97BEA88FF856502FC23F979EE1EF89F" box="[113,121,1789,1812]" italics="true" pageId="21" pageNumber="466">s</emphasis>
.
<emphasis id="B97BEA88FF856502FCD2F979EEEFF89F" box="[128,136,1789,1812]" italics="true" pageId="21" pageNumber="466">s</emphasis>
.
</taxonomicNameLabel>
This group, first depicted in
<bibRefCitation id="EF9E4B6BFF856502FD86F978EED9F8BF" author="Korshunova T &amp; Martynov A" pageId="21" pageNumber="466" pagination="0242103" refId="ref25588" refString="Korshunova T, Martynov A. Consolidated data on the phylogeny and evolution of the family Tritoniidae (Gastropoda: Nudibranchia) contribute to genera reassessment and clarify the taxonomic status of the neuroscience models Tritonia and Tochuina. PLoS One 2020; 15: e 0242103. hups: // doi. org / 10.1371 / journal. pone. 0242103" type="journal article" year="2020">Korshunova and Martynov (2020)</bibRefCitation>
and further focused in
<bibRefCitation id="EF9E4B6BFF856502FD98F898ECC7F8BF" author="Moles J &amp; Berning MI &amp; Hooker Y" box="[458,672,1820,1844]" pageId="21" pageNumber="466" pagination="107209" refId="ref26342" refString="Moles J, Berning MI, Hooker Y, et al. Due South: the evolutionary history of sub-Antarctic and Antarctic Tritoniidae nudibranchs. Molecular Phylogenetics and Evolution 2021; 162: 107209." type="journal article" year="2021">
Moles
<emphasis id="B97BEA88FF856502FE47F898EC23F8BF" box="[533,580,1820,1844]" italics="true" pageId="21" pageNumber="466">et al</emphasis>
. (2021)
</bibRefCitation>
, corresponds to the resurrected
<taxonomicName id="4C0F4D19FF856502FD2DF8BFEC01F8D8" authority="Odhner 1963" authorityName="Odhner" authorityYear="1963" box="[383,614,1851,1875]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502FD2DF8BFEFAAF8D8" box="[383,461,1851,1875]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
<bibRefCitation id="EF9E4B6BFF856502FD86F8BFEC01F8D8" author="Odhner NH" box="[468,614,1851,1875]" pageId="21" pageNumber="466" pagination="48 - 62" refId="ref26607" refString="Odhner NH. On the taxonomy of the family Tritoniidae (Mollusca: Opisthobranchia). The Veliger 1963; 6: 48 - 62." type="journal article" year="1963">Odhner 1963</bibRefCitation>
</taxonomicName>
, based on the
<typeStatus id="54B48838FF856502FC23F8DFEEC7F8F8" box="[113,160,1883,1907]" pageId="21" pageNumber="466">type</typeStatus>
species
<taxonomicName id="4C0F4D19FF856502FCAFF8DFEFDAF8F9" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Bergh" baseAuthorityYear="1884" box="[253,445,1882,1906]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="challengeriana">
<emphasis id="B97BEA88FF856502FCAFF8DFEFDAF8F9" box="[253,445,1882,1906]" italics="true" pageId="21" pageNumber="466">My. challengeriana</emphasis>
</taxonomicName>
. The species included in this group are characterized by a unique combination of features within
<taxonomicName id="4C0F4D19FF856502FCEDF81DEF5DF83A" authorityName="Lamarck" authorityYear="1809" box="[191,314,1945,1969]" pageId="21" pageNumber="462" rank="subFamily" subFamily="Tritoniinae">Tritoniinae</taxonomicName>
: moderate size (up to
<quantity id="4CF79B7FFF856502FE61F81DECE1F83A" box="[563,646,1945,1969]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="21" pageNumber="466" unit="mm" value="80.0">80 mm</quantity>
); tricuspid rachidian tooth with minor folds; denticulate masticatory borders with conical denticles; penis thin and flagelliform or conical at the distal end and digitiform towards the proximal end; and a distinct capsule and mucous gland (
<bibRefCitation id="EF9E4B6BFF856502F976FF6AEBCAFE8D" author="Wagele H" box="[1316,1453,238,262]" pageId="21" pageNumber="466" pagination="21 - 46" refId="ref27940" refString="Wagele H. The morphology and taxonomy of the Antarctic species of Tritonia Cuvier, 1797 (Nudibranchia: Dendronotoidea). Zoological Journal of the Linnean Society 1995; 113: 21 - 46." type="journal article" year="1995">Wägele 1995</bibRefCitation>
, Ballesteros and Ávila 2006,
<bibRefCitation id="EF9E4B6BFF856502F81AFE89EA88FEAE" author="Rossi ME &amp; Avila C &amp; Moles J" box="[1096,1263,269,293]" pageId="21" pageNumber="466" pagination="559 - 573" refId="ref27227" refString="Rossi ME, Avila C, Moles J. Orange is the new white: taxonomic revision of Tritonia species (Gastropoda: Nudibranchia) from the Weddell Sea and Bouvet Island. Polar Biology 2021; 44: 559 - 573." type="journal article" year="2021">
Rossi
<emphasis id="B97BEA88FF856502F8D4FE8AEAC8FEAE" box="[1158,1199,269,293]" italics="true" pageId="21" pageNumber="466">et al</emphasis>
. 2021
</bibRefCitation>
). In addition, they are distinguished by their unique Antarctic, sub-Antarctic and Magellanic distribution.
</paragraph>
<paragraph id="8BB0369AFF856502FF17FEEFEBF8FBBF" blockId="21.[810,1459,144,1390]" pageId="21" pageNumber="466">
Externally,
<taxonomicName id="4C0F4D19FF856502FFE8FEEFEA62FE08" authorityName="Odhner" authorityYear="1963" box="[954,1029,363,387]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502FFE8FEEFEA62FE08" box="[954,1029,363,387]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F4D19FF856502F86EFEE8EAEBFE08" authorityName="Cuvier" authorityYear="1798" box="[1084,1164,364,387]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502F86EFEE8EAEBFE08" box="[1084,1164,364,387]" italics="true" pageId="21" pageNumber="466">Tritonia</emphasis>
</taxonomicName>
are similar: both genera include large species with several arborescent gills and a bilobed oral veil with several processes. Some of the internal morpho-anatomical characters found in
<taxonomicName id="4C0F4D19FF856502F83EFE4DEAD0FE6A" authorityName="Odhner" authorityYear="1963" box="[1132,1207,457,481]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502F83EFE4DEAD0FE6A" box="[1132,1207,457,481]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
</taxonomicName>
are also found individually in a few species of the heterogeneous
<taxonomicName id="4C0F4D19FF856502F8A4FE6DEB21FD8B" authorityName="Cuvier" authorityYear="1798" box="[1270,1350,489,512]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502F8A4FE6DEB21FD8B" box="[1270,1350,489,512]" italics="true" pageId="21" pageNumber="466">Tritonia</emphasis>
</taxonomicName>
, but their combination is unique to
<taxonomicName id="4C0F4D19FF856502F862FD8CEA1CFDAB" authorityName="Odhner" authorityYear="1963" box="[1072,1147,520,544]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502F862FD8CEA1CFDAB" box="[1072,1147,520,544]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
</taxonomicName>
within the
<taxonomicName id="4C0F4D19FF856502F8BDFD8CEB04FDAB" baseAuthorityName="sensu Odhner" baseAuthorityYear="1963" box="[1263,1379,520,544]" class="Gastropoda" family="Tritoniidae" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="family">Tritoniidae</taxonomicName>
. For example,
<taxonomicName id="4C0F4D19FF856502FF21FDACEA53FDB4" authorityName="Pallas" authorityYear="1788" box="[883,1076,552,575]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="tetraquetra">
<emphasis id="B97BEA88FF856502FF21FDACEA53FDB4" box="[883,1076,552,575]" italics="true" pageId="21" pageNumber="466">Tritonia tetraquetra</emphasis>
</taxonomicName>
also has a denticulate rachidian tooth like
<taxonomicName id="4C0F4D19FF856502FF06FDC2EDF8FDD5" authorityName="Odhner" authorityYear="1963" box="[852,927,582,606]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502FF06FDC2EDF8FDD5" box="[852,927,582,606]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
</taxonomicName>
, but it is larger than
<taxonomicName id="4C0F4D19FF856502F83FFDC2EB6BFDD5" box="[1133,1292,582,606]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B97BEA88FF856502F83FFDC2EADFFDD5" box="[1133,1208,582,606]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
species.
</taxonomicName>
(up to
<quantity id="4CF79B7FFF856502F906FDC2EBCCFDD5" box="[1364,1451,582,606]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="466" unit="mm" value="300.0">300 mm</quantity>
) and its denticulate masticatory border is characterized by rod-shaped denticles instead of conical denticles (
<bibRefCitation id="EF9E4B6BFF856502F967FD01ED9EFD37" author="Korshunova T &amp; Martynov A" pageId="21" pageNumber="466" pagination="0242103" refId="ref25588" refString="Korshunova T, Martynov A. Consolidated data on the phylogeny and evolution of the family Tritoniidae (Gastropoda: Nudibranchia) contribute to genera reassessment and clarify the taxonomic status of the neuroscience models Tritonia and Tochuina. PLoS One 2020; 15: e 0242103. hups: // doi. org / 10.1371 / journal. pone. 0242103" type="journal article" year="2020">Korshunova and Martynov 2020</bibRefCitation>
). The penis is also an important feature in distinguishing the two genera:
<taxonomicName id="4C0F4D19FF856502F832FD40EAD7FD50" authorityName="Cuvier" authorityYear="1798" box="[1120,1200,708,731]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502F832FD40EAD7FD50" box="[1120,1200,708,731]" italics="true" pageId="21" pageNumber="466">Tritonia</emphasis>
</taxonomicName>
is characterized by a large penis that is elongate or conical and has distal characters in a few species, while
<taxonomicName id="4C0F4D19FF856502FFEBFC86EA63FC91" authorityName="Odhner" authorityYear="1963" box="[953,1028,770,794]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502FFEBFC86EA63FC91" box="[953,1028,770,794]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
</taxonomicName>
is characterized by a thin penis that is completely flagelliform (
<bibRefCitation id="EF9E4B6BFF856502FFA4FCA5EAAAFCB2" author="Rossi ME &amp; Avila C &amp; Moles J" box="[1014,1229,801,826]" pageId="21" pageNumber="466" pagination="559 - 573" refId="ref27227" refString="Rossi ME, Avila C, Moles J. Orange is the new white: taxonomic revision of Tritonia species (Gastropoda: Nudibranchia) from the Weddell Sea and Bouvet Island. Polar Biology 2021; 44: 559 - 573." type="journal article" year="2021">
Rossi
<emphasis id="B97BEA88FF856502F860FCA6EA3EFCB2" box="[1074,1113,801,825]" italics="true" pageId="21" pageNumber="466">et al</emphasis>
. 2021: 564
</bibRefCitation>
) or conical at the distal end and digitiform towards the proximal end (
<bibRefCitation id="EF9E4B6BFF856502F8A8FCC5EBC1FCD2" author="Wagele H" box="[1274,1446,833,857]" pageId="21" pageNumber="466" pagination="21 - 46" refId="ref27940" refString="Wagele H. The morphology and taxonomy of the Antarctic species of Tritonia Cuvier, 1797 (Nudibranchia: Dendronotoidea). Zoological Journal of the Linnean Society 1995; 113: 21 - 46." type="journal article" year="1995">Wägele 1995: 37</bibRefCitation>
), with no distal features. Furthermore, most studies described
<taxonomicName id="4C0F4D19FF856502FF78FCFBEDABFC1C" box="[810,972,895,919]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B97BEA88FF856502FF78FCFBED12FC1C" box="[810,885,895,919]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
species.
</taxonomicName>
with a conspicuous capsule and mucous gland (
<bibRefCitation id="EF9E4B6BFF856502FF67FC1BED80FC3C" author="Wagele H" box="[821,999,927,951]" pageId="21" pageNumber="466" pagination="21 - 46" refId="ref27940" refString="Wagele H. The morphology and taxonomy of the Antarctic species of Tritonia Cuvier, 1797 (Nudibranchia: Dendronotoidea). Zoological Journal of the Linnean Society 1995; 113: 21 - 46." type="journal article" year="1995">Wägele 1995: 37</bibRefCitation>
; Ballesteros and Ávila 2006: 134). Although further morphological studies are needed to beưer understand this clade, our phylogenetic analyses, species delimitation analyses and interspecific uncorrected
<emphasis id="B97BEA88FF856502F8C3FC7AEAF9FB9E" box="[1169,1182,1022,1045]" italics="true" pageId="21" pageNumber="466">p</emphasis>
-distances strongly support the resurrection of the cohesive, well-supported clade
<taxonomicName id="4C0F4D19FF856502F91CFB98EBFEFBBF" authorityName="Odhner" authorityYear="1963" box="[1358,1433,1052,1076]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502F91CFB98EBFEFBBF" box="[1358,1433,1052,1076]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB0369AFF856502FF17FBBFED08FAE5" blockId="21.[810,1459,144,1390]" pageId="21" pageNumber="466">
Currently,
<taxonomicName id="4C0F4D19FF856502FFE6FBB8EABAFBD8" box="[948,1245,1083,1107]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="variety" species="challengeriana" variety="vorax">
<emphasis id="B97BEA88FF856502FFE6FBB8EDB3FBD8" box="[948,980,1084,1107]" italics="true" pageId="21" pageNumber="466">My</emphasis>
.
<emphasis id="B97BEA88FF856502FFB3FBBFEA0AFBD8" box="[993,1133,1083,1107]" italics="true" pageId="21" pageNumber="466">challengeriana</emphasis>
,
<emphasis id="B97BEA88FF856502F828FBB8EABAFBD8" box="[1146,1245,1084,1107]" italics="true" pageId="21" pageNumber="466">My. vorax</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F4D19FF856502F943FBB8EBEAFBD8" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Ballesteros and Avila" baseAuthorityYear="2006" box="[1297,1421,1083,1107]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="dantarti">
<emphasis id="B97BEA88FF856502F943FBB8EBEAFBD8" box="[1297,1421,1083,1107]" italics="true" pageId="21" pageNumber="466">My. dantarti</emphasis>
</taxonomicName>
are the three valid species of
<taxonomicName id="4C0F4D19FF856502F868FBDFEAE2FBF8" authorityName="Odhner" authorityYear="1963" box="[1082,1157,1115,1139]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502F868FBDFEAE2FBF8" box="[1082,1157,1115,1139]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
</taxonomicName>
found in the South Atlantic Ocean. However, recent studies have confirmed that the species diversity of
<taxonomicName id="4C0F4D19FF856502FFF1FB1DED89FB3A" authorityName="Odhner" authorityYear="1963" box="[931,1006,1177,1201]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF856502FFF1FB1DED89FB3A" box="[931,1006,1177,1201]" italics="true" pageId="21" pageNumber="466">Myrella</emphasis>
</taxonomicName>
is richer than presented in our phylogenetic analyses, including a complex of unnamed species previously identified as
<taxonomicName id="4C0F4D19FF856502FFE1FB5DEB23FB7B" authority="(Moles et al. 2021)" baseAuthorityName="Moles" baseAuthorityYear="2021" box="[947,1348,1240,1264]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="challengeriana">
<emphasis id="B97BEA88FF856502FFE1FB5DEA0BFB7B" box="[947,1132,1240,1264]" italics="true" pageId="21" pageNumber="466">My. challengeriana</emphasis>
(
<bibRefCitation id="EF9E4B6BFF856502F8D2FB5CEB5FFB7B" author="Moles J &amp; Berning MI &amp; Hooker Y" box="[1152,1336,1240,1264]" pageId="21" pageNumber="466" pagination="107209" refId="ref26342" refString="Moles J, Berning MI, Hooker Y, et al. Due South: the evolutionary history of sub-Antarctic and Antarctic Tritoniidae nudibranchs. Molecular Phylogenetics and Evolution 2021; 162: 107209." type="journal article" year="2021">
Moles
<emphasis id="B97BEA88FF856502F89BFB5DEA92FB7B" box="[1225,1269,1240,1264]" italics="true" pageId="21" pageNumber="466">et al</emphasis>
. 2021
</bibRefCitation>
)
</taxonomicName>
. Although the original description of
<taxonomicName id="4C0F4D19FF856502F866FB7CEAD9FA84" box="[1076,1214,1272,1295]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="antarctica">
<emphasis id="B97BEA88FF856502F866FB7CEAD9FA84" box="[1076,1214,1272,1295]" italics="true" pageId="21" pageNumber="466">My. antarctica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F4D19FF856502F8BFFB7CEB0FFA84" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Bergh" baseAuthorityYear="1898" box="[1261,1384,1271,1295]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="466" phylum="Mollusca" rank="species" species="australis">
<emphasis id="B97BEA88FF856502F8BFFB7CEB0FFA84" box="[1261,1384,1271,1295]" italics="true" pageId="21" pageNumber="466">My. australis</emphasis>
</taxonomicName>
fits our proposed diagnosis, these species are included with caution due to their uncertain taxonomic validity (
<bibRefCitation id="EF9E4B6BFF856502F8FFFAB2EB51FAC5" author="Schrodl M" box="[1197,1334,1334,1358]" pageId="21" pageNumber="466" refId="ref27482" refString="Schrodl M. Sea slugs of southern South America - systematics, biogeography and biology of Chilean and Magellanic Nudipleura (Mollusca: Opisthobranchia). Hackenheim: ConchBooks, 2003." type="book" year="2003">Schrödl 2003</bibRefCitation>
,
<bibRefCitation id="EF9E4B6BFF856502F913FAB2ED39FAE6" author="Moles J &amp; Berning MI &amp; Hooker Y" pageId="21" pageNumber="466" pagination="107209" refId="ref26342" refString="Moles J, Berning MI, Hooker Y, et al. Due South: the evolutionary history of sub-Antarctic and Antarctic Tritoniidae nudibranchs. Molecular Phylogenetics and Evolution 2021; 162: 107209." type="journal article" year="2021">
Moles
<emphasis id="B97BEA88FF856502F9D7FAB3EBCAFAC5" box="[1413,1453,1334,1358]" italics="true" pageId="21" pageNumber="466">et al</emphasis>
. 2021
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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