153 lines
19 KiB
XML
153 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="B53025763D530C18BC5512710C71B297" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Ornithocheirus xyphorhynchus Seeley 1870, (nomen dubium)" docType="treatment" docVersion="5" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="49" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="49" updateTime="1634706563434" updateUser="ExternalLinkService">
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<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rodrigues, Taissa</mods:namePart>
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<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
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<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2013</mods:date>
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<mods:number>2013-06-12</mods:number>
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<mods:detail type="volume">
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<mods:number>308</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>112</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
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<treatment ID-GBIF-Taxon="152044994" LSID="urn:lsid:plazi:treatment:B53025763D530C18BC5512710C71B297" httpUri="http://treatment.plazi.org/id/B53025763D530C18BC5512710C71B297" lastPageNumber="49" pageId="48" pageNumber="49">
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<subSubSection pageId="48" pageNumber="49" type="nomenclature">
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName LSID="B5302576-3D53-0C18-BC55-12710C71B297" authority="Seeley, 1870" authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Ornithocheirus xyphorhynchus" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="xyphorhynchus" status="(nomen dubium)">Ornithocheirus xyphorhynchus Seeley, 1870</taxonomicName>
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<taxonomicNameLabel pageId="48" pageNumber="49">(nomen dubium)</taxonomicNameLabel>
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. A-B Ornithocheirus oxyrhinus, holotype CAMSM B 54612 (Albian, Cambridge Greensand), anterior part of the rostrum. A ventral view B respective line drawing. C-E Ornithocheirus scaphorhynchus, holotype CAMSM B 54441 (Albian, Cambridge Greensand), anterior part of the rostrum C ventral view D respective line drawing E line drawing in posterior view. F-I Ornithocheirus tenuirostris, holotype CAMSM B 54584 (Albian, Cambridge Greensand), anterior part of the rostrum F right lateral view G respective line drawing H ventral view I respective line drawing. J-K Ornithocheirus xyphorhynchus, holotype (Albian, Cambridge Greensand), anterior part of the rostrum J lateral view K dorsal view. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge, sul - sulcus. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. J and K from Seeley 1881." httpUri="https://binary.pensoft.net/fig/22087" pageId="48" pageNumber="49">Fig. 19J-K</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="reference_group">
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="xyphorhynchus" status="(nomen dubium)">Ornithocheirus xyphorhynchus</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
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: p. 117
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="xyphorhynchus" status="(nomen dubium)">Ornithocheirus xyphorhynchus</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Geological Magazine" pageId="69" pageNumber="70" pagination="13 - 20" publicationUrl="10.1017/S0016756800113007" refId="B80" refString="Seeley, HG, 1881. On evidence of two ornithosaurians referable to the genus Ornithocheirus, from the Upper Greensand of Cambridge, preserved in the collection of W. Reed, Esq., F.G.S. Geological Magazine 8: 13 - 20, 10.1017/S0016756800113007" title="On evidence of two ornithosaurians referable to the genus Ornithocheirus, from the Upper Greensand of Cambridge, preserved in the collection of W. Reed, Esq., F. G. S." url="10.1017/S0016756800113007" volume="8" year="1881">Seeley 1881</bibRefCitation>
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: p. 18, plate I, fig. 2
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="xyphorhynchus" status="(nomen dubium)">Ornithocheirus xyphorhynchus</taxonomicName>
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Seeley:
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
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: p. 538
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="xyphorhynchus" status="(nomen dubium)">Ornithocheirus xyphorhynchus</taxonomicName>
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Seeley:
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<bibRefCitation author="Arthaber, G" journalOrPublisher="Palaeontologische Zeitschrift" pageId="64" pageNumber="65" pagination="1 - 47" refId="B2" refString="Arthaber, G, 1922. Ueber Entwicklung, Ausbildung und Austerben der Flugsaurier. Palaeontologische Zeitschrift 4: 1 - 47" title="Ueber Entwicklung, Ausbildung und Austerben der Flugsaurier." volume="4" year="1922">Arthaber 1922</bibRefCitation>
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: p. 17
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Anhanguera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="cuvieri" status="(nomen dubium)">Anhanguera cuvieri</taxonomicName>
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(Bowerbank):
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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: table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="holotype">
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<paragraph pageId="48" pageNumber="49">Holotype.</paragraph>
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<paragraph pageId="48" pageNumber="49">
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anterior portion of the rostrum (collection data could not be recovered) (
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. A-B Ornithocheirus oxyrhinus, holotype CAMSM B 54612 (Albian, Cambridge Greensand), anterior part of the rostrum. A ventral view B respective line drawing. C-E Ornithocheirus scaphorhynchus, holotype CAMSM B 54441 (Albian, Cambridge Greensand), anterior part of the rostrum C ventral view D respective line drawing E line drawing in posterior view. F-I Ornithocheirus tenuirostris, holotype CAMSM B 54584 (Albian, Cambridge Greensand), anterior part of the rostrum F right lateral view G respective line drawing H ventral view I respective line drawing. J-K Ornithocheirus xyphorhynchus, holotype (Albian, Cambridge Greensand), anterior part of the rostrum J lateral view K dorsal view. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge, sul - sulcus. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. J and K from Seeley 1881." httpUri="https://binary.pensoft.net/fig/22087" pageId="48" pageNumber="49">Fig. 19J-K</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="type locality">
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<paragraph pageId="48" pageNumber="49">Type locality.</paragraph>
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<paragraph pageId="48" pageNumber="49">Cambridge, Cambridgeshire, England.</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="type horizon">
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<paragraph pageId="48" pageNumber="49">Type horizon.</paragraph>
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<paragraph pageId="48" pageNumber="49">Cambridge Greensand (Cenomanian; fossils Albian in age).</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="description">
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<paragraph pageId="48" pageNumber="49">Description.</paragraph>
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<paragraph pageId="48" pageNumber="49">
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The holotype of
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="xyphorhynchus">
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<emphasis italics="true" pageId="48" pageNumber="49">Ornithocheirus xyphorhynchus</emphasis>
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</taxonomicName>
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was illustrated by
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Geological Magazine" pageId="69" pageNumber="70" pagination="13 - 20" publicationUrl="10.1017/S0016756800113007" refId="B80" refString="Seeley, HG, 1881. On evidence of two ornithosaurians referable to the genus Ornithocheirus, from the Upper Greensand of Cambridge, preserved in the collection of W. Reed, Esq., F.G.S. Geological Magazine 8: 13 - 20, 10.1017/S0016756800113007" title="On evidence of two ornithosaurians referable to the genus Ornithocheirus, from the Upper Greensand of Cambridge, preserved in the collection of W. Reed, Esq., F. G. S." url="10.1017/S0016756800113007" volume="8" year="1881">Seeley (1881</bibRefCitation>
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: plate I,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ornithocheirus simus. A-D referred specimen CAMSM 54429 (Albian, Cambridge Greensand), anterior part of the rostrum A right lateral view B respective line drawing C anterior view D respective line drawing E-H referred specimen CAMSM 54677 (Albian, Cambridge Greensand), anterior part of the rostrum E right lateral view F respective line drawing G anterior view H respective line drawing. Abbreviations: m - maxillae, pm - premaxillae. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22070" pageId="48" pageNumber="49">Figs 2a and 2b</figureCitation>
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), and, as the type material of
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<taxonomicName lsidName="" pageId="48" pageNumber="49" rank="species" species="reedi">
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<emphasis italics="true" pageId="48" pageNumber="49">
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<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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reedi
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</emphasis>
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</taxonomicName>
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, belonged to the collection of W. Reed of York at the time when it was first described. It could not be found in the collections of the Natural History Museum, the Sedgwick Museum of Earth Sciences, or the Manchester Museum during visits in October 2009. Therefore, we base our remarks on the descriptions and illustrations provided by
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870</bibRefCitation>
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,
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Geological Magazine" pageId="69" pageNumber="70" pagination="13 - 20" publicationUrl="10.1017/S0016756800113007" refId="B80" refString="Seeley, HG, 1881. On evidence of two ornithosaurians referable to the genus Ornithocheirus, from the Upper Greensand of Cambridge, preserved in the collection of W. Reed, Esq., F.G.S. Geological Magazine 8: 13 - 20, 10.1017/S0016756800113007" title="On evidence of two ornithosaurians referable to the genus Ornithocheirus, from the Upper Greensand of Cambridge, preserved in the collection of W. Reed, Esq., F. G. S." url="10.1017/S0016756800113007" volume="8" year="1881">1881</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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The species was based on a fragmentary mandible, lacking the tip. It did not have a dentary crest (
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
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).
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin (2001)</bibRefCitation>
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recently referred it to
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<taxonomicName class="Reptilia" family="Anhangueridae" genus="Anhanguera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="48" pageNumber="49">Anhanguera</emphasis>
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</taxonomicName>
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[=
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" genus="Cimoliopterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="48" pageNumber="49">Cimoliopterus</emphasis>
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</taxonomicName>
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]
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<emphasis italics="true" pageId="48" pageNumber="49">cuvieri</emphasis>
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. However, the latter is known by a cranial material from the Chalk Formation, whereas
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="xyphorhynchus">
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<emphasis italics="true" pageId="48" pageNumber="49">Ornithocheirus xyphorhynchus</emphasis>
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</taxonomicName>
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is known only by a partial lower jaw from the Cambridge Greensand. As both species are known by material which is not directly comparable, the proposed synonymy can be rejected.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |