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<document id="08B0B359EFA76BAB838038726AA080F6" ID-DOI="10.11646/zootaxa.3941.3.1" ID-GBIF-Dataset="67bc3c9d-ec99-4b47-81df-3fca7bd584ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="243860" ID-ZooBank="9907D081-A413-44F1-ABF6-76CC13DC8813" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461026017375" checkinUser="plazi" docAuthor="Wang, Cheng-Bin &amp; Zhou, Hong-Zhang" docDate="2015" docId="03EC87E6FF93880D9DBF19B0FC1A0847" docLanguage="en" docName="zt03941p338.pdf" docOrigin="Zootaxa 3941 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptomaphaginus perreaui Wang &amp; Zhou, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="324" masterDocId="FFD5FF9EFF85881A9D281927FFF50E4C" masterDocTitle="Taxonomy of the genus Ptomaphaginus Portevin (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from China, with description of eleven new species" masterLastPageNumber="338" masterPageNumber="301" pageNumber="323" updateTime="1698572131055" updateUser="plazi">
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<mods:title id="FC0F256492908C38F232D913D299D820">Taxonomy of the genus Ptomaphaginus Portevin (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from China, with description of eleven new species</mods:title>
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<mods:namePart id="65AA339492342BFEB3A88CBC1F5F197B">Wang, Cheng-Bin</mods:namePart>
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<mods:namePart id="5FCB669B27F0E71CB0D9382BEC26F5F2">Zhou, Hong-Zhang</mods:namePart>
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<mods:date id="8E72157F3F9B48F7B6A6B2677282119C">2015</mods:date>
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<treatment id="03EC87E6FF93880D9DBF19B0FC1A0847" ID-DOI="http://doi.org/10.5281/zenodo.5691676" ID-GBIF-Taxon="119606322" ID-Zenodo-Dep="5691676" LSID="urn:lsid:plazi:treatment:03EC87E6FF93880D9DBF19B0FC1A0847" httpUri="http://treatment.plazi.org/id/03EC87E6FF93880D9DBF19B0FC1A0847" lastPageId="23" lastPageNumber="324" pageId="22" pageNumber="323">
<subSubSection id="C35F657BFF93880C9DBF19B0FEDA0E9F" pageId="22" pageNumber="323" type="nomenclature">
<paragraph id="8BFA36F0FF93880C9DBF19B0FDDD0EFE" blockId="22.[151,552,151,211]" box="[151,552,151,178]" pageId="22" pageNumber="323">
<heading id="D0B2819CFF93880C9DBF19B0FDDD0EFE" bold="true" box="[151,552,151,178]" fontSize="11" level="1" pageId="22" pageNumber="323" reason="1">
<emphasis id="B931EAE2FF93880C9DBF19B0FDDD0EFE" bold="true" box="[151,552,151,178]" pageId="22" pageNumber="323">
<taxonomicName id="4C454D73FF93880C9DBF19B0FE360EFD" ID-CoL="6WLPJ" box="[151,451,151,177]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="perreaui" status="sp. nov.">
<emphasis id="B931EAE2FF93880C9DBF19B0FE360EFD" bold="true" box="[151,451,151,177]" italics="true" pageId="22" pageNumber="323">Ptomaphaginus perreaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2025799FF93880C9CE319BFFDDD0EFE" box="[459,552,152,178]" pageId="22" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA36F0FF93880C9DBF199DFEDA0E9F" blockId="22.[151,552,151,211]" box="[151,303,186,211]" pageId="22" pageNumber="323">
(
<figureCitation id="137E2A75FF93880C9DB7199DFF0D0E9F" box="[159,248,186,211]" captionStart="FIGURE 13" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[176,1408,380,1874]" captionTargetId="figure@22.[151,1436,371,1895]" captionTargetPageId="22" captionText="FIGURE 13. Ptomaphaginus perreaui sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, abdominal segment IX (ventral view); G, aedeagus (dorsal view); H, aedeagus (ventral view); I, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243873/files/figure.png" pageId="22" pageNumber="323">Figs. 13</figureCitation>
AI)
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF93880C9DBF1825FDE60F73" pageId="22" pageNumber="323" type="materials_examined">
<paragraph id="8BFA36F0FF93880C9DBF1825FDE60F73" blockId="22.[151,1436,258,319]" pageId="22" pageNumber="323">
<emphasis id="B931EAE2FF93880C9DBF1825FD440F57" bold="true" box="[151,689,258,283]" pageId="22" pageNumber="323">
<typeStatus id="54FE8852FF93880C9DBF1825FF270F57" box="[151,210,258,283]" pageId="22" pageNumber="323">Type</typeStatus>
material.
<typeStatus id="54FE8852FF93880C9C7F1825FE3D0F57" box="[343,456,258,283]" pageId="22" pageNumber="323" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3527660FF93880C9CF11825FDCD0F57" box="[473,568,258,283]" name="China" pageId="22" pageNumber="323">CHINA</collectingCountry>
, Yunnan:
</emphasis>
♂, Jinghong forest park,
<quantity id="4CBD9B15FF93880C9EC41824FBCF0F56" box="[1004,1082,259,283]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="323" unit="m" value="800.0">800 m</quantity>
, rainforest valley, pitfall trap,
<date id="FFFB1030FF93880C9DBF1801FEFB0F73" box="[151,270,294,319]" pageId="22" pageNumber="323" value="2004-02-26">26.II.2004</date>
, Jie Wu leg. (IZ-CAS).
</paragraph>
</subSubSection>
<caption id="DF3A6678FF93880C9DBF1E5AFC0F099C" httpUri="https://zenodo.org/record/243873/files/figure.png" pageId="22" pageNumber="323" targetBox="[176,1408,380,1874]" targetPageId="22">
<paragraph id="8BFA36F0FF93880C9DBF1E5AFC0F099C" blockId="22.[151,1436,1917,2000]" pageId="22" pageNumber="323">
<emphasis id="B931EAE2FF93880C9DBF1E5AFD3B09DF" bold="true" box="[151,718,1917,1939]" pageId="22" pageNumber="323">
FIGURE 13.
<taxonomicName id="4C454D73FF93880C9C031E5AFDC409DF" box="[299,561,1917,1939]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="perreaui" status="sp. nov.">
<emphasis id="B931EAE2FF93880C9C031E5AFDC409DF" bold="true" box="[299,561,1917,1939]" italics="true" pageId="22" pageNumber="323">Ptomaphaginus perreaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2025799FF93880C9F131E5AFD7B09DF" box="[571,654,1917,1939]" pageId="22" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
♂: A
</emphasis>
, antenna (dorsal view);
<emphasis id="B931EAE2FF93880C9EF81E5AFC1409DF" bold="true" box="[976,993,1917,1939]" pageId="22" pageNumber="323">B</emphasis>
, proleg (dorsal view);
<emphasis id="B931EAE2FF93880C99FB1E5AFB1009DF" bold="true" box="[1235,1253,1917,1939]" pageId="22" pageNumber="323">C</emphasis>
, protibia (ventral view);
<emphasis id="B931EAE2FF93880C9DC81EBBFF0709FE" bold="true" box="[224,242,1948,1970]" pageId="22" pageNumber="323">D</emphasis>
, pronotum (dorsal view);
<emphasis id="B931EAE2FF93880C9F2B1EBBFDE109FE" bold="true" box="[515,532,1948,1970]" pageId="22" pageNumber="323">E</emphasis>
, elytral apex (dorsoapical view);
<emphasis id="B931EAE2FF93880C9E591EBBFC8B09FE" bold="true" box="[881,894,1948,1970]" pageId="22" pageNumber="323">F</emphasis>
, abdominal segment IX (ventral view);
<emphasis id="B931EAE2FF93880C98081EBBFAC409FE" bold="true" box="[1312,1329,1948,1970]" pageId="22" pageNumber="323">G</emphasis>
, aedeagus (dorsal view);
<emphasis id="B931EAE2FF93880C9C021E9DFECB099C" bold="true" box="[298,318,1978,2000]" pageId="22" pageNumber="323">H</emphasis>
, aedeagus (ventral view);
<emphasis id="B931EAE2FF93880C9F621E9DFDA1099C" bold="true" box="[586,596,1978,2000]" pageId="22" pageNumber="323">I</emphasis>
, aedeagus (lateral view). Scales: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C35F657BFF92880D9DEF19B0FE6B0A8B" pageId="23" pageNumber="324" type="description">
<paragraph id="8BFA36F0FF92880D9DEF19B0FF240F51" blockId="23.[151,1437,151,1547]" pageId="23" pageNumber="324">
<emphasis id="B931EAE2FF92880D9DEF19B0FEAC0EFC" bold="true" box="[199,345,151,176]" pageId="23" pageNumber="324">Description.</emphasis>
<emphasis id="B931EAE2FF92880D9C4819B0FE6F0EFC" box="[352,410,151,176]" italics="true" pageId="23" pageNumber="324">Male</emphasis>
. EBL:
<quantity id="4CBD9B15FF92880D9CC519B0FDA00EE3" box="[493,597,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="23" pageNumber="324" unit="mm" value="2.05">2.05 mm</quantity>
. Length of different body parts: HL: AL: PL: ELL = 0.35: 0.52: 0.50:
<quantity id="4CBD9B15FF92880D9DBF199BFF0B0E98" box="[151,254,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="23" pageNumber="324" unit="mm" value="1.13">1.13 mm</quantity>
; width: HW: EW: PW: ELW = 0.59: 0.08: 0.83:
<quantity id="4CBD9B15FF92880D9E13199BFC570E98" box="[827,930,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="23" pageNumber="324" unit="mm" value="0.91">0.91 mm</quantity>
. Proportion of antennomeres from base to tip in µm (length × width): 82 × 28, 78 × 35, 41 × 32, 28 × 35, 28 × 39, 22 × 46, 40 × 58, 13 × 56, 41 × 64, 40 × 67, 78 × 65.
</paragraph>
<paragraph id="8BFA36F0FF92880D9DEF1800FE2C0F9C" blockId="23.[151,1437,151,1547]" pageId="23" pageNumber="324">Habitus elongated oval, relatively convex and lustreless. Moderately pigmented: mostly chestnut brown; occiput blakish brown; mouthparts, basal six antennomeres and apical half of ultimate antennomere, base of pronotum, about apical 1/3 of elytra, and tarsi yellowish. Dorsum continually clothed with fine, recumbent and sallow pubescence. Insertions of pubescence on dorsal surfaces of pronotum, scutellum, elytra and femora align along transverse striolations.</paragraph>
<paragraph id="8BFA36F0FF92880D9DE718FBFA930C73" blockId="23.[151,1437,151,1547]" pageId="23" pageNumber="324">
Head moderately short and convex, finely punctured, without visible transverse striolations, HW/HL = 1.67; anterior margin round. Compound eyes well developed, EW/HW = 0.13. Antennae short and condensed (
<figureCitation id="137E2A75FF92880D984718D8FF460C73" captionStart="FIGURE 13" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[176,1408,380,1874]" captionTargetId="figure@22.[151,1436,371,1895]" captionTargetPageId="22" captionText="FIGURE 13. Ptomaphaginus perreaui sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, abdominal segment IX (ventral view); G, aedeagus (dorsal view); H, aedeagus (ventral view); I, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243873/files/figure.png" pageId="23" pageNumber="324">Fig. 13</figureCitation>
A), AL/HW = 0.89; 4th antennomere as long as 5th; 6th transverse; 9th and 10th wider than long; 11th pear-like.
</paragraph>
<paragraph id="8BFA36F0FF92880D9DEF1B6CFAA80CE7" blockId="23.[151,1437,151,1547]" pageId="23" pageNumber="324">
Pronotum transverse and convex (
<figureCitation id="137E2A75FF92880D9F491B6DFD4C0C2F" box="[609,697,586,611]" captionStart="FIGURE 13" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[176,1408,380,1874]" captionTargetId="figure@22.[151,1436,371,1895]" captionTargetPageId="22" captionText="FIGURE 13. Ptomaphaginus perreaui sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, abdominal segment IX (ventral view); G, aedeagus (dorsal view); H, aedeagus (ventral view); I, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243873/files/figure.png" pageId="23" pageNumber="324">Fig. 13</figureCitation>
D), widest at base, PW/PL = 1.65. Sides almost rectilinearly narrowing from posterior to anterior; hind corners drawn out and bluntly rounded. Posterior margin with distinct postero-lateral emargination. Surface transversely striolated, SP: 3134, interspaces as wide as that on elytra.
</paragraph>
<paragraph id="8BFA36F0FF92880D9DEF1B91FD730D5B" blockId="23.[151,1437,151,1547]" pageId="23" pageNumber="324">
Elytra of medium length and convex, widest directly after base, ELL/EW = 1.23. Sides feebly curved, gradually narrowing from base to apices; apices nearly rounded (
<figureCitation id="137E2A75FF92880D9EA21BFDFC2B0CBF" box="[906,990,730,755]" captionStart="FIGURE 13" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[176,1408,380,1874]" captionTargetId="figure@22.[151,1436,371,1895]" captionTargetPageId="22" captionText="FIGURE 13. Ptomaphaginus perreaui sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, abdominal segment IX (ventral view); G, aedeagus (dorsal view); H, aedeagus (ventral view); I, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243873/files/figure.png" pageId="23" pageNumber="324">Fig. 13</figureCitation>
E). Surface transversely striolated, SE: 6164. Metathoracic wings fully developed.
</paragraph>
<paragraph id="8BFA36F0FF92880D9DEF1A05FCAB0DCE" blockId="23.[151,1437,151,1547]" pageId="23" pageNumber="324">
Prolegs robust, with basal three protarsomeres moderately expanded (
<figureCitation id="137E2A75FF92880D9EDE1A05FBB90D77" box="[1014,1100,802,827]" captionStart="FIGURE 13" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[176,1408,380,1874]" captionTargetId="figure@22.[151,1436,371,1895]" captionTargetPageId="22" captionText="FIGURE 13. Ptomaphaginus perreaui sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, abdominal segment IX (ventral view); G, aedeagus (dorsal view); H, aedeagus (ventral view); I, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243873/files/figure.png" pageId="23" pageNumber="324">Fig. 13</figureCitation>
B): TW/BTW = 1.37. Spinal arrangement on ventral side of protibia as shown in
<figureCitation id="137E2A75FF92880D9FF71A61FCC50D13" box="[735,816,838,863]" captionStart="FIGURE 13" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[176,1408,380,1874]" captionTargetId="figure@22.[151,1436,371,1895]" captionTargetPageId="22" captionText="FIGURE 13. Ptomaphaginus perreaui sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, abdominal segment IX (ventral view); G, aedeagus (dorsal view); H, aedeagus (ventral view); I, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243873/files/figure.png" pageId="23" pageNumber="324">Fig. 13</figureCitation>
C. Profemora wider than protibiae, and hairs on ventral side similar to that of protibiae. Mesotibiae moderately arcuate.
</paragraph>
<paragraph id="8BFA36F0FF92880D9DEF1AA9FC1F0D87" blockId="23.[151,1437,151,1547]" pageId="23" pageNumber="324">
Ventrite VII simple. Ventrite VIII regularly rounded posteriorly. Segment IX: spiculum gastrale elongate and moderately wide, sides nearly parallel, posterior end subrounded (
<figureCitation id="137E2A75FF92880D9E521A95FC390D87" box="[890,972,946,971]" captionStart="FIGURE 13" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[176,1408,380,1874]" captionTargetId="figure@22.[151,1436,371,1895]" captionTargetPageId="22" captionText="FIGURE 13. Ptomaphaginus perreaui sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, abdominal segment IX (ventral view); G, aedeagus (dorsal view); H, aedeagus (ventral view); I, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243873/files/figure.png" pageId="23" pageNumber="324">Fig. 13</figureCitation>
F).
</paragraph>
<paragraph id="8BFA36F0FF92880D9DEF1AF1FD1B0AEF" blockId="23.[151,1437,151,1547]" pageId="23" pageNumber="324">
Aedeagus oval in dorsal view (
<figureCitation id="137E2A75FF92880D9F1F1AF1FD790DA3" box="[567,652,982,1007]" captionStart="FIGURE 13" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[176,1408,380,1874]" captionTargetId="figure@22.[151,1436,371,1895]" captionTargetPageId="22" captionText="FIGURE 13. Ptomaphaginus perreaui sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, abdominal segment IX (ventral view); G, aedeagus (dorsal view); H, aedeagus (ventral view); I, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243873/files/figure.png" pageId="23" pageNumber="324">Fig. 13</figureCitation>
G): median lobe gently narrowed towards apex; opening of genital orifice trended to cut right margin of median lobe; right apical expansion golfclub-like at apical part; parameres narrow, firmly attached to median lobe. Ventral operculum distinctly and widely emarginate, each short lobe obliquely truncated apically (
<figureCitation id="137E2A75FF92880D9CF71D65FDC50A17" box="[479,560,1090,1115]" captionStart="FIGURE 13" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[176,1408,380,1874]" captionTargetId="figure@22.[151,1436,371,1895]" captionTargetPageId="22" captionText="FIGURE 13. Ptomaphaginus perreaui sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, abdominal segment IX (ventral view); G, aedeagus (dorsal view); H, aedeagus (ventral view); I, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243873/files/figure.png" pageId="23" pageNumber="324">Fig. 13</figureCitation>
H). In lateral view, median lobe slightly bent ventrad at about apical 1/3, apex a little thinned, and three ventrally-oriented setae inserted below apex as well as two just before middle (
<figureCitation id="137E2A75FF92880D981A1D41FA710A33" box="[1330,1412,1126,1151]" captionStart="FIGURE 13" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[176,1408,380,1874]" captionTargetId="figure@22.[151,1436,371,1895]" captionTargetPageId="22" captionText="FIGURE 13. Ptomaphaginus perreaui sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, abdominal segment IX (ventral view); G, aedeagus (dorsal view); H, aedeagus (ventral view); I, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243873/files/figure.png" pageId="23" pageNumber="324">
Fig.
<date id="FFFB1030FF92880D984E1D40FA710A33" box="[1382,1412,1127,1151]" pageId="23" pageNumber="324" value="2013-01">13</date>
</figureCitation>
I). Internal stylus wide, multiannulate at about basal 1/3.
</paragraph>
<paragraph id="8BFA36F0FF92880D9DEF1D89FE6B0A8B" blockId="23.[151,1437,151,1547]" box="[199,414,1198,1223]" pageId="23" pageNumber="324">
<emphasis id="B931EAE2FF92880D9DEF1D89FEEE0A8B" box="[199,283,1198,1223]" italics="true" pageId="23" pageNumber="324">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF92880D9DEF1DF5FDD40AA7" box="[199,545,1234,1259]" pageId="23" pageNumber="324" type="distribution">
<paragraph id="8BFA36F0FF92880D9DEF1DF5FDD40AA7" blockId="23.[151,1437,151,1547]" box="[199,545,1234,1259]" pageId="23" pageNumber="324">
<emphasis id="B931EAE2FF92880D9DEF1DF5FE950AA7" bold="true" box="[199,352,1234,1259]" pageId="23" pageNumber="324">Distribution.</emphasis>
<collectingCountry id="F3527660FF92880D9C4F1DF5FE5F0AA7" box="[359,426,1234,1259]" name="China" pageId="23" pageNumber="324">China</collectingCountry>
(Yunnan).
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF92880D9DEF1DD1FACC0B7F" pageId="23" pageNumber="324" type="etymology">
<paragraph id="8BFA36F0FF92880D9DEF1DD1FACC0B7F" blockId="23.[151,1437,151,1547]" pageId="23" pageNumber="324">
<emphasis id="B931EAE2FF92880D9DEF1DD1FEBB0B43" bold="true" box="[199,334,1270,1295]" pageId="23" pageNumber="324">Etymology.</emphasis>
The specific epithet is dedicated to Dr. Michel Perreau (Université Paris 7, Paris,
<collectingCountry id="F3527660FF92880D980F1DD0FA8D0B43" box="[1319,1400,1271,1295]" name="France" pageId="23" pageNumber="324">France</collectingCountry>
), a famous taxonomist on
<taxonomicName id="4C454D73FF92880D9CBF1C3DFDFF0B7F" box="[407,522,1306,1331]" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
, for his excellent work on Chinese fauna and generous help to our study.
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF92880D9DEF1C19FC1A0847" pageId="23" pageNumber="324" type="discussion">
<paragraph id="8BFA36F0FF92880D9DEF1C19FC1A0847" blockId="23.[151,1437,151,1547]" pageId="23" pageNumber="324">
<emphasis id="B931EAE2FF92880D9DEF1C19FECE0B1B" bold="true" box="[199,315,1342,1367]" pageId="23" pageNumber="324">Remarks.</emphasis>
Obviously, this new species has the smallest body size of all species included here, EBL only approximately equal to 2.00 mm and its aedeagus is similar to
<taxonomicName id="4C454D73FF92880D9EBD1C43FB1C0B37" authority="Perreau, 1991" authorityName="Perreau" authorityYear="1991" box="[917,1257,1378,1403]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="bengalicola">
<emphasis id="B931EAE2FF92880D9EBD1C43FC560B37" box="[917,931,1380,1403]" italics="true" pageId="23" pageNumber="324">P</emphasis>
.
<emphasis id="B931EAE2FF92880D9E9F1C45FBCE0B37" box="[951,1083,1378,1403]" italics="true" pageId="23" pageNumber="324">bengalicola</emphasis>
<bibRefCitation id="EFD44B01FF92880D99601C44FB1C0B37" author="Perreau" box="[1096,1257,1379,1403]" pageId="23" pageNumber="324" refString="Perreau, M. (1991) Contribution a la connaissance des Cholevidae de l'Inde du Nord et du Pakistan (Coleoptera). Revue Suisse de Zoologie, 98 (3), 555 - 566." type="journal article" year="1991">Perreau, 1991</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3527660FF92880D98121C45FA830B37" box="[1338,1398,1378,1403]" name="India" pageId="23" pageNumber="324">India</collectingCountry>
,
<taxonomicName id="4C454D73FF92880D98AF1C43FDEA0BD3" authority="Szymczakowski, 1974" authorityName="Szymczakowski" authorityYear="1974" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="bucculentus">
<emphasis id="B931EAE2FF92880D98AF1C43FA600B37" box="[1415,1429,1380,1403]" italics="true" pageId="23" pageNumber="324">P</emphasis>
.
<emphasis id="B931EAE2FF92880D9DBF1CA1FEE80BD3" box="[151,285,1414,1439]" italics="true" pageId="23" pageNumber="324">bucculentus</emphasis>
<bibRefCitation id="EFD44B01FF92880D9C0B1CA1FDEA0BD3" author="Szymczakowski" box="[291,543,1414,1439]" pageId="23" pageNumber="324" refString="Szymczakowski, W. (1974) Nouvelles remarques sur les Catopidae (Coleoptera) de la region orientale. Acta Zoologica Cracoviensia, 19 (10), 197 - 216." type="journal article" year="1974">Szymczakowski, 1974</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3527660FF92880D9F4B1CA1FD5C0BD3" box="[611,681,1414,1439]" name="Nepal" pageId="23" pageNumber="324">Nepal</collectingCountry>
,
<taxonomicName id="4C454D73FF92880D9F9C1CAFFBC60BD3" authority="Perreau, 1991" authorityName="Perreau" authorityYear="1991" box="[692,1075,1414,1439]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="cherrapunjeensis">
<emphasis id="B931EAE2FF92880D9F9C1CAFFD370BD3" box="[692,706,1416,1439]" italics="true" pageId="23" pageNumber="324">P</emphasis>
.
<emphasis id="B931EAE2FF92880D9FF81CA1FC640BD3" box="[720,913,1414,1439]" italics="true" pageId="23" pageNumber="324">cherrapunjeensis</emphasis>
<bibRefCitation id="EFD44B01FF92880D9EBF1CAFFBC60BD3" author="Perreau" box="[919,1075,1415,1439]" pageId="23" pageNumber="324" refString="Perreau, M. (1991) Contribution a la connaissance des Cholevidae de l'Inde du Nord et du Pakistan (Coleoptera). Revue Suisse de Zoologie, 98 (3), 555 - 566." type="journal article" year="1991">Perreau, 1991</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3527660FF92880D995E1CA1FB470BD3" box="[1142,1202,1414,1439]" name="India" pageId="23" pageNumber="324">India</collectingCountry>
,
<taxonomicName id="4C454D73FF92880D99961CAFFF3A0B8F" authority="Perreau, 1988" authorityName="Perreau" authorityYear="1988" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B931EAE2FF92880D99961CAFFB390BD3" box="[1214,1228,1416,1439]" italics="true" pageId="23" pageNumber="324">P</emphasis>
.
<emphasis id="B931EAE2FF92880D99F21CAFFACC0BD3" box="[1242,1337,1416,1439]" italics="true" pageId="23" pageNumber="324">minimus</emphasis>
<bibRefCitation id="EFD44B01FF92880D98171CAFFF3A0B8F" author="Perreau" pageId="23" pageNumber="324" refString="Perreau, M. (1988) Les Cholevidae himalayiens du Museum d'Histoire naturelle de Geneve (Coleoptera). Revue Suisse de Zoologie, 95 (4), 1005 - 1018." type="journal article" year="1988">Perreau, 1988</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3527660FF92880D9C3E1C8DFEAC0B8F" box="[278,345,1450,1475]" name="Nepal" pageId="23" pageNumber="324">Nepal</collectingCountry>
and
<taxonomicName id="4C454D73FF92880D9CBA1C8BFD220B8F" authority="Perreau, 1988" authorityName="Perreau" authorityYear="1988" box="[402,727,1450,1475]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="palpaloides">
<emphasis id="B931EAE2FF92880D9CBA1C8BFE550B8F" box="[402,416,1452,1475]" italics="true" pageId="23" pageNumber="324">P</emphasis>
.
<emphasis id="B931EAE2FF92880D9C871C8DFDC40B8F" box="[431,561,1450,1475]" italics="true" pageId="23" pageNumber="324">palpaloides</emphasis>
<bibRefCitation id="EFD44B01FF92880D9F121C8CFD220B8F" author="Perreau" box="[570,727,1451,1475]" pageId="23" pageNumber="324" refString="Perreau, M. (1988) Les Cholevidae himalayiens du Museum d'Histoire naturelle de Geneve (Coleoptera). Revue Suisse de Zoologie, 95 (4), 1005 - 1018." type="journal article" year="1988">Perreau, 1988</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3527660FF92880D9E361C8DFC910B8F" box="[798,868,1450,1475]" name="Nepal" pageId="23" pageNumber="324">Nepal</collectingCountry>
, but it can be distinguished from those mentioned by the combination of its minute size, spiculum gastrale elongate and lateral sides parallel, ventral operculum distinctly and widely emarginate, and aedeagal shape when viewed laterally.
</paragraph>
</subSubSection>
</treatment>
</document>