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<document id="043E2BC5CB83388B843F2D4CC7AB3430" ID-CLB-Dataset="53392" ID-DOI="10.1080/00222933.2023.2198736" ID-GBIF-Dataset="4a877809-d44c-4c07-8f2a-869a6aab577b" ID-ISSN="1464-5262" ID-Zenodo-Dep="7975445" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1685084614132" checkinUser="plazi" docAuthor="Cáceres, Juan Sebastián D. &amp; Grossi, Paschoal Coelho" docDate="2023" docId="972787AFFF9A5805FED65C27FB78FDF7" docLanguage="en" docName="JNATHIST.57.9-12.557-595.pdf" docOrigin="Journal of Natural History 57 (9 - 12)" docSource="http://dx.doi.org/10.1080/00222933.2023.2198736" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Aegognathus aguirei Arnaud and Bomans 2007" docType="treatment" docVersion="3" lastPageNumber="589" masterDocId="6B1EFFD7FF875825FFB2597CFFFCFFE5" masterDocTitle="Taxonomic revision of Aegognathus Leuthner, 1883 (Coleoptera: Lucanidae)" masterLastPageNumber="595" masterPageNumber="557" pageNumber="586" updateTime="1685125870407" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C8153400A3BCB9CFDEC058BD3ED3E6E6">Taxonomic revision of Aegognathus Leuthner, 1883 (Coleoptera: Lucanidae)</mods:title>
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<mods:namePart id="D71AB1C7BC7BC492E60F309799467084">Cáceres, Juan Sebastián D.</mods:namePart>
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<mods:namePart id="2AAE3A0AF61978ACA487B7EF23612241">Grossi, Paschoal Coelho</mods:namePart>
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<mods:title id="A9E3AA8AC03A75CA8039D35BCD8871D9">Journal of Natural History</mods:title>
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<mods:date id="7B8E277FF5951490E7FDB1D7F55D9F96">2023</mods:date>
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<mods:number id="541EAA0252F718B64FC0969892EE34CA">2023-05-18</mods:number>
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<mods:number id="ED8C7CB7C7B58F1DC9C1772B6041367B">57</mods:number>
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<mods:number id="0AF575FAFD4D1A79A7BCDD7D04A1CF94">9 - 12</mods:number>
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<treatment id="972787AFFF9A5805FED65C27FB78FDF7" ID-DOI="http://doi.org/10.5281/zenodo.7975487" ID-Zenodo-Dep="7975487" LSID="urn:lsid:plazi:treatment:972787AFFF9A5805FED65C27FB78FDF7" httpUri="http://treatment.plazi.org/id/972787AFFF9A5805FED65C27FB78FDF7" lastPageId="32" lastPageNumber="589" pageId="29" pageNumber="586">
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<paragraph id="1F3136B9FF9A5838FED65C27FC43FA93" blockId="29.[356,959,1371,1398]" box="[356,959,1371,1398]" pageId="29" pageNumber="586">
<heading id="447981D5FF9A5838FED65C27FC43FA93" box="[356,959,1371,1398]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="586" reason="5">
<taxonomicName id="D88E4D3AFF9A5838FED65C27FC43FA93" ID-CoL="65947" authority="Arnaud and Bomans, 2007" authorityName="Arnaud and Bomans" authorityYear="2007" box="[356,959,1371,1398]" class="Insecta" family="Lucanidae" genus="Aegognathus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="586" phylum="Arthropoda" rank="species" species="aguirei">
<emphasis id="2DFAEAABFF9A5838FED65C27FD8CFA93" bold="true" box="[356,624,1371,1398]" italics="true" pageId="29" pageNumber="586">Aegognathus aguirei</emphasis>
<bibRefCitation id="7B1F4B48FF9A5838FDCA5C27FC43FA93" author="Arnaud P &amp; Bomans H" box="[632,959,1371,1398]" pageId="29" pageNumber="586" pagination="2 - 5" refId="ref16360" refString="Arnaud P, Bomans H. 2007. Descriptions de trois nouvelles especes de Coleopteres Lucanidae du Perou. Besoiro. 16: 2 - 5." type="journal article" year="2007">Arnaud and Bomans, 2007</bibRefCitation>
</taxonomicName>
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</paragraph>
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<subSubSection id="57946532FF9A5838FDA15CDFFCECFA58" box="[531,784,1443,1469]" pageId="29" pageNumber="586" type="description">
<paragraph id="1F3136B9FF9A5838FDA15CDFFCECFA58" blockId="29.[531,784,1443,1469]" box="[531,784,1443,1469]" pageId="29" pageNumber="586">
(
<figureCitation id="87B52A3CFF9A5838FDA95CDFFD5AFA58" box="[539,678,1443,1469]" captionStart="Figure 5" captionStartId="24.[160,225,1462,1485]" captionTargetBox="[165,1152,144,1429]" captionTargetId="figure-3@24.[163,1153,142,1430]" captionTargetPageId="24" captionText="Figure 5. Dorsal habitus of Aegognathus damasoi Arnaud and Bomans, 2006: (a) Male and (b) female (CERPE).Aegognathus aguirei Arnaud and Bomans, 2007: (c) male and (d) female (CERPE).Aegognathus dulima Cáceres, Ríos-Málaver and Grossi, 2019: (e) holotype male (CERPE). Aegognathus arnaudi Cáceres, Ríos-Málaver and Grossi, 2019: (f) holotype male and (g) paratype female (CERPE). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7975459" httpUri="https://zenodo.org/record/7975459/files/figure.png" pageId="29" pageNumber="586">Figure 5c–d</figureCitation>
: 1J,
<specimenCount id="0988FD30FF9A5838FD575CDFFCF4FA59" box="[741,776,1443,1469]" pageId="29" pageNumber="586" type="female">1♀</specimenCount>
)
</paragraph>
</subSubSection>
<subSubSection id="57946532FF9A5838FF125C9DFB7FFA1F" box="[160,1155,1505,1530]" pageId="29" pageNumber="586" type="reference_group">
<paragraph id="1F3136B9FF9A5838FF125C9DFB7FFA1F" blockId="29.[160,1155,1505,1530]" box="[160,1155,1505,1530]" pageId="29" pageNumber="586">
<treatmentCitationGroup id="3F9E1197FF9A5838FF125C9DFB7FFA1F" box="[160,1155,1505,1530]" pageId="29" pageNumber="586">
<treatmentCitation id="9E2F10A8FF9A5838FF125C9DFD30FA1F" author="Arnaud P &amp; Bomans H" box="[160,716,1505,1530]" page="2" pageId="29" pageNumber="586" year="2007">
<taxonomicName id="D88E4D3AFF9A5838FF125C9DFD30FA1F" ID-CoL="65947" authority="Arnaud and Bomans, 2007: 2" authorityName="Arnaud and Bomans" authorityPageNumber="2" authorityYear="2007" box="[160,716,1505,1530]" class="Insecta" family="Lucanidae" genus="Aegognathus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="586" phylum="Arthropoda" rank="species" species="aguirei">
<emphasis id="2DFAEAABFF9A5838FF125C9DFE7EFA1F" box="[160,386,1505,1530]" italics="true" pageId="29" pageNumber="586">Aegognathus aguirei</emphasis>
<bibRefCitation id="7B1F4B48FF9A5838FE3B5C9DFD30FA1F" author="Arnaud P &amp; Bomans H" box="[393,716,1505,1530]" pageId="29" pageNumber="586" pagination="2 - 5" refId="ref16360" refString="Arnaud P, Bomans H. 2007. Descriptions de trois nouvelles especes de Coleopteres Lucanidae du Perou. Besoiro. 16: 2 - 5." type="journal article" year="2007">Arnaud and Bomans, 2007: 2</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original combination and description)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="57946532FF9A583BFF125F49FF0EFEF5" lastPageId="30" lastPageNumber="587" pageId="29" pageNumber="586" type="materials_examined">
<paragraph id="1F3136B9FF9A5838FF125F49FE7CF9AA" blockId="29.[160,1156,1589,1685]" box="[160,384,1589,1615]" pageId="29" pageNumber="586">
<emphasis id="2DFAEAABFF9A5838FF125F49FE7CF9AA" bold="true" box="[160,384,1589,1615]" italics="true" pageId="29" pageNumber="586">Examined material</emphasis>
</paragraph>
<paragraph id="1F3136B9FF9A5838FF125F25FC89F970" blockId="29.[160,1156,1589,1685]" pageId="29" pageNumber="586">
<materialsCitation id="AFE63CE4FF9A5838FF125F25FEDCF995" box="[160,288,1625,1650]" collectingDate="2006-11" collectionCode="A, CPFA" collectorName="Damaso" country="Peru" elevation="1800" location="Chalcos" pageId="29" pageNumber="586" specimenCount="1" specimenCount-female="1" stateProvince="Ayacucho" typeStatus="holotype">
<typeStatus id="C035881BFF9A5838FF125F25FEFBF997" box="[160,263,1625,1650]" pageId="29" pageNumber="586">Holotype</typeStatus>
J
</materialsCitation>
<materialsCitation id="AFE63CE4FF9A5838FE945F25FED3F970" collectingDate="2006-11" collectionCode="A, CPFA" collectorName="Damaso" country="Peru" elevation="1800" location="Chalcos" pageId="29" pageNumber="586" specimenCount="1" specimenCount-female="1" stateProvince="Ayacucho" typeStatus="holotype">
and
<typeStatus id="C035881BFF9A5838FEE55F25FE4DF997" box="[343,433,1625,1650]" pageId="29" pageNumber="586">allotype</typeStatus>
<specimenCount id="0988FD30FF9A5838FE045F25FE36F995" box="[438,458,1625,1648]" pageId="29" pageNumber="586" type="female">♀</specimenCount>
:
<collectingCountry id="67997629FF9A5838FE675F25FDF7F997" box="[469,523,1625,1650]" name="Peru" pageId="29" pageNumber="586">Peru</collectingCountry>
,
<collectingRegion id="DD4AF85BFF9A5838FDA65F25FD79F997" box="[532,645,1625,1650]" country="Peru" name="Ayacucho" pageId="29" pageNumber="586">Ayacucho</collectingRegion>
, dist.
<location id="1A516062FF9A5838FD725F25FCE5F997" LSID="urn:lsid:plazi:treatment:972787AFFF9A5805FED65C27FB78FDF7:1A516062FF9A5838FD725F25FCE5F997" box="[704,793,1625,1650]" country="Peru" name="Chalcos" pageId="29" pageNumber="586" stateProvince="Ayacucho">Chalcos</location>
,
<quantity id="D8769B5CFF9A5838FC905F25FC8AF997" box="[802,886,1625,1650]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="586" unit="m" value="1800.0">
<elevation id="94A3D18AFF9A5838FC905F25FC8AF997" box="[802,886,1625,1650]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="586" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="6B301079FF9A5838FC325F25FC2EF997" box="[896,978,1625,1650]" pageId="29" pageNumber="586" value="2006-11">
<collectingDate id="7B74E991FF9A5838FC325F25FC2EF997" box="[896,978,1625,1650]" pageId="29" pageNumber="586" value="2006-11">xi.2006</collectingDate>
</date>
,
<collectionCode id="799FAE7CFF9A5838FC685F25FC11F997" box="[986,1005,1625,1650]" pageId="29" pageNumber="586">A</collectionCode>
.
<collectorName id="B27B536FFF9A5838FC445F25FBADF997" box="[1014,1105,1625,1650]" pageId="29" pageNumber="586">Damaso</collectorName>
rec., in coll.
<collectionCode id="799FAE7CFF9A5838FF435F00FED3F970" box="[241,303,1660,1685]" pageId="29" pageNumber="586">CPFA</collectionCode>
</materialsCitation>
.
<typeStatus id="C035881BFF9A5838FE8B5F00FE5AF970" box="[313,422,1660,1685]" pageId="29" pageNumber="586">Paratypes</typeStatus>
4J,
<specimenCount id="0988FD30FF9A5838FE6E5F00FE01F976" box="[476,509,1660,1685]" pageId="29" pageNumber="586" type="female">4♀</specimenCount>
, same data as
<typeStatus id="C035881BFF9A5838FD175F00FCF0F970" box="[677,780,1660,1685]" pageId="29" pageNumber="586">holotype</typeStatus>
, in
<collectionCode id="799FAE7CFF9A5838FC815F00FC8DF970" box="[819,881,1660,1685]" pageId="29" pageNumber="586">CPFA</collectionCode>
.
</paragraph>
<paragraph id="1F3136B9FF99583BFF1259F2FE7BFF4D" blockId="30.[160,1155,142,272]" box="[160,391,142,168]" pageId="30" pageNumber="587">
<emphasis id="2DFAEAABFF99583BFF1259F2FE7BFF4D" bold="true" box="[160,391,142,168]" italics="true" pageId="30" pageNumber="587">Additional material</emphasis>
</paragraph>
<paragraph id="1F3136B9FF99583BFF1259CEFF0EFEF5" blockId="30.[160,1155,142,272]" pageId="30" pageNumber="587">
<materialsCitation id="AFE63CE4FF99583BFF1259CEFDAFFF2E" box="[160,595,178,203]" country="Peru" location="Ayacucho" pageId="30" pageNumber="587" specimenCount="1" stateProvince="Ayacucho">
<collectingCountry id="67997629FF99583BFF1259CEFF2AFF2E" box="[160,214,178,203]" name="Peru" pageId="30" pageNumber="587">Peru</collectingCountry>
,
<collectingRegion id="DD4AF85BFF99583BFF5759CEFEABFF2E" box="[229,343,178,203]" country="Peru" name="Ayacucho" pageId="30" pageNumber="587">Ayacucho</collectingRegion>
, Sucre, Chalcos Distr
</materialsCitation>
.,
<quantity id="D8769B5CFF99583BFDD759CEFD3CFF2E" box="[613,704,178,203]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="30" pageNumber="587" unit="m" value="1400.0">1400 m</quantity>
,
<date id="6B301079FF99583BFD6259CEFCBFFF2E" box="[720,835,178,203]" pageId="30" pageNumber="587" value="2007-11-18">
18
<materialsCitation id="AFE63CE4FF99583BFD5E59CEFCFDFF2E" box="[748,769,178,203]" country="Peru" isTailEnumeration="true" location="Ayacucho" pageId="30" pageNumber="587" specimenCount="1" stateProvince="Ayacucho">.xi</materialsCitation>
.2007
</date>
,
<collectionCode id="799FAE7CFF99583BFCE059CEFC99FF2E" box="[850,869,178,203]" pageId="30" pageNumber="587">A</collectionCode>
. Damaso coll
<materialsCitation id="AFE63CE4FF99583BFBB459CEFD73FF08" collectionCode="UNSM" country="Peru" location="District of Chalcos" pageId="30" pageNumber="587" specimenCount="1" specimenCount-female="1" stateProvince="Ayacucho">
. –
<specimenCount id="0988FD30FF99583BFB9C59CEFBACFF2C" box="[1070,1104,178,203]" pageId="30" pageNumber="587" type="female">1♀</specimenCount>
, 3J (
<collectionCode id="799FAE7CFF99583BFF1459A8FF0DFF08" box="[166,241,212,237]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="30" pageNumber="587">UNSM</collectionCode>
);
<collectingCountry id="67997629FF99583BFEB059A8FEC4FF08" box="[258,312,212,237]" name="Peru" pageId="30" pageNumber="587">Peru</collectingCountry>
,
<collectingRegion id="DD4AF85BFF99583BFEF059A8FE4FFF08" box="[322,435,212,237]" country="Peru" name="Ayacucho" pageId="30" pageNumber="587">Ayacucho</collectingRegion>
,
<location id="1A516062FF99583BFE0F59A8FD73FF08" LSID="urn:lsid:plazi:treatment:972787AFFF9A5805FED65C27FB78FDF7:1A516062FF99583BFE0F59A8FD73FF08" box="[445,655,212,237]" country="Peru" name="District of Chalcos" pageId="30" pageNumber="587" stateProvince="Ayacucho">District of Chalcos</location>
</materialsCitation>
.,
<quantity id="D8769B5CFF99583BFD2E59A8FD0FFF08" box="[668,755,212,237]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="30" pageNumber="587" unit="m" value="1800.0">1800 m</quantity>
,
<date id="6B301079FF99583BFD4C59A8FC96FF08" box="[766,874,212,237]" pageId="30" pageNumber="587" value="2007-02-19">
19
<materialsCitation id="AFE63CE4FF99583BFCA959A8FCDBFF08" box="[795,807,212,237]" collectionCode="UNSM" country="Peru" isTailEnumeration="true" location="District of Chalcos" pageId="30" pageNumber="587" specimenCount="1" stateProvince="Ayacucho">.ii</materialsCitation>
.2007
</date>
, D
<materialsCitation id="AFE63CE4FF99583BFC3B59A8FBEEFF08" box="[905,1042,212,237]" collectionCode="UNSM" collectorName="Aguirre" country="Peru" isTailEnumeration="true" location="District of Chalcos" pageId="30" pageNumber="587" specimenCount="1" stateProvince="Ayacucho">
.
<collectorName id="B27B536FFF99583BFC2659A8FC1BFF08" box="[916,999,212,237]" pageId="30" pageNumber="587">Aguirre</collectorName>
col
</materialsCitation>
<materialsCitation id="AFE63CE4FF99583BFBA059A8FF12FEF5" collectionCode="EPGC" country="Peru" pageId="30" pageNumber="587" specimenCount="2" specimenCount-female="2" stateProvince="Ayacucho">
. –
<specimenCount id="0988FD30FF99583BFB8059A8FBA8FF09" box="[1074,1108,212,237]" pageId="30" pageNumber="587" type="female">2♀</specimenCount>
, 2J (
<collectionCode id="799FAE7CFF99583BFF17598BFF14FEF5" box="[165,232,247,272]" pageId="30" pageNumber="587">EPGC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="57946532FF99583BFF125840FC6CFD55" pageId="30" pageNumber="587" type="diagnosis">
<paragraph id="1F3136B9FF99583BFF125840FEE8FEB3" blockId="30.[160,1156,316,688]" box="[160,276,316,342]" pageId="30" pageNumber="587">
<emphasis id="2DFAEAABFF99583BFF125840FEE8FEB3" bold="true" box="[160,276,316,342]" italics="true" pageId="30" pageNumber="587">Diagnosis</emphasis>
</paragraph>
<paragraph id="1F3136B9FF99583BFF125823FC6CFD55" blockId="30.[160,1156,316,688]" pageId="30" pageNumber="587">
Males dorsally dark red with carmine to bluish tones and black spots, ventrally black. Head shape trapezoidal, wider than elytra and larger compared with other
<taxonomicName id="D88E4D3AFF99583BFC2558FEFB7CFE7E" box="[919,1152,386,411]" pageId="30" pageNumber="587">
<emphasis id="2DFAEAABFF99583BFC2558FEFBDBFE7E" box="[919,1063,386,411]" italics="true" pageId="30" pageNumber="587">Aegognathus</emphasis>
species
</taxonomicName>
; temporal process rounded and somewhat lamellate; mandibles almost straight and wide. Pronotum wider than elytra. Elytral surface with coarse, dense to contiguous punctures, setae varying from dendritic to smooth. Gena with two oblique carinae besides the one at theposterior margin, present in all members of the genus; abdominal ventrite II laterally wrinkled. Dorsal cross bar of aedeagus slightly lobed posteriorly. Females with normal size heads, showing darker tones along whole body; body surface with coarse, dense to contiguous punctation; elytra with distinct yellowish setae laterally; gena lacking oblique carinae, instead with a strong and conspicuous, coarse punctation.
</paragraph>
</subSubSection>
<subSubSection id="57946532FF99583AFF125BA0FDD4F97B" lastPageId="31" lastPageNumber="588" pageId="30" pageNumber="587" type="description">
<paragraph id="1F3136B9FF99583BFF125BA0FE82FD13" blockId="30.[160,1156,732,1728]" box="[160,382,732,758]" pageId="30" pageNumber="587">
<emphasis id="2DFAEAABFF99583BFF125BA0FE82FD13" bold="true" box="[160,382,732,758]" italics="true" pageId="30" pageNumber="587">Description – male</emphasis>
</paragraph>
<paragraph id="1F3136B9FF99583AFF125B83FB78FD4A" blockId="30.[160,1156,732,1728]" lastBlockId="31.[160,1156,142,687]" lastPageId="31" lastPageNumber="588" pageId="30" pageNumber="587">
<emphasis id="2DFAEAABFF99583BFF125B83FF28FCFD" bold="true" box="[160,212,767,792]" pageId="30" pageNumber="587">
<emphasis id="2DFAEAABFF99583BFF125B83FF31FCFD" bold="true" box="[160,205,767,792]" italics="true" pageId="30" pageNumber="587">Size</emphasis>
:
</emphasis>
<emphasis id="2DFAEAABFF99583BFF695B83FEA1FCFD" box="[219,349,767,792]" italics="true" pageId="30" pageNumber="587">Total length</emphasis>
.
<quantity id="D8769B5CFF99583BFEDB5B83FE08FCFD" box="[361,500,767,792]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" metricValueMax="1.6" metricValueMin="1.56" pageId="30" pageNumber="587" unit="mm" value="15.8" valueMax="16.0" valueMin="15.6">15.6–16 mm</quantity>
.
<emphasis id="2DFAEAABFF99583BFE4D5B83FD85FCFD" box="[511,633,767,792]" italics="true" pageId="30" pageNumber="587">Total width</emphasis>
.
<quantity id="D8769B5CFF99583BFD345B83FD18FCFD" box="[646,740,767,792]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="30" pageNumber="587" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
.
<emphasis id="2DFAEAABFF99583BFD5D5B83FCC7FCFD" bold="true" box="[751,827,767,792]" italics="true" pageId="30" pageNumber="587">Colour</emphasis>
: In dorsal view dark red, with carmine to bluish tones and black spots. Head mainly black except for a dark carmine region medially. Pronotum dark red with two black spots. Elytra carmine with bluish reflections along disc, darker close to margins. Legs seem black at first glance but on closer examination are dark red. Ventrally mainly black with some dark red regions along the gula and coxae.
<emphasis id="2DFAEAABFF99583BFE915AD1FEA3FC23" bold="true" box="[291,351,941,966]" italics="true" pageId="30" pageNumber="587">Head</emphasis>
: Shape somewhat trapezoidal, emarginate anteriorly, wider than elytra and larger compared with other
<taxonomicName id="D88E4D3AFF99583BFDB35AB3FD1AFC0D" box="[513,742,975,1000]" pageId="30" pageNumber="587">
<emphasis id="2DFAEAABFF99583BFDB35AB3FD72FC0D" box="[513,654,975,1000]" italics="true" pageId="30" pageNumber="587">Aegognathus</emphasis>
species
</taxonomicName>
; surface shagreened, mainly glabrous, anterior margin with an interrupted band of yellowish setae; temporal process distinct, rounded and somewhat lamellate. Mandibles as long as the head and the anterior portion of pronotum, less curved compared with other members of the genus, wide, with two rounded basal teeth, one located more over the anterior third, rounded apically.
<emphasis id="2DFAEAABFF99583BFBB95D26FB81FB96" bold="true" box="[1035,1149,1114,1139]" italics="true" pageId="30" pageNumber="587">Pronotum</emphasis>
: Wider than elytra; surface shagreened with some moderate to somewhat large punctures along the posterior region, with yellowish short setae along lateral and posterior margins; anterior angles rounded and somewhat convex; sides convergent posteriorly; posterior angles rounded.
<emphasis id="2DFAEAABFF99583BFEEB5D99FE60FB1B" bold="true" box="[345,412,1253,1278]" italics="true" pageId="30" pageNumber="587">Elytra</emphasis>
: Surface strongly punctured, punctures coarse, dense to somewhat contiguous at some striae, with yellowish dendritic setae mainly distributed along disc, becoming smooth and suberect over lateral and posterior regions, setae more distinct closer to margins. Epipleuron with large to coarse punctures, with short yellowish setae.
<emphasis id="2DFAEAABFF99583BFF125C13FF28FA6D" bold="true" box="[160,212,1391,1416]" italics="true" pageId="30" pageNumber="587">Legs</emphasis>
: Protibiae externally serrate with variable number of teeth but presenting at least two strong ones at apex, one internal tooth and one apical spur. Mesotibiae with two median teeth, when two present the second one reduced, four apical teeth and two apical spurs. Metatibiae with one median tooth, apically as described for mesotibiae.
<emphasis id="2DFAEAABFF99583BFC795CABFBEBFA15" bold="true" box="[971,1047,1495,1520]" italics="true" pageId="30" pageNumber="587">Venter</emphasis>
: Mentum trapezoidal with a weak transverse groove present in some individuals; surface punctured, punctures moderate to large, with yellowish curved to somewhat appressed smooth setae. Gena with two oblique carinae, also with a curved one along the posterior margin, strongly but not densely punctured, punctures large over posterior region, coarse anteriorly, glabrous to poorly covered by setae. Prosternum strongly punctate, punctures large to coarse, dense, mainly distributed along lateral portions, with yellowish appressed setae. Mesoventrite punctate, punctures moderate to large, dense, mainly distributed medially, with yellowish appressed to suberect setae. Metaventrite punctate mainly along anterior margin and lateral portions, punctures large to somewhat coarse, with yellowish curved setae; discrimen present as a continuous suture, becoming a distinct groove posteriorly. Abdominal ventrites mainly punctate laterally, punctures large to somewhat coarse, densely distributed at lateral portions, becoming sparse medially, with yellowish curved to appressed setae, and a somewhat continuous discrete longitudinal carina over lateral margins of every ventrite, ventrite II wrinkled.
<emphasis id="2DFAEAABFF98583AFD2558FDFD03FE7F" bold="true" box="[663,767,385,410]" italics="true" pageId="31" pageNumber="588">Genitalia</emphasis>
(
<figureCitation id="87B52A3CFF98583AFCBF58FDFC8EFE7F" box="[781,882,385,410]" captionStart="Figure 8" captionStartId="28.[160,225,1699,1722]" captionTargetBox="[175,1150,145,1651]" captionTargetId="figure-3@28.[163,1153,142,1667]" captionTargetPageId="28" captionText="Figure 8. Ventral view of A. damasoi (a) male aedeagus and (b) female genitalia. Aegognathus aguirei (c) male aedeagus and (d) female genitalia. (e) Aegognathus dulima male aedeagus. Aegognathus arnaudi: (f) male aedeagus and (g) female genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7975465" httpUri="https://zenodo.org/record/7975465/files/figure.png" pageId="31" pageNumber="588">Figure 8c</figureCitation>
): Aedeagus symmetrical, subtriangular, somewhat convex along the proximal region and slightly divergent distally. Basal piece subtriangular with slightly convex sides; slightly convex in lateral view, disc translucent, 1.0 times longer than the parameres. Median lobe almost cylindrical, slightly convex distally, laterally punctured, in dorsal view with ′X̍-shaped cross bar struts that are slightly lobed posteriorly. Parameres rounded over the posterior margin, setose, almost straight in lateral view. Everted sac 1.5 times longer than the basal piece and parameres together, with two strap-like sclerites that end at the beginning of the apical third, apex subconical with concave lateral margins over the median portion, ending in the gonopore.
</paragraph>
<paragraph id="1F3136B9FF98583AFF125BA0FE69FD13" blockId="31.[160,1157,732,1694]" box="[160,405,732,758]" pageId="31" pageNumber="588">
<emphasis id="2DFAEAABFF98583AFF125BA0FE69FD13" bold="true" box="[160,405,732,758]" italics="true" pageId="31" pageNumber="588">Description – female</emphasis>
</paragraph>
<paragraph id="1F3136B9FF98583AFF125B83FDD4F97B" blockId="31.[160,1157,732,1694]" pageId="31" pageNumber="588">
<emphasis id="2DFAEAABFF98583AFF125B83FF2AFCFD" bold="true" box="[160,214,767,792]" pageId="31" pageNumber="588">
<emphasis id="2DFAEAABFF98583AFF125B83FF33FCFD" bold="true" box="[160,207,767,792]" italics="true" pageId="31" pageNumber="588">Size</emphasis>
:
</emphasis>
<emphasis id="2DFAEAABFF98583AFF535B83FE95FCFD" box="[225,361,767,792]" italics="true" pageId="31" pageNumber="588">Total length</emphasis>
.
<quantity id="D8769B5CFF98583AFEC85B83FDF0FCFD" box="[378,524,767,792]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" metricValueMax="1.56" metricValueMin="1.2" pageId="31" pageNumber="588" unit="mm" value="13.8" valueMax="15.6" valueMin="12.0">12–15.6 mm</quantity>
.
<emphasis id="2DFAEAABFF98583AFDA85B83FD60FCFD" box="[538,668,767,792]" italics="true" pageId="31" pageNumber="588">Total width</emphasis>
.
<quantity id="D8769B5CFF98583AFD195B83FCCAFCFD" box="[683,822,767,792]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.5" metricValueMin="4.7" pageId="31" pageNumber="588" unit="mm" value="5.1" valueMax="5.5" valueMin="4.7">4.7–5.5 mm</quantity>
.
<emphasis id="2DFAEAABFF98583AFCF65B83FC6FFCFD" bold="true" box="[836,915,767,792]" italics="true" pageId="31" pageNumber="588">Colour</emphasis>
: In dorsal view dark red, with darker tones along the head, and two large spots at pronotum. Elytra with dark red and black longitudinal interleaved lines. Legs presenting dark red tones distally. Ventrally black with a few dark red regions.
<emphasis id="2DFAEAABFF98583AFCBC5A1BFCB0FC65" bold="true" box="[782,844,871,896]" italics="true" pageId="31" pageNumber="588">Head</emphasis>
: Smaller than male heads, about a half, not including the mandibles; surface highly punctate, punctures coarse, dense to contiguous with minute yellowish setae, also with some slightly carinate areas close to the anterior angles, and over the disc; anterior margin emarginate with an interrupted line of yellowish suberect setae; temporal process minute; mandibles reduced, being as long as the first half of the head, with acuminate apex and a median laminar tooth, asymmetrical, laminar teeth with three acuminate projections, more distinct at left mandible.
<emphasis id="2DFAEAABFF98583AFDBC5D26FD78FB96" bold="true" box="[526,644,1114,1139]" italics="true" pageId="31" pageNumber="588">Pronotum</emphasis>
: Surface strongly punctate, punctures large to coarse, densely distributed along margins and somewhat contiguous at disc, with yellowish erect setae, mainly visible laterally; lateral margins somewhat convergent and rounded anteriorly, posterior margin as wide as elytra.
<emphasis id="2DFAEAABFF98583AFC305DBEFC3BFB3E" bold="true" box="[898,967,1218,1243]" italics="true" pageId="31" pageNumber="588">Elytra</emphasis>
: Surface highly punctate, punctures coarse, contiguous at disc and densely distributed along margins, with yellowish, curved, dendritic setae, more distinct along distal portions; weak irregular longitudinal costae present over each elytra.
<emphasis id="2DFAEAABFF98583AFC9F5C56FC9EFAA6" bold="true" box="[813,866,1322,1347]" italics="true" pageId="31" pageNumber="588">Legs</emphasis>
: As in males except for presenting two to three median teeth at mesotibiae and one at metatibiae.
<emphasis id="2DFAEAABFF98583AFB9D5C31FB82FA83" bold="true" box="[1071,1150,1357,1382]" italics="true" pageId="31" pageNumber="588">Venter</emphasis>
: As in males except for presenting more distinct punctation along whole venter; mentum and gena with coarse punctures, gena not carinate; discrimen present as a continuous suture at the metaventrite. Abdominal ventrites almost regular punctate, with yellowish appressed setae, and a somewhat continuous discrete longitudinal carina along lateral margins of every ventrite.
<emphasis id="2DFAEAABFF98583AFCF05C86FC52F9F6" bold="true" box="[834,942,1530,1555]" italics="true" pageId="31" pageNumber="588">Genitalia</emphasis>
(
<figureCitation id="87B52A3CFF98583AFC765C86FBCBF9F6" box="[964,1079,1530,1555]" captionStart="Figure 8" captionStartId="28.[160,225,1699,1722]" captionTargetBox="[175,1150,145,1651]" captionTargetId="figure-3@28.[163,1153,142,1667]" captionTargetPageId="28" captionText="Figure 8. Ventral view of A. damasoi (a) male aedeagus and (b) female genitalia. Aegognathus aguirei (c) male aedeagus and (d) female genitalia. (e) Aegognathus dulima male aedeagus. Aegognathus arnaudi: (f) male aedeagus and (g) female genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7975465" httpUri="https://zenodo.org/record/7975465/files/figure.png" pageId="31" pageNumber="588">Figure 8d</figureCitation>
): Styli symmetrical, almost quadrate, with a slightly concave apex and fine setae. Distal coxite with convex internal sides, externally somewhat sinuous. Proximal coxite almost straight, distally with a concave portion. Lateral paraproct with a truncate lateral extension, basally straight.
</paragraph>
</subSubSection>
<subSubSection id="57946532FFA75805FF1259F2FC2EFF2E" box="[160,978,142,203]" pageId="32" pageNumber="589" type="distribution">
<paragraph id="1F3136B9FFA75805FF1259F2FED0FF4D" blockId="32.[160,978,142,203]" box="[160,300,142,168]" pageId="32" pageNumber="589">
<emphasis id="2DFAEAABFFA75805FF1259F2FED0FF4D" bold="true" box="[160,300,142,168]" italics="true" pageId="32" pageNumber="589">Distribution</emphasis>
</paragraph>
<paragraph id="1F3136B9FFA75805FF1259CEFC2EFF2E" blockId="32.[160,978,142,203]" box="[160,978,178,203]" pageId="32" pageNumber="589">
Southern
<collectingCountry id="67997629FFA75805FEBC59CEFEB8FF2E" box="[270,324,178,203]" name="Peru" pageId="32" pageNumber="589">Peru</collectingCountry>
, Central Cordillera. Yungas province (
<bibRefCitation id="7B1F4B48FFA75805FD5A59CEFC3AFF2E" author="Morrone JJ &amp; Escalante T &amp; Rodrigiuez-Tapia G &amp; Carmona A &amp; Arana M &amp; Mercado-Gomez JD" box="[744,966,178,203]" pageId="32" pageNumber="589" pagination="1 - 5" refId="ref17309" refString="Morrone JJ, Escalante T, Rodrigiuez-Tapia G, Carmona A, Arana M, Mercado-Gomez JD. 2022. Biogeographic regionalization of the Neotropical region: new map and shapefile. Anais da Academia Brasileira de Ciencias. 94 (1): 1 - 5. doi: 10.1590 / 0001 - 3765202220211167." type="journal article" year="2022">Morrone et al. 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="57946532FFA75805FF125879FB78FDF7" pageId="32" pageNumber="589" type="discussion">
<paragraph id="1F3136B9FFA75805FF125879FEFAFEFA" blockId="32.[160,1156,261,530]" box="[160,262,261,287]" pageId="32" pageNumber="589">
<emphasis id="2DFAEAABFFA75805FF125879FEFAFEFA" bold="true" box="[160,262,261,287]" italics="true" pageId="32" pageNumber="589">Remarks</emphasis>
</paragraph>
<paragraph id="1F3136B9FFA75805FF125855FB78FDF7" blockId="32.[160,1156,261,530]" pageId="32" pageNumber="589">
<taxonomicName id="D88E4D3AFFA75805FF125855FEF4FEA7" authorityName="Arnaud and Bomans" authorityYear="2007" box="[160,264,297,322]" class="Insecta" family="Lucanidae" genus="Aegognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="589" phylum="Arthropoda" rank="species" species="aguirei">
<emphasis id="2DFAEAABFFA75805FF125855FEF4FEA7" box="[160,264,297,322]" italics="true" pageId="32" pageNumber="589">A. aguirei</emphasis>
</taxonomicName>
shows colour patterns similar to those shown by several species of the genus, but it is probably most easily recognisable by its males with wider head, mandibles not curved and wide, temporal process somewhat lamellate, and a gena distinctly carinate. In the case of female specimens, the correct identification is more difficult.
<emphasis id="2DFAEAABFFA75805FC4158EDFD98FE28" italics="true" pageId="32" pageNumber="589">
<taxonomicName id="D88E4D3AFFA75805FC4158EDFEFDFE28" authorityName="Van de Poll" authorityYear="1886" class="Insecta" family="Lucanidae" genus="Aegognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="589" phylum="Arthropoda" rank="species" species="leuthneri">Aegognathus leuthneri</taxonomicName>
,
<taxonomicName id="D88E4D3AFFA75805FEBC58C8FE93FE28" authorityName="Arnaud and Bomans" authorityYear="2006" box="[270,367,436,461]" class="Insecta" family="Lucanidae" genus="Aegognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="589" phylum="Arthropoda" rank="species" species="similis">A. similis</taxonomicName>
,
<taxonomicName id="D88E4D3AFFA75805FECF58C8FE06FE28" authorityName="Arnaud and Bomans" authorityYear="2006" box="[381,506,436,461]" class="Insecta" family="Lucanidae" genus="Aegognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="589" phylum="Arthropoda" rank="species" species="damasoi">A. damasoi</taxonomicName>
,
<taxonomicName id="D88E4D3AFFA75805FDB558C8FD98FE28" authorityName="Arnaud and Bomans" authorityYear="2004" box="[519,612,436,461]" class="Insecta" family="Lucanidae" genus="Aegognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="589" phylum="Arthropoda" rank="species" species="soulai">A. soulai</taxonomicName>
</emphasis>
and
<taxonomicName id="D88E4D3AFFA75805FD1358C8FCF6FE28" authorityName="Arnaud and Bomans" authorityYear="2007" box="[673,778,436,461]" class="Insecta" family="Lucanidae" genus="Aegognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="589" phylum="Arthropoda" rank="species" species="aguirei">
<emphasis id="2DFAEAABFFA75805FD1358C8FCF6FE28" box="[673,778,436,461]" italics="true" pageId="32" pageNumber="589">A. aguirei</emphasis>
</taxonomicName>
are all very similar, and the best characteristic to separate
<taxonomicName id="D88E4D3AFFA75805FE7E58AAFDCAFE0A" authorityName="Arnaud and Bomans" authorityYear="2007" box="[460,566,470,495]" class="Insecta" family="Lucanidae" genus="Aegognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="589" phylum="Arthropoda" rank="species" species="aguirei">
<emphasis id="2DFAEAABFFA75805FE7E58AAFDCAFE0A" box="[460,566,470,495]" italics="true" pageId="32" pageNumber="589">A. aguirei</emphasis>
</taxonomicName>
female from females of other species is likely the strong coarse punctation at the gena, that replaces the oblique carinae present in males.
</paragraph>
</subSubSection>
</treatment>
</document>
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