287 lines
27 KiB
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287 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.987.53648" ID-GBIF-Dataset="e14abbdb-e0a3-4c35-ad98-1cfd0cee46b5" ID-PMC="PMC7666096" ID-Pensoft-Pub="1313-2970-987-135" ID-Pensoft-UUID="A304608B226C5889BE8AF26E618669A3" ID-PubMed="33223888" ID-ZooBank="861F9C146E6D4222A41FFC56F978C2E8" ModsDocID="1313-2970-987-135" checkinTime="1604753978193" checkinUser="pensoft" docAuthor="Inoue, Shota, Nomura, Shuhei & Yin, Zi-Wei" docDate="2020" docId="190D674F79B358378011AF4CAE51FF2B" docLanguage="en" docName="ZooKeys 987: 135-156" docOrigin="ZooKeys 987" docSource="http://dx.doi.org/10.3897/zookeys.987.53648" docTitle="Pseudophanias excavatus Inoue, Nomura & Yin 2020, sp. nov." docType="treatment" docUuid="E9C17AEF-CEA2-45DB-A952-52905F0943BE" docUuidSource="ZooBank" docVersion="4" id="A304608B226C5889BE8AF26E618669A3" lastPageNumber="135" masterDocId="A304608B226C5889BE8AF26E618669A3" masterDocTitle="Three new species of Pseudophanias Raffray from Japan and Taiwan Island, and synonymy of Chandleriella Hlavac with Pseudophanias (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="156" masterPageNumber="135" pageNumber="135" updateTime="1668169777972" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Three new species of Pseudophanias Raffray from Japan and Taiwan Island, and synonymy of Chandleriella Hlavac with Pseudophanias (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Inoue, Shota</mods:namePart>
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<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka 819 - 0395, Japan & The Kyushu University Museum, Fukuoka 812 - 8581, Japan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7005-6531</mods:nameIdentifier>
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<mods:nameIdentifier type="email">pselaphineman@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nomura, Shuhei</mods:namePart>
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<mods:affiliation>Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1, Amakubo, Tsukuba-shi, Ibaraki 305 - 0005, Japan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yin, Zi-Wei</mods:namePart>
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<mods:affiliation>Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai 200234, China</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>987</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>135</mods:start>
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<mods:end>156</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.987.53648</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.987.53648</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-987-135</mods:identifier>
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<mods:identifier type="ZooBank">861F9C146E6D4222A41FFC56F978C2E8</mods:identifier>
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<mods:identifier type="Pensoft-UUID">A304608B226C5889BE8AF26E618669A3</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="168821845" LSID="urn:lsid:zoobank.org:act:E9C17AEF-CEA2-45DB-A952-52905F0943BE" httpUri="http://treatment.plazi.org/id/190D674F79B358378011AF4CAE51FF2B" lastPageNumber="135" pageId="0" pageNumber="135">
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<subSubSection pageId="0" pageNumber="135" type="nomenclature">
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<paragraph pageId="0" pageNumber="135">
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<taxonomicName LSID="190D674F-79B3-5837-8011-AF4CAE51FF2B" authority="Inoue, Nomura & Yin" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus" status="sp. nov.">Pseudophanias excavatus Inoue, Nomura & Yin</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="135">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dorsal habitus of Pseudophanias excavatus A male B female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.987.53648.figure9" httpUri="https://binary.pensoft.net/fig/472464" pageId="0" pageNumber="135">Figs 9</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Morphological details of Pseudophanias excavatus A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135">, 10</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">, 11</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male antenna of Pseudophanias excavatus by SEM images A lateral view B ventral view C antennomere 11 in ventral view D antennomeres 5 - 10 in ventral view E special setae on antennomere VIII in ventral view. Scale bars: 0.2 mm (A, B); 0.1 mm (C, D); 0.01 mm (E)." figureDoi="10.3897/zookeys.987.53648.figure12" httpUri="https://binary.pensoft.net/fig/472467" pageId="0" pageNumber="135">, 12</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male genitalia of Pseudophanias excavatus A ventral view B lateral view C dorsal view D tergite VIII E sternite VIII. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.987.53648.figure13" httpUri="https://binary.pensoft.net/fig/472468" pageId="0" pageNumber="135">, 13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="135" type="type material">
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<paragraph pageId="0" pageNumber="135">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="135">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Holotype</emphasis>
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(NSMT): ♂, "Tengshih (1400 m, litter) / Kaosiung Hsien / [M-Taiwan] /
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<normalizedToken originalValue="台湾高雄縣藤枝">台湾高雄縣藤枝</normalizedToken>
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/ 20-22. iv. 2001, H. Sugaya leg. // HOLOTYPE (red) /♂,
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<taxonomicName authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
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<emphasis italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
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</taxonomicName>
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sp. nov., / det. Inoue, Nomura & Yin, 2020"
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<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Paratypes</emphasis>
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: (NSMT, KUM, NMNS) 1 ♂, 3 ♀, same data as holotype; 6 ♂, 1 ♀, same data as holotype, but 29-30 IV 2001; 1 ♂, 2 ♀, same data as holotype, but 30 IV 2001. Each paratype pinned with the following label: "PARATYPE (yellow) / ♂ (or ♀),
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<taxonomicName authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
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<emphasis italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
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</taxonomicName>
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sp. nov., / det. Inoue, Nomura & Yin, 2020".
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="135" type="diagnosis">
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<paragraph pageId="0" pageNumber="135">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="135">
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This species is readily distinguished from other members of
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<taxonomicName authorityName="Raffray" authorityYear="1890" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="135">Pseudophanias</emphasis>
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</taxonomicName>
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by the clasping formed antennae in the male, frontal sulcus indistinct, and the rounded pronotum.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="135" type="description">
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<paragraph pageId="0" pageNumber="135">Description.</paragraph>
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<paragraph pageId="0" pageNumber="135">
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<emphasis bold="true" pageId="0" pageNumber="135">Male</emphasis>
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(Figs
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dorsal habitus of Pseudophanias excavatus A male B female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.987.53648.figure9" httpUri="https://binary.pensoft.net/fig/472464" pageId="0" pageNumber="135">9A</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Morphological details of Pseudophanias excavatus A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135">10A</figureCitation>
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). Body length 2.21-2.48 mm. Dorsal surface with dense setae.
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</paragraph>
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<paragraph pageId="0" pageNumber="135">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Head</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11A</figureCitation>
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) as long as wide, HL 0.45-0.50 mm, HW 0.43-0.50 mm, densely punctate, with dense, long setae; frontal rostrum broad, with frontal fovea nude; antennal tubercles distinct; vertex flat, with pair of glabrous foveae; eyes prominent, small; occiput with dense setae; postocular margin two times longer than eyes; small areas just posterior to U-shaped setose sulci finely punctate.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Antennae</emphasis>
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(Figs
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Morphological details of Pseudophanias excavatus A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135">10B</figureCitation>
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,
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male antenna of Pseudophanias excavatus by SEM images A lateral view B ventral view C antennomere 11 in ventral view D antennomeres 5 - 10 in ventral view E special setae on antennomere VIII in ventral view. Scale bars: 0.2 mm (A, B); 0.1 mm (C, D); 0.01 mm (E)." figureDoi="10.3897/zookeys.987.53648.figure12" httpUri="https://binary.pensoft.net/fig/472467" pageId="0" pageNumber="135">12A-E</figureCitation>
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) strongly modified, 0.98-1.12 mm in length; antennomeres 1 elongate, as long as 2-4 combined; 2-4 each transverse, successively shorter; 5-11 strongly excavated on ventral side, with tufts of setae on ventral surface, modified to form clasping, each excavated on ventral surface; 5-9 each two times wider than 4, each transverse; 9 longer than 8; 10 as long as 9, asymmetrical; outer side strongly produced ventrally in 9; 11 enlarged, with glabrous areas on ventral side. Maxillary palpi (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11B</figureCitation>
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) symmetrical; palpomeres 1 minute; palpomeres 2 elongate, narrowed in basal halves; palpomeres 3 small, widest at apices; palpomeres 4 fusiform.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Pronotum</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11C</figureCitation>
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) about as long as wide, PL 0.47-0.55 mm, PW 0.50-0.55 mm, broadly rounded, widest at middle, finely punctate on dorsal surface, with a median and pair of lateral antebasal foveae; antebasal area strongly punctate.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Metaventrite</emphasis>
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(Figs
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Morphological details of Pseudophanias excavatus A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135">10D</figureCitation>
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,
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11D</figureCitation>
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) finely punctate, strongly convex, but area just above metaventral apex roundly impressed; that impression 2/3 as long as metaventral length, occupying 1/4 metaventral width; anterior margin of that impression sharply rounded, distinct.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Elytra</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11E</figureCitation>
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) much wider than long, EL 0.57-0.65 mm, EW 0.90-0.98 mm, trapezoidal, finely punctate, each elytron with two basal fovea; discal stria shallow, extending from basal fovea placed middle to posterior 1/3.
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<emphasis bold="true" pageId="0" pageNumber="135">
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<emphasis italics="true" pageId="0" pageNumber="135">Legs</emphasis>
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.
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</emphasis>
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All legs moderately short; each femora broadest near middle; protibiae, mesotibiae with dense yellow setae at apices; protibiae, metatibiae moderately straight; mesotibiae slightly arcuate at apical fourth; tarsi (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11G</figureCitation>
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) each with tarsomere 2 half as long as tarsomere 3; tarsal claws (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11H</figureCitation>
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) asymmetrical; anterior claws long, posterior claws thin, short.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Abdomen</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11F</figureCitation>
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) much wider than long, widest at tergite IV, AL 0.64-0.81 mm, AW 0.97-1.05 mm, lacking discal carinae; tergite IV longest, twice as long as V, with setose depression at base. Tergite and sternite VIII as in Fig.
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male genitalia of Pseudophanias excavatus A ventral view B lateral view C dorsal view D tergite VIII E sternite VIII. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.987.53648.figure13" httpUri="https://binary.pensoft.net/fig/472468" pageId="0" pageNumber="135">13D, E</figureCitation>
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.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Aedeagus</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male genitalia of Pseudophanias excavatus A ventral view B lateral view C dorsal view D tergite VIII E sternite VIII. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.987.53648.figure13" httpUri="https://binary.pensoft.net/fig/472468" pageId="0" pageNumber="135">13A-C</figureCitation>
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) 0.60-0.63 mm in length, well-sclerotized, slightly asymmetrical in dorsal and ventral view, tubular in lateral view; parameres symmetrical, extremely elongate, reaching apical third, each with five setae at apex; median lobe roundly curved, C-shaped in lateral view; apical part widely opened, narrowed towards basal part to connect ovoidal dorsal diaphragm; apical lobe extending downward, curved to form S-shaped in lateral view, widely opened at base, strongly produced laterally at apex; endophallus indistinct.
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</paragraph>
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<paragraph pageId="0" pageNumber="135">
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<emphasis bold="true" pageId="0" pageNumber="135">Female</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dorsal habitus of Pseudophanias excavatus A male B female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.987.53648.figure9" httpUri="https://binary.pensoft.net/fig/472464" pageId="0" pageNumber="135">9B</figureCitation>
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). BL 2.21-2.27 mm; HL 0.44-0.48 mm; HW 0.44-0.48 mm; PL 0.47-0.55 mm; PW 0.46-0.51 mm; EL 0.56-0.63 mm; EW 0.90-0.94 mm; AL 0.67-0.70 mm; AW 0.94-0.96 mm. Antennae (Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Morphological details of Pseudophanias excavatus A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135">10C</figureCitation>
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) with antennomeres 11 simple, successively widened towards apices; antennomeres 2-10 each transverse; 11 ovoid, largest. Metaventrite convex, lacking metaventral impression.
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</paragraph>
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<caption doi="10.3897/zookeys.987.53648.figure9" httpUri="https://binary.pensoft.net/fig/472464" pageId="0" pageNumber="135" start="Figure 9" startId="F9">
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<paragraph pageId="0" pageNumber="135">
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<emphasis bold="true" pageId="0" pageNumber="135">Figure 9.</emphasis>
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Dorsal habitus of
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<taxonomicName authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
male
|
|
<emphasis bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
female. Scale bar: 1.0 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135" start="Figure 10" startId="F10">
|
|
<paragraph pageId="0" pageNumber="135">
|
|
<emphasis bold="true" pageId="0" pageNumber="135">Figure 10.</emphasis>
|
|
Morphological details of
|
|
<taxonomicName authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
habitus in lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
male antenna
|
|
<emphasis bold="true" pageId="0" pageNumber="135">C</emphasis>
|
|
female antenna
|
|
<emphasis bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
male metaventrite. Scale bars: 1.0 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
); 0.5 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
); 0.2 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="135">B, C</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135" start="Figure 11" startId="F11">
|
|
<paragraph pageId="0" pageNumber="135">
|
|
<emphasis bold="true" pageId="0" pageNumber="135">Figure 11.</emphasis>
|
|
SEM images of
|
|
<taxonomicName authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
|
|
</taxonomicName>
|
|
, male
|
|
<emphasis bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
head
|
|
<emphasis bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
maxillary palp
|
|
<emphasis bold="true" pageId="0" pageNumber="135">C</emphasis>
|
|
pronotum
|
|
<emphasis bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
metaventrite
|
|
<emphasis bold="true" pageId="0" pageNumber="135">E</emphasis>
|
|
elytra
|
|
<emphasis bold="true" pageId="0" pageNumber="135">F</emphasis>
|
|
abdomen
|
|
<emphasis bold="true" pageId="0" pageNumber="135">G</emphasis>
|
|
protarsus
|
|
<emphasis bold="true" pageId="0" pageNumber="135">H</emphasis>
|
|
protarsal claws. Scale bars: 0.2 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="135">A, C-F</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="135">B, G</emphasis>
|
|
); 0.05 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="135">H</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.987.53648.figure12" httpUri="https://binary.pensoft.net/fig/472467" pageId="0" pageNumber="135" start="Figure 12" startId="F12">
|
|
<paragraph pageId="0" pageNumber="135">
|
|
<emphasis bold="true" pageId="0" pageNumber="135">Figure 12.</emphasis>
|
|
Male antenna of
|
|
<taxonomicName authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
|
|
</taxonomicName>
|
|
by SEM images
|
|
<emphasis bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
ventral view
|
|
<emphasis bold="true" pageId="0" pageNumber="135">C</emphasis>
|
|
antennomere 11 in ventral view
|
|
<emphasis bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
antennomeres 5-10 in ventral view
|
|
<emphasis bold="true" pageId="0" pageNumber="135">E</emphasis>
|
|
special setae on antennomere VIII in ventral view. Scale bars: 0.2 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="135">A, B</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="135">C, D</emphasis>
|
|
); 0.01 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="135">E</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.987.53648.figure13" httpUri="https://binary.pensoft.net/fig/472468" pageId="0" pageNumber="135" start="Figure 13" startId="F13">
|
|
<paragraph pageId="0" pageNumber="135">
|
|
<emphasis bold="true" pageId="0" pageNumber="135">Figure 13.</emphasis>
|
|
Male genitalia of
|
|
<taxonomicName authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
ventral view
|
|
<emphasis bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="135">C</emphasis>
|
|
dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
tergite VIII
|
|
<emphasis bold="true" pageId="0" pageNumber="135">E</emphasis>
|
|
sternite VIII. Scale bars: 0.1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="135" type="etymology">
|
|
<paragraph pageId="0" pageNumber="135">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="135">The specific epithet refers to the strongly excavated antennae in the male of the new species.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="135" type="distribution">
|
|
<paragraph pageId="0" pageNumber="135">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="135">China (Taiwan).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="135" type="biology_ecology">
|
|
<paragraph pageId="0" pageNumber="135">Biology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="135">This species was collected from leaf litter.</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |