128 lines
19 KiB
XML
128 lines
19 KiB
XML
<document id="9104E4B18DE9F7D608DD3833B66680AD" ID-DOI="10.5281/zenodo.5174192" ID-ISSN="1942-1354" ID-Zenodo-Dep="5174192" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628335183268" checkinUser="felipe" docAuthor="Mawdsley, Jonathan R." docDate="2012" docId="2B55EA35FFEAFFCDFF30B9BFFCF0FDD8" docLanguage="en" docName="InsectaMundi.0228.1-11.pdf" docOrigin="Insecta Mundi 2012 (228)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Graphipterus distinctus Peringuey 2012, new status" docType="treatment" docVersion="4" lastPageNumber="7" masterDocId="D76C924DFFEDFFC5FFF1BC3EFFE9FFC2" masterDocTitle="A taxonomic review of the ancora species group of Graphipterus Latreille (Coleoptera: Carabidae)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="6" updateTime="1699045096987" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
|
||
<mods:mods id="874BDE19653988791609A9B981845703" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="45C2A3B0F7625FD58479519FDFE9519A">
|
||
<mods:title id="6A9B79063CEF5F076BFE339739227DA7">A taxonomic review of the ancora species group of Graphipterus Latreille (Coleoptera: Carabidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="0E945908DDB6E1DCFDBFF4B3018B5993" type="personal">
|
||
<mods:role id="DE72DA94A31F80FAB882C18D04AC1A03">
|
||
<mods:roleTerm id="9C833E907146090715F0E7B31028DAB2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1EE0FD8C31BC019E083E966DFDBEB12C">Mawdsley, Jonathan R.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="EE85292CDC12E73162E5DD47B01AA381">text</mods:typeOfResource>
|
||
<mods:relatedItem id="627F7BD7718CCB15344BA8FC9362D975" type="host">
|
||
<mods:titleInfo id="71C6FF7B527E655B299731AD97C4FAD5">
|
||
<mods:title id="7D7ACA58CD44C43F5107077A698391E9">Insecta Mundi</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="04A66F2227EEAD48F286ACE2DF29752D">
|
||
<mods:date id="89BD63EFAEE4745DC15D789933F3AFCE">2012</mods:date>
|
||
<mods:detail id="F166D1C0843B856E5D274A9EA9981C91" type="pubDate">
|
||
<mods:number id="4963F49C04791D7EF3897ACE99FDEB23">2012-04-06</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="3590728496F6B81DD736493DD67393F8" type="volume">
|
||
<mods:number id="3856D5C5A92886AB0C4A030B6F99532E">2012</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="AB0CA563657EDEF11D0A008BA6001F32" type="issue">
|
||
<mods:number id="821C6CA46F95EEDE05889BDFD1223403">228</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="E7EE60FC90F32B87A7CF41747F53B858" unit="page">
|
||
<mods:start id="50E43BDACA94E1B20571C5F2EEB3B614">1</mods:start>
|
||
<mods:end id="E9F74C18B0EE06776EC23F9244519BB7">11</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="2AB9B91B5139E6E3F306BAEC9408C0C6">journal article</mods:classification>
|
||
<mods:identifier id="E2C76FCD20209D8DA05845ABE0BCF34E" type="DOI">10.5281/zenodo.5174192</mods:identifier>
|
||
<mods:identifier id="51F7380F9B33458C8D21F3C2B9FFE909" type="ISSN">1942-1354</mods:identifier>
|
||
<mods:identifier id="B9C9FF4D57B1BA4D3859B607015A8AC1" type="Zenodo-Dep">5174192</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="2B55EA35FFEAFFCDFF30B9BFFCF0FDD8" ID-DOI="http://doi.org/10.5281/zenodo.5187076" ID-Zenodo-Dep="5187076" LSID="urn:lsid:plazi:treatment:2B55EA35FFEAFFCDFF30B9BFFCF0FDD8" httpUri="http://treatment.plazi.org/id/2B55EA35FFEAFFCDFF30B9BFFCF0FDD8" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
|
||
<subSubSection id="EBE608A8FFEAFFC2FF30B9BFFCAEFA58" box="[193,839,1409,1434]" pageId="7" pageNumber="6" type="nomenclature">
|
||
<paragraph id="A3435B23FFEAFFC2FF30B9BFFCAEFA58" blockId="7.[193,839,1409,1466]" box="[193,839,1409,1434]" pageId="7" pageNumber="6">
|
||
<heading id="F80BEC4FFFEAFFC2FF30B9BFFCAEFA58" bold="true" box="[193,839,1409,1434]" fontSize="10" level="2" pageId="7" pageNumber="6" reason="3">
|
||
<emphasis id="91888731FFEAFFC2FF30B9BFFCAEFA58" bold="true" box="[193,839,1409,1434]" pageId="7" pageNumber="6">
|
||
<taxonomicName id="64FC20A0FFEAFFC2FF30B9BFFD4CFA58" ID-CoL="3H973" authority="Peringuey" authorityName="Peringuey" authorityYear="2012" box="[193,677,1409,1434]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="distinctus" status="stat. nov.">
|
||
<emphasis id="91888731FFEAFFC2FF30B9BFFDE5FA5B" bold="true" box="[193,524,1409,1433]" italics="true" pageId="7" pageNumber="6">Graphipterus distinctus</emphasis>
|
||
Péringuey
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="8ABB3A4AFFEAFFC2FD43B9BCFCAEFA58" box="[690,839,1410,1434]" pageId="7" pageNumber="6">new status</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EBE608A8FFEAFFC2FF30B99CFEC3FA78" box="[193,298,1442,1466]" pageId="7" pageNumber="6" type="description">
|
||
<paragraph id="A3435B23FFEAFFC2FF30B99CFEC3FA78" blockId="7.[193,839,1409,1466]" box="[193,298,1442,1466]" pageId="7" pageNumber="6">
|
||
<figureCitation id="3BC747A6FFEAFFC2FF30B99CFEC3FA78" box="[193,298,1442,1466]" captionStart="Figures 1–9" captionStartId="5.[192,288,1044,1066]" captionTargetBox="[192,1440,192,1024]" captionTargetId="figure-218@5.[192,1440,192,1024]" captionTargetPageId="5" captionText="Figures 1–9. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 1) G. ancora Dejean [G. ancora ancora Dejean], Hanover, Northern Cape Province, RSA. 2) G. distinctus Péringuey [G. fasciatus distinctus Péringuey], Shilouvane, Limpopo Province, RSA. 3) G. fasciatus Chaudoir [G. fasciatus fasciatus Chaudoir], Umfolozi, KwaZulu/Natal Province, RSA. 4) G. fritschi Chaudoir, Pietersburg, Limpopo Province, RSA. 5) G. westwoodi Brême [G. ancora aurantiacus Basilewsky], Bulawayo, Zimbabwe. 6) G. westwoodi [G. ancora egressus Péringuey], Ohrigstad, Mpumalanga Province, RSA. 7) G. westwoodi [G. ancora subegregius Basilewsky, paratype], Tuli District, Botswana. 8) G. westwoodi [G. ancora transitans Péringuey], Metsimaklaba, Botswana. 9) G. westwoodi [G. ancora westwoodi Brême], Swartruggens Marico, North West Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174194" httpUri="https://zenodo.org/record/5174194/files/figure.png" pageId="7" pageNumber="6">Figure 2</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EBE608A8FFEAFFC2FF30B9DCFE84F9D8" pageId="7" pageNumber="6" type="reference_group">
|
||
<paragraph id="A3435B23FFEAFFC2FF30B9DCFE84F9D8" blockId="7.[193,1442,1506,1562]" pageId="7" pageNumber="6">
|
||
<treatmentCitationGroup id="83EC7C0DFFEAFFC2FF30B9DCFE84F9D8" pageId="7" pageNumber="6">
|
||
<treatmentCitation id="225D7D32FFEAFFC2FF30B9DCFD18FA38" author="Peringuey, L." box="[193,753,1506,1530]" page=")" pageId="7" pageNumber="6" year="1898">
|
||
<taxonomicName id="64FC20A0FFEAFFC2FF30B9DCFD18FA38" ID-CoL="3H973" authority="Peringuey (1898: 340)" authorityName="Peringuey" authorityPageNumber="340" authorityYear="1898" box="[193,753,1506,1530]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="distinctus">
|
||
<emphasis id="91888731FFEAFFC2FF30B9DCFE0AFA38" box="[193,483,1506,1530]" italics="true" pageId="7" pageNumber="6">Graphipterus distinctus</emphasis>
|
||
<bibRefCitation id="C76D26D2FFEAFFC2FE19B9DCFD18FA38" author="Peringuey, L." box="[488,753,1506,1530]" pageId="7" pageNumber="6" pagination="315 - 378" refId="ref8077" refString="Peringuey, L. 1898. Descriptive catalogue of the Coleoptera of South Africa, Family Carabidae, First supplement. Transactions of the South African Philosophical Society 10: 315 - 378." type="journal article" year="1898">Péringuey (1898: 340)</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(
|
||
<typeStatus id="7C47E581FFEAFFC2FD0FB9DCFC99FA38" box="[766,880,1506,1530]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
|
||
, Transvaal, Lydenburg District, South African Museum).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EBE608A8FFEAFFCDFF30BA7FFAE2FEF8" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6" type="diagnosis">
|
||
<paragraph id="A3435B23FFEAFFCDFF30BA7FFAE2FEF8" blockId="7.[193,1442,1601,1914]" lastBlockId="8.[192,1440,194,314]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
|
||
<emphasis id="91888731FFEAFFC2FF30BA7FFEA0F99B" bold="true" box="[193,329,1601,1625]" pageId="7" pageNumber="6">Diagnosis</emphasis>
|
||
. Smaller adults, length
|
||
<quantity id="6404F6C6FFEAFFC2FD85BA7CFD10F99B" box="[628,761,1602,1626]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="7" pageNumber="6" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">11–13 mm</quantity>
|
||
. Pattern of elytral vestiture distinctive (
|
||
<figureCitation id="3BC747A6FFEAFFC2FB18BA7CFABAF998" box="[1257,1363,1602,1626]" captionStart="Figures 1–9" captionStartId="5.[192,288,1044,1066]" captionTargetBox="[192,1440,192,1024]" captionTargetId="figure-218@5.[192,1440,192,1024]" captionTargetPageId="5" captionText="Figures 1–9. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 1) G. ancora Dejean [G. ancora ancora Dejean], Hanover, Northern Cape Province, RSA. 2) G. distinctus Péringuey [G. fasciatus distinctus Péringuey], Shilouvane, Limpopo Province, RSA. 3) G. fasciatus Chaudoir [G. fasciatus fasciatus Chaudoir], Umfolozi, KwaZulu/Natal Province, RSA. 4) G. fritschi Chaudoir, Pietersburg, Limpopo Province, RSA. 5) G. westwoodi Brême [G. ancora aurantiacus Basilewsky], Bulawayo, Zimbabwe. 6) G. westwoodi [G. ancora egressus Péringuey], Ohrigstad, Mpumalanga Province, RSA. 7) G. westwoodi [G. ancora subegregius Basilewsky, paratype], Tuli District, Botswana. 8) G. westwoodi [G. ancora transitans Péringuey], Metsimaklaba, Botswana. 9) G. westwoodi [G. ancora westwoodi Brême], Swartruggens Marico, North West Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174194" httpUri="https://zenodo.org/record/5174194/files/figure.png" pageId="7" pageNumber="6">Figure 2</figureCitation>
|
||
), with the elytra covered in contrasting black and yellowish-grey pubescence. Basal half of elytron with a black linear band of pubescence along suture and a triangular patch of black pubescence completely surrounded by yellowish-grey pubescence; apical half of elytron predominantly covered in black pubescence but with a small ovate patch of yellowish-grey pubescence along the suture at apex, an angled median band of yellowish-grey pubescence which is contiguous with the patches of yellowish-grey pubescence on the basal half, and a narrow lateral band of yellowish-grey pubescence which connects the apical and median patches of yellowish-grey pubescence. The pronotal vestiture is likewise distinctive in that the median longitudinal band of black pubescence is more or less equal in width from base to apex. The shape of the patches of pubescence will readily separate individuals of this species from those of
|
||
<taxonomicName id="64FC20A0FFE5FFCDFF31BCFCFEB8FF18" authorityName="Chaudoir" authorityYear="1870" box="[192,337,194,218]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="fasciatus">
|
||
<emphasis id="91888731FFE5FFCDFF31BCFCFEB8FF18" box="[192,337,194,218]" italics="true" pageId="8" pageNumber="7">G. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
, which has much more extensive areas of black pubescence on the elytra, and in which the median longitudinal band of dark pubescence on the pronotum becomes distinctly wider from base to apex (
|
||
<figureCitation id="3BC747A6FFE5FFCDFEFBBD3CFE9CFED8" box="[266,373,258,282]" captionStart="Figures 1–9" captionStartId="5.[192,288,1044,1066]" captionTargetBox="[192,1440,192,1024]" captionTargetId="figure-218@5.[192,1440,192,1024]" captionTargetPageId="5" captionText="Figures 1–9. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 1) G. ancora Dejean [G. ancora ancora Dejean], Hanover, Northern Cape Province, RSA. 2) G. distinctus Péringuey [G. fasciatus distinctus Péringuey], Shilouvane, Limpopo Province, RSA. 3) G. fasciatus Chaudoir [G. fasciatus fasciatus Chaudoir], Umfolozi, KwaZulu/Natal Province, RSA. 4) G. fritschi Chaudoir, Pietersburg, Limpopo Province, RSA. 5) G. westwoodi Brême [G. ancora aurantiacus Basilewsky], Bulawayo, Zimbabwe. 6) G. westwoodi [G. ancora egressus Péringuey], Ohrigstad, Mpumalanga Province, RSA. 7) G. westwoodi [G. ancora subegregius Basilewsky, paratype], Tuli District, Botswana. 8) G. westwoodi [G. ancora transitans Péringuey], Metsimaklaba, Botswana. 9) G. westwoodi [G. ancora westwoodi Brême], Swartruggens Marico, North West Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174194" httpUri="https://zenodo.org/record/5174194/files/figure.png" pageId="8" pageNumber="7">Figure 2</figureCitation>
|
||
). Small adults of
|
||
<taxonomicName id="64FC20A0FFE5FFCDFDBDBD3CFD2DFEDB" authorityName="Dejean" authorityYear="1831" box="[588,708,258,281]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="ancora">
|
||
<emphasis id="91888731FFE5FFCDFDBDBD3CFD2DFEDB" box="[588,708,258,281]" italics="true" pageId="8" pageNumber="7">G. ancora</emphasis>
|
||
</taxonomicName>
|
||
are also similar but that species is generally larger in body size (length
|
||
<quantity id="6404F6C6FFE5FFCDFEA7BD1CFE3EFEF8" box="[342,471,290,314]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="8" pageNumber="7" unit="mm" value="15.5" valueMax="18.0" valueMin="13.0">13–18 mm</quantity>
|
||
) and the elytral vestiture is coarser and more irregular (
|
||
<figureCitation id="3BC747A6FFE5FFCDFB63BD1CFB13FEF8" box="[1170,1274,290,314]" captionStart="Figures 1–9" captionStartId="5.[192,288,1044,1066]" captionTargetBox="[192,1440,192,1024]" captionTargetId="figure-218@5.[192,1440,192,1024]" captionTargetPageId="5" captionText="Figures 1–9. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 1) G. ancora Dejean [G. ancora ancora Dejean], Hanover, Northern Cape Province, RSA. 2) G. distinctus Péringuey [G. fasciatus distinctus Péringuey], Shilouvane, Limpopo Province, RSA. 3) G. fasciatus Chaudoir [G. fasciatus fasciatus Chaudoir], Umfolozi, KwaZulu/Natal Province, RSA. 4) G. fritschi Chaudoir, Pietersburg, Limpopo Province, RSA. 5) G. westwoodi Brême [G. ancora aurantiacus Basilewsky], Bulawayo, Zimbabwe. 6) G. westwoodi [G. ancora egressus Péringuey], Ohrigstad, Mpumalanga Province, RSA. 7) G. westwoodi [G. ancora subegregius Basilewsky, paratype], Tuli District, Botswana. 8) G. westwoodi [G. ancora transitans Péringuey], Metsimaklaba, Botswana. 9) G. westwoodi [G. ancora westwoodi Brême], Swartruggens Marico, North West Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174194" httpUri="https://zenodo.org/record/5174194/files/figure.png" pageId="8" pageNumber="7">Figure 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EBE608A8FFE5FFCDFF31BD5FFCF0FDD8" pageId="8" pageNumber="7" type="distribution">
|
||
<paragraph id="A3435B23FFE5FFCDFF31BD5FFC27FE58" blockId="8.[192,1439,353,410]" pageId="8" pageNumber="7">
|
||
<emphasis id="91888731FFE5FFCDFF31BD5FFE9DFEBB" bold="true" box="[192,372,353,377]" pageId="8" pageNumber="7">Distribution.</emphasis>
|
||
<collectingCountry id="DBEB1BB3FFE5FFCDFE8ABD5CFDF5FEB8" box="[379,540,354,378]" name="Mozambique" pageId="8" pageNumber="7">Mozambique</collectingCountry>
|
||
,
|
||
<collectingCountry id="DBEB1BB3FFE5FFCDFDD9BD5CFCBEFEB8" box="[552,855,354,378]" name="South Africa" pageId="8" pageNumber="7">Republic of South Africa</collectingCountry>
|
||
(
|
||
<collectingRegion id="613895C1FFE5FFCDFC96BD5CFC3AFEB8" box="[871,979,354,378]" country="South Africa" name="Limpopo" pageId="8" pageNumber="7">Limpopo</collectingRegion>
|
||
and
|
||
<collectingRegion id="613895C1FFE5FFCDFBE3BD5CFB5FFEB8" box="[1042,1206,354,378]" country="South Africa" name="Mpumalanga" pageId="8" pageNumber="7">Mpumalanga</collectingRegion>
|
||
Provinces),
|
||
<collectingCountry id="DBEB1BB3FFE5FFCDFABCBD5CFF13FE58" name="Swaziland" pageId="8" pageNumber="7">Swaziland</collectingCountry>
|
||
. For a list of collecting localities see
|
||
<bibRefCitation id="C76D26D2FFE5FFCDFD4BBDBCFC23FE58" author="Basilewsky, P." box="[698,970,386,410]" pageId="8" pageNumber="7" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky (1977: 93)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A3435B23FFE5FFCDFF31BDFFFCF0FDD8" blockId="8.[192,1440,449,538]" pageId="8" pageNumber="7">
|
||
<emphasis id="91888731FFE5FFCDFF31BDFFFE51FE18" bold="true" box="[192,440,449,474]" pageId="8" pageNumber="7">Taxonomic Notes.</emphasis>
|
||
This species was considered a subspecies of
|
||
<taxonomicName id="64FC20A0FFE5FFCDFC19BDFCFB92FE18" authorityName="Chaudoir" authorityYear="1870" box="[1000,1147,450,474]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="fasciatus">
|
||
<emphasis id="91888731FFE5FFCDFC19BDFCFB92FE18" box="[1000,1147,450,474]" italics="true" pageId="8" pageNumber="7">G. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="C76D26D2FFE5FFCDFB5BBDFCFA81FE18" author="Burgeon, L." box="[1194,1384,450,474]" pageId="8" pageNumber="7" pagination="273 - 351" refId="ref7492" refString="Burgeon, L. 1929. Monographie du genre Graphipterus Latr. Annales de la Societe Entomologique du Belgique 69: 273 - 351." type="journal article" year="1929">Burgeon (1929)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="C76D26D2FFE5FFCDFF31BDDCFE77FE38" author="Basilewsky, P." box="[192,414,482,506]" pageId="8" pageNumber="7" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky (1977)</bibRefCitation>
|
||
but the lack of any intergrade specimens between these two rather distinctive forms suggests that two separate species are indicated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |