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<document id="71C5B6F1737219F858CAE160A92DB0C6" ID-CLB-Dataset="302084" ID-DOI="10.11646/phytotaxa.220.1.11" ID-GBIF-Dataset="3be1850e-a52b-4da1-81cd-31cd6130f33e" ID-ISSN="1179-3163" ID-Zenodo-Dep="13634850" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725328652167" checkinUser="felipe" docAuthor="Palacios, Walter A." docDate="2015" docId="03EC878935405479FF59E1AC8FDC4A28" docLanguage="en" docName="phytotaxa.220.1.11.pdf" docOrigin="Phytotaxa 220 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.220.1.11" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Amyris karlitae W. Palacios 2015, sp. nov." docType="treatment" docVersion="4" lastPageNumber="99" masterDocId="FFD5FFF135405478FFD1E4688E484F23" masterDocTitle="A new species of Amyris (Rutaceae) from southern Ecuador" masterLastPageNumber="100" masterPageNumber="98" pageNumber="98" updateTime="1725334322766" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="72EFF8823B0EDC8C3912815D54A02E74">A new species of Amyris (Rutaceae) from southern Ecuador</mods:title>
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<mods:namePart id="3C6E84E18EA05D054B052681B6382E6E">Palacios, Walter A.</mods:namePart>
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<mods:title id="92875ECD29EC8A75968444E7210F8766">Phytotaxa</mods:title>
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<mods:date id="06D10AEC66099A876EBEA8D6CF1B635D">2015</mods:date>
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<treatment id="03EC878935405479FF59E1AC8FDC4A28" ID-DOI="http://doi.org/10.5281/zenodo.13634852" ID-GBIF-Taxon="240097026" ID-Zenodo-Dep="13634852" LSID="urn:lsid:plazi:treatment:03EC878935405479FF59E1AC8FDC4A28" httpUri="http://treatment.plazi.org/id/03EC878935405479FF59E1AC8FDC4A28" lastPageId="1" lastPageNumber="99" pageId="0" pageNumber="98">
<subSubSection id="C35F651435405478FF59E1AC8C694AFC" box="[136,545,1476,1503]" pageId="0" pageNumber="98" type="nomenclature">
<paragraph id="8BFA369F35405478FF59E1AC8C694AFC" blockId="0.[136,545,1476,1503]" box="[136,545,1476,1503]" pageId="0" pageNumber="98">
<heading id="D0B281F335405478FF59E1AC8C694AFC" box="[136,545,1476,1503]" fontSize="11" level="2" pageId="0" pageNumber="98" reason="3">
<taxonomicName id="4C454D1C35405478FF59E1AC8F884AFC" ID-CoL="7R5Y2" authority="W.Palacios" authorityName="W. Palacios" authorityYear="2015" box="[136,448,1476,1503]" class="Magnoliopsida" family="Rutaceae" genus="Amyris" kingdom="Plantae" order="Sapindales" pageId="0" pageNumber="98" phylum="Tracheophyta" rank="species" species="karlitae" status="sp. nov.">
<emphasis id="B931EA8D35405478FF59E1AC8F734AFC" bold="true" box="[136,315,1476,1503]" italics="true" pageId="0" pageNumber="98">Amyris karlitae</emphasis>
W.Palacios
</taxonomicName>
,
<taxonomicNameLabel id="A20257F635405478FE1AE1AE8C694AFC" box="[459,545,1478,1503]" pageId="0" pageNumber="98" rank="species">
<emphasis id="B931EA8D35405478FE1AE1AE8FAC4AFC" box="[459,484,1478,1503]" italics="true" pageId="0" pageNumber="98">sp</emphasis>
.
<emphasis id="B931EA8D35405478FE23E1AE8C524AFC" box="[498,538,1478,1503]" italics="true" pageId="0" pageNumber="98">nov</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C35F651435405478FF59E2788AB8496A" pageId="0" pageNumber="98" type="diagnosis">
<paragraph id="8BFA369F35405478FF59E2788AB8496A" blockId="0.[136,1452,1552,1681]" pageId="0" pageNumber="98">
<emphasis id="B931EA8D35405478FF59E2788EBB4906" bold="true" box="[136,243,1552,1573]" pageId="0" pageNumber="98">Diagnosis:</emphasis>
<taxonomicName id="4C454D1C35405478FEDAE2788FD74906" authorityName="W. Palacios" authorityYear="2015" box="[267,415,1552,1573]" class="Magnoliopsida" family="Rutaceae" genus="Amyris" kingdom="Plantae" order="Sapindales" pageId="0" pageNumber="98" phylum="Tracheophyta" rank="species" species="karlitae">
<emphasis id="B931EA8D35405478FEDAE2788FD74906" box="[267,415,1552,1573]" italics="true" pageId="0" pageNumber="98">Amyris karlitae</emphasis>
</taxonomicName>
is distinct from other species of the genus by a combination of characters: unifoliolate leaves, 611 ×
<quantity id="4CBD9B7A35405478FA8EE2788BE04906" box="[1375,1448,1552,1573]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="0" pageNumber="98" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">46 cm</quantity>
, pinnate venation, and emarginate or acuminate apex and stamens 8(or 10), alternating short and long. (
<figureCitation id="137E2A1A35405478FB47E25C8AAF496A" box="[1174,1255,1588,1609]" captionStart="FIGURE 1" captionStartId="1.[136,229,585,606]" captionTargetBox="[164,1423,187,557]" captionTargetId="figure-17@1.[164,1423,187,557]" captionTargetPageId="1" captionText="FIGURE 1. Amyris karlitae. A. habit (bar = 1 cm); B. leaf abaxial side, notice emarginated tip (bar = 2 cm); C. flower (bar = 2 mm). Photographs by W.A. Palacios." figureDoi="http://doi.org/10.5281/zenodo.13634854" httpUri="https://zenodo.org/record/13634854/files/figure.png" pageId="0" pageNumber="98">Figure 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35F651435405478FF59E2308A0E473C" pageId="0" pageNumber="98" type="materials_examined">
<paragraph id="8BFA369F35405478FF59E2308DD649B2" blockId="0.[136,1452,1552,1681]" pageId="0" pageNumber="98">
<emphasis id="B931EA8D35405478FF59E2308EF2494E" bold="true" box="[136,186,1624,1645]" pageId="0" pageNumber="98">Type</emphasis>
:—
<collectingCountry id="F352760F35405478FF07E2308F18494E" box="[214,336,1624,1645]" name="Ecuador" pageId="0" pageNumber="98">ECUADOR</collectingCountry>
.
<collectingRegion id="4981F87D35405478FE89E2308FC0494E" box="[344,392,1624,1645]" country="Ecuador" name="Loja" pageId="0" pageNumber="98">Loja</collectingRegion>
: Cantón Macará, Sabiango, Achima-Cerro Jatumpamba, sitio El Fondo, aprox.
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,
<date id="FFFB105F35405478FB12E2308B7F494E" box="[1219,1335,1624,1645]" pageId="0" pageNumber="98" value="2010-08">agosto 2010</date>
<emphasis id="B931EA8D35405478FAEAE2308EB149B2" italics="true" pageId="0" pageNumber="98">
<collectionCode id="ED54AE5A35405478FAEAE2308B18494E" box="[1339,1360,1624,1645]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="0" pageNumber="98" type="Herbarium">W</collectionCode>
. Palacios 17324
</emphasis>
(
<typeStatus id="54FE883D35405478FED7E2148F1449B2" box="[262,348,1660,1681]" pageId="0" pageNumber="98" type="holotype">holotype</typeStatus>
<collectionCode id="ED54AE5A35405478FEB0E2148FE249B2" box="[353,426,1660,1681]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13634" lsid="urn:lsid:biocol.org:col:13634" name="Museo Ecuatoriano de Ciencias Naturales" pageId="0" pageNumber="98" type="Herbarium">QCNE</collectionCode>
!,
<typeStatus id="54FE883D35405478FE66E2148FB449B2" box="[439,508,1660,1681]" pageId="0" pageNumber="98" type="isotype">isotype</typeStatus>
to be distributed to
<collectionCode id="ED54AE5A35405478FD6CE2148CBA49B2" box="[701,754,1660,1681]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15650" lsid="urn:lsid:biocol.org:col:15650" name="Pontificia Universidad Católica del Ecuador" pageId="0" pageNumber="98" type="Herbarium">QCA</collectionCode>
,
<collectionCode id="ED54AE5A35405478FD2AE2148D6C49B2" box="[763,804,1660,1681]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="0" pageNumber="98" type="Herbarium">MO</collectionCode>
,
<collectionCode id="ED54AE5A35405478FCFCE2148D2B49B2" box="[813,867,1660,1681]" country="Ethiopia" httpUri="http://grbio.org/cool/5dxx-xet2" name="Addis Ababa University, Department of Biology" pageId="0" pageNumber="98">AAU</collectionCode>
,
<collectionCode id="ED54AE5A35405478FCBCE2148DDC49B2" box="[877,916,1660,1681]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="0" pageNumber="98" type="Herbarium">NY</collectionCode>
).
</paragraph>
<paragraph id="8BFA369F35405478FF59E2AA8A0E473C" blockId="0.[136,1452,1730,2079]" pageId="0" pageNumber="98">
Tree to
<quantity id="4CBD9B7A35405478FF30E2AA8F5349F8" box="[225,283,1730,1755]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="0" pageNumber="98" unit="m" value="18.0">18 m</quantity>
tall; dbh to
<quantity id="4CBD9B7A35405478FE76E2AA8FBA49F8" box="[423,498,1730,1755]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="98" unit="cm" value="50.0">50 cm</quantity>
. Inner bark creamy-yellow and very aromatic. Terminal twigs clustered, cylindrical, browns. Leaves unifoliolate, alternate, spiral, 611 ×
<quantity id="4CBD9B7A35405478FD3AE28E8D0B49DC" box="[747,835,1766,1791]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="0" pageNumber="98" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">46 cm</quantity>
, elliptic, ovate-lanceolate or oblong-elliptic, glabrous, shiny above; apex acuminate or emarginate and in this case slightly recurved; base obtuse; secondary veins 1013 pairs, more or less convergent, inconspicuous, forked towards the margin; intersecondary and secondary nerves very similar; tertiary nerves inconspicuous; margin entire; glandular dots translucent on leaves; petioles 13(4) cm long, up to half as long as the lamina, thickened and curved at the apex. Inflorescence an axillary or terminal panicle, up to
<quantity id="4CBD9B7A35405478FF59E3F28E884890" box="[136,192,1946,1971]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="0" pageNumber="98" unit="cm" value="8.0">8 cm</quantity>
long, with lateral branches up to
<quantity id="4CBD9B7A35405478FDEAE3F28CC04890" box="[571,648,1946,1971]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="0" pageNumber="98" unit="cm" value="2.5">2.5 cm</quantity>
long; bracteoles ovate,
<quantity id="4CBD9B7A35405478FC47E3F28A404890" box="[918,1032,1946,1971]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="0" pageNumber="98" unit="mm" value="0.85" valueMax="1.0" valueMin="0.7">0.71 mm</quantity>
long. Flowers hermaphrodite, glanddotted; calyx patelliform, 4-lobed, approx.
<quantity id="4CBD9B7A35405478FDA3E3D68CFD48F4" box="[626,693,1982,2007]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="98" unit="mm" value="1.0">1 mm</quantity>
long, lobes broadly ovate, shortly fused at the base; petals 4, free, elliptic or oblong-elliptic,
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long, white; stamens 8(or 10), alternating short and long; disc glandular, orange, prominent between the stamens and pistil; pedicels
<quantity id="4CBD9B7A35405478FD1BEC6E8D74473C" box="[714,828,2054,2079]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="0" pageNumber="98" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.53 mm</quantity>
long, articulate at base.
</paragraph>
</subSubSection>
<caption id="DF3A661735415479FF59E6218FFB4DA1" ID-DOI="http://doi.org/10.5281/zenodo.13634854" ID-Zenodo-Dep="13634854" httpUri="https://zenodo.org/record/13634854/files/figure.png" pageId="1" pageNumber="99" startId="1.[136,229,585,606]" targetBox="[164,1423,187,557]" targetPageId="1" targetType="figure">
<paragraph id="8BFA369F35415479FF59E6218FFB4DA1" blockId="1.[136,1452,585,642]" pageId="1" pageNumber="99">
<emphasis id="B931EA8D35415479FF59E6218EB14D7D" bold="true" box="[136,249,585,606]" pageId="1" pageNumber="99">FIGURE 1</emphasis>
.
<taxonomicName id="4C454D1C35415479FEDCE6218FEB4D7D" authorityName="W. Palacios" authorityYear="2015" box="[269,419,585,606]" class="Magnoliopsida" family="Rutaceae" genus="Amyris" kingdom="Plantae" order="Sapindales" pageId="1" pageNumber="99" phylum="Tracheophyta" rank="species" species="karlitae">
<emphasis id="B931EA8D35415479FEDCE6218FEB4D7D" box="[269,419,585,606]" italics="true" pageId="1" pageNumber="99">Amyris karlitae</emphasis>
</taxonomicName>
. A. habit (bar = 1 cm); B. leaf abaxial side, notice emarginated tip (bar = 2 cm); C. flower (bar = 2 mm). Photographs by W.A. Palacios.
</paragraph>
</caption>
<subSubSection id="C35F651435415479FF6CE6BE8DE44DCC" box="[189,940,726,751]" pageId="1" pageNumber="99" type="etymology">
<paragraph id="8BFA369F35415479FF6CE6BE8DE44DCC" blockId="1.[136,1452,726,1291]" box="[189,940,726,751]" pageId="1" pageNumber="99">
<emphasis id="B931EA8D35415479FF6CE6BE8F004DCC" bold="true" box="[189,328,726,751]" pageId="1" pageNumber="99">Etymology:</emphasis>
—Specific epithet is dedicated to my daughter Karlita.
</paragraph>
</subSubSection>
<subSubSection id="C35F651435415479FF6CE6928FDC4A28" pageId="1" pageNumber="99" type="distribution">
<paragraph id="8BFA369F35415479FF6CE6928F9C4CC8" blockId="1.[136,1452,726,1291]" pageId="1" pageNumber="99">
<emphasis id="B931EA8D35415479FF6CE6928FA04C30" bold="true" box="[189,488,762,787]" pageId="1" pageNumber="99">Distribution and habitat:</emphasis>
<taxonomicName id="4C454D1C35415479FDD5E6928CF84C30" authorityName="W. Palacios" authorityYear="2015" box="[516,688,762,787]" class="Magnoliopsida" family="Rutaceae" genus="Amyris" kingdom="Plantae" order="Sapindales" pageId="1" pageNumber="99" phylum="Tracheophyta" rank="species" species="karlitae">
<emphasis id="B931EA8D35415479FDD5E6928CF84C30" box="[516,688,762,787]" italics="true" pageId="1" pageNumber="99">Amyris karlitae</emphasis>
</taxonomicName>
grows between 1400 and
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, where trees are laden with mosses in semideciduous forest. Typical trees include
<taxonomicName id="4C454D1C35415479FD42E7768A734C14" authority="Diels (1906: 398)" authorityName="Diels" authorityPageNumber="398" authorityYear="1906" box="[659,1083,798,823]" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="99" phylum="Tracheophyta" rank="species" species="neotropica">
<emphasis id="B931EA8D35415479FD42E7768D224C14" box="[659,874,798,823]" italics="true" pageId="1" pageNumber="99">Juglans neotropica</emphasis>
<bibRefCitation id="EFD44B6E35415479FCA3E7768A734C14" author="Diels, F." box="[882,1083,798,823]" pageId="1" pageNumber="99" refId="ref1780" refString="Diels, F. (1906) Juglans in Peruvia Amazonica Collecta. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 37: 398." type="journal volume" year="1906">Diels (1906: 398)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C454D1C35415479FB99E7768F924C78" authority="Ruiz &amp; Pavon (1802: 17) Kuntze (1898: 195)" authorityName="Ruiz &amp; Pavon (1802: 17) Kuntze" authorityYear="1898" class="Magnoliopsida" family="Sapotaceae" genus="Pouteria" kingdom="Plantae" order="Ericales" pageId="1" pageNumber="99" phylum="Tracheophyta" rank="species" species="lucuma">
<emphasis id="B931EA8D35415479FB99E7768B4B4C14" box="[1096,1283,798,823]" italics="true" pageId="1" pageNumber="99">Pouteria lucuma</emphasis>
<bibRefCitation id="EFD44B6E35415479FADAE7768EB44C78" author="Ruiz, H. &amp; Pavon, J." pageId="1" pageNumber="99" refId="ref2044" refString="Ruiz, H. &amp; Pavon, J. (1802) Flora Peruviana, et Chilensis 3. Typis Gabrielis de Sancha, Madrid, 95 pp." type="book" year="1802">Ruiz &amp; Pavon (1802: 17)</bibRefCitation>
<bibRefCitation id="EFD44B6E35415479FED3E72A8F924C78" author="Kuntze, C." box="[258,474,834,859]" pageId="1" pageNumber="99" refId="ref1841" refString="Kuntze, C. (1898) Revisio Generum Plantarum. Vol. 3. A. Felix, Leipzig, 195 pp." type="book" year="1898">Kuntze (1898: 195)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C454D1C35415479FDDFE72A8AE14C78" authority="Blake (1922: 419) Robinson (1976: 37)" authorityName="Blake (1922: 419) Robinson" authorityYear="1976" box="[526,1193,834,859]" class="Magnoliopsida" family="Asteraceae" genus="Monactis" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="99" phylum="Tracheophyta" rank="species" species="holwayae">
<emphasis id="B931EA8D35415479FDDFE72A8CAF4C78" box="[526,743,834,859]" italics="true" pageId="1" pageNumber="99">Monactis holwayae</emphasis>
<bibRefCitation id="EFD44B6E35415479FD23E72A8DF34C78" author="Blake, S." box="[754,955,834,859]" pageId="1" pageNumber="99" pagination="414 - 430" refId="ref1598" refString="Blake, S. (1922) New South American Asteraceae collected by E. W. D. Hollway. Botanical Gazette 74: 414 - 430. http: // dx. doi. org / 10.1086 / 333127" type="journal article" year="1922">Blake (1922: 419)</bibRefCitation>
<bibRefCitation id="EFD44B6E35415479FC10E72A8AE14C78" author="Robinson, H." box="[961,1193,834,859]" pageId="1" pageNumber="99" pagination="33 - 45" refId="ref2011" refString="Robinson, H. (1976) Monactis holwayae, studies in the Heliantheae (Asteraceae). VII. Notes on the genus Monactis. Phytologia 34: 33 - 45." type="journal article" year="1976">Robinson (1976: 37)</bibRefCitation>
</taxonomicName>
. This
<typeStatus id="54FE883D35415479FB38E72A8B514C78" box="[1257,1305,834,859]" pageId="1" pageNumber="99">type</typeStatus>
of forest that is part of the Huancabamba bioregion, located between
<collectingCountry id="F352760F35415479FCD3E70E8D174C5C" box="[770,863,870,895]" name="Ecuador" pageId="1" pageNumber="99">Ecuador</collectingCountry>
and
<collectingCountry id="F352760F35415479FC49E70E8D864C5C" box="[920,974,870,895]" name="Peru" pageId="1" pageNumber="99">Perú</collectingCountry>
, is very scarce and ecologically important because of the presence of several endemic species (
<bibRefCitation id="EFD44B6E35415479FD33E7E28D364C80" author="Lozano, P." box="[738,894,906,931]" pageId="1" pageNumber="99" pagination="29 - 50" refId="ref1865" refString="Lozano, P. (2002) Los tipos de bosques en el sur del Ecuador. In: Aguirre M., A., Madsen, J. E., Cotton, E. &amp; Balslev, H. (Eds.) Botanica AustroEcuatoriana - Estudios sobre los recursos vegetales en las provincias de El Oro, Loja y Zamora Chinchipe. Editorial Abya Yala, Quito, pp. 29 - 50." type="book chapter" year="2002">Lozano 2002</bibRefCitation>
). The area receives rainfall between January and April, and corresponds to the
<emphasis id="B931EA8D35415479FE34E7C68ABE4CE4" box="[485,1270,942,967]" italics="true" pageId="1" pageNumber="99">ecosistema bosque montano bajo semideciduo de Catamayo-Alamor</emphasis>
(Ministerio del Ambiente del
<collectingCountry id="F352760F35415479FEF6E7BA8FCC4CC8" box="[295,388,978,1003]" name="Ecuador" pageId="1" pageNumber="99">Ecuador</collectingCountry>
2013).
</paragraph>
<paragraph id="8BFA369F35415479FF6CE79E8D9E4B74" blockId="1.[136,1452,726,1291]" pageId="1" pageNumber="99">
<emphasis id="B931EA8D35415479FF6CE79E8C194B2C" bold="true" box="[189,593,1014,1039]" pageId="1" pageNumber="99">Etnobotany &amp; vernacular names:</emphasis>
—The common name of
<taxonomicName id="4C454D1C35415479FCA1E79E8DAF4B2C" authorityName="W. Palacios" authorityYear="2015" box="[880,999,1014,1039]" class="Magnoliopsida" family="Rutaceae" genus="Amyris" kingdom="Plantae" order="Sapindales" pageId="1" pageNumber="99" phylum="Tracheophyta" rank="species" species="karlitae">
<emphasis id="B931EA8D35415479FCA1E79E8DAF4B2C" box="[880,999,1014,1039]" italics="true" pageId="1" pageNumber="99">A. karlitae</emphasis>
</taxonomicName>
is “guallache serrano” and local people appreciate the high quality and rot-resistant wood. The inner bark and wood are aromatic and expel a delicious citrus scent; wood retains the smell even after several years after having been cut.
</paragraph>
<paragraph id="8BFA369F35415479FF6CE00A8B224B58" blockId="1.[136,1452,726,1291]" box="[189,1386,1122,1147]" pageId="1" pageNumber="99">
<emphasis id="B931EA8D35415479FF6CE00A8F0B4B58" bold="true" box="[189,323,1122,1147]" pageId="1" pageNumber="99">Phenology:</emphasis>
—The flowering period occurs between August and September. Fruits have not been observed.
</paragraph>
<paragraph id="8BFA369F35415479FF6CE0EE8FDC4A28" blockId="1.[136,1452,726,1291]" pageId="1" pageNumber="99">
<emphasis id="B931EA8D35415479FF6CE0EE8C184BBC" bold="true" box="[189,592,1158,1183]" pageId="1" pageNumber="99">
Specimens examined (
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):
</emphasis>
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<collectingCountry id="F352760F35415479FDBCE0EE8CBF4BBC" box="[621,759,1158,1183]" name="Ecuador" pageId="1" pageNumber="99">ECUADOR</collectingCountry>
.
<collectingRegion id="4981F87D35415479FCD3E0EE8D724BBC" box="[770,826,1158,1183]" country="Ecuador" name="Loja" pageId="1" pageNumber="99">Loja</collectingRegion>
:
<collectingCounty id="629B4E1335415479FC95E0EE8DD34BBC" box="[836,923,1158,1183]" pageId="1" pageNumber="99">Macará</collectingCounty>
,
<collectingMunicipality id="6B9EACE535415479FC77E0EE8A594BBC" box="[934,1041,1158,1183]" pageId="1" pageNumber="99">Sabiango</collectingMunicipality>
,
<location id="8E9A604435415479FBCAE0EE8A3F4BBC" LSID="urn:lsid:plazi:treatment:03EC878935405479FF59E1AC8FDC4A28:8E9A604435415479FBCAE0EE8A3F4BBC" box="[1051,1143,1158,1183]" country="Ecuador" county="Macara" municipality="Sabiango" name="Achima" pageId="1" pageNumber="99" stateProvince="Loja">Achima</location>
,
<location id="8E9A604435415479FB53E0EE8B124BBC" LSID="urn:lsid:plazi:treatment:03EC878935405479FF59E1AC8FDC4A28:8E9A604435415479FB53E0EE8B124BBC" box="[1154,1370,1158,1183]" country="Ecuador" county="Macara" municipality="Sabiango" name="Cerro Jatumpamba" pageId="1" pageNumber="99" stateProvince="Loja">Cerro Jatumpamba</location>
, aprox.
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<elevation id="0068D1AC35415479FF59E0C28E964BE0" box="[136,222,1194,1219]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="99" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
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<collectingDate id="EFBFE9B735415479FF38E0C28FD44BE0" box="[233,412,1194,1219]" pageId="1" pageNumber="99" value="2009-08-29">29 August 2009</collectingDate>
</date>
,
<emphasis id="B931EA8D35415479FE76E0C28C3F4BE0" box="[423,631,1194,1219]" italics="true" pageId="1" pageNumber="99">
<collectionCode id="ED54AE5A35415479FE76E0C28FF74BE0" box="[423,447,1194,1219]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="99" type="Herbarium">W</collectionCode>
. Palacios 16872
</emphasis>
(
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!)
</materialsCitation>
;
<materialsCitation id="3B2D3CC235415479FD23E0C28FD84A28" ID-GBIF-Occurrence="4927585302" collectingDate="2014-08" collectionCode="W, QCNE, AAU, MO, QCA" country="Ecuador" pageId="1" pageNumber="99" specimenCount="1">
<date id="FFFB105F35415479FD23E0C28DCC4BE0" box="[754,900,1194,1219]" pageId="1" pageNumber="99" value="2014-08">
<collectingDate id="EFBFE9B735415479FD23E0C28DCC4BE0" box="[754,900,1194,1219]" pageId="1" pageNumber="99" value="2014-08">August 2014</collectingDate>
</date>
,
<emphasis id="B931EA8D35415479FC5EE0C28A174BE0" box="[911,1119,1194,1219]" italics="true" pageId="1" pageNumber="99">
<collectionCode id="ED54AE5A35415479FC5EE0C28DEF4BE0" box="[911,935,1194,1219]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="99" type="Herbarium">W</collectionCode>
. Palacios 17609
</emphasis>
(
<collectionCode id="ED54AE5A35415479FBBFE0C28A8D4BE0" box="[1134,1221,1194,1219]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13634" lsid="urn:lsid:biocol.org:col:13634" name="Museo Ecuatoriano de Ciencias Naturales" pageId="1" pageNumber="99" type="Herbarium">QCNE</collectionCode>
!),
<emphasis id="B931EA8D35415479FB0AE0C28BE34BE0" box="[1243,1451,1194,1219]" italics="true" pageId="1" pageNumber="99">
<collectionCode id="ED54AE5A35415479FB0AE0C28ABB4BE0" box="[1243,1267,1194,1219]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="99" type="Herbarium">W</collectionCode>
. Palacios 17610
</emphasis>
(
<collectionCode id="ED54AE5A35415479FF41E0A68EAF4BC4" box="[144,231,1230,1255]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13634" lsid="urn:lsid:biocol.org:col:13634" name="Museo Ecuatoriano de Ciencias Naturales" pageId="1" pageNumber="99" type="Herbarium">QCNE</collectionCode>
!),
<emphasis id="B931EA8D35415479FF2FE0A68F984BC4" box="[254,464,1230,1255]" italics="true" pageId="1" pageNumber="99">
<collectionCode id="ED54AE5A35415479FF2FE0A68F5E4BC4" box="[254,278,1230,1255]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="99" type="Herbarium">W</collectionCode>
. Palacios 17612
</emphasis>
(
<collectionCode id="ED54AE5A35415479FE0EE0A68C7E4BC4" box="[479,566,1230,1255]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13634" lsid="urn:lsid:biocol.org:col:13634" name="Museo Ecuatoriano de Ciencias Naturales" pageId="1" pageNumber="99" type="Herbarium">QCNE</collectionCode>
!),
<emphasis id="B931EA8D35415479FD9CE0A68D574BC4" box="[589,799,1230,1255]" italics="true" pageId="1" pageNumber="99">
<collectionCode id="ED54AE5A35415479FD9CE0A68C2D4BC4" box="[589,613,1230,1255]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="99" type="Herbarium">W</collectionCode>
. Palacios 17616
</emphasis>
(
<collectionCode id="ED54AE5A35415479FCFFE0A68DCD4BC4" box="[814,901,1230,1255]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13634" lsid="urn:lsid:biocol.org:col:13634" name="Museo Ecuatoriano de Ciencias Naturales" pageId="1" pageNumber="99" type="Herbarium">QCNE</collectionCode>
!). Duplicates of these specimens to be distributed to
<collectionCode id="ED54AE5A35415479FF72E09A8EAA4A28" box="[163,226,1266,1291]" country="Ethiopia" httpUri="http://grbio.org/cool/5dxx-xet2" name="Addis Ababa University, Department of Biology" pageId="1" pageNumber="99">AAU</collectionCode>
,
<collectionCode id="ED54AE5A35415479FF3FE09A8F534A28" box="[238,283,1266,1291]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="99" type="Herbarium">MO</collectionCode>
and
<collectionCode id="ED54AE5A35415479FE80E09A8FD84A28" box="[337,400,1266,1291]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15650" lsid="urn:lsid:biocol.org:col:15650" name="Pontificia Universidad Católica del Ecuador" pageId="1" pageNumber="99" type="Herbarium">QCA</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>