treatments-xml/data/03/A6/87/03A68780FFC1FFA48F3E022A79A8110D.xml

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<mods:title id="AEACF32BB103393D9C7B183DEE796D3F">The land snail genus Pincerna Preston, 1907 (Gastropoda: Alycaeidae) from Vietnam and Laos, with description of a new species</mods:title>
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<heading id="D0F881FAFFC1FFA98F3E022A7DAF1419" bold="true" box="[862,1447,604,629]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="369" reason="2">
<taxonomicName id="4C0F4D15FFC1FFA98F3E022A7DAF1419" ID-CoL="7SQ47" authority="(Bavay &amp; Dautzenberg, 1900)" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[862,1447,604,629]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="vanbuensis">
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<emphasis id="B97BEA84FFC1FFA98F3E022A7C4F1419" bold="true" box="[862,1095,604,628]" italics="true" pageId="5" pageNumber="369">Pincerna vanbuensis</emphasis>
(Bavay &amp; Dautzenberg, 1900)
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(
<figureCitation id="13342A13FFC1FFA98FBA020B7C2D14F8" box="[986,1061,637,661]" captionStart="Fig" captionStartId="2.[113,148,1941,1963]" captionTargetBox="[113,1452,311,1926]" captionTargetId="figure-11@2.[113,1452,311,1926]" captionTargetPageId="2" captionText="Fig. 1. Distribution of Pincerna species in Vietnam and Laos. Triangle: Pincerna anceyi; Grey circle: Pincerna costulosus; Black circles: Pincerna mouhoti; Squares: Pincerna vanbuensis; Star: Pincerna clausa, new species. The localities are detailed in Table 1." figureDoi="http://doi.org/10.5281/zenodo.7174166" httpUri="https://zenodo.org/record/7174166/files/figure.png" pageId="5" pageNumber="369">Figs. 1</figureCitation>
, 
<figureCitation id="13342A13FFC1FFA98855020B7C7D14F8" box="[1077,1141,637,661]" captionStart="Fig" captionStartId="4.[113,148,2035,2057]" captionTargetBox="[134,1429,188,2020]" captionTargetId="figure-12@4.[134,1429,188,2020]" captionTargetPageId="4" captionText="Fig. 3. Shells (apertural view, dorsal view, ventral view, and side view) and operculum (outer surface, inner surface, and side view) of Pincerna vanbuensis (Bavay &amp; Dautzenberg, 1900). A, syntype MNHN-IM-2000-31798; B, E, specimen from Thuận Châu, SƠn La, Vietnam, ZMHU; C, specimen from Khoune, Xieng Khouang, Laos, ZMHU; D, specimen from Xiengkho, Houaphanh, Laos, ZMHU." figureDoi="http://doi.org/10.5281/zenodo.7174170" httpUri="https://zenodo.org/record/7174170/files/figure.png" pageId="5" pageNumber="369">3A–E</figureCitation>
, 
<figureCitation id="13342A13FFC1FFA988E3020A7CAC14F9" box="[1155,1188,636,660]" captionStart="Fig" captionStartId="8.[113,149,1150,1172]" captionTargetBox="[113,1451,188,1135]" captionTargetId="figure-34@8.[113,1451,188,1135]" captionTargetPageId="8" captionText="Fig. 6. Synoptic view of Vietnamese and Laos Pincerna species. A, MNHN-IM-2000-31797; B, NHMUK 20170120; C, MNHN- IM-2000-31786; D, VNMN-IZ 002.304; E, MNHN-IM-2000-31798." figureDoi="http://doi.org/10.5281/zenodo.7174176" httpUri="https://zenodo.org/record/7174176/files/figure.png" pageId="5" pageNumber="369">6E</figureCitation>
; 
<tableCitation id="C68D032DFFC1FFA988D0020B7D0514F8" box="[1200,1293,637,661]" captionStart="Table 1" captionStartId="9.[136,191,189,211]" captionText="Table 1. Shell size variation of Pincerna spp. specimen collections in this study" pageId="5" pageNumber="369">Tables 1</tableCitation>
, 
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)
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<treatmentCitation id="0AAE1087FFC1FFA98F2302CD7DCA14BD" author="Bavay A &amp; Dautzenberg P" box="[835,1474,699,721]" page="120" pageId="5" pageNumber="369" year="1900">
<taxonomicName id="4C0F4D15FFC1FFA98F2302CD7DCA14BD" ID-CoL="7PYPJ" authority="Bavay &amp; Dautzenberg, 1900 a: 120" authorityName="Bavay &amp; Dautzenberg" authorityPageNumber="120" authorityYear="1900" box="[835,1474,699,721]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="vanbuensis" subGenus="Dioryx">
<emphasis id="B97BEA84FFC1FFA98F2302CD7B9214BD" box="[835,922,699,720]" italics="true" pageId="5" pageNumber="369">Alycaeus</emphasis>
(
<emphasis id="B97BEA84FFC1FFA98FC902CA7BE414BC" box="[937,1004,700,721]" italics="true" pageId="5" pageNumber="369">Dioryx</emphasis>
) 
<emphasis id="B97BEA84FFC1FFA98F9C02CD7C6F14BD" box="[1020,1127,699,720]" italics="true" pageId="5" pageNumber="369">vanbuensis</emphasis>
<materialsCitation id="3B673CCBFFC1FFA9880E02CA7DCA14BD" box="[1134,1474,699,721]" country="Vietnam" location="SOn La Province" pageId="5" pageNumber="369" specimenCount="1" stateProvince="Son La">
<bibRefCitation id="EF9E4B67FFC1FFA9880E02CA7DCA14BD" author="Bavay A &amp; Dautzenberg P" box="[1134,1474,699,721]" pageId="5" pageNumber="369" pagination="108 - 122" refId="ref5893" refString="Bavay A &amp; Dautzenberg P (1900 a) Diagnoses de Coquilles Nouvelles de l'Indo-Chine. Journal de Conchyliologie, 48: 108 - 122." type="journal article" year="1900">Bavay &amp; Dautzenberg, 1900a: 120</bibRefCitation>
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(
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locality: Van-Bu = 
<collectingRegion id="49CBF874FFC1FFA9880202AE7D011483" box="[1122,1289,728,751]" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La Province</collectingRegion>
, 
<collectingCountry id="F3187606FFC1FFA9897402AE7D631483" box="[1300,1387,728,750]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
).
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<treatmentCitationGroup id="AB1F11B8FFC1FFA98F2302837C5D154A" pageId="5" pageNumber="369">
<taxonomicName id="4C0F4D15FFC1FFA98F2302837C781567" ID-CoL="7PYPJ" authorityName="Bavay &amp; Dautzenberg" authorityYear="1900" box="[835,1136,757,778]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="vanbuensis" subGenus="Dioryx">
<emphasis id="B97BEA84FFC1FFA98F2302837B931567" box="[835,923,757,778]" italics="true" pageId="5" pageNumber="369">Alycaeus</emphasis>
(
<emphasis id="B97BEA84FFC1FFA98FCD02837BF91567" box="[941,1009,757,778]" italics="true" pageId="5" pageNumber="369">Dioryx</emphasis>
) 
<emphasis id="B97BEA84FFC1FFA9886302837C781567" box="[1027,1136,757,778]" italics="true" pageId="5" pageNumber="369">vanbuensis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AAE1087FFC1FFA988EE02837B87154A" author="Bavay A &amp; Dautzenberg P" page="455" pageId="5" pageNumber="369" year="1900">
<bibRefCitation id="EF9E4B67FFC1FFA988EE02837B87154A" author="Bavay A &amp; Dautzenberg P" pageId="5" pageNumber="369" pagination="435 - 460" refId="ref5919" refString="Bavay A &amp; Dautzenberg P (1900 b) Description de Coquilles Nouvelles de l'Indo-Chine. Journal de Conchyliologie, 48: 435 - 460." type="journal article" year="1900">Bavay &amp; Dautzenberg, 1900b: 455</bibRefCitation>
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, pl. 11, figs. 19–21.
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<treatmentCitationGroup id="AB1F11B8FFC1FFA98F2303597DCB1529" box="[835,1475,814,836]" pageId="5" pageNumber="369">
<taxonomicName id="4C0F4D15FFC1FFA98F2303597BFB152E" ID-CoL="7R37L" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[835,1011,814,836]" class="Gastropoda" family="Alycaeidae" genus="Dioryx" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="B97BEA84FFC1FFA98F2303597BFB152E" box="[835,1011,814,836]" italics="true" pageId="5" pageNumber="369">Dioryx vanbuensis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AAE1087FFC1FFA9886803587CBB1529" author="Kobelt W" box="[1032,1203,814,836]" page="340" pageId="5" pageNumber="369" year="1902">
<bibRefCitation id="EF9E4B67FFC1FFA9886803587CBB1529" author="Kobelt W" box="[1032,1203,814,836]" pageId="5" pageNumber="369" refId="ref6654" refString="Kobelt W (1902) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlander und Sohn, Berlin, 662 pp." type="book" year="1902">Kobelt, 1902: 340</bibRefCitation>
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; 
<treatmentCitation id="0AAE1087FFC1FFA988DC03597D7D1529" author="Do DS &amp; Nguyen THT &amp; Do VN" box="[1212,1397,814,836]" page="120" pageId="5" pageNumber="369" year="2015">
<bibRefCitation id="EF9E4B67FFC1FFA988DC03597D7D1529" author="Do DS &amp; Nguyen THT &amp; Do VN" box="[1212,1397,814,836]" pageId="5" pageNumber="369" pagination="117 - 132" refId="ref6084" refString="Do DS, Nguyen THT &amp; Do VN (2015) A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica, 25 (4): 117 - 132." type="journal article" year="2015">Do et al., 2015: 120</bibRefCitation>
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, fig. 2C.
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<paragraph id="8BB03696FFC1FFA98F23033D7BD915F7" blockId="5.[835,1475,699,952]" pageId="5" pageNumber="369">
<treatmentCitationGroup id="AB1F11B8FFC1FFA98F23033D7BD915F7" pageId="5" pageNumber="369">
<taxonomicName id="4C0F4D15FFC1FFA98F23033D7C06150D" ID-CoL="7PYPJ" authorityName="Bavay &amp; Dautzenberg" authorityYear="1900" box="[835,1038,843,864]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="B97BEA84FFC1FFA98F23033D7C06150D" box="[835,1038,843,864]" italics="true" pageId="5" pageNumber="369">Alycaeus vanbuensis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AAE1087FFC1FFA9884C033D7D80150D" author="Fischer H &amp; Dautzenberg P" box="[1068,1416,843,865]" page="432" pageId="5" pageNumber="369" year="1904">
<bibRefCitation id="EF9E4B67FFC1FFA9884C033D7D80150D" author="Fischer H &amp; Dautzenberg P" box="[1068,1416,843,865]" pageId="5" pageNumber="369" pagination="390 - 450" refId="ref6307" refString="Fischer H &amp; Dautzenberg P (1904) Catalogue des mollusques terrestres et fluviatiles de l'IndoChine orientale cites jusqu' a ce jour. In: Mission Pavie, Etudes diverses. Volume 3, pp. 390 - 450." type="journal article" year="1904">Fischer &amp; Dautzenberg, 1904: 432</bibRefCitation>
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; PállGergely et al., 2017: 10, fig. 3C; 
<treatmentCitation id="0AAE1087FFC1FFA988D6031E7DB61510" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" box="[1206,1470,872,894]" httpUri="http://treatment.plazi.org/id/B7B508E8191A6819ED663418CE1EED79" page="13" pageId="5" pageNumber="369" year="2019">
<bibRefCitation id="EF9E4B67FFC1FFA988D6031E7DB61510" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" box="[1206,1470,872,894]" pageId="5" pageNumber="369" pagination="1 - 166" refId="ref6528" refString="Inkhavilay K, Sutcharit C, Bantaowong U, Chanabun R, Siriwut W, Srisonchai R, Pholyotha A, Jirapatrasilp P &amp; Panha S (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys, 834: 1 - 166." type="journal article" year="2019">Inkhavilay et al., 2019: 13</bibRefCitation>
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, figs 4C–E.
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<paragraph id="8BB03696FFC1FFA98F2303D47D5B15DA" blockId="5.[835,1475,699,952]" box="[835,1363,930,952]" pageId="5" pageNumber="369">
<taxonomicName id="4C0F4D15FFC1FFA98F2303D47C0615DA" ID-CoL="7SQ47" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[835,1038,930,951]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="B97BEA84FFC1FFA98F2303D47C0615DA" box="[835,1038,930,951]" italics="true" pageId="5" pageNumber="369">Pincerna vanbuensis</emphasis>
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– Páll-Gergely et al., 2020: 184.
</paragraph>
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<emphasis id="B97BEA84FFC1FFA98F2303967C2D1595" bold="true" box="[835,1061,992,1016]" pageId="5" pageNumber="369">Material examined.</emphasis>
<materialsCitation id="3B673CCBFFC1FFA9884F03967D6D1255" collectionCode="R" collectorName="Dr. &amp; Van Bu" country="Vietnam" location="Tonkin" pageId="5" pageNumber="369" specimenCode="RBINS 525448" specimenCount="2" typeStatus="syntype">
Type material. 
<collectingCountry id="F3187606FFC1FFA988B803967D201595" box="[1240,1320,992,1016]" name="Vietnam" pageId="5" pageNumber="369">Tonkin</collectingCountry>
, Van-Bu, leg. 
<collectorName id="26FA5340FFC1FFA98F2304767B6F127A" box="[835,871,1024,1047]" pageId="5" pageNumber="369">Dr.</collectorName>
<collectionCode id="ED1EAE53FFC1FFA98F0D04767B8A127A" box="[877,898,1024,1047]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="369">R</collectionCode>
. Bavay, MNHN-IM-2000-31798 (
<typeStatus id="54B48834FFC1FFA9889204767D401275" box="[1266,1352,1024,1048]" pageId="5" pageNumber="369" type="syntype">syntype</typeStatus>
) (
<figureCitation id="13342A13FFC1FFA9893E04767DB51275" box="[1374,1469,1024,1048]" captionStart="Fig" captionStartId="4.[113,148,2035,2057]" captionTargetBox="[134,1429,188,2020]" captionTargetId="figure-12@4.[134,1429,188,2020]" captionTargetPageId="4" captionText="Fig. 3. Shells (apertural view, dorsal view, ventral view, and side view) and operculum (outer surface, inner surface, and side view) of Pincerna vanbuensis (Bavay &amp; Dautzenberg, 1900). A, syntype MNHN-IM-2000-31798; B, E, specimen from Thuận Châu, SƠn La, Vietnam, ZMHU; C, specimen from Khoune, Xieng Khouang, Laos, ZMHU; D, specimen from Xiengkho, Houaphanh, Laos, ZMHU." figureDoi="http://doi.org/10.5281/zenodo.7174170" httpUri="https://zenodo.org/record/7174170/files/figure.png" pageId="5" pageNumber="369">Figs. 3A</figureCitation>
, 
<figureCitation id="13342A13FFC1FFA98F2304567B6C1255" box="[835,868,1056,1080]" captionStart="Fig" captionStartId="8.[113,149,1150,1172]" captionTargetBox="[113,1451,188,1135]" captionTargetId="figure-34@8.[113,1451,188,1135]" captionTargetPageId="8" captionText="Fig. 6. Synoptic view of Vietnamese and Laos Pincerna species. A, MNHN-IM-2000-31797; B, NHMUK 20170120; C, MNHN- IM-2000-31786; D, VNMN-IZ 002.304; E, MNHN-IM-2000-31798." figureDoi="http://doi.org/10.5281/zenodo.7174176" httpUri="https://zenodo.org/record/7174176/files/figure.png" pageId="5" pageNumber="369">6E</figureCitation>
); 
<collectorName id="26FA5340FFC1FFA98F1A04567BD9125A" box="[890,977,1056,1079]" pageId="5" pageNumber="369">Van Bu</collectorName>
, 
<collectingCountry id="F3187606FFC1FFA98FBD04567C251255" box="[989,1069,1056,1080]" name="Vietnam" pageId="5" pageNumber="369">Tonkin</collectingCountry>
, 
<specimenCode id="DBA99EEDFFC1FFA9885A04567CE91255" box="[1082,1249,1056,1080]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="5" pageNumber="369" type="Museum">RBINS 525448</specimenCode>
(
<typeStatus id="54B48834FFC1FFA9889104567D561255" box="[1265,1374,1056,1080]" pageId="5" pageNumber="369" type="paratype">paratypes</typeStatus>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB03696FFC1FFA98F2304167D841195" blockId="5.[835,1475,1120,2040]" pageId="5" pageNumber="369">
<emphasis id="B97BEA84FFC1FFA98F2304167CA91215" bold="true" box="[835,1185,1120,1144]" pageId="5" pageNumber="369">Additional material examined.</emphasis>
<materialsCitation id="3B673CCBFFC1FFA988C804167B6C129A" collectionCode="ZMHU" collectorName="D. S. Do" country="Vietnam" county="Thu" elevation="1239" latitude="21.356945" location="Co" longLatPrecision="21" longitude="103.52056" municipality="Chau District" pageId="5" pageNumber="369" specimenCode="ZMHU/ 20" specimenCount="1" stateProvince="Son La">
<specimenCode id="DBA99EEDFFC1FFA988C804167D2B1215" box="[1192,1315,1120,1144]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 20</specimenCode>
, 
<collectingCountry id="F3187606FFC1FFA9894F04167D861215" box="[1327,1422,1120,1144]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
, 
<collectingRegion id="49CBF874FFC1FFA989F904167BC012F5" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La Province</collectingRegion>
, 
<collectingCounty id="62D14E1AFFC1FFA98FB104F67BF412F5" box="[977,1020,1152,1176]" pageId="5" pageNumber="369">Thu</collectingCounty>
ận 
<collectingMunicipality id="6BD4ACECFFC1FFA9887C04F67CA512F5" box="[1052,1197,1152,1176]" pageId="5" pageNumber="369">Châu District</collectingMunicipality>
, 
<location id="8ED0604DFFC1FFA988D704F67CDE12F5" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA988D704F67CDE12F5" box="[1207,1238,1152,1176]" country="Vietnam" county="Thu" latitude="21.356945" longLatPrecision="21" longitude="103.52056" municipality="Chau District" name="Co" pageId="5" pageNumber="369" stateProvince="Son La">Co</location>
Mạ 
<location id="8ED0604DFFC1FFA9896604F67D7012F5" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA9896604F67D7012F5" box="[1286,1400,1152,1176]" country="Vietnam" county="Thu" latitude="21.356945" longLatPrecision="21" longitude="103.52056" municipality="Chau District" name="Commune" pageId="5" pageNumber="369" stateProvince="Son La">Commune</location>
, 
<location id="8ED0604DFFC1FFA989E204F67BE412DA" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA989E204F67BE412DA" country="Vietnam" county="Thu" latitude="21.356945" longLatPrecision="21" longitude="103.52056" municipality="Chau District" name="Copia Nature Reserve" pageId="5" pageNumber="369" stateProvince="Son La">Copia Nature Reserve</location>
, steep limestone slope with dense disturbed vegetation, 
<geoCoordinate id="EE3B5051FFC1FFA98FDC04C87C3412BA" box="[956,1084,1214,1239]" degrees="21" direction="north" minutes="21" orientation="latitude" pageId="5" pageNumber="369" precision="15" seconds="25" value="21.356945">21°21′25″N</geoCoordinate>
, 
<geoCoordinate id="EE3B5051FFC1FFA9882504C87CC712BA" box="[1093,1231,1214,1239]" degrees="103" direction="east" minutes="31" orientation="longitude" pageId="5" pageNumber="369" precision="15" seconds="14" value="103.52056">103°31′14″E</geoCoordinate>
, 
<elevation id="0022D1A5FFC1FFA988B804B67D5412B5" box="[1240,1372,1216,1240]" metricMagnitude="3" metricUnit="m" metricValue="1.239" pageId="5" pageNumber="369" unit="m" value="1239.0">
<quantity id="4CF79B73FFC1FFA988B804B67D2E12BA" box="[1240,1318,1216,1240]" metricMagnitude="3" metricUnit="m" metricValue="1.239" pageId="5" pageNumber="369" unit="m" value="1239.0">1239 m</quantity>
a.s.l.
</elevation>
, col. 
<collectorName id="26FA5340FFC1FFA989F304B67B6C129A" pageId="5" pageNumber="369">D.S. Do</collectorName>
</materialsCitation>
&amp; 
<materialsCitation id="3B673CCBFFC1FFA98FED04967CCE1295" box="[909,1222,1248,1272]" collectingDate="2013-06-08" collectionCode="T" country="Vietnam" location="Nguyen" pageId="5" pageNumber="369" specimenCount="1" stateProvince="Son La">
<collectionCode id="ED1EAE53FFC1FFA98FED04967BA8129A" box="[909,928,1248,1271]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="369">T</collectionCode>
.H. 
<location id="8ED0604DFFC1FFA98FA804967C171295" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA98FA804967C171295" box="[968,1055,1248,1272]" country="Vietnam" name="Nguyen" pageId="5" pageNumber="369" stateProvince="Son La">Nguyen</location>
, 
<date id="FFB11056FFC1FFA9884D04967CCE1295" box="[1069,1222,1248,1272]" pageId="5" pageNumber="369" value="2013-06-08">
<collectingDate id="EFF5E9BEFFC1FFA9884D04967CCE1295" box="[1069,1222,1248,1272]" pageId="5" pageNumber="369" value="2013-06-08">08 June 2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B673CCBFFC1FFA988B404967CBA1375" collectionCode="ZMHU" collectorName="Mai &amp; District" country="Vietnam" location="SOn La Province" pageId="5" pageNumber="369" specimenCode="ZMHU/ 11" specimenCount="1" stateProvince="Son La">
<specimenCode id="DBA99EEDFFC1FFA988B404967D581295" box="[1236,1360,1248,1272]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 11</specimenCode>
, 
<collectingCountry id="F3187606FFC1FFA9893F04967DB61295" box="[1375,1470,1248,1272]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
, 
<collectingRegion id="49CBF874FFC1FFA98F2305767BFD1375" box="[835,1013,1280,1306]" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La Province</collectingRegion>
, 
<collectorName id="26FA5340FFC1FFA98F9F05767C221375" box="[1023,1066,1280,1304]" pageId="5" pageNumber="369">Mai</collectorName>
SƠn 
<collectorName id="26FA5340FFC1FFA9883F05767CBA1375" box="[1119,1202,1280,1304]" pageId="5" pageNumber="369">District</collectorName>
</materialsCitation>
, 
<materialsCitation id="3B673CCBFFC1FFA988DB05767DAB133A" collectionCode="MU" collectorName="D. S. Do" country="Vietnam" elevation="779" latitude="21.303055" location="Na Hoi Village" longLatPrecision="21" longitude="104.03778" municipality="Commune" pageId="5" pageNumber="369" specimenCount="1" stateProvince="Son La">
<collectionCode id="ED1EAE53FFC1FFA988DB05767CED1377" box="[1211,1253,1280,1306]" country="USA" name="Midwestern University" pageId="5" pageNumber="369">MƯ</collectionCode>
ờng Bằng 
<collectingMunicipality id="6BD4ACECFFC1FFA9892C05767DB61375" box="[1356,1470,1280,1304]" pageId="5" pageNumber="369">Commune</collectingMunicipality>
, 
<location id="8ED0604DFFC1FFA98F2305567BFE1355" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA98F2305567BFE1355" box="[835,1014,1312,1336]" country="Vietnam" latitude="21.303055" longLatPrecision="21" longitude="104.03778" municipality="Commune" name="Na Hoi Village" pageId="5" pageNumber="369" stateProvince="Son La">Nà Hoi Village</location>
, limestone hill with degraded regrowth, 
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, 
<geoCoordinate id="EE3B5051FFC1FFA98FB905487C61133A" box="[985,1129,1342,1367]" degrees="104" direction="east" minutes="02" orientation="longitude" pageId="5" pageNumber="369" precision="15" seconds="16" value="104.03778">104°02′16″E</geoCoordinate>
, 
<elevation id="0022D1A5FFC1FFA9881905367D0A1335" box="[1145,1282,1344,1368]" metricMagnitude="2" metricUnit="m" metricValue="7.79" pageId="5" pageNumber="369" unit="m" value="779.0">
<quantity id="4CF79B73FFC1FFA9881905367CCA133A" box="[1145,1218,1344,1368]" metricMagnitude="2" metricUnit="m" metricValue="7.79" pageId="5" pageNumber="369" unit="m" value="779.0">779 m</quantity>
a.s.l.
</elevation>
, col. 
<collectorName id="26FA5340FFC1FFA9892605367DAB133A" box="[1350,1443,1344,1368]" pageId="5" pageNumber="369">D.S. Do</collectorName>
</materialsCitation>
&amp; 
<materialsCitation id="3B673CCBFFC1FFA98F2405167CA31315" box="[836,1195,1376,1400]" collectingDate="2013-11-22" collectionCode="T" country="Vietnam" location="Nguyen" pageId="5" pageNumber="369" specimenCount="1" stateProvince="Son La">
<collectionCode id="ED1EAE53FFC1FFA98F2405167B5F131A" box="[836,855,1376,1399]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="369">T</collectionCode>
.L. 
<location id="8ED0604DFFC1FFA98F1805167BC71315" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA98F1805167BC71315" box="[888,975,1376,1400]" country="Vietnam" name="Nguyen" pageId="5" pageNumber="369" stateProvince="Son La">Nguyen</location>
, 
<date id="FFB11056FFC1FFA98FBA05167CA31315" box="[986,1195,1376,1400]" pageId="5" pageNumber="369" value="2013-11-22">
<collectingDate id="EFF5E9BEFFC1FFA98FBA05167CA31315" box="[986,1195,1376,1400]" pageId="5" pageNumber="369" value="2013-11-22">22 November 2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B673CCBFFC1FFA988D505167CF313BA" collectionCode="ZMHU" collectorName="D. S. Do" country="Vietnam" county="Thu" elevation="605" latitude="21.373056" location="Bon Ph" longLatPrecision="21" longitude="103.77195" municipality="Chau District" pageId="5" pageNumber="369" specimenCode="ZMHU/ 23" specimenCount="1" stateProvince="Son La">
<specimenCode id="DBA99EEDFFC1FFA988D505167D241315" box="[1205,1324,1376,1400]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 23</specimenCode>
(
<figureCitation id="13342A13FFC1FFA9895A05167DBE1315" box="[1338,1462,1376,1400]" captionStart="Fig" captionStartId="4.[113,148,2035,2057]" captionTargetBox="[134,1429,188,2020]" captionTargetId="figure-12@4.[134,1429,188,2020]" captionTargetPageId="4" captionText="Fig. 3. Shells (apertural view, dorsal view, ventral view, and side view) and operculum (outer surface, inner surface, and side view) of Pincerna vanbuensis (Bavay &amp; Dautzenberg, 1900). A, syntype MNHN-IM-2000-31798; B, E, specimen from Thuận Châu, SƠn La, Vietnam, ZMHU; C, specimen from Khoune, Xieng Khouang, Laos, ZMHU; D, specimen from Xiengkho, Houaphanh, Laos, ZMHU." figureDoi="http://doi.org/10.5281/zenodo.7174170" httpUri="https://zenodo.org/record/7174170/files/figure.png" pageId="5" pageNumber="369">Figs. 3B, E</figureCitation>
), 
<collectingCountry id="F3187606FFC1FFA98F2305F67BAA13F5" box="[835,930,1408,1432]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
, 
<collectingRegion id="49CBF874FFC1FFA98FCC05F67C5713F5" box="[940,1119,1408,1434]" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La Province</collectingRegion>
, 
<collectingCounty id="62D14E1AFFC1FFA9880905F67C9C13F5" box="[1129,1172,1408,1432]" pageId="5" pageNumber="369">Thu</collectingCounty>
ận 
<collectingMunicipality id="6BD4ACECFFC1FFA988D305F67D4C13F5" box="[1203,1348,1408,1432]" pageId="5" pageNumber="369">Châu District</collectingMunicipality>
, 
<location id="8ED0604DFFC1FFA9892D05F67D9413F5" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA9892D05F67D9413F5" box="[1357,1436,1408,1432]" country="Vietnam" county="Thu" latitude="21.373056" longLatPrecision="21" longitude="103.77195" municipality="Chau District" name="Bon Ph" pageId="5" pageNumber="369" stateProvince="Son La">Bon Ph</location>
ặng 
<location id="8ED0604DFFC1FFA98F2305D67BBD13D5" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA98F2305D67BBD13D5" box="[835,949,1440,1464]" country="Vietnam" county="Thu" latitude="21.373056" longLatPrecision="21" longitude="103.77195" municipality="Chau District" name="Commune" pageId="5" pageNumber="369" stateProvince="Son La">Commune</location>
, Đốm HƯ 
<location id="8ED0604DFFC1FFA9884205D67C9F13D5" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA9884205D67C9F13D5" box="[1058,1175,1440,1466]" country="Vietnam" county="Thu" latitude="21.373056" longLatPrecision="21" longitude="103.77195" municipality="Chau District" name="On Village" pageId="5" pageNumber="369" stateProvince="Son La">Ợn Village</location>
, 
<geoCoordinate id="EE3B5051FFC1FFA988C405E87D2D13DA" box="[1188,1317,1438,1463]" degrees="21" direction="north" minutes="22" orientation="latitude" pageId="5" pageNumber="369" precision="15" seconds="23" value="21.373056">21°22′23″N</geoCoordinate>
, 
<geoCoordinate id="EE3B5051FFC1FFA9895205E87DB613DA" box="[1330,1470,1438,1463]" degrees="103" direction="east" minutes="46" orientation="longitude" pageId="5" pageNumber="369" precision="15" seconds="19" value="103.77195">103°46′19″E</geoCoordinate>
, 
<quantity id="4CF79B73FFC1FFA98F2305B67B8F13BA" box="[835,903,1472,1496]" metricMagnitude="2" metricUnit="m" metricValue="6.05" pageId="5" pageNumber="369" unit="m" value="605.0">
<elevation id="0022D1A5FFC1FFA98F2305B67B8F13BA" box="[835,903,1472,1496]" metricMagnitude="2" metricUnit="m" metricValue="6.05" pageId="5" pageNumber="369" unit="m" value="605.0">605 m</elevation>
</quantity>
a.s.l, limestone karst, col. 
<collectorName id="26FA5340FFC1FFA988C405B67CF313BA" box="[1188,1275,1472,1496]" pageId="5" pageNumber="369">D.S. Do</collectorName>
</materialsCitation>
&amp; 
<materialsCitation id="3B673CCBFFC1FFA9897D05B67BDB1395" collectingDate="2013-08-20" collectionCode="T" country="Vietnam" location="Luong" pageId="5" pageNumber="369" specimenCount="1" stateProvince="Son La">
<collectionCode id="ED1EAE53FFC1FFA9897D05B67D3813BA" box="[1309,1328,1472,1495]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="369">T</collectionCode>
.D. 
<location id="8ED0604DFFC1FFA9893405B67D9413B5" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA9893405B67D9413B5" box="[1364,1436,1472,1496]" country="Vietnam" name="Luong" pageId="5" pageNumber="369" stateProvince="Son La">Luong</location>
, 
<date id="FFB11056FFC1FFA989C705B67BDB1395" pageId="5" pageNumber="369" value="2013-08-20">
<collectingDate id="EFF5E9BEFFC1FFA989C705B67BDB1395" pageId="5" pageNumber="369" value="2013-08-20">20 August 2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B673CCBFFC1FFA98F8105967D691035" collectingDate="2017-09-02" collectionCode="ZMHU" collectorName="D. S. Do &amp; X. H. Nguyen" country="Vietnam" county="Yen Chau District" elevation="815" latitude="21.141666" location="Commune" longLatPrecision="21" longitude="104.17472" municipality="Yen" pageId="5" pageNumber="369" specimenCode="ZMHU/ 5" specimenCount="1" stateProvince="Son La">
<specimenCode id="DBA99EEDFFC1FFA98F8105967C581395" box="[993,1104,1504,1528]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 5</specimenCode>
, 
<collectingCountry id="F3187606FFC1FFA9883F05967CB61395" box="[1119,1214,1504,1528]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
, 
<collectingRegion id="49CBF874FFC1FFA988AC05967D801395" box="[1228,1416,1504,1530]" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La Province</collectingRegion>
, 
<collectingCounty id="62D14E1AFFC1FFA989F505967BDE1075" pageId="5" pageNumber="369">Yên Châu District</collectingCounty>
, 
<collectingMunicipality id="6BD4ACECFFC1FFA98F8206767C061075" box="[994,1038,1536,1560]" pageId="5" pageNumber="369">Yên</collectingMunicipality>
SƠn 
<location id="8ED0604DFFC1FFA9882806767CB21075" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA9882806767CB21075" box="[1096,1210,1536,1560]" country="Vietnam" county="Yen Chau District" latitude="21.141666" longLatPrecision="21" longitude="104.17472" municipality="Yen" name="Commune" pageId="5" pageNumber="369" stateProvince="Son La">Commune</location>
, 
<location id="8ED0604DFFC1FFA988A606767CE51075" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA988A606767CE51075" box="[1222,1261,1536,1560]" country="Vietnam" county="Yen Chau District" latitude="21.141666" longLatPrecision="21" longitude="104.17472" municipality="Yen" name="Chi" pageId="5" pageNumber="369" stateProvince="Son La">Chi</location>
Đảy 
<location id="8ED0604DFFC1FFA9894806767D691075" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA9894806767D691075" box="[1320,1377,1536,1560]" country="Vietnam" county="Yen Chau District" latitude="21.141666" longLatPrecision="21" longitude="104.17472" municipality="Yen" name="Cave" pageId="5" pageNumber="369" stateProvince="Son La">Cave</location>
, primary forest on limestone, 
<geoCoordinate id="EE3B5051FFC1FFA9887D06687C96105A" box="[1053,1182,1566,1591]" degrees="21" direction="north" minutes="08" orientation="latitude" pageId="5" pageNumber="369" precision="15" seconds="30" value="21.141666">21°08′30″N</geoCoordinate>
, 
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, 
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a.s.l.
</elevation>
, col. 
<collectorName id="26FA5340FFC1FFA98F1206367BC2103A" box="[882,970,1600,1624]" pageId="5" pageNumber="369">D.S. Do</collectorName>
&amp; 
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, 
<date id="FFB11056FFC1FFA988EC06367D691035" box="[1164,1377,1600,1624]" pageId="5" pageNumber="369" value="2017-09-02">
<collectingDate id="EFF5E9BEFFC1FFA988EC06367D691035" box="[1164,1377,1600,1624]" pageId="5" pageNumber="369" value="2017-09-02">02 September 2017</collectingDate>
</date>
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; 
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<specimenCode id="DBA99EEDFFC1FFA9890C06367B5A1015" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 4</specimenCode>
, 
<collectingCountry id="F3187606FFC1FFA98F3D06167BB41015" box="[861,956,1632,1656]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
, 
<collectingRegion id="49CBF874FFC1FFA98FA606167C9C1015" box="[966,1172,1632,1656]" country="Vietnam" name="Hoa Binh" pageId="5" pageNumber="369">Hòa Bình Province</collectingRegion>
, 
<collectingCounty id="62D14E1AFFC1FFA988FD06167D571015" box="[1181,1375,1632,1656]" pageId="5" pageNumber="369">Mai Châu District</collectingCounty>
, 
<collectingMunicipality id="6BD4ACECFFC1FFA9890806167D991015" box="[1384,1425,1632,1656]" pageId="5" pageNumber="369">Tân</collectingMunicipality>
SƠn 
<location id="8ED0604DFFC1FFA98F2306F67BBD10F5" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA98F2306F67BBD10F5" box="[835,949,1664,1688]" country="Vietnam" county="Mai Chau District" latitude="20.17472" longLatPrecision="21" longitude="104.97139" municipality="Tan" name="Commune" pageId="5" pageNumber="369" stateProvince="Hoa Binh">Commune</location>
, 
<location id="8ED0604DFFC1FFA98FA106F67C7310F5" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA98FA106F67C7310F5" box="[961,1147,1664,1688]" country="Vietnam" county="Mai Chau District" latitude="20.17472" longLatPrecision="21" longitude="104.97139" municipality="Tan" name="Bo Liem Village" pageId="5" pageNumber="369" stateProvince="Hoa Binh">Bó Liêm Village</location>
, under rocks in old secondary forest, 
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, 
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, 
<elevation id="0022D1A5FFC1FFA988A606D67D4710D5" box="[1222,1359,1696,1720]" metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" pageId="5" pageNumber="369" unit="m" value="553.0">
<quantity id="4CF79B73FFC1FFA988A606D67D0710DA" box="[1222,1295,1696,1720]" metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" pageId="5" pageNumber="369" unit="m" value="553.0">553 m</quantity>
a.s.l.
</elevation>
, col. 
<collectorName id="26FA5340FFC1FFA989F206D67B6D10BA" pageId="5" pageNumber="369">D.S. Do</collectorName>
&amp; 
<collectorName id="26FA5340FFC1FFA98FEA06B67C1E10B5" box="[906,1046,1728,1752]" pageId="5" pageNumber="369">C. Vilachark</collectorName>
, 
<date id="FFB11056FFC1FFA9884206B67CD110B5" box="[1058,1241,1728,1752]" pageId="5" pageNumber="369" value="2018-01-07">
<collectingDate id="EFF5E9BEFFC1FFA9884206B67CD110B5" box="[1058,1241,1728,1752]" pageId="5" pageNumber="369" value="2018-01-07">07 January 2018</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B673CCBFFC1FFA9888506B67CAD115A" collectionCode="ZMHU" collectorName="D. S. Do" country="Vietnam" elevation="297" latitude="20.353611" location="National Park" longLatPrecision="21" longitude="105.90611" municipality="Cuc" pageId="5" pageNumber="369" specimenCode="ZMHU/ 9" specimenCount="1" stateProvince="Ninh Binh">
<specimenCode id="DBA99EEDFFC1FFA9888506B67D5B10B5" box="[1253,1363,1728,1752]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 9</specimenCode>
, 
<collectingCountry id="F3187606FFC1FFA9890006B67DB710B5" box="[1376,1471,1728,1752]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
, 
<collectingRegion id="49CBF874FFC1FFA98F2306967C161095" box="[835,1054,1760,1784]" country="Vietnam" name="Ninh Binh" pageId="5" pageNumber="369">Ninh Bình Province</collectingRegion>
, 
<collectingMunicipality id="6BD4ACECFFC1FFA9884906967C5C1095" box="[1065,1108,1760,1784]" pageId="5" pageNumber="369">Cúc</collectingMunicipality>
PhƯƠng 
<location id="8ED0604DFFC1FFA988D706967D471095" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA988D706967D471095" box="[1207,1359,1760,1784]" country="Vietnam" latitude="20.353611" longLatPrecision="21" longitude="105.90611" municipality="Cuc" name="National Park" pageId="5" pageNumber="369" stateProvince="Ninh Binh">National Park</location>
, limestone outcrops in shale bedrock of primary forest, 
<geoCoordinate id="EE3B5051FFC1FFA9895B06887DB6117A" box="[1339,1470,1790,1815]" degrees="20" direction="north" minutes="21" orientation="latitude" pageId="5" pageNumber="369" precision="15" seconds="13" value="20.353611">20°21′13″N</geoCoordinate>
, 
<geoCoordinate id="EE3B5051FFC1FFA98F2307687BC6115A" box="[835,974,1822,1847]" degrees="105" direction="east" minutes="54" orientation="longitude" pageId="5" pageNumber="369" precision="15" seconds="22" value="105.90611">105°54′22″E</geoCoordinate>
, 
<quantity id="4CF79B73FFC1FFA98FB707567C13115A" box="[983,1051,1824,1848]" metricMagnitude="2" metricUnit="m" metricValue="2.9699999999999998" pageId="5" pageNumber="369" unit="m" value="297.0">
<elevation id="0022D1A5FFC1FFA98FB707567C13115A" box="[983,1051,1824,1848]" metricMagnitude="2" metricUnit="m" metricValue="2.9699999999999998" pageId="5" pageNumber="369" unit="m" value="297.0">297 m</elevation>
</quantity>
, col. 
<collectorName id="26FA5340FFC1FFA9883007567CAD115A" box="[1104,1189,1824,1848]" pageId="5" pageNumber="369">D.S. Do</collectorName>
</materialsCitation>
&amp; 
<materialsCitation id="3B673CCBFFC1FFA988A407567B741135" collectingDate="2016-08-19" collectionCode="T" country="Vietnam" location="Nguyen" pageId="5" pageNumber="369" specimenCount="1" stateProvince="Ninh Binh">
<collectionCode id="ED1EAE53FFC1FFA988A407567CDF115A" box="[1220,1239,1824,1847]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="369">T</collectionCode>
.L. 
<location id="8ED0604DFFC1FFA9889707567D451155" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA9889707567D451155" box="[1271,1357,1824,1848]" country="Vietnam" name="Nguyen" pageId="5" pageNumber="369" stateProvince="Ninh Binh">Nguyen</location>
, 
<date id="FFB11056FFC1FFA9893607567B741135" pageId="5" pageNumber="369" value="2016-08-19">
<collectingDate id="EFF5E9BEFFC1FFA9893607567B741135" pageId="5" pageNumber="369" value="2016-08-19">19 August 2016</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B673CCBFFC1FFA98FE607367C7011D5" collectingDate="2018-07-26" collectionCode="ZMHU" collectorName="C. Vilachark" country="Laos" county="Khoune District" latitude="19.533054" location="Pa Kha Village" longLatPrecision="21" longitude="103.516945" municipality="Nam Phan Commune" pageId="5" pageNumber="369" specimenCode="ZMHU/ 26" specimenCount="1" stateProvince="Xiangkhouang">
<specimenCode id="DBA99EEDFFC1FFA98FE607367BF41135" box="[902,1020,1856,1880]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 26</specimenCode>
(
<figureCitation id="13342A13FFC1FFA9886907367C531135" box="[1033,1115,1856,1880]" captionStart="Fig" captionStartId="4.[113,148,2035,2057]" captionTargetBox="[134,1429,188,2020]" captionTargetId="figure-12@4.[134,1429,188,2020]" captionTargetPageId="4" captionText="Fig. 3. Shells (apertural view, dorsal view, ventral view, and side view) and operculum (outer surface, inner surface, and side view) of Pincerna vanbuensis (Bavay &amp; Dautzenberg, 1900). A, syntype MNHN-IM-2000-31798; B, E, specimen from Thuận Châu, SƠn La, Vietnam, ZMHU; C, specimen from Khoune, Xieng Khouang, Laos, ZMHU; D, specimen from Xiengkho, Houaphanh, Laos, ZMHU." figureDoi="http://doi.org/10.5281/zenodo.7174170" httpUri="https://zenodo.org/record/7174170/files/figure.png" pageId="5" pageNumber="369">Fig. 3C</figureCitation>
), 
<collectingCountry id="F3187606FFC1FFA9880E07367CAC113A" box="[1134,1188,1856,1879]" name="Laos" pageId="5" pageNumber="369">Laos</collectingCountry>
, 
<collectingRegion id="49CBF874FFC1FFA988CE07367DB61135" box="[1198,1470,1856,1880]" country="Laos" name="Xiangkhouang" pageId="5" pageNumber="369">Xieng Khouang Province</collectingRegion>
, 
<collectingCounty id="62D14E1AFFC1FFA98F2307167BF21115" box="[835,1018,1888,1912]" pageId="5" pageNumber="369">Khoune District</collectingCounty>
, 
<collectingMunicipality id="6BD4ACECFFC1FFA9886907167CF51115" box="[1033,1277,1888,1912]" pageId="5" pageNumber="369">Nam Phan Commune</collectingMunicipality>
, 
<location id="8ED0604DFFC1FFA9896C07167DB51115" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA9896C07167DB51115" box="[1292,1469,1888,1912]" country="Laos" county="Khoune District" latitude="19.533054" longLatPrecision="21" longitude="103.516945" municipality="Nam Phan Commune" name="Pa Kha Village" pageId="5" pageNumber="369" stateProvince="Xiangkhouang">Pa Kha Village</location>
, limestone area near village, 
<geoCoordinate id="EE3B5051FFC1FFA9881507087CFE11FA" box="[1141,1270,1918,1943]" degrees="19" direction="north" minutes="31" orientation="latitude" pageId="5" pageNumber="369" precision="15" seconds="59" value="19.533054">19°31′59″N</geoCoordinate>
, 
<geoCoordinate id="EE3B5051FFC1FFA9896307087D8711FA" box="[1283,1423,1918,1943]" degrees="103" direction="east" minutes="31" orientation="longitude" pageId="5" pageNumber="369" precision="15" seconds="01" value="103.516945">103°31′01″E</geoCoordinate>
, col. 
<collectorName id="26FA5340FFC1FFA98F2307D67BDC11D5" box="[835,980,1952,1976]" pageId="5" pageNumber="369">C. Vilachark</collectorName>
, 
<date id="FFB11056FFC1FFA98F8207D67C7011D5" box="[994,1144,1952,1976]" pageId="5" pageNumber="369" value="2018-07-26">
<collectingDate id="EFF5E9BEFFC1FFA98F8207D67C7011D5" box="[994,1144,1952,1976]" pageId="5" pageNumber="369" value="2018-07-26">26 July 2018</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B673CCBFFC1FFA988E707D67D801195" collectingDate="2018-08-20" collectionCode="ZMHU" collectorName="P. S. L. Ti Khun" country="Laos" location="Na Tong Village" municipality="Xiengkho District" pageId="5" pageNumber="369" specimenCode="ZMHU/ 18" specimenCount="1" stateProvince="Houaphan">
<specimenCode id="DBA99EEDFFC1FFA988E707D67D0B11D5" box="[1159,1283,1952,1976]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 18</specimenCode>
(
<figureCitation id="13342A13FFC1FFA9897507D67D6611D5" box="[1301,1390,1952,1976]" captionStart="Fig" captionStartId="4.[113,148,2035,2057]" captionTargetBox="[134,1429,188,2020]" captionTargetId="figure-12@4.[134,1429,188,2020]" captionTargetPageId="4" captionText="Fig. 3. Shells (apertural view, dorsal view, ventral view, and side view) and operculum (outer surface, inner surface, and side view) of Pincerna vanbuensis (Bavay &amp; Dautzenberg, 1900). A, syntype MNHN-IM-2000-31798; B, E, specimen from Thuận Châu, SƠn La, Vietnam, ZMHU; C, specimen from Khoune, Xieng Khouang, Laos, ZMHU; D, specimen from Xiengkho, Houaphanh, Laos, ZMHU." figureDoi="http://doi.org/10.5281/zenodo.7174170" httpUri="https://zenodo.org/record/7174170/files/figure.png" pageId="5" pageNumber="369">Fig. 3D</figureCitation>
), 
<collectingCountry id="F3187606FFC1FFA989E607D67DB511DA" box="[1414,1469,1952,1975]" name="Laos" pageId="5" pageNumber="369">Laos</collectingCountry>
, 
<collectingRegion id="49CBF874FFC1FFA98F2307B67C2111B5" box="[835,1065,1984,2008]" country="Laos" name="Houaphan" pageId="5" pageNumber="369">Houaphanh Province</collectingRegion>
, 
<collectingMunicipality id="6BD4ACECFFC1FFA9885507B67CF111B5" box="[1077,1273,1984,2008]" pageId="5" pageNumber="369">Xiengkho District</collectingMunicipality>
, 
<location id="8ED0604DFFC1FFA9896507B67DB611B5" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA48F3E022A79A8110D:8ED0604DFFC1FFA9896507B67DB611B5" box="[1285,1470,1984,2008]" country="Laos" municipality="Xiengkho District" name="Na Tong Village" pageId="5" pageNumber="369" stateProvince="Houaphan">Na Tong Village</location>
, limestone hills, col. 
<collectorName id="26FA5340FFC1FFA9884107967CC21195" box="[1057,1226,2016,2040]" pageId="5" pageNumber="369">P.S.L. Ti Khun</collectorName>
, 
<date id="FFB11056FFC1FFA988B607967D801195" box="[1238,1416,2016,2040]" pageId="5" pageNumber="369" value="2018-08-20">
<collectingDate id="EFF5E9BEFFC1FFA988B607967D801195" box="[1238,1416,2016,2040]" pageId="5" pageNumber="369" value="2018-08-20">20 August 2018</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF70661EFFC2FFAA8C11087979C51E2C" ID-DOI="http://doi.org/10.5281/zenodo.7174172" ID-Zenodo-Dep="7174172" httpUri="https://zenodo.org/record/7174172/files/figure.png" pageId="6" pageNumber="370" startId="6.[113,148,2063,2085]" targetBox="[126,1440,188,2047]" targetPageId="6">
<paragraph id="8BB03696FFC2FFAA8C11087979C51E2C" blockId="6.[113,1453,2063,2113]" pageId="6" pageNumber="370">
Fig. 4. Shell (apertural view, dorsal view, ventral view, and side view) of 
<taxonomicName id="4C0F4D15FFC2FFAA8F2308667BE91E49" authority="Sang &amp; Son, 2022" authorityName="Sang &amp; Son" authorityYear="2022" box="[835,993,2063,2085]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="370" phylum="Mollusca" rank="species" species="clausa" status="sp. nov.">
<emphasis id="B97BEA84FFC2FFAA8F2308667BE91E49" box="[835,993,2063,2085]" italics="true" pageId="6" pageNumber="370">Pincerna clausa</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A24857FFFFC2FFAA8F8E08667C6D1E48" box="[1006,1125,2063,2085]" pageId="6" pageNumber="370" rank="species">new species</taxonomicNameLabel>
. A, holotype VNMN-IZ 002.304; B, C, paratypes ZMHU. MOL 044.
</paragraph>
</caption>
<caption id="DF70661EFFC3FFAB8CE808797ABD1E2F" ID-DOI="http://doi.org/10.5281/zenodo.7174174" ID-Zenodo-Dep="7174174" httpUri="https://zenodo.org/record/7174174/files/figure.png" pageId="7" pageNumber="371" startId="7.[136,171,2063,2085]" targetBox="[148,1462,188,2047]" targetPageId="7">
<paragraph id="8BB03696FFC3FFAB8CE808797ABD1E2F" blockId="7.[136,1474,2063,2114]" pageId="7" pageNumber="371">
Fig. 5. Details of the shell of 
<taxonomicName id="4C0F4D15FFC3FFAB8DC908667A4F1E49" authority="Sang &amp; Son, 2022" authorityName="Sang &amp; Son" authorityYear="2022" box="[425,583,2063,2085]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="7" pageNumber="371" phylum="Mollusca" rank="species" species="clausa" status="sp. nov.">
<emphasis id="B97BEA84FFC3FFAB8DC908667A4F1E49" box="[425,583,2063,2085]" italics="true" pageId="7" pageNumber="371">Pincerna clausa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24857FFFFC3FFAB8E2E08797A8B1E49" box="[590,643,2063,2084]" pageId="7" pageNumber="371" rank="species">n. sp.</taxonomicNameLabel>
(holotype VNMN-IZ 002.304). A–B, protoconch; C, R2 and sutural tube; D, R3 and boundary between R2 and R3; E, aperture; F, umbilicus.
</paragraph>
</caption>
<caption id="DF70661EFFCCFFA48C1104087B1A12DD" ID-DOI="http://doi.org/10.5281/zenodo.7174176" ID-Zenodo-Dep="7174176" httpUri="https://zenodo.org/record/7174176/files/figure.png" pageId="8" pageNumber="372" startId="8.[113,149,1150,1172]" targetBox="[113,1451,188,1135]" targetPageId="8">
<paragraph id="8BB03696FFCCFFA48C1104087B1A12DD" blockId="8.[113,1451,1150,1201]" pageId="8" pageNumber="372">
Fig. 6. Synoptic view of Vietnamese and Laos 
<taxonomicName id="4C0F4D15FFCCFFA48E3E04097B2612FE" box="[606,814,1150,1172]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="372" phylum="Mollusca" rank="species" species="undefined-a">
<emphasis id="B97BEA84FFCCFFA48E3E04097AB112F9" box="[606,697,1151,1172]" italics="true" pageId="8" pageNumber="372">Pincerna</emphasis>
species. A
</taxonomicName>
, MNHN-IM-2000-31797; B, NHMUK 20170120; C, MNHN- IM-2000-31786; D, VNMN-IZ 002.304; E, MNHN-IM-2000-31798.
</paragraph>
</caption>
<subSubSection id="C315651DFFCCFFA48C11049E7AF9134D" pageId="8" pageNumber="372" type="diagnosis">
<paragraph id="8BB03696FFCCFFA48C11049E7AF9134D" blockId="8.[113,753,1256,1312]" pageId="8" pageNumber="372">
<emphasis id="B97BEA84FFCCFFA48C11049E78EE136D" bold="true" box="[113,230,1256,1280]" pageId="8" pageNumber="372">Diagnosis.</emphasis>
<taxonomicName id="4C0F4D15FFCCFFA48C8D049F79C5136D" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[237,461,1256,1280]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="372" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="B97BEA84FFCCFFA48C8D049F79C5136D" box="[237,461,1256,1280]" italics="true" pageId="8" pageNumber="372">Pincerna vanbuensis</emphasis>
</taxonomicName>
is characterised by a small sized, globular shell with irregular, clearly visible ribs on R3.
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFCCFFA48C11053E7A4E13AD" pageId="8" pageNumber="372" type="distribution">
<paragraph id="8BB03696FFCCFFA48C11053E7A4E13AD" blockId="8.[113,753,1352,1472]" pageId="8" pageNumber="372">
<emphasis id="B97BEA84FFCCFFA48C11053E790F130D" bold="true" box="[113,263,1352,1376]" pageId="8" pageNumber="372">Distribution.</emphasis>
This species is known from 
<collectingCountry id="F3187606FFCCFFA48E34053E7AB9130D" box="[596,689,1352,1376]" name="Vietnam" pageId="8" pageNumber="372">Vietnam</collectingCountry>
(
<collectingRegion id="49CBF874FFCCFFA48EA3053F789913ED" country="Vietnam" name="Son La" pageId="8" pageNumber="372">SƠn La</collectingRegion>
, 
<collectingRegion id="49CBF874FFCCFFA48CFB051E790C13ED" box="[155,260,1384,1408]" country="Vietnam" name="Hoa Binh" pageId="8" pageNumber="372">Hòa Bình</collectingRegion>
, 
<collectingRegion id="49CBF874FFCCFFA48D70051E799D13ED" box="[272,405,1384,1408]" country="Vietnam" name="Quang Ninh" pageId="8" pageNumber="372">Quang Ninh</collectingRegion>
, 
<collectingRegion id="49CBF874FFCCFFA48DC0051E7A1813ED" box="[416,528,1384,1408]" country="Vietnam" name="Ninh Binh" pageId="8" pageNumber="372">Ninh Binh</collectingRegion>
Provinces) and 
<collectingCountry id="F3187606FFCCFFA48EDD051F7AF913ED" box="[701,753,1385,1408]" name="Laos" pageId="8" pageNumber="372">Laos</collectingCountry>
(
<collectingRegion id="49CBF874FFCCFFA48C1905FE791F13CD" box="[121,279,1416,1440]" country="Laos" name="Louangnamtha" pageId="8" pageNumber="372">Luang Namtha</collectingRegion>
and Luang Phrabang Provinces) (
<bibRefCitation id="EF9E4B67FFCCFFA48E1905FE793C13AD" author="Vermeulen JJ &amp; Maassen WJM" pageId="8" pageNumber="372" refId="ref7660" refString="Vermeulen JJ &amp; Maassen WJM (2003) The non-marine mollusk fauna of the Pu Luong, Cuc Phuong, Phu Ly, and Ha Long regions in northern Vietnam. A survey for the Vietnam Programme of FFI (Flora and Fauna International), 35 pp." type="book" year="2003">Vermeulen &amp; Maassen, 2003</bibRefCitation>
; 
<bibRefCitation id="EF9E4B67FFCCFFA48D2105DE7A3313AD" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" box="[321,571,1448,1472]" pageId="8" pageNumber="372" pagination="1 - 166" refId="ref6528" refString="Inkhavilay K, Sutcharit C, Bantaowong U, Chanabun R, Siriwut W, Srisonchai R, Pholyotha A, Jirapatrasilp P &amp; Panha S (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys, 834: 1 - 166." type="journal article" year="2019">Inkhavilay et al., 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFCCFFA48C11059E79A8110D" pageId="8" pageNumber="372" type="discussion">
<paragraph id="8BB03696FFCCFFA48C11059E79A8110D" blockId="8.[113,753,1512,1888]" pageId="8" pageNumber="372">
<emphasis id="B97BEA84FFCCFFA48C11059E78EA106D" bold="true" box="[113,226,1512,1536]" pageId="8" pageNumber="372">Remarks.</emphasis>
Its 
<typeStatus id="54B48834FFCCFFA48D73059F794A106C" box="[275,322,1513,1537]" pageId="8" pageNumber="372">type</typeStatus>
locality Van-bu most probably refers to “Vạn Bú” Province, which was established in 1895 and remained until 1904. This species clearly differs from 
<taxonomicName id="4C0F4D15FFCCFFA48EB9065F78BF100D" baseAuthorityName="Mabille" baseAuthorityYear="1887" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="372" phylum="Mollusca" rank="species" species="anceyi">
<emphasis id="B97BEA84FFCCFFA48EB9065F78BF100D" italics="true" pageId="8" pageNumber="372">P. anceyi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0F4D15FFCCFFA48C92063F7970100D" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[242,376,1608,1632]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="372" phylum="Mollusca" rank="species" species="costulosa">
<emphasis id="B97BEA84FFCCFFA48C92063F7970100D" box="[242,376,1608,1632]" italics="true" pageId="8" pageNumber="372">P. costulosa</emphasis>
</taxonomicName>
because its R2 is relatively long, while its R1 possesses clearly visible and regular spaced ribs. The operculum of 
<specimenCount id="9D09FD1FFFCCFFA48D0606FE7A0710CD" box="[358,527,1672,1696]" pageId="8" pageNumber="372" type="generic">three specimens</specimenCount>
have been examined. The inner surface of operculum is smooth, glossy and has no central nipple. The outer surface is relatively flat, with indistinct appressed multilamellae. 
<taxonomicName id="4C0F4D15FFCCFFA48D93069F7ADD116D" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[499,725,1768,1792]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="372" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="B97BEA84FFCCFFA48D93069F7ADD116D" box="[499,725,1768,1792]" italics="true" pageId="8" pageNumber="372">Pincerna vanbuensis</emphasis>
</taxonomicName>
is a very variable species in terms of shell size, sculpture of R1 and R3, number of R2 ribs and the extension of the columella (see 
<tableCitation id="C68D032DFFCCFFA48D77073E797C110D" box="[279,372,1864,1888]" captionStart="Table 1" captionStartId="9.[136,191,189,211]" captionText="Table 1. Shell size variation of Pincerna spp. specimen collections in this study" pageId="8" pageNumber="372">Tables 1</tableCitation>
, 
<tableCitation id="C68D032DFFCCFFA48DE4073E799C110D" box="[388,404,1864,1888]" captionStart="Table 2" captionStartId="9.[136,191,1135,1157]" captionText="Table 2. Comparison of diagnostic conchological characters among Pincerna species in this study." pageId="8" pageNumber="372">2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>