357 lines
54 KiB
XML
357 lines
54 KiB
XML
<document id="20C68EE9C3FCB0D963326EAB1C9BF639" ID-CLB-Dataset="27324" ID-DOI="10.11646/zootaxa.4585.2.5" ID-GBIF-Dataset="086cbbca-58b1-4012-a9f4-3bfdb5ffbe93" ID-ISSN="1175-5326" ID-Zenodo-Dep="2637366" ID-ZooBank="6E61D5EE-E3AD-44EC-9AB3-C7B3E303BF92" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555050046501" checkinUser="plazi" docAuthor="Husana, Daniel Edison M. & Ng, Peter K. L." docDate="2019" docId="C6560C5A6B23FF87E7FEFBB1FCB8F874" docLanguage="en" docName="zootaxa.4585.2.5.pdf" docOrigin="Zootaxa 4585 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sundathelphusa quirino Husana & Ng 2019, sp. nov." docType="treatment" docUuid="AACEE47B-0E19-4AE6-8D82-11C1C48B861E" docUuidSource="ZooBank" docVersion="10" lastPageNumber="329" masterDocId="3A6F74226B28FF89E769FFADFF8EFF9E" masterDocTitle="On the identity of Sundathelphusa philippina (von Martens, 1868) (Decapoda: Brachyura: Gecarcinucidae) from the Philippines, with descriptions of two new species" masterLastPageNumber="331" masterPageNumber="315" pageNumber="326" updateTime="1698719825494" updateUser="ExternalLinkService">
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<mods:mods id="3DB0259F446A325578B9BFDFD1E3046B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C0A3002E7924C2330388F6B4726B6155">
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<mods:title id="15310DC9995FE782241647104FEF50CF">On the identity of Sundathelphusa philippina (von Martens, 1868) (Decapoda: Brachyura: Gecarcinucidae) from the Philippines, with descriptions of two new species</mods:title>
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</mods:titleInfo>
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<mods:name id="E0A6F02BD84321E6D87058125ED63ADE" type="personal">
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<mods:roleTerm id="E4FF30AC7FBE87E3C8C53A78B27A102D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AA14A61AFD071FE633269D5AD340C616">Husana, Daniel Edison M.</mods:namePart>
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</mods:name>
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<mods:name id="448462CE40D79EAD1B2603F0D7A649EE" type="personal">
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<mods:role id="900C58FE9EDC9E3D9005F614C3C78150">
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<mods:roleTerm id="4EB1CD1D8CF8365654FED420B4B8E3AB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7CBC29C7853A608CBFE5685419084A1A">Ng, Peter K. L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="BDA340A012147B0C30C418B41AC95B1A">text</mods:typeOfResource>
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<mods:titleInfo id="F9058DC27320181AE06C4BE77DB0502F">
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<mods:title id="2DCA13F4036A1BA1EB4DF829218A1D70">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="2EF24A20499B7080110EAE33F25C8EF4">
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<mods:date id="38BF057B3D6C5D815FA922E6691DAA52">2019</mods:date>
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<mods:detail id="A7CAE83DDD2955F864C00C2AB07A7C21" type="pubDate">
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<mods:number id="6883C4D59AD3CDF0C9A1A161E0A370E5">2019-04-12</mods:number>
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</mods:detail>
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<mods:detail id="F9F9621A3E4CE79957A7E16794781D7F" type="volume">
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<mods:number id="D1D38926D591B874E7832ED9C6B16A9D">4585</mods:number>
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</mods:detail>
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<mods:detail id="02B8F7AFB12380E4C53369AF5ED53705" type="issue">
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<mods:number id="417C8C0E38351513C1AECB6437266CCB">2</mods:number>
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<mods:start id="25F163DAD92A710D9F89AE1D04DE73C5">315</mods:start>
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<mods:end id="ADE31DDB37B5FCEEC387F4E6EE41E18C">331</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="32438C186E90C6A6591E67FBA1344951">journal article</mods:classification>
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<mods:identifier id="94B929083BD908013DEBE41DC12A0E61" type="CLB-Dataset">27324</mods:identifier>
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<mods:identifier id="FF957D8F07A90CE99A7AA84D2E60DF55" type="DOI">10.11646/zootaxa.4585.2.5</mods:identifier>
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<mods:identifier id="DF1DDD4B889ACD37421A94269219C6A8" type="GBIF-Dataset">086cbbca-58b1-4012-a9f4-3bfdb5ffbe93</mods:identifier>
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<mods:identifier id="3207F69F38DA8879EDB853D3199CF7D0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6AB37FD7E68650CE3C11B7225C3EA71F" type="Zenodo-Dep">2637366</mods:identifier>
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<mods:identifier id="E60F550187392657A0BEDF86DA2AF30B" type="ZooBank">6E61D5EE-E3AD-44EC-9AB3-C7B3E303BF92</mods:identifier>
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</mods:mods>
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<treatment id="C6560C5A6B23FF87E7FEFBB1FCB8F874" ID-DOI="http://doi.org/10.5281/zenodo.5466963" ID-GBIF-Taxon="156192032" ID-Zenodo-Dep="5466963" LSID="urn:lsid:zoobank.org:act:AACEE47B-0E19-4AE6-8D82-11C1C48B861E" httpUri="http://treatment.plazi.org/id/C6560C5A6B23FF87E7FEFBB1FCB8F874" lastPageId="14" lastPageNumber="329" pageId="11" pageNumber="326">
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<subSubSection id="06E5EEC76B23FF82E7FEFBB1FDAFFBA8" box="[151,545,1052,1078]" pageId="11" pageNumber="326" type="nomenclature">
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<paragraph id="4E40BD4C6B23FF82E7FEFBB1FDAFFBA8" blockId="11.[151,545,1052,1111]" box="[151,545,1052,1078]" pageId="11" pageNumber="326">
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<heading id="15080A206B23FF82E7FEFBB1FDAFFBA8" bold="true" box="[151,545,1052,1078]" fontSize="11" level="1" pageId="11" pageNumber="326" reason="1">
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||
<emphasis id="7C8B615E6B23FF82E7FEFBB1FDAFFBA8" bold="true" box="[151,545,1052,1078]" pageId="11" pageNumber="326">
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||
<taxonomicName id="89FFC6CF6B23FF82E7FEFBB1FE33FBA8" ID-CoL="84GBN" authority="Husana & Ng, 2019" authorityName="Husana & Ng" authorityYear="2019" box="[151,445,1052,1078]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="quirino" status="sp. nov.">
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<emphasis id="7C8B615E6B23FF82E7FEFBB1FE33FBA8" bold="true" box="[151,445,1052,1078]" italics="true" pageId="11" pageNumber="326">Sundathelphusa quirino</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="67B8DC256B23FF82E6ADFBB1FDAFFBA8" box="[452,545,1052,1078]" pageId="11" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="06E5EEC76B23FF82E7FEFB93FED3FBC9" box="[151,349,1086,1111]" pageId="11" pageNumber="326" type="description">
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<paragraph id="4E40BD4C6B23FF82E7FEFB93FED3FBC9" blockId="11.[151,545,1052,1111]" box="[151,349,1086,1111]" pageId="11" pageNumber="326">
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||
(
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<figureCitation id="D6C4A1C96B23FF82E7F6FB93FE85FBC9" box="[159,267,1086,1111]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="12.[151,250,1881,1903]" captionStartId-1="13.[151,250,1892,1914]" captionTargetBox-0="[150,1436,697,1859]" captionTargetBox-1="[174,1378,213,1867]" captionTargetId-0="figure@12.[150,1436,697,1859]" captionTargetId-1="figure@13.[169,1384,193,1873]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." captionText-1="FIGURE 8. Sundathelphusa quirino sp. nov. A–H, holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino; I–M, paratype ♂ (47.6 × 36.9 mm), ZRC 2017.1063, Quirino. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G, I, J = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri-0="https://zenodo.org/record/2637380/files/figure.png" httpUri-1="https://zenodo.org/record/2637382/files/figure.png" pageId="11" pageNumber="326">Figs. 7–8</figureCitation>
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;
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<figureCitation id="D6C4A1C96B23FF82E67EFB92FEDBFBC9" box="[279,341,1087,1111]" captionStart="FIGURE 9" captionStartId="14.[151,250,1020,1042]" captionTargetBox="[161,1428,171,999]" captionTargetId="figure@14.[161,1428,171,999]" captionTargetPageId="14" captionText="FIGURE 9. Right third and fourth ambulatory legs. A, B, Sundathelphusa philippina (von Martens, 1868), ♂ (51.3 × 40.4 mm), ZRC 2009.0409, Luzon; C, D, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; E, F, S. quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, C, E, third ambulatory leg; B, D, F, fourth ambulatory leg. Scales: A–F = 5.0 mm." httpUri="https://zenodo.org/record/2637384/files/figure.png" pageId="11" pageNumber="326">9E, F</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="06E5EEC76B23FF82E7FEFB2BFA8FFACD" pageId="11" pageNumber="326" type="materials_examined">
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||
<paragraph id="4E40BD4C6B23FF82E7FEFB2BFA8FFACD" blockId="11.[151,1437,1158,1975]" pageId="11" pageNumber="326">
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||
<emphasis id="7C8B615E6B23FF82E7FEFB2BFE0EFB01" bold="true" box="[151,384,1158,1183]" pageId="11" pageNumber="326">Material examined</emphasis>
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.
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<materialsCitation id="FE97B7116B23FF82E6FDFB2BFBD7FB5D" ID-GBIF-Occurrence="2235592135" collectingDate="2009-04-17" collectorName="D. E. Husana" county="Luzon Island" location="Aglipay Cave" municipality="Quirino province" pageId="11" pageNumber="326" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<typeStatus id="914403EE6B23FF82E6FDFB2BFD8FFB01" box="[404,513,1158,1183]" pageId="11" pageNumber="326" type="holotype">Holotype</typeStatus>
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||
:
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||
<specimenCount id="58F976C56B23FF82E578FB2AFDA8FB00" box="[529,550,1159,1182]" pageId="11" pageNumber="326" type="male">♂</specimenCount>
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||
(49.3 ×
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||
<quantity id="890710A96B23FF82E5F0FB2BFC89FB01" box="[665,775,1158,1183]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="11" pageNumber="326" unit="mm" value="37.5">37.5 mm</quantity>
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||
), NMCR 50701,
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||
<location id="4B20EB976B23FF82E4B2FB2BFBF1FB01" LSID="urn:lsid:plazi:treatment:C6560C5A6B23FF87E7FEFBB1FCB8F874:4B20EB976B23FF82E4B2FB2BFBF1FB01" box="[987,1151,1158,1183]" county="Luzon Island" municipality="Quirino province" name="Aglipay Cave" pageId="11" pageNumber="326">Aglipay Cave</location>
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||
,
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||
<location id="4B20EB976B23FF82E3F8FB2BFABAFB00" LSID="urn:lsid:plazi:treatment:C6560C5A6B23FF87E7FEFBB1FCB8F874:4B20EB976B23FF82E3F8FB2BFABAFB00" box="[1169,1332,1158,1183]" county="Luzon Island" municipality="Quirino province" name="Aglipay town" pageId="11" pageNumber="326">Aglipay town</location>
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||
,
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||
<collectingMunicipality id="AE2427366B23FF82E22CFB2BFF72FB5D" pageId="11" pageNumber="326">Quirino province</collectingMunicipality>
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||
,
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||
<collectingCounty id="A721C5C06B23FF82E66EFB06FE13FB5D" box="[263,413,1194,1219]" pageId="11" pageNumber="326">Luzon Island</collectingCounty>
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||
, 16˚28.439'N 121˚36.577'E, coll.
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||
<collectorName id="E30AD89A6B23FF82E473FB06FC22FB5C" box="[794,940,1195,1218]" pageId="11" pageNumber="326">D.E. Husana</collectorName>
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||
,
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||
<date id="3A419B8C6B23FF82E4DEFB06FBD7FB5D" box="[951,1113,1194,1219]" pageId="11" pageNumber="326" value="2009-04-17">
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||
<collectingDate id="2A0562646B23FF82E4DEFB06FBD7FB5D" box="[951,1113,1194,1219]" pageId="11" pageNumber="326" value="2009-04-17">17 April 2009</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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||
<typeStatus id="914403EE6B23FF82E30DFB06FB58FB5D" box="[1124,1238,1195,1219]" pageId="11" pageNumber="326" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="58F976C56B23FF82E389FB06FA84FB5C" box="[1248,1290,1195,1219]" pageId="11" pageNumber="326" type="male">1 ♂</specimenCount>
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||
(44.0 ×
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||
<quantity id="890710A96B23FF82E202FB06FF49FB79" metricMagnitude="-2" metricUnit="m" metricValue="3.42" pageId="11" pageNumber="326" unit="mm" value="34.2">34.2 mm</quantity>
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||
),
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||
<materialsCitation id="FE97B7116B23FF82E7B2FB62FB04FB79" ID-GBIF-Occurrence="2235592132" box="[219,1162,1230,1255]" collectingDate="2009-04-17" collectionCode="ZRC" collectorName="D. E. Husana" county="Luzon Island" location="Aglipay Cave" municipality="Quirino province" pageId="11" pageNumber="326" specimenCode="ZRC 2017.1063" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
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||
<specimenCount id="58F976C56B23FF82E7B2FB62FE92FB78" box="[219,284,1231,1255]" pageId="11" pageNumber="326" type="female">2 ♀♀</specimenCount>
|
||
(45.3 ×
|
||
<quantity id="890710A96B23FF82E6E9FB63FE66FB78" box="[384,488,1230,1255]" metricMagnitude="-2" metricUnit="m" metricValue="3.54" pageId="11" pageNumber="326" unit="mm" value="35.4">35.4 mm</quantity>
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||
, 52.8 × 41.0 mm),
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<specimenCode id="1E5915376B23FF82E5ACFB62FCF0FB79" box="[709,894,1230,1255]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="326">ZRC 2017.1063</specimenCode>
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||
, same data as holotype
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="FE97B7116B23FF82E3FEFB62FA0CFA95" ID-GBIF-Occurrence="2235592105" collectingDate="1999-02-03" collectionCode="ZRC" collectorName="L. Liao" county="Luzon" location="Diduyan River" municipality="Dingasan" pageId="11" pageNumber="326" specimenCode="ZRC 2017.1257" specimenCount="1" specimenCount-female="1" stateProvince="Quirino" typeStatus="paratype">
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||
<specimenCount id="58F976C56B23FF82E3FEFB62FB4DFB78" box="[1175,1219,1231,1255]" pageId="11" pageNumber="326" type="female">1 ♀</specimenCount>
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||
(27.4 × 23.0 mm),
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||
<specimenCode id="1E5915376B23FF82E7FEFB5EFEC0FA95" box="[151,334,1266,1291]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="326">ZRC 2017.1257</specimenCode>
|
||
,
|
||
<location id="4B20EB976B23FF82E630FB5FFD8FFA95" LSID="urn:lsid:plazi:treatment:C6560C5A6B23FF87E7FEFBB1FCB8F874:4B20EB976B23FF82E630FB5FFD8FFA95" box="[345,513,1266,1291]" county="Luzon" municipality="Dingasan" name="Diduyan River" pageId="11" pageNumber="326" stateProvince="Quirino">Diduyan River</location>
|
||
,
|
||
<location id="4B20EB976B23FF82E565FB5FFD0BFA95" LSID="urn:lsid:plazi:treatment:C6560C5A6B23FF87E7FEFBB1FCB8F874:4B20EB976B23FF82E565FB5FFD0BFA95" box="[524,645,1266,1291]" county="Luzon" municipality="Dingasan" name="Sitio Tubo" pageId="11" pageNumber="326" stateProvince="Quirino">Sitio Tubo</location>
|
||
,
|
||
<collectingMunicipality id="AE2427366B23FF82E5F9FB5FFD73FA95" box="[656,765,1266,1291]" pageId="11" pageNumber="326">Dingasan</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="8C3B73AE6B23FF82E461FB5FFC43FA95" box="[776,973,1266,1291]" country="Philippines" name="Quirino" pageId="11" pageNumber="326">Quirino Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="A721C5C06B23FF82E4B1FB5EFBACFA94" box="[984,1058,1267,1290]" pageId="11" pageNumber="326">Luzon</collectingCounty>
|
||
, coll.
|
||
<collectorName id="E30AD89A6B23FF82E30FFB5EFB35FA95" box="[1126,1211,1266,1291]" pageId="11" pageNumber="326">L. Liao</collectorName>
|
||
,
|
||
<date id="3A419B8C6B23FF82E3AFFB5EFA0CFA95" box="[1222,1410,1266,1291]" pageId="11" pageNumber="326" value="1999-02-03">
|
||
<collectingDate id="2A0562646B23FF82E3AFFB5EFA0CFA95" box="[1222,1410,1266,1291]" pageId="11" pageNumber="326" value="1999-02-03">3 February 1999</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="58F976C56B23FF82E2E7FB5EFF22FAB0" pageId="11" pageNumber="326" type="male">1 ♂</specimenCount>
|
||
(47.6 ×
|
||
<quantity id="890710A96B23FF82E665FABBFEFBFAB1" box="[268,373,1302,1327]" metricMagnitude="-2" metricUnit="m" metricValue="3.69" pageId="11" pageNumber="326" unit="mm" value="36.9">36.9 mm</quantity>
|
||
),
|
||
<specimenCount id="58F976C56B23FF82E6E0FABAFE3DFAB0" box="[393,435,1303,1327]" pageId="11" pageNumber="326" type="male">1 ♂</specimenCount>
|
||
(33.1 ×
|
||
<quantity id="890710A96B23FF82E57DFABBFDF7FAB1" box="[532,633,1302,1327]" metricMagnitude="-2" metricUnit="m" metricValue="2.61" pageId="11" pageNumber="326" unit="mm" value="26.1">26.1 mm</quantity>
|
||
) (only carapace intact),
|
||
<materialsCitation id="FE97B7116B23FF82E4E2FABAFB73FACD" ID-GBIF-Occurrence="2235592114" collectingDate="1979-07" collectionCode="RMNH" collectorName="F. Alanday & All" country="Philippines" county="Luzon" location="Disiluat Cave" municipality="San Leonardo" pageId="11" pageNumber="326" specimenCode="RMNH 32925" specimenCount="1" specimenCount-female="1" stateProvince="Quirino" typeStatus="paratype">
|
||
<specimenCount id="58F976C56B23FF82E4E2FABAFC38FAB0" box="[907,950,1303,1327]" pageId="11" pageNumber="326" type="female">1 ♀</specimenCount>
|
||
(crushed),
|
||
<specimenCode id="1E5915376B23FF82E35FFABAFB54FAB1" box="[1078,1242,1302,1327]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="326" type="Museum">RMNH 32925</specimenCode>
|
||
,
|
||
<location id="4B20EB976B23FF82E38FFABBFA0CFAB1" LSID="urn:lsid:plazi:treatment:C6560C5A6B23FF87E7FEFBB1FCB8F874:4B20EB976B23FF82E38FFABBFA0CFAB1" box="[1254,1410,1302,1327]" country="Philippines" county="Luzon" municipality="San Leonardo" name="Disiluat Cave" pageId="11" pageNumber="326" stateProvince="Quirino">Disiluat Cave</location>
|
||
,
|
||
<quantity id="890710A96B23FF82E2E7FABAFF35FACD" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="326" unit="km" value="8.0">8 km</quantity>
|
||
from
|
||
<collectingMunicipality id="AE2427366B23FF82E796FA96FE10FACD" box="[255,414,1338,1363]" pageId="11" pageNumber="326">San Leonardo</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="8C3B73AE6B23FF82E6C0FA97FD8DFACD" box="[425,515,1338,1363]" country="Philippines" name="Quirino" pageId="11" pageNumber="326">Quirino</collectingRegion>
|
||
,
|
||
<collectingCounty id="A721C5C06B23FF82E567FA96FDD6FACC" box="[526,600,1339,1362]" pageId="11" pageNumber="326">Luzon</collectingCounty>
|
||
, coll.
|
||
<collectorName id="E30AD89A6B23FF82E5F5FA96FC99FACD" box="[668,791,1338,1363]" pageId="11" pageNumber="326">F. Alanday</collectorName>
|
||
,
|
||
<date id="3A419B8C6B23FF82E44BFA97FC1DFACD" box="[802,915,1338,1363]" pageId="11" pageNumber="326" value="1979-07">
|
||
<collectingDate id="2A0562646B23FF82E44BFA97FC1DFACD" box="[802,915,1338,1363]" pageId="11" pageNumber="326" value="1979-07">July 1979</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="E30AD89A6B23FF82E4F7FA97FC4CFACD" box="[926,962,1338,1363]" pageId="11" pageNumber="326">All</collectorName>
|
||
locations in the
|
||
<collectingCountry id="36E8FDDC6B23FF82E312FA97FB73FACD" box="[1147,1277,1338,1363]" name="Philippines" pageId="11" pageNumber="326">Philippines</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06E5EEC76B23FF87E7AEFAF3FBDCFA50" lastPageId="14" lastPageNumber="329" pageId="11" pageNumber="326" type="description">
|
||
<paragraph id="4E40BD4C6B23FF82E7AEFAF3FF47F829" blockId="11.[151,1437,1158,1975]" pageId="11" pageNumber="326">
|
||
<emphasis id="7C8B615E6B23FF82E7AEFAF3FEDFFAE9" bold="true" box="[199,337,1374,1399]" pageId="11" pageNumber="326">Description</emphasis>
|
||
. Carapace trapezoidal in shape, widest at anterior quarter, dorsal surface distinctly convex longitudinally, dorsoventrally inflated, regions distinct (
|
||
<figureCitation id="D6C4A1C96B23FF82E42BFA2FFC27FA05" box="[834,937,1410,1435]" captionStart="FIGURE 7" captionStartId="12.[151,250,1881,1903]" captionTargetBox="[150,1436,697,1859]" captionTargetId="figure@12.[150,1436,697,1859]" captionTargetPageId="12" captionText="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="11" pageNumber="326">Fig. 7A</figureCitation>
|
||
). Frontal region sloping anteroventrally; anterolateral regions inflated dorsolaterally, smooth; cervical grooves deep; H-shaped gastric groove deep; epigastric cristae, distinct, edges rough, separated by distinct median furrow; postorbital cristae, sharp but low; epigastric and postorbital cristae confluent; epibranchial teeth and postorbital cristae not confluent, separated by gaps (
|
||
<figureCitation id="D6C4A1C96B23FF82E7B6F9BFFEB1F9B5" box="[223,319,1554,1579]" captionStart="FIGURE 7" captionStartId="12.[151,250,1881,1903]" captionTargetBox="[150,1436,697,1859]" captionTargetId="figure@12.[150,1436,697,1859]" captionTargetPageId="12" captionText="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="11" pageNumber="326">Fig. 7A</figureCitation>
|
||
). Frontal margin broadly protruded, two lobes clearly separated with broad median concavity; external orbital tooth produced anteriorly, outer margin longer than inner margin; epibranchial tooth distinct, triangular, well separated from external orbital tooth, tapering anteriorly, surface and margin flat; anterolateral margin convex, with very low crest, not distinct from lateral view, not clearly demarcated from posterolateral margin; posterolateral margin gently concave, converging gradually towards posterior margin of carapace (
|
||
<figureCitation id="D6C4A1C96B23FF82E206F90FFF32F941" captionStart="FIGURE 7" captionStartId="12.[151,250,1881,1903]" captionTargetBox="[150,1436,697,1859]" captionTargetId="figure@12.[150,1436,697,1859]" captionTargetPageId="12" captionText="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="11" pageNumber="326">Fig. 7A</figureCitation>
|
||
). Frontal median triangle complete; dorsal and lateral margins distinct, smooth, confluent; dorsal margin more produced anteriorly than lateral margins (
|
||
<figureCitation id="D6C4A1C96B23FF82E500F947FD31F89D" box="[617,703,1770,1795]" captionStart="FIGURE 8" captionStartId="13.[151,250,1892,1914]" captionTargetBox="[174,1378,213,1867]" captionTargetId="figure@13.[169,1384,193,1873]" captionTargetPageId="13" captionText="FIGURE 8. Sundathelphusa quirino sp. nov. A–H, holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino; I–M, paratype ♂ (47.6 × 36.9 mm), ZRC 2017.1063, Quirino. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G, I, J = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637382/files/figure.png" pageId="11" pageNumber="326">Fig. 8F</figureCitation>
|
||
); orbit well demarcated; supraorbital margin smooth; infraorbital margin, smooth, sinuous, protruded anteriorly; outer edge reaching and fused with anterolateral margin; suborbital and subbranchial regions covered with scattered oblique long, short striae; pterygostomial region smooth with oblique ridges on upper outer part (
|
||
<figureCitation id="D6C4A1C96B23FF82E55DF8FBFD01F8F1" box="[564,655,1878,1903]" captionStart="FIGURE 7" captionStartId="12.[151,250,1881,1903]" captionTargetBox="[150,1436,697,1859]" captionTargetId="figure@12.[150,1436,697,1859]" captionTargetPageId="12" captionText="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="11" pageNumber="326">Fig. 7C</figureCitation>
|
||
). Posterior margin of epistome with three lobes, median lobe large, subtriangular, notch absent; lateral lobes wider and protruded, placed more anteriorly than median lobe (
|
||
<figureCitation id="D6C4A1C96B23FF82E25AF8D7FA19F80D" box="[1331,1431,1914,1939]" captionStart="FIGURE 7" captionStartId="12.[151,250,1881,1903]" captionTargetBox="[150,1436,697,1859]" captionTargetId="figure@12.[150,1436,697,1859]" captionTargetPageId="12" captionText="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="11" pageNumber="326">Figs. 7C</figureCitation>
|
||
;
|
||
<figureCitation id="D6C4A1C96B23FF82E7FEF832FF32F829" box="[151,188,1951,1975]" captionStart="FIGURE 8" captionStartId="13.[151,250,1892,1914]" captionTargetBox="[174,1378,213,1867]" captionTargetId="figure@13.[169,1384,193,1873]" captionTargetPageId="13" captionText="FIGURE 8. Sundathelphusa quirino sp. nov. A–H, holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino; I–M, paratype ♂ (47.6 × 36.9 mm), ZRC 2017.1063, Quirino. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G, I, J = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637382/files/figure.png" pageId="11" pageNumber="326">8G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4E40BD4C6B24FF85E7AEFF35FE45FE82" blockId="12.[151,1437,151,645]" pageId="12" pageNumber="327">
|
||
Eyes well developed, occupying entire orbit (
|
||
<figureCitation id="D6C4A1C96B24FF85E5B1FF3AFCD4FF2E" box="[728,858,151,176]" captionStart="FIGURE 7" captionStartId="12.[151,250,1881,1903]" captionTargetBox="[150,1436,697,1859]" captionTargetId="figure@12.[150,1436,697,1859]" captionTargetPageId="12" captionText="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="12" pageNumber="327">Fig. 7A, C</figureCitation>
|
||
). Ischium of third maxilliped rectangular, bearing distinct submedian sulcus closer to mesial margin; merus quadrate, anteroexternal angle convex, anterior margin slightly concave; tip of exopod reaches midpoint of outer margin of merus, with long flagellum reaching beyond mesial margin (
|
||
<figureCitation id="D6C4A1C96B24FF85E62CFEA9FE30FE82" box="[325,446,260,285]" captionStart="FIGURE 7" captionStartId="12.[151,250,1881,1903]" captionTargetBox="[150,1436,697,1859]" captionTargetId="figure@12.[150,1436,697,1859]" captionTargetPageId="12" captionText="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="12" pageNumber="327">Fig. 7B, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4E40BD4C6B24FF85E7AEFE8AFB7CFE32" blockId="12.[151,1437,151,645]" pageId="12" pageNumber="327">
|
||
Adult chelipeds relatively stout, subequal, stronger in males; dorsal margin of merus serrated, dorsal margin with distinct subdistal tooth; carpus with strong distal inner angle, flattened dorsoventrally, laterally fringed with proximal teeth; palm with minutely granulated outer surface; ventral margin granulated; fingers robust, cutting edges with teeth of various sizes, largest medially, smaller on distal and proximal parts (
|
||
<figureCitation id="D6C4A1C96B24FF85E318FE39FB6BFE32" box="[1137,1253,404,429]" captionStart="FIGURE 7" captionStartId="12.[151,250,1881,1903]" captionTargetBox="[150,1436,697,1859]" captionTargetId="figure@12.[150,1436,697,1859]" captionTargetPageId="12" captionText="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="12" pageNumber="327">Fig. 7E, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4E40BD4C6B24FF85E7AEFE1AFDB8FD1A" blockId="12.[151,1437,151,645]" pageId="12" pageNumber="327">
|
||
Ambulatory legs relatively longer (
|
||
<figureCitation id="D6C4A1C96B24FF85E534FE1AFD4AFE4E" box="[605,708,439,464]" captionStart="FIGURE 7" captionStartId="12.[151,250,1881,1903]" captionTargetBox="[150,1436,697,1859]" captionTargetId="figure@12.[150,1436,697,1859]" captionTargetPageId="12" captionText="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="12" pageNumber="327">Figs. 7A</figureCitation>
|
||
;
|
||
<figureCitation id="D6C4A1C96B24FF85E5BAFE15FC9BFE4E" box="[723,789,440,464]" captionStart="FIGURE 9" captionStartId="14.[151,250,1020,1042]" captionTargetBox="[161,1428,171,999]" captionTargetId="figure@14.[161,1428,171,999]" captionTargetPageId="14" captionText="FIGURE 9. Right third and fourth ambulatory legs. A, B, Sundathelphusa philippina (von Martens, 1868), ♂ (51.3 × 40.4 mm), ZRC 2009.0409, Luzon; C, D, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; E, F, S. quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, C, E, third ambulatory leg; B, D, F, fourth ambulatory leg. Scales: A–F = 5.0 mm." httpUri="https://zenodo.org/record/2637384/files/figure.png" pageId="12" pageNumber="327">9E, F</figureCitation>
|
||
), second leg longest; anterior margin of merus serrated, without subdistal tooth or spine, posterior margins smooth on all legs; carpus short, with longitudinal submedian ridge on dorsal and ventral surfaces of all legs except on fourth leg that lacks ventral ridge, with barely visible dorsal ridge, widened distally, outer margins smooth; propodus with rows of spines on inner margins, outer margins smooth; dactylus with rows of short spines on all margins, spines of both outer and inner margins of dactylus almost equal in length (
|
||
<figureCitation id="D6C4A1C96B24FF85E6F7FDC1FE72FD1A" box="[414,508,620,645]" captionStart="FIGURE 7" captionStartId="12.[151,250,1881,1903]" captionTargetBox="[150,1436,697,1859]" captionTargetId="figure@12.[150,1436,697,1859]" captionTargetPageId="12" captionText="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="12" pageNumber="327">Figs 7A</figureCitation>
|
||
;
|
||
<figureCitation id="D6C4A1C96B24FF85E56EFDC1FDA6FD1A" box="[519,552,620,644]" captionStart="FIGURE 9" captionStartId="14.[151,250,1020,1042]" captionTargetBox="[161,1428,171,999]" captionTargetId="figure@14.[161,1428,171,999]" captionTargetPageId="14" captionText="FIGURE 9. Right third and fourth ambulatory legs. A, B, Sundathelphusa philippina (von Martens, 1868), ♂ (51.3 × 40.4 mm), ZRC 2009.0409, Luzon; C, D, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; E, F, S. quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, C, E, third ambulatory leg; B, D, F, fourth ambulatory leg. Scales: A–F = 5.0 mm." httpUri="https://zenodo.org/record/2637384/files/figure.png" pageId="12" pageNumber="327">9F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="1A80EDC46B24FF85E7FEF8F4FF75F832" httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="12" pageNumber="327" startId="12.[151,250,1881,1903]" targetBox="[150,1436,697,1859]" targetPageId="12">
|
||
<paragraph id="4E40BD4C6B24FF85E7FEF8F4FF75F832" blockId="12.[151,1436,1881,1964]" pageId="12" pageNumber="327">
|
||
<emphasis id="7C8B615E6B24FF85E7FEF8F4FE9CF8F0" bold="true" box="[151,274,1881,1903]" pageId="12" pageNumber="327">FIGURE 7.</emphasis>
|
||
<taxonomicName id="89FFC6CF6B24FF85E671F8F4FD85F8F1" authority="Husana & Ng, 2019" authorityName="Husana & Ng" authorityYear="2019" box="[280,523,1881,1903]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="327" phylum="Arthropoda" rank="species" species="quirino" status="sp. nov.">
|
||
<emphasis id="7C8B615E6B24FF85E671F8F4FD85F8F1" box="[280,523,1881,1903]" italics="true" pageId="12" pageNumber="327">Sundathelphusa quirino</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="67B8DC256B24FF85E578F8F4FDEFF8F0" box="[529,609,1881,1902]" pageId="12" pageNumber="327" rank="species">
|
||
<emphasis id="7C8B615E6B24FF85E578F8F4FDEFF8F0" bold="true" box="[529,609,1881,1902]" pageId="12" pageNumber="327">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1A80EDC46B25FF84E7FEF8C9FD3FF86B" httpUri="https://zenodo.org/record/2637382/files/figure.png" pageId="13" pageNumber="328" startId="13.[151,250,1892,1914]" targetBox="[174,1378,213,1867]" targetPageId="13">
|
||
<paragraph id="4E40BD4C6B25FF84E7FEF8C9FD3FF86B" blockId="13.[151,1436,1892,2037]" pageId="13" pageNumber="328">
|
||
<emphasis id="7C8B615E6B25FF84E7FEF8C9FE80F8E7" bold="true" box="[151,270,1892,1914]" pageId="13" pageNumber="328">FIGURE 8</emphasis>
|
||
.
|
||
<taxonomicName id="89FFC6CF6B25FF84E675F8C9FD9EF8E4" authority="Husana & Ng, 2019" authorityName="Husana & Ng" authorityYear="2019" box="[284,528,1892,1914]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="328" phylum="Arthropoda" rank="species" species="quirino" status="sp. nov.">
|
||
<emphasis id="7C8B615E6B25FF84E675F8C9FD9EF8E4" box="[284,528,1892,1914]" italics="true" pageId="13" pageNumber="328">Sundathelphusa quirino</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="67B8DC256B25FF84E571F8C8FDE2F8E4" box="[536,620,1893,1914]" pageId="13" pageNumber="328" rank="species">
|
||
<emphasis id="7C8B615E6B25FF84E571F8C8FDE2F8E4" bold="true" box="[536,620,1893,1914]" pageId="13" pageNumber="328">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A–H, holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino; I–M, paratype ♂ (47.6 × 36.9 mm), ZRC 2017.1063, Quirino. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G, I, J = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1A80EDC46B26FF87E7FEFC51FE0FFBF0" httpUri="https://zenodo.org/record/2637384/files/figure.png" pageId="14" pageNumber="329" startId="14.[151,250,1020,1042]" targetBox="[161,1428,171,999]" targetPageId="14">
|
||
<paragraph id="4E40BD4C6B26FF87E7FEFC51FE0FFBF0" blockId="14.[151,1436,1020,1134]" pageId="14" pageNumber="329">
|
||
<emphasis id="7C8B615E6B26FF87E7FEFC51FE9BFB8F" bold="true" box="[151,277,1020,1042]" pageId="14" pageNumber="329">FIGURE 9.</emphasis>
|
||
Right third and fourth ambulatory legs. A, B,
|
||
<taxonomicName id="89FFC6CF6B26FF87E468FC51FB7CFB8C" authority="(von Martens, 1868)" authorityName="von Martens" authorityYear="1868" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[769,1266,1020,1042]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="329" phylum="Arthropoda" rank="species" species="philippina">
|
||
<emphasis id="7C8B615E6B26FF87E468FC51FC2DFB8C" box="[769,931,1020,1042]" italics="true" pageId="14" pageNumber="329">Sundathelphusa</emphasis>
|
||
<emphasis id="7C8B615E6B26FF87E4C2FC51FB9DFB8C" box="[939,1043,1020,1042]" italics="true" pageId="14" pageNumber="329">philippina</emphasis>
|
||
(von Martens, 1868)
|
||
</taxonomicName>
|
||
, ♂ (51.3 × 40.4 mm), ZRC 2009.0409, Luzon; C, D,
|
||
<taxonomicName id="89FFC6CF6B26FF87E562FBB1FDDAFBAF" authority="Husana & Ng, 2019" authorityName="Husana & Ng" authorityYear="2019" box="[523,596,1051,1073]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="329" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
|
||
<emphasis id="7C8B615E6B26FF87E562FBB1FDDAFBAF" box="[523,596,1051,1073]" italics="true" pageId="14" pageNumber="329">S. cebu</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7C8B615E6B26FF87E533FBB6FD22FBAE" bold="true" box="[602,684,1051,1072]" pageId="14" pageNumber="329">
|
||
<taxonomicNameLabel id="67B8DC256B26FF87E533FBB6FD22FBAE" box="[602,684,1051,1072]" pageId="14" pageNumber="329" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; E, F,
|
||
<taxonomicName id="89FFC6CF6B26FF87E27DFBB1FAF6FBAF" authority="Husana & Ng, 2019" authorityName="Husana & Ng" authorityYear="2019" box="[1300,1400,1052,1073]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="329" phylum="Arthropoda" rank="species" species="quirino" status="sp. nov.">
|
||
<emphasis id="7C8B615E6B26FF87E27DFBB1FAF6FBAF" box="[1300,1400,1052,1073]" italics="true" pageId="14" pageNumber="329">S. quirino</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7C8B615E6B26FF87E214FBB6FF4BFBD1" bold="true" pageId="14" pageNumber="329">
|
||
<taxonomicNameLabel id="67B8DC256B26FF87E214FBB6FF4BFBD1" pageId="14" pageNumber="329" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, C, E, third ambulatory leg; B, D, F, fourth ambulatory leg. Scales: A–F = 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4E40BD4C6B26FF87E7AEFB38FDABFAFC" blockId="14.[151,1437,1173,2027]" pageId="14" pageNumber="329">
|
||
Male sternopleonal cavity reaching to level of proximal quarter of coxae of chelipeds. Adult male pleon narrow, T-shaped; somite 1 very short, proximal and distal margins sinuous; somite 2 transversely subrectangular; somites 3–5 narrow gradually; lateral margins of somite 3 convex, lateral margins of somite 4 slightly convex with angle at distal end, lateral margins of somite 5 concave; somite 6 length equal to width at widest point, lateral margins concave; telson subtriangular, length equal at the broadest distal margin, lateral margin convex medially, rounded distally (
|
||
<figureCitation id="D6C4A1C96B26FF87E632FAE7FE4EFAFC" box="[347,448,1354,1379]" captionStart="FIGURE 7" captionStartId="12.[151,250,1881,1903]" captionTargetBox="[150,1436,697,1859]" captionTargetId="figure@12.[150,1436,697,1859]" captionTargetPageId="12" captionText="FIGURE 7. Sundathelphusa quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, overall habitus; B, left third maxilliped; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, outer view of right chela; F, outer view of left chela." httpUri="https://zenodo.org/record/2637380/files/figure.png" pageId="14" pageNumber="329">Figs. 7D</figureCitation>
|
||
;
|
||
<figureCitation id="D6C4A1C96B26FF87E6A5FAE7FD97FAFC" box="[460,537,1354,1378]" captionStart="FIGURE 8" captionStartId="13.[151,250,1892,1914]" captionTargetBox="[174,1378,213,1867]" captionTargetId="figure@13.[169,1384,193,1873]" captionTargetPageId="13" captionText="FIGURE 8. Sundathelphusa quirino sp. nov. A–H, holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino; I–M, paratype ♂ (47.6 × 36.9 mm), ZRC 2017.1063, Quirino. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G, I, J = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637382/files/figure.png" pageId="14" pageNumber="329">8H, M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4E40BD4C6B26FF87E7AEFAC3FBDCFA50" blockId="14.[151,1437,1173,2027]" pageId="14" pageNumber="329">
|
||
G1 relatively slender, almost straight but slightly bended outward; outer margin of subterminal segment slightly concave; terminal segment slightly curved outward, tapering, cylindrical, slightly setose (
|
||
<figureCitation id="D6C4A1C96B26FF87E3B5FA3FFA93FA34" box="[1244,1309,1426,1451]" captionStart="FIGURE 8" captionStartId="13.[151,250,1892,1914]" captionTargetBox="[174,1378,213,1867]" captionTargetId="figure@13.[169,1384,193,1873]" captionTargetPageId="13" captionText="FIGURE 8. Sundathelphusa quirino sp. nov. A–H, holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino; I–M, paratype ♂ (47.6 × 36.9 mm), ZRC 2017.1063, Quirino. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G, I, J = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637382/files/figure.png" pageId="14" pageNumber="329">Fig. 8</figureCitation>
|
||
A–D, I–L). G2 longer than G1, distal segment long, about half length of basal segment (
|
||
<figureCitation id="D6C4A1C96B26FF87E487FA18FBCBFA50" box="[1006,1093,1461,1486]" captionStart="FIGURE 8" captionStartId="13.[151,250,1892,1914]" captionTargetBox="[174,1378,213,1867]" captionTargetId="figure@13.[169,1384,193,1873]" captionTargetPageId="13" captionText="FIGURE 8. Sundathelphusa quirino sp. nov. A–H, holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino; I–M, paratype ♂ (47.6 × 36.9 mm), ZRC 2017.1063, Quirino. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G, I, J = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637382/files/figure.png" pageId="14" pageNumber="329">Fig. 8E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06E5EEC76B26FF87E7AEFA77FDB7F988" pageId="14" pageNumber="329" type="etymology">
|
||
<paragraph id="4E40BD4C6B26FF87E7AEFA77FDB7F988" blockId="14.[151,1437,1173,2027]" pageId="14" pageNumber="329">
|
||
<emphasis id="7C8B615E6B26FF87E7AEFA77FEC8FA6D" bold="true" box="[199,326,1498,1523]" pageId="14" pageNumber="329">Etymology</emphasis>
|
||
. The species is named after the province of
|
||
<collectingRegion id="8C3B73AE6B26FF87E422FA77FC2CFA6D" box="[843,930,1498,1523]" country="Philippines" name="Quirino" pageId="14" pageNumber="329">Quirino</collectingRegion>
|
||
where all the materials were collected. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06E5EEC76B26FF87E7AEF98FFCB8F874" pageId="14" pageNumber="329" type="discussion">
|
||
<paragraph id="4E40BD4C6B26FF87E7AEF98FFCEDF970" blockId="14.[151,1437,1173,2027]" pageId="14" pageNumber="329">
|
||
<emphasis id="7C8B615E6B26FF87E7AEF98FFEBAF9A5" bold="true" box="[199,308,1570,1595]" pageId="14" pageNumber="329">Remarks</emphasis>
|
||
. The
|
||
<collectingRegion id="8C3B73AE6B26FF87E61FF98FFE43F9A5" box="[374,461,1570,1595]" country="Philippines" name="Quirino" pageId="14" pageNumber="329">Quirino</collectingRegion>
|
||
specimens, while superficially resembling
|
||
<taxonomicName id="89FFC6CF6B26FF87E4D1F98EFBC6F9A5" authority="Husana & Ng, 2019" authorityName="Husana & Ng" authorityYear="2019" box="[952,1096,1570,1595]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="329" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
|
||
<emphasis id="7C8B615E6B26FF87E4D1F98EFBC6F9A5" box="[952,1096,1570,1595]" italics="true" pageId="14" pageNumber="329">S. philippina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="67B8DC256B26FF87E338F98EFB6EF9A5" box="[1105,1248,1570,1595]" pageId="14" pageNumber="329" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
, differ markedly in having proportionately more slender and longer ambulatory merus; more inflated anterolateral region, the proximal part of the telson and distal region of pleonal somite 6 are proportionately wider, and the G1 has a relatively straighter distal segment (
|
||
<figureCitation id="D6C4A1C96B26FF87E54EF920FDFCF938" box="[551,626,1677,1702]" captionStart="FIGURE 8" captionStartId="13.[151,250,1892,1914]" captionTargetBox="[174,1378,213,1867]" captionTargetId="figure@13.[169,1384,193,1873]" captionTargetPageId="13" captionText="FIGURE 8. Sundathelphusa quirino sp. nov. A–H, holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino; I–M, paratype ♂ (47.6 × 36.9 mm), ZRC 2017.1063, Quirino. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G, I, J = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637382/files/figure.png" pageId="14" pageNumber="329">Figs. 8</figureCitation>
|
||
A–D, I–L; 9E, F) (vs. relatively shorter and stouter ambulatory legs, less convex anterolateral region, less wide telson and somite 6, and a more curved G1;
|
||
<figureCitation id="D6C4A1C96B26FF87E324F91FFB15F954" box="[1101,1179,1714,1739]" captionStart="FIGURE 2" captionStartId="4.[151,250,1799,1821]" captionTargetBox="[198,1324,189,1779]" captionTargetId="figure@4.[192,1328,183,1779]" captionTargetPageId="4" captionText="FIGURE 2. Sundathelphusa philippina (von Martens, 1868). A–G, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; H– K, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar; L–O, U, ♂ (33.8 × 27.2 mm), ZRC 2017.1059, Leyte; P–T, V, ♂ (51.3 × 40.2 mm), ZRC 2009.409, Luzon; W, ♂ (33.7 × 26.8 mm), ZRC 2009.409, Luzon. A, H, dorsal view of right G1; B, I, ventral view of right G1; C, J, dorsal view of terminal segment of right G1; D, K, ventral view of terminal segment of right G1; L, P, ventral view of left G1; M, Q, dorsal view of left G1; N, R, ventral view of terminal segment of left G1; O, S, dorsal view of terminal segment of left G1; E, male pleonal somites 3–6 and telson; F, frontal median triangle; G, posterior margin of epistome; T, right G2; U–W, male pleonal somites 4–6 and telson. Scales: A, B, H, I, L, M, P, Q = 1.0 mm; U, W = 2.0 mm; E = 5.0 mm; F, G, V = 3.0 mm; C, D, J, K, N, O, R, S = 0.5 mm." httpUri="https://zenodo.org/record/2637370/files/figure.png" pageId="14" pageNumber="329">Figs. 2</figureCitation>
|
||
A–D, F–S; 9A, B). As such, they are here referred to a new species,
|
||
<taxonomicName id="89FFC6CF6B26FF87E5FDF97BFC8DF970" authority="Husana & Ng, 2019" authorityName="Husana & Ng" authorityYear="2019" box="[660,771,1750,1774]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="329" phylum="Arthropoda" rank="species" species="quirino" status="sp. nov.">
|
||
<emphasis id="7C8B615E6B26FF87E5FDF97BFC8DF970" box="[660,771,1750,1774]" italics="true" pageId="14" pageNumber="329">S. quirino</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7C8B615E6B26FF87E460F978FCEDF970" bold="true" box="[777,867,1749,1774]" pageId="14" pageNumber="329">
|
||
<taxonomicNameLabel id="67B8DC256B26FF87E460F978FCEDF970" box="[777,867,1749,1774]" pageId="14" pageNumber="329" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4E40BD4C6B26FF87E7AEF957FCEAF8E0" blockId="14.[151,1437,1173,2027]" pageId="14" pageNumber="329">
|
||
The
|
||
<typeStatus id="914403EE6B26FF87E792F956FEA5F88D" box="[251,299,1787,1811]" pageId="14" pageNumber="329">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="89FFC6CF6B26FF87E6D8F956FDAFF88C" authority="Husana & Ng, 2019" authorityName="Husana & Ng" authorityYear="2019" box="[433,545,1787,1810]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="329" phylum="Arthropoda" rank="species" species="quirino" status="sp. nov.">
|
||
<emphasis id="7C8B615E6B26FF87E6D8F956FDAFF88C" box="[433,545,1787,1810]" italics="true" pageId="14" pageNumber="329">S. quirino</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7C8B615E6B26FF87E542F957FD08F88D" bold="true" box="[555,646,1786,1811]" pageId="14" pageNumber="329">
|
||
<taxonomicNameLabel id="67B8DC256B26FF87E542F957FD08F88D" box="[555,646,1786,1811]" pageId="14" pageNumber="329" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is quite far from the known distribution range of
|
||
<taxonomicName id="89FFC6CF6B26FF87E3ADF956FADBF88D" authority="Husana & Ng, 2019" authorityName="Husana & Ng" authorityYear="2019" box="[1220,1365,1786,1811]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="329" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
|
||
<emphasis id="7C8B615E6B26FF87E3ADF956FADBF88D" box="[1220,1365,1786,1811]" italics="true" pageId="14" pageNumber="329">S. philippina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="67B8DC256B26FF87E237F956FF53F8A8" pageId="14" pageNumber="329" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
and are separated by at least two geographic barriers. The Sierra Madre-Caraballo-Cordillera mountain ranges separates the
|
||
<typeStatus id="914403EE6B26FF87E6E8F8EEFE3FF8C5" box="[385,433,1859,1883]" pageId="14" pageNumber="329">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="89FFC6CF6B26FF87E55DF8EEFD2AF8C4" authority="Husana & Ng, 2019" authorityName="Husana & Ng" authorityYear="2019" box="[564,676,1859,1882]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="329" phylum="Arthropoda" rank="species" species="quirino" status="sp. nov.">
|
||
<emphasis id="7C8B615E6B26FF87E55DF8EEFD2AF8C4" box="[564,676,1859,1882]" italics="true" pageId="14" pageNumber="329">S. quirino</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7C8B615E6B26FF87E5C5F8EFFC88F8C5" bold="true" box="[684,774,1858,1883]" pageId="14" pageNumber="329">
|
||
<taxonomicNameLabel id="67B8DC256B26FF87E5C5F8EFFC88F8C5" box="[684,774,1858,1883]" pageId="14" pageNumber="329" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the far north of Luzon Island while the San Bernardino Strait in the south separates the big islands of Luzon and Samar.
|
||
</paragraph>
|
||
<paragraph id="4E40BD4C6B26FF87E7AEF827FCB8F874" blockId="14.[151,1437,1173,2027]" pageId="14" pageNumber="329">
|
||
While some of the specimens of
|
||
<taxonomicName id="89FFC6CF6B26FF87E52BF826FD3DF83C" authority="Husana & Ng, 2019" authorityName="Husana & Ng" authorityYear="2019" box="[578,691,1931,1954]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="329" phylum="Arthropoda" rank="species" species="quirino" status="sp. nov.">
|
||
<emphasis id="7C8B615E6B26FF87E52BF826FD3DF83C" box="[578,691,1931,1954]" italics="true" pageId="14" pageNumber="329">S. quirino</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7C8B615E6B26FF87E5D5F827FC99F83D" bold="true" box="[700,791,1930,1955]" pageId="14" pageNumber="329">
|
||
<taxonomicNameLabel id="67B8DC256B26FF87E5D5F827FC99F83D" box="[700,791,1930,1955]" pageId="14" pageNumber="329" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were collected from caves, none of the material has the typical characters associated with stygobionts (see discussion for
|
||
<taxonomicName id="89FFC6CF6B26FF87E41CF803FB8BF858" authorityName="Husana & Ng" authorityYear="2019" box="[885,1029,1965,1990]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="329" phylum="Arthropoda" rank="species" species="philippina">
|
||
<emphasis id="7C8B615E6B26FF87E41CF803FB8BF858" box="[885,1029,1965,1990]" italics="true" pageId="14" pageNumber="329">S. philippina</emphasis>
|
||
</taxonomicName>
|
||
) and the species is probably at most a facultative cavernicole (see
|
||
<bibRefCitation id="2A6EC0BD6B26FF87E68DF87EFDC6F874" author="Ng, P. K. L." box="[484,584,2002,2027]" pageId="14" pageNumber="329" pagination="91 - 97" refId="ref9480" refString="Ng, P. K. L. (2013) Stygothelphusa cranbrooki, a new species of cave crab from Gua Sireh, Sarawak, Malaysia (Crustacea: Decapoda: Brachyura: Gecarcinucidae). Raffles Bulletin of Zoology, 29 (Supplement), 91 - 97." type="journal article" year="2013">Ng 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2A6EC0BD6B26FF87E53DF87EFCA4F874" author="Husana, D. E. M. & Yamamuro, M. & Ng, P. K. L." box="[596,810,2002,2027]" pageId="14" pageNumber="329" pagination="565 - 574" refId="ref8690" refString="Husana, D. E. M., Yamamuro, M. & Ng, P. K. L. (2014) Two new species of freshwater crabs of the genus Sundathelphusa Bott, 1969 (Decapoda: Brachyura: Gecarcinucidae) from caves in Luzon, Philippines. Zootaxa, 3815 (4), 565 - 574. https: // doi. org / 10.11646 / zootaxa. 3815.4.6" type="journal article" year="2014">
|
||
Husana
|
||
<emphasis id="7C8B615E6B26FF87E5C7F87EFD69F875" box="[686,743,2002,2027]" italics="true" pageId="14" pageNumber="329">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |