254 lines
22 KiB
XML
254 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1142.90097" ID-Pensoft-Pub="1313-2970-1142-1" ID-Pensoft-UUID="1D4BDF04F72B59B9984E00D0B4FB338D" ID-ZooBank="A1129EE50F9941CFB73AE771B66E2486" ModsDocID="1313-2970-1142-1" checkinTime="1674655519517" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Tumpeesuwan, Sakboworn, Inkhavilay, Khamla, Prasankok, Pongpun, Jeratthitikul, Ekgachai, Panha, Somsak & Sutcharit, Chirasak" docDate="2023" docId="09AEBD7307735DBEAA2D351792EEC068" docLanguage="en" docName="ZooKeys 1142: 1-144" docOrigin="ZooKeys 1142" docPubDate="2023-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1142.90097" docTitle="Rhiostoma strubelli Moellendorff 1899" docType="treatment" docVersion="1" id="1D4BDF04F72B59B9984E00D0B4FB338D" lastPageNumber="1" masterDocId="1D4BDF04F72B59B9984E00D0B4FB338D" masterDocTitle="Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1674655519517" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prasankok, Pongpun</mods:namePart>
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<mods:affiliation>School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation>Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-24</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1142</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>144</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1142.90097</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1142.90097</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1142-1</mods:identifier>
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<mods:identifier type="ZooBank">A1129EE50F9941CFB73AE771B66E2486</mods:identifier>
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<mods:identifier type="Pensoft-UUID">1D4BDF04F72B59B9984E00D0B4FB338D</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:09AEBD7307735DBEAA2D351792EEC068" httpUri="http://treatment.plazi.org/id/09AEBD7307735DBEAA2D351792EEC068" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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15.
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<taxonomicName ID-CoL="7SZXJ" LSID="09AEBD73-0773-5DBE-AA2D-351792EEC068" authority="Moellendorff, 1899" authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma strubelli" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="strubelli">
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Rhiostoma strubelli
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1899
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Distribution map of Rhiostoma asiphon - species group." figureDoi="10.3897/zookeys.1142.90097.figure34" httpUri="https://binary.pensoft.net/fig/799892" pageId="0" pageNumber="1">Figs 34</figureCitation>
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<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shell of Rhiostoma strubelli A lectotype SMF 130513 from S-Shan Staaten: Kalow B-E specimens CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos and E with long breathing device F specimen CUMZ 10047 from Xishuangbanna Botanical Garden, Yunnan, China." figureDoi="10.3897/zookeys.1142.90097.figure36" httpUri="https://binary.pensoft.net/fig/799894" pageId="0" pageNumber="1">, 36</figureCitation>
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<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Shell of Rhiostoma platymorpha sp. nov. A holotype CUMZ 4498 / 1 from Tham Muang On, Mae On, Chiang Mai B paratype CUMZ 4763 from type locality C, D paratypes CUMZ 4498 / 2 from type locality E, F specimens CUMZ 4783 from Tham Bua Tong, Mae Tang, Chiang Mai (with F albinistic shell form)." figureDoi="10.3897/zookeys.1142.90097.figure41" httpUri="https://binary.pensoft.net/fig/799899" pageId="0" pageNumber="1">, 41B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName family="Cyclophoridae" genus="Rhiostoms" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoms strubelli" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="strubelli">Rhiostoms strubelli</taxonomicName>
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1899: 166. Type locality: Kalow, 5000',
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<normalizedToken originalValue="südliche">suedliche</normalizedToken>
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Shan Staaten [Kalaw, Taunggyi District, Shan State, Myanmar].
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="1" pagination="1 - 663" refId="B71" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich 16: 1 - 663" title="Cyclophoridae." volume="16" year="1902">Kobelt 1902</bibRefCitation>
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: 179. Kobelt 1911: 763, pl. 113, fig. 11.
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<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch 1956</bibRefCitation>
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: 174.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The original description did not explicitly indicate the number of specimens, and was without illustration or explicit designation of the name-bearing type, although one set of measurements was given in
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<bibRefCitation author="Moellendorff, OF von" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="165 - 166" refId="B99" refString="Moellendorff, OF von, 1899. Neue arten aus Hinterindien. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 31: 165 - 166" title="Neue arten aus Hinterindien." volume="31" year="1899">
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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(1899)
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</bibRefCitation>
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. Later, Kobelt (1911: pl. 113, fig. 11) illustrated this species for the first time, and then
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<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch (1956)</bibRefCitation>
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recognised this figured specimen (SMF 130513) as the
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<normalizedToken originalValue="“holotypus”">"holotypus"</normalizedToken>
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. This SMF collection lot contains only one specimen; therefore, the subsequent use of the term
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<normalizedToken originalValue="“holotypus”">"holotypus"</normalizedToken>
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in
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<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch (1956)</bibRefCitation>
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constitutes a valid lectotype designation.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
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(designation in
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<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch (1956)</bibRefCitation>
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) SMF 130513 (Fig.
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<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shell of Rhiostoma strubelli A lectotype SMF 130513 from S-Shan Staaten: Kalow B-E specimens CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos and E with long breathing device F specimen CUMZ 10047 from Xishuangbanna Botanical Garden, Yunnan, China." figureDoi="10.3897/zookeys.1142.90097.figure36" httpUri="https://binary.pensoft.net/fig/799894" pageId="0" pageNumber="1">36A</figureCitation>
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) from S-Shan Staaten: Kalow, 5000'.
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</paragraph>
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<caption doi="10.3897/zookeys.1142.90097.figure36" httpUri="https://binary.pensoft.net/fig/799894" pageId="0" pageNumber="1" start="Figure 36" startId="F36">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 36.</emphasis>
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Shell of
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<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma strubelli" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="strubelli">
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<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma strubelli</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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lectotype SMF 130513 from S-Shan Staaten: Kalow
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<emphasis bold="true" pageId="0" pageNumber="1">B-E</emphasis>
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specimens CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos and
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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with long breathing device
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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specimen CUMZ 10047 from Xishuangbanna Botanical Garden, Yunnan, China.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingCountry name="China">China</collectingCountry>
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</emphasis>
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: Xishuangbanna Tropical Botanical Garden, Mengla County, Xishuangbanna Prefecture,
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<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
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: CUMZ 10047 (7 shells; Fig.
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<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shell of Rhiostoma strubelli A lectotype SMF 130513 from S-Shan Staaten: Kalow B-E specimens CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos and E with long breathing device F specimen CUMZ 10047 from Xishuangbanna Botanical Garden, Yunnan, China." figureDoi="10.3897/zookeys.1142.90097.figure36" httpUri="https://binary.pensoft.net/fig/799894" pageId="0" pageNumber="1">36F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingCountry name="Laos">Laos</collectingCountry>
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</emphasis>
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: Khao Pa Huak, Vieng Phuka District,
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<collectingRegion country="Laos" name="Louangnamtha">Luang Nam Tha Province</collectingRegion>
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: CUMZ 10200 (29 shells; Figs
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<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shell of Rhiostoma strubelli A lectotype SMF 130513 from S-Shan Staaten: Kalow B-E specimens CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos and E with long breathing device F specimen CUMZ 10047 from Xishuangbanna Botanical Garden, Yunnan, China." figureDoi="10.3897/zookeys.1142.90097.figure36" httpUri="https://binary.pensoft.net/fig/799894" pageId="0" pageNumber="1">36B-E</figureCitation>
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,
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<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Radula morphology A Rhiostoma asiphon CUMZ 4826 from Angthong Islands, Surat Thani B Rhiostoma strubelli specimen CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos C Rhiostoma breviocollar sp. nov. paratypes CUMZ 4491 from type locality D Rhiostoma anceyi sp. nov. paratype CUMZ 4494 / 2 from type locality E Rhiostoma furfurosum sp. nov. paratype CUMZ 4705 / 2 from type locality F Rhiostoma platymorpha sp. nov. paratype CUMZ 4763 from type locality." figureDoi="10.3897/zookeys.1142.90097.figure42" httpUri="https://binary.pensoft.net/fig/799900" pageId="0" pageNumber="1">42B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell small, thin, and depressed; detached whorl shorter than apertural width. Shell with brownish zigzag patterns. Breathing device short and incomplete tube shape.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma strubelli" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="strubelli">
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<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma strubelli</emphasis>
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</taxonomicName>
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can be distinguished from
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<taxonomicName lsidName="R. haughtoni" pageId="0" pageNumber="1" rank="species" species="haughtoni">
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<emphasis italics="true" pageId="0" pageNumber="1">R. haughtoni</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="R. tigrina" pageId="0" pageNumber="1" rank="species" species="tigrina">
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<emphasis italics="true" pageId="0" pageNumber="1">R. tigrina</emphasis>
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</taxonomicName>
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sp. nov. by having a smaller shell, very short detached whorl, incomplete tubular shape of breathing device, and apex with dark colouration. In contrast,
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<taxonomicName lsidName="R. haughtoni" pageId="0" pageNumber="1" rank="species" species="haughtoni">
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<emphasis italics="true" pageId="0" pageNumber="1">R. haughtoni</emphasis>
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</taxonomicName>
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has a canal-shaped breathing device with a monochrome shell, while
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<taxonomicName lsidName="R. tigrina" pageId="0" pageNumber="1" rank="species" species="tigrina">
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<emphasis italics="true" pageId="0" pageNumber="1">R. tigrina</emphasis>
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</taxonomicName>
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sp. nov. has a long, detached whorl and short to long breathing device with complete tube shape.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Shell</emphasis>
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.
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</emphasis>
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Shell small, cW 17.3-22.9 mm, cH 8.5-12.0 mm, thin, and sub-discoidal shell; detached-whorl length 2.0-7.0 mm. Apex acute and dark colouration; spire slightly elevated. Whorls 4 to 5, convex, increasing regularly; suture wide and shallow; last whorl rounded. Shell surface with fine growth lines. Periostracum thin, corneous, and transparent. Shell colour with brown zigzag pattern, and narrow dark brown spiral band on periphery. Detached whorl shorter than apertural width. Peristome circular and double; lip thickened and slightly expanded. Aperture opened sub-laterally. Breathing device with incomplete tube shape; outer lip protruded into a nearly closed tube; inner lip with deep incision or small hole inside aperture. Umbilicus widely opened and deep. Operculum calcareous, low cup-shaped, and multispiral (Fig.
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<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shell of Rhiostoma strubelli A lectotype SMF 130513 from S-Shan Staaten: Kalow B-E specimens CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos and E with long breathing device F specimen CUMZ 10047 from Xishuangbanna Botanical Garden, Yunnan, China." figureDoi="10.3897/zookeys.1142.90097.figure36" httpUri="https://binary.pensoft.net/fig/799894" pageId="0" pageNumber="1">36</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Radula</emphasis>
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.
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</emphasis>
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Teeth arrangement and shape are similar to
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<taxonomicName lsidName="R. asiphon" pageId="0" pageNumber="1" rank="species" species="asiphon">
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<emphasis italics="true" pageId="0" pageNumber="1">R. asiphon</emphasis>
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</taxonomicName>
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. Central tooth with large central cusp and two lateral cusps on each side. Lateral teeth having four cusps; central cusp large, triangular, and flanked by two small inner cusps and one outer cusp. Marginal teeth each consisting of three pointed cusps (Fig.
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<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Radula morphology A Rhiostoma asiphon CUMZ 4826 from Angthong Islands, Surat Thani B Rhiostoma strubelli specimen CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos C Rhiostoma breviocollar sp. nov. paratypes CUMZ 4491 from type locality D Rhiostoma anceyi sp. nov. paratype CUMZ 4494 / 2 from type locality E Rhiostoma furfurosum sp. nov. paratype CUMZ 4705 / 2 from type locality F Rhiostoma platymorpha sp. nov. paratype CUMZ 4763 from type locality." figureDoi="10.3897/zookeys.1142.90097.figure42" httpUri="https://binary.pensoft.net/fig/799900" pageId="0" pageNumber="1">42B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species is known from Kalaw, Shan State, Myanmar. This study recorded specimens from Luang Namtha Province, Laos, and Yunnan in China (Fig.
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<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Distribution map of Rhiostoma asiphon - species group." figureDoi="10.3897/zookeys.1142.90097.figure34" httpUri="https://binary.pensoft.net/fig/799892" pageId="0" pageNumber="1">34</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species is previously known only from the holotype from the south of Shan State, Myanmar, which is close to northern Thailand and Laos. However, no
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<taxonomicName authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
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</taxonomicName>
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specimens collected from northern Thailand could be identified as this species. The specimen from Luang Namtha Province, Laos and Yunnan Province, China are very similar to the lectotype in having short detached whorl and breathing device with an incomplete tube shape. The only difference is in having darker irregular blotches on the shell than the type specimen, and this is considered as intrapopulation variation.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |