275 lines
41 KiB
XML
275 lines
41 KiB
XML
<document id="7787A3384F6C1D26C067E4E1F87623EC" ID-CLB-Dataset="256669" ID-DOI="10.5852/ejt.2023.864.2093" ID-GBIF-Dataset="7f562ba5-17db-46bf-93e8-4d1ba49c2800" ID-ISSN="2118-9773" ID-Zenodo-Dep="7872174" ID-ZooBank="D47B6B04-B2AD-4FDD-B7C4-B71CA6A5BB84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1682513062582" checkinUser="felipe" docAuthor="Brunke, Adam James" docDate="2023" docId="6C38C875FFBE5B5CFDAFFBE7E707F883" docLanguage="en" docName="ejt-864_117-145.pdf" docOrigin="European Journal of Taxonomy 864" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Malaisdius Brunke, 2023, gen. nov." docType="treatment" docUuid="6947E75F-7462-4642-A8D4-2AA3EA25E22C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="127" masterDocId="9001B00DFFB75B56FFDAFFF6E270FF89" masterDocTitle="Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar" masterLastPageNumber="145" masterPageNumber="117" pageNumber="126" updateTime="1699514916083" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2D88586A1348327E44C4B4E20FF22D49">Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar</mods:title>
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<mods:name id="EF7A5FB968F7D9BB3A70EC003E83FAF6" type="personal">
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<mods:namePart id="6201DE0B9D2655D87F5540631879BFE2">Brunke, Adam James</mods:namePart>
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<mods:typeOfResource id="1842F87BA9AEF15E67D37158CDCCF124">text</mods:typeOfResource>
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<mods:title id="4F7D85D5940AA2DC2C8B6069D5CDC671">European Journal of Taxonomy</mods:title>
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<mods:part id="D9B450B39E065F606BFFCCC1DED1A1B1">
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<mods:date id="32BCB1F6032AF08AFE3503567AB6BBAD">2023</mods:date>
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<mods:number id="7CB134B19C6781DD4191CFC0FEFB8C93">2023-04-20</mods:number>
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<treatment id="6C38C875FFBE5B5CFDAFFBE7E707F883" ID-DOI="http://doi.org/10.5281/zenodo.7872290" ID-GBIF-Taxon="209579388" ID-Zenodo-Dep="7872290" LSID="urn:lsid:zoobank.org:act:6947E75F-7462-4642-A8D4-2AA3EA25E22C" httpUri="http://treatment.plazi.org/id/6C38C875FFBE5B5CFDAFFBE7E707F883" lastPageId="10" lastPageNumber="127" pageId="9" pageNumber="126">
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<subSubSection id="AC8B2AE8FFBE5B5FFDAFFBE7E1CFFBA2" box="[629,959,1040,1067]" pageId="9" pageNumber="126" type="nomenclature">
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<paragraph id="E42E7963FFBE5B5FFDAFFBE7E1CFFBA2" blockId="9.[368,1219,1040,1138]" box="[629,959,1040,1067]" pageId="9" pageNumber="126">
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<heading id="BF66CE0FFFBE5B5FFDAFFBE7E1CFFBA2" box="[629,959,1040,1067]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="126" reason="2">
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Genus
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<taxonomicName id="239102E0FFBE5B5FFD1DFBE6E13DFBA2" authorityName="Brunke" authorityYear="2023" box="[711,845,1040,1067]" class="Insecta" family="Staphylinidae" genus="Malaisdius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="D6E5A571FFBE5B5FFD1DFBE6E13DFBA2" bold="true" box="[711,845,1040,1067]" italics="true" pageId="9" pageNumber="126">Malaisdius</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CDD6180AFFBE5B5FFC8EFBE7E1CFFBA2" box="[852,959,1041,1067]" pageId="9" pageNumber="126" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="AC8B2AE8FFBE5B5FFEAAFBC2E19FFBF8" pageId="9" pageNumber="126" type="description">
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<paragraph id="E42E7963FFBE5B5FFEAAFBC2E6B3FBC6" blockId="9.[368,1219,1040,1138]" box="[368,1219,1076,1103]" pageId="9" pageNumber="126">
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<uri id="90007561FFBE5B5FFEAAFBC2E6B3FBC6" box="[368,1219,1076,1103]" pageId="9" pageNumber="126">
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urn:lsid:zoobank.org:act:
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<uuid id="903743B6FFBE5B5FFD54FBC2E6B3FBC6" box="[654,1219,1076,1103]" pageId="9" pageNumber="126">6947E75F-7462-4642-A8D4-2AA3EA25E22C</uuid>
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</uri>
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</paragraph>
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<paragraph id="E42E7963FFBE5B5FFD9FFBA1E19FFBF8" blockId="9.[368,1219,1040,1138]" box="[581,1007,1111,1138]" pageId="9" pageNumber="126">
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<figureCitation id="7CAA65E6FFBE5B5FFD9FFBA1E0B8FBF8" box="[581,712,1111,1137]" captionStart="Fig" captionStartId="16.[189,228,1872,1896]" captionTargetBox="[190,1396,472,1833]" captionTargetId="figure-78@16.[189,1398,471,1834]" captionTargetPageId="16" captionText="Fig. 2. A–E. Dorsal habitus. A. Malaisdius ruficeps (Scheerpeltz, 1965), ♂, paratype (NHW). B. Malaisdius smetanai gen. et. sp. nov., ♂, holotype (NME). C. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965, ♂, holotype (NHRS). D. Q. (M.) piceolineatus Scheerpeltz, 1965, ♂ holotype (NHRS). E. Q. (M.) impressithorax Scheerpeltz, 1965, ♀ holotype (NHRS). F. Forebody of Q. (M.) impressithorax Scheerpeltz, 1965. Abbreviation: Pf = posterior frontal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872182" httpUri="https://zenodo.org/record/7872182/files/figure.png" pageId="9" pageNumber="126">Figs 2A–B</figureCitation>
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,
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<figureCitation id="7CAA65E6FFBE5B5FFD0EFBA1E163FBF8" box="[724,787,1111,1137]" captionStart="Fig" captionStartId="21.[189,228,1904,1928]" captionTargetBox="[191,1395,503,1860]" captionTargetId="figure-97@21.[189,1398,499,1864]" captionTargetPageId="21" captionText="Fig. 4. A–B. Indoquedius dispersepunctatus(Scheerpeltz, 1965). A. Elytra. B. Dorsal abdomen. C–D. I. recticollis (Scheerpeltz, 1965). C. Elytra. D. Dorsal abdomen. E–F. Malaisdius smetanai gen. et sp. nov. E. Head. F. Elytra. Abbreviations: Af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supraantennal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872188" httpUri="https://zenodo.org/record/7872188/files/figure.png" pageId="9" pageNumber="126">4E–F</figureCitation>
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,
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<figureCitation id="7CAA65E6FFBE5B5FFCC5FBA1E118FBFB" box="[799,872,1111,1138]" captionStart="Fig" captionStartId="22.[189,228,1840,1864]" captionTargetBox="[192,1395,440,1798]" captionTargetId="figure-53@22.[189,1398,436,1802]" captionTargetPageId="22" captionText="Fig. 5. A. Malaisdius ruficeps (Scheerpeltz, 1965), dorsal abdomen. B. M. smetanai gen. et sp. nov., dorsal abdomen. C–E. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965. C. Antenna. D. Pronotum (white arrow: marginal punctures removed from marginal bead; black arrow: anterior angles with marginal bead entirely visible in dorsal view). E. Elytra and basal abdomen. F. Q. (M.) masasatoi Smetana, 2007, pronotum (arrows indicating punctures of the dorsal row). Abbreviation: Pf = posterior frontal puncture. Scale bars: A–B, D–E = 1 mm; C, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7872192" httpUri="https://zenodo.org/record/7872192/files/figure.png" pageId="9" pageNumber="126">5A–B</figureCitation>
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,
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<figureCitation id="7CAA65E6FFBE5B5FFCAEFBA1E1CFFBF8" box="[884,959,1111,1137]" captionStart="Fig" captionStartId="24.[189,228,1872,1896]" captionTargetBox="[229,1350,707,1815]" captionTargetId="figure-144@24.[223,1364,704,1834]" captionTargetPageId="24" captionText="Fig. 6. A, F, H–I, K–L.Aedeagus, in situ. A, F, H, K. Ventral view (inset showing paired teeth). I, L. Lateral view. B–D. Median lobe. B. Ventral view. C–D. Lateral view. E, G, J, M. Underside of paramere. A–E. Malaisdius ruficeps (Scheerpeltz, 1965). F–G. M. smetanai gen. et sp. nov. H–J. Quedius (Microsaurus) piceolineatus Scheerpeltz, 1965. K–M. Q. (M.) masasatoi Smetana, 2007. Scale bars: A–D, F, H–I, K–L = 0.5 mm; E, G, J, M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872196" httpUri="https://zenodo.org/record/7872196/files/figure.png" pageId="9" pageNumber="126">6A–G</figureCitation>
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,
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<figureCitation id="7CAA65E6FFBE5B5FFC11FBA1E19FFBF8" box="[971,1007,1111,1137]" captionStart="Fig" captionStartId="25.[188,227,1936,1960]" captionTargetBox="[192,1395,533,1891]" captionTargetId="figure-75@25.[189,1398,529,1894]" captionTargetPageId="25" captionText="Fig. 7. Female tergite X. A. Indoquedius dispersepunctatus (Scheerpeltz, 1965). B. I. parallelicollis (Scheerpeltz, 1965). C. I. recticollis (Scheerpeltz, 1965). D. Malaisdius ruficeps (Scheerpeltz, 1965). E. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965. F. Q. (M.) impressithorax Scheerpeltz, 1965. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872200" httpUri="https://zenodo.org/record/7872200/files/figure.png" pageId="9" pageNumber="126">7D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8B2AE8FFBE5B5FFF67FB6EE014FB6B" pageId="9" pageNumber="126" type="type_taxon">
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<paragraph id="E42E7963FFBE5B5FFF67FB6EE328FB3B" blockId="9.[189,344,1176,1202]" box="[189,344,1176,1202]" pageId="9" pageNumber="126">
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<heading id="BF66CE0FFFBE5B5FFF67FB6EE328FB3B" bold="true" box="[189,344,1176,1202]" fontSize="11" level="3" pageId="9" pageNumber="126" reason="3">
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<emphasis id="D6E5A571FFBE5B5FFF67FB6EE328FB3B" bold="true" box="[189,344,1176,1202]" pageId="9" pageNumber="126">
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<typeStatus id="3B2AC7C1FFBE5B5FFF67FB6EE28AFB3B" box="[189,250,1176,1202]" pageId="9" pageNumber="126">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E42E7963FFBE5B5FFF67FB31E014FB6B" blockId="9.[189,612,1223,1250]" box="[189,612,1223,1250]" pageId="9" pageNumber="126">
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<taxonomicName id="239102E0FFBE5B5FFF67FB31E010FB6B" ID-CoL="4R3V3" authority="Scheerpeltz, 1965" authorityName=", O. Scheerpeltz" authorityYear="1965" baseAuthorityName=", O. Scheerpeltz" baseAuthorityYear="1965" box="[189,608,1223,1250]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="species" species="ruficeps">
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<emphasis id="D6E5A571FFBE5B5FFF67FB31E3F3FB68" box="[189,387,1223,1249]" italics="true" pageId="9" pageNumber="126">Quedius ruficeps</emphasis>
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<bibRefCitation id="80000492FFBE5B5FFE50FB31E010FB6B" author="Scheerpeltz O." box="[394,608,1223,1250]" pageId="9" pageNumber="126" pagination="93 - 371" refId="ref13560" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Arkiv for Zoologi 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8B2AE8FFBE5B5FFF67FAFEE6DEF988" pageId="9" pageNumber="126" type="diagnosis">
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<paragraph id="E42E7963FFBE5B5FFF67FAFEE346FAAB" blockId="9.[189,310,1288,1314]" box="[189,310,1288,1314]" pageId="9" pageNumber="126">
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<heading id="BF66CE0FFFBE5B5FFF67FAFEE346FAAB" bold="true" box="[189,310,1288,1314]" fontSize="11" level="3" pageId="9" pageNumber="126" reason="3">
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<emphasis id="D6E5A571FFBE5B5FFF67FAFEE346FAAB" bold="true" box="[189,310,1288,1314]" pageId="9" pageNumber="126">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E42E7963FFBE5B5FFF67FAC1E6DEF988" blockId="9.[187,1400,1335,1537]" pageId="9" pageNumber="126">
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Within the
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<taxonomicName id="239102E0FFBE5B5FFE99FACEE3A8FAD8" authorityName="Dejean" authorityYear="1833" box="[323,472,1336,1361]" pageId="9" pageNumber="144" rank="subGenus" subGenus="Microsaurus">
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<emphasis id="D6E5A571FFBE5B5FFE99FACEE3A8FAD8" box="[323,472,1336,1361]" italics="true" pageId="9" pageNumber="126">Microsaurus</emphasis>
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</taxonomicName>
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lineage,
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<taxonomicName id="239102E0FFBE5B5FFD99FAC1E0B5FAD8" box="[579,709,1335,1361]" class="Insecta" family="Staphylinidae" genus="Malaisdius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="D6E5A571FFBE5B5FFD99FAC1E0B5FAD8" box="[579,709,1335,1361]" italics="true" pageId="9" pageNumber="126">Malaisdius</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CDD6180AFFBE5B5FFD14FAC1E14AFAD8" box="[718,826,1335,1361]" pageId="9" pageNumber="126" rank="genus">gen. nov.</taxonomicNameLabel>
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can be recognized by the following: elytral disc distinctive, uneven and with deeply impressed, foveate and setose punctures (
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<figureCitation id="7CAA65E6FFBE5B5FFB58FAACE691FAFD" box="[1154,1249,1370,1396]" captionStart="Fig" captionStartId="21.[189,228,1904,1928]" captionTargetBox="[191,1395,503,1860]" captionTargetId="figure-97@21.[189,1398,499,1864]" captionTargetPageId="21" captionText="Fig. 4. A–B. Indoquedius dispersepunctatus(Scheerpeltz, 1965). A. Elytra. B. Dorsal abdomen. C–D. I. recticollis (Scheerpeltz, 1965). C. Elytra. D. Dorsal abdomen. E–F. Malaisdius smetanai gen. et sp. nov. E. Head. F. Elytra. Abbreviations: Af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supraantennal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872188" httpUri="https://zenodo.org/record/7872188/files/figure.png" pageId="9" pageNumber="126">Fig. 4F</figureCitation>
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); abdominal tergites III–VI with distinct, coarsely punctate basal impressions, disc otherwise impunctate or nearly so, punctures elongate (
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<figureCitation id="7CAA65E6FFBE5B5FFE72FA56E05AFA32" box="[424,554,1440,1467]" captionStart="Fig" captionStartId="22.[189,228,1840,1864]" captionTargetBox="[192,1395,440,1798]" captionTargetId="figure-53@22.[189,1398,436,1802]" captionTargetPageId="22" captionText="Fig. 5. A. Malaisdius ruficeps (Scheerpeltz, 1965), dorsal abdomen. B. M. smetanai gen. et sp. nov., dorsal abdomen. C–E. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965. C. Antenna. D. Pronotum (white arrow: marginal punctures removed from marginal bead; black arrow: anterior angles with marginal bead entirely visible in dorsal view). E. Elytra and basal abdomen. F. Q. (M.) masasatoi Smetana, 2007, pronotum (arrows indicating punctures of the dorsal row). Abbreviation: Pf = posterior frontal puncture. Scale bars: A–B, D–E = 1 mm; C, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7872192" httpUri="https://zenodo.org/record/7872192/files/figure.png" pageId="9" pageNumber="126">Fig. 5A–B</figureCitation>
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). In the latter two features, species of
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<taxonomicName id="239102E0FFBE5B5FFC30FA56E61CFA33" box="[1002,1132,1440,1466]" class="Insecta" family="Staphylinidae" genus="Malaisdius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="genus" status="gen. nov.">
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||
<emphasis id="D6E5A571FFBE5B5FFC30FA56E61CFA33" box="[1002,1132,1440,1466]" italics="true" pageId="9" pageNumber="126">Malaisdius</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="CDD6180AFFBE5B5FFBA9FA57E6AEFA32" box="[1139,1246,1441,1467]" pageId="9" pageNumber="126" rank="genus">gen. nov.</taxonomicNameLabel>
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superficially resemble some members of cyrtoquediine genus
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<taxonomicName id="239102E0FFBE5B5FFD21FA32E638FA57" authority="Chevrolat, 1842" authorityName="Chevrolat" authorityYear="1842" box="[763,1096,1476,1502]" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E5A571FFBE5B5FFD21FA32E1F4FA57" box="[763,900,1476,1502]" italics="true" pageId="9" pageNumber="126">Bolitogyrus</emphasis>
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Chevrolat, 1842
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</taxonomicName>
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though asetose punctures are always present in
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<taxonomicName id="239102E0FFBE5B5FFE61FA11E034F988" authorityName="Chevrolat" authorityYear="1842" box="[443,580,1511,1537]" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E5A571FFBE5B5FFE61FA11E034F988" box="[443,580,1511,1537]" italics="true" pageId="9" pageNumber="126">Bolitogyrus</emphasis>
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</taxonomicName>
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and the forebody is entirely without microsculpture.
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8B2AE8FFBE5B5FFF67F9DEE1DAF952" pageId="9" pageNumber="126" type="etymology">
|
||
<paragraph id="E42E7963FFBE5B5FFF67F9DEE334F9CB" blockId="9.[189,324,1576,1602]" box="[189,324,1576,1602]" pageId="9" pageNumber="126">
|
||
<heading id="BF66CE0FFFBE5B5FFF67F9DEE334F9CB" bold="true" box="[189,324,1576,1602]" fontSize="11" level="3" pageId="9" pageNumber="126" reason="3">
|
||
<emphasis id="D6E5A571FFBE5B5FFF67F9DEE334F9CB" bold="true" box="[189,324,1576,1602]" pageId="9" pageNumber="126">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E42E7963FFBE5B5FFF67F9A1E1DAF952" blockId="9.[188,1399,1623,1755]" pageId="9" pageNumber="126">
|
||
This genus is dedicated to the Swedish explorer and entomologist René Malaise, who collected a diversity of rove beetles at Kambaiti Pass,
|
||
<collectingRegion id="2655B781FFBE5B5FFD62F98CE160F91D" box="[696,784,1658,1684]" country="Myanmar" name="Kachin" pageId="9" pageNumber="126">Kachin</collectingRegion>
|
||
,
|
||
<collectingCountry id="9C8639F3FFBE5B5FFCC6F98DE1E0F91C" box="[796,912,1659,1685]" name="Myanmar" pageId="9" pageNumber="126">Myanmar</collectingCountry>
|
||
. Malaise is also famous for the invention of the insect trap that bears his name, which has revolutionized our knowledge of insect biodiversity and was first used on a significant scale in Kambaiti (
|
||
<bibRefCitation id="80000492FFBE5B5FFD20F936E1EEF952" author="Sjoberg F." box="[762,926,1728,1755]" pageId="9" pageNumber="126" refId="ref13617" refString="Sjoberg F. 2014. The Fly Trap [English translation]. Pantheon Books, New York." type="book" year="2014">Sjöberg 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC8B2AE8FFBE5B5CFF67F8F4E33DFB8E" lastPageId="10" lastPageNumber="127" pageId="9" pageNumber="126" type="description">
|
||
<paragraph id="E42E7963FFBE5B5FFF67F8F4E33EF895" blockId="9.[189,334,1794,1820]" box="[189,334,1794,1820]" pageId="9" pageNumber="126">
|
||
<heading id="BF66CE0FFFBE5B5FFF67F8F4E33EF895" bold="true" box="[189,334,1794,1820]" fontSize="11" level="3" pageId="9" pageNumber="126" reason="3">
|
||
<emphasis id="D6E5A571FFBE5B5FFF67F8F4E33EF895" bold="true" box="[189,334,1794,1820]" pageId="9" pageNumber="126">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E42E7963FFBE5B5CFF67F8C6E33DFB8E" blockId="9.[188,1399,1840,2008]" lastBlockId="10.[188,1400,265,1031]" lastPageId="10" lastPageNumber="127" pageId="9" pageNumber="126">
|
||
Medium-sized, glossy beetles, with bright coloration. With the character states of
|
||
<taxonomicName id="239102E0FFBE5B5FFB62F8C6E76FF8C3" authorityName="Kraatz" authorityYear="1857" box="[1208,1311,1840,1866]" pageId="9" pageNumber="126" rank="tribe" tribe="Quediini">Quediini</taxonomicName>
|
||
(sensu
|
||
<bibRefCitation id="80000492FFBE5B5FFF67F8A2E3E5F8E7" author="Brunke A. J. & Hansen A. K. & Salnitska M. & Kypke J. L. & Predeus A. V. & Escalona H. E. & Chapados J. T. & Eyres J. & Richter R. & Smetana A. & Slipinski S. A. & Zwick A. & Hajek J. & Leschen R. A. B. & Solodovnikov A. & Dettman J. R." box="[189,405,1876,1902]" pageId="9" pageNumber="126" pagination="396 - 421" refId="ref13357" refString="Brunke A. J., Hansen A. K., Salnitska M., Kypke J. L., Predeus A. V., Escalona H. E., Chapados J. T., Eyres J., Richter R., Smetana A., Slipinski S. A., Zwick A., Hajek J., Leschen R. A. B., Solodovnikov A. & Dettman J. R. 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle megaradiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46: 396 - 421. https: // doi. org / 10.1111 / syen. 12468" type="journal article" year="2021">
|
||
Brunke
|
||
<emphasis id="D6E5A571FFBE5B5FFEC3F8A2E323F8E7" box="[281,339,1876,1902]" italics="true" pageId="9" pageNumber="126">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
) and the following: antennomere 3 longer than 2, without dense setation or tomentose pubescence; outer antennomeres (6–10 or 8–10) weakly transverse and slightly asymmetrical; antennae inserted far from eyes, about 2.0 × the width of antennal sclerite; head with eyes large, strongly convex, protruding from lateral head margin, temples distinct, about half the length of eyes; eyes subparallel, inner margin forming acute angle with supra-antennal ridge; basal puncture doubled, interocular punctures absent, genal puncture present but extremely small, rudimentary; frons not well developed anterolaterad of antennal insertions; labrum notched medially creating two lobes; apical maxillary and labial palpomeres fusiform and glabrous; infraorbital ridge complete to base of mandibles; gular sutures convergent towards neck, narrowly spaced posteriad; mandibles with small dorsolateral groove; right mandible with two teeth, distal tooth on lower plane compared to proximal; pronotum shield-shaped, slightly explanate; marginal punctures touching marginal bead, dorsal rows with three punctures, sublateral rows with only one puncture, therefore not extended posteriad of large lateral puncture (
|
||
<specimenCount id="F297B2EAFFBD5B5CFF1FFDD5E33EFDB5" box="[197,334,546,573]" pageId="10" pageNumber="127" type="female">two female</specimenCount>
|
||
specimens with puncture-like elements (without sockets) giving superficial appearance of 2–3 punctures in the sublateral row), with single large lateral puncture; hypomeron strongly inflexed, hypomeron not visible; basisternum with pair of small, pale macrosetae, with well-developed longitudinal carina; scutellum impunctate; elytron with subbasal ridge complete, forming scutellar collar; row of humeral spines present but short (2–4 spines); disc of elytra without microsculpture between the punctures, surface uneven, punctures irregularly and sparsely distributed, or in very loose rows, punctures bearing setae and situated in broad depressions; foretibia without lateral spines but with apical spur; metatibia with only two slender spines on lateral face; metatarsomeres with disc setose; prototergal glands likely absent, with only vague, shallow impression lacking marginal setae; abdominal tergites III–VI with distinct basal depressions bearing elongate punctures, remainder of disc impunctate or nearly so; abdominal sternite III transverse basal line acutely and sharply produced posteriad at middle; males without modifications on sternite VII; aedeagus asymmetrical in known species, with well-developed paramere, paramere without peg setae; aedeagus without large copulatory sclerites in internal sac.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC8B2AE8FFBD5B5CFF67FBDBE72EFBFE" box="[189,1374,1069,1143]" pageId="10" pageNumber="127" type="distribution">
|
||
<paragraph id="E42E7963FFBD5B5CFF67FBDBE326FBCE" blockId="10.[189,342,1069,1095]" box="[189,342,1069,1095]" pageId="10" pageNumber="127">
|
||
<heading id="BF66CE0FFFBD5B5CFF67FBDBE326FBCE" bold="true" box="[189,342,1069,1095]" fontSize="11" level="3" pageId="10" pageNumber="127" reason="3">
|
||
<emphasis id="D6E5A571FFBD5B5CFF67FBDBE326FBCE" bold="true" box="[189,342,1069,1095]" pageId="10" pageNumber="127">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E42E7963FFBD5B5CFF67FBAAE72EFBFE" blockId="10.[189,1374,1116,1143]" box="[189,1374,1116,1143]" pageId="10" pageNumber="127">
|
||
<taxonomicName id="239102E0FFBD5B5CFF67FBAAE34FFBFF" box="[189,319,1116,1142]" class="Insecta" family="Staphylinidae" genus="Malaisdius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="D6E5A571FFBD5B5CFF67FBAAE34FFBFF" box="[189,319,1116,1142]" italics="true" pageId="10" pageNumber="127">Malaisdius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CDD6180AFFBD5B5CFE9DFBABE3C2FBFE" box="[327,434,1117,1143]" pageId="10" pageNumber="127" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is known from the Himalaya of
|
||
<collectingCountry id="9C8639F3FFBD5B5CFCF5FBAAE105FBFF" box="[815,885,1116,1142]" name="Nepal" pageId="10" pageNumber="127">Nepal</collectingCountry>
|
||
and the mountains of northern
|
||
<collectingCountry id="9C8639F3FFBD5B5CFB3CFBABE72AFBFE" box="[1254,1370,1117,1143]" name="Myanmar" pageId="10" pageNumber="127">Myanmar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC8B2AE8FFBD5B5CFF67FB6BE631FB6E" box="[189,1089,1181,1255]" pageId="10" pageNumber="127" type="biology_ecology">
|
||
<paragraph id="E42E7963FFBD5B5CFF67FB6BE34FFB3E" blockId="10.[189,319,1181,1207]" box="[189,319,1181,1207]" pageId="10" pageNumber="127">
|
||
<heading id="BF66CE0FFFBD5B5CFF67FB6BE34FFB3E" bold="true" box="[189,319,1181,1207]" fontSize="11" level="3" pageId="10" pageNumber="127" reason="3">
|
||
<emphasis id="D6E5A571FFBD5B5CFF67FB6BE34FFB3E" bold="true" box="[189,319,1181,1207]" pageId="10" pageNumber="127">Bionomics</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E42E7963FFBD5B5CFF67FB3AE631FB6E" blockId="10.[189,1089,1228,1255]" box="[189,1089,1228,1255]" pageId="10" pageNumber="127">
|
||
Nothing is currently known about the natural history of
|
||
<taxonomicName id="239102E0FFBD5B5CFC97FB3AE1BFFB6F" box="[845,975,1228,1254]" class="Insecta" family="Staphylinidae" genus="Malaisdius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="D6E5A571FFBD5B5CFC97FB3AE1BFFB6F" box="[845,975,1228,1254]" italics="true" pageId="10" pageNumber="127">Malaisdius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CDD6180AFFBD5B5CFC0CFB3BE631FB6E" box="[982,1089,1229,1255]" pageId="10" pageNumber="127" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC8B2AE8FFBD5B5CFF67FAF8E185FA15" pageId="10" pageNumber="127" type="discussion">
|
||
<paragraph id="E42E7963FFBD5B5CFF67FAF8E35FFAA1" blockId="10.[189,303,1294,1320]" box="[189,303,1294,1320]" pageId="10" pageNumber="127">
|
||
<heading id="BF66CE0FFFBD5B5CFF67FAF8E35FFAA1" bold="true" box="[189,303,1294,1320]" fontSize="11" level="3" pageId="10" pageNumber="127" reason="3">
|
||
<emphasis id="D6E5A571FFBD5B5CFF67FAF8E35FFAA1" bold="true" box="[189,303,1294,1320]" pageId="10" pageNumber="127">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E42E7963FFBD5B5CFF67FACAE185FA15" blockId="10.[189,1399,1340,1437]" pageId="10" pageNumber="127">
|
||
According to phylogenomic analyses (
|
||
<bibRefCitation id="80000492FFBD5B5CFD5CFACAE11BFADF" author="Brunke A. J. & Hansen A. K. & Salnitska M. & Kypke J. L. & Predeus A. V. & Escalona H. E. & Chapados J. T. & Eyres J. & Richter R. & Smetana A. & Slipinski S. A. & Zwick A. & Hajek J. & Leschen R. A. B. & Solodovnikov A. & Dettman J. R." box="[646,875,1340,1366]" pageId="10" pageNumber="127" pagination="396 - 421" refId="ref13357" refString="Brunke A. J., Hansen A. K., Salnitska M., Kypke J. L., Predeus A. V., Escalona H. E., Chapados J. T., Eyres J., Richter R., Smetana A., Slipinski S. A., Zwick A., Hajek J., Leschen R. A. B., Solodovnikov A. & Dettman J. R. 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle megaradiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46: 396 - 421. https: // doi. org / 10.1111 / syen. 12468" type="journal article" year="2021">
|
||
Brunke
|
||
<emphasis id="D6E5A571FFBD5B5CFD33FACBE153FADF" box="[745,803,1340,1366]" italics="true" pageId="10" pageNumber="127">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
), the sister group of
|
||
<taxonomicName id="239102E0FFBD5B5CFBBEFACAE696FADF" box="[1124,1254,1340,1366]" class="Insecta" family="Staphylinidae" genus="Malaisdius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="D6E5A571FFBD5B5CFBBEFACAE696FADF" box="[1124,1254,1340,1366]" italics="true" pageId="10" pageNumber="127">Malaisdius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CDD6180AFFBD5B5CFB34FACBE72AFADE" box="[1262,1370,1341,1367]" pageId="10" pageNumber="127" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is the more generalized Masasatoi group (see below), currently included within polyphyletic
|
||
<taxonomicName id="239102E0FFBD5B5CFACFFAA9E316FA14" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="subGenus" subGenus="Microsaurus">
|
||
<emphasis id="D6E5A571FFBD5B5CFACFFAA9E707FAF0" box="[1301,1399,1375,1401]" italics="true" pageId="10" pageNumber="127">Quedius</emphasis>
|
||
(
|
||
<emphasis id="D6E5A571FFBD5B5CFF1DFA75E32CFA15" box="[199,348,1411,1436]" italics="true" pageId="10" pageNumber="127">Microsaurus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
but will be treated as a separate genus in a future study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC8B2AE8FFBD5B5CFF67FA30E090FA6B" box="[189,736,1477,1506]" pageId="10" pageNumber="127" type="multiple">
|
||
<paragraph id="E42E7963FFBD5B5CFF67FA30E090FA6B" blockId="10.[189,736,1477,1506]" box="[189,736,1477,1506]" pageId="10" pageNumber="127">
|
||
<emphasis id="D6E5A571FFBD5B5CFF67FA30E090FA6B" bold="true" box="[189,736,1477,1506]" italics="true" pageId="10" pageNumber="127">
|
||
Key to the species of
|
||
<taxonomicName id="239102E0FFBD5B5CFE0AFA33E012FA6B" box="[464,610,1477,1506]" class="Insecta" family="Staphylinidae" genus="Malaisdius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="genus" status="gen. nov.">Malaisdius</taxonomicName>
|
||
<taxonomicNameLabel id="CDD6180AFFBD5B5CFDB3FA30E090FA6B" box="[617,736,1478,1506]" pageId="10" pageNumber="127" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC8B2AE8FFBD5B5CFF67FA0CE707F883" pageId="10" pageNumber="127" type="key">
|
||
<keyStep id="5F6561C6FFBD5B5CFF67FA0CE707F883" pageId="10" pageNumber="127">
|
||
<key id="90002E52FFBD5B5CFF67FA0CE707F883" pageId="10" pageNumber="127">
|
||
<paragraph id="E42E7963FFBD5B5CFF67FA0CE707F9F7" blockId="10.[189,1399,1529,1802]" pageId="10" pageNumber="127">
|
||
<keyLead id="5F60DA56FFBD5B5CFF67FA0CE707F9F7" pageId="10" pageNumber="127">
|
||
1. Body bicolored, head and pronotum orange-red, sharply contrasting with elytra and abdomen (
|
||
<figureCitation id="7CAA65E6FFBD5B5CFF29F9EBE323F9BE" box="[243,339,1565,1591]" captionStart="Fig" captionStartId="16.[189,228,1872,1896]" captionTargetBox="[190,1396,472,1833]" captionTargetId="figure-78@16.[189,1398,471,1834]" captionTargetPageId="16" captionText="Fig. 2. A–E. Dorsal habitus. A. Malaisdius ruficeps (Scheerpeltz, 1965), ♂, paratype (NHW). B. Malaisdius smetanai gen. et. sp. nov., ♂, holotype (NME). C. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965, ♂, holotype (NHRS). D. Q. (M.) piceolineatus Scheerpeltz, 1965, ♂ holotype (NHRS). E. Q. (M.) impressithorax Scheerpeltz, 1965, ♀ holotype (NHRS). F. Forebody of Q. (M.) impressithorax Scheerpeltz, 1965. Abbreviation: Pf = posterior frontal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872182" httpUri="https://zenodo.org/record/7872182/files/figure.png" pageId="10" pageNumber="127">Fig. 2A</figureCitation>
|
||
); legs entirely pale (
|
||
<figureCitation id="7CAA65E6FFBD5B5CFD9DF9EBE0D7F9BE" box="[583,679,1565,1591]" captionStart="Fig" captionStartId="16.[189,228,1872,1896]" captionTargetBox="[190,1396,472,1833]" captionTargetId="figure-78@16.[189,1398,471,1834]" captionTargetPageId="16" captionText="Fig. 2. A–E. Dorsal habitus. A. Malaisdius ruficeps (Scheerpeltz, 1965), ♂, paratype (NHW). B. Malaisdius smetanai gen. et. sp. nov., ♂, holotype (NME). C. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965, ♂, holotype (NHRS). D. Q. (M.) piceolineatus Scheerpeltz, 1965, ♂ holotype (NHRS). E. Q. (M.) impressithorax Scheerpeltz, 1965, ♀ holotype (NHRS). F. Forebody of Q. (M.) impressithorax Scheerpeltz, 1965. Abbreviation: Pf = posterior frontal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872182" httpUri="https://zenodo.org/record/7872182/files/figure.png" pageId="10" pageNumber="127">Fig. 2A</figureCitation>
|
||
); elytral punctures irregularly placed; paramere in basal half subequal in width to median lobe, apex not hooked (
|
||
<figureCitation id="7CAA65E6FFBD5B5CFC89F9B6E1C1F9D2" box="[851,945,1600,1627]" captionStart="Fig" captionStartId="24.[189,228,1872,1896]" captionTargetBox="[229,1350,707,1815]" captionTargetId="figure-144@24.[223,1364,704,1834]" captionTargetPageId="24" captionText="Fig. 6. A, F, H–I, K–L.Aedeagus, in situ. A, F, H, K. Ventral view (inset showing paired teeth). I, L. Lateral view. B–D. Median lobe. B. Ventral view. C–D. Lateral view. E, G, J, M. Underside of paramere. A–E. Malaisdius ruficeps (Scheerpeltz, 1965). F–G. M. smetanai gen. et sp. nov. H–J. Quedius (Microsaurus) piceolineatus Scheerpeltz, 1965. K–M. Q. (M.) masasatoi Smetana, 2007. Scale bars: A–D, F, H–I, K–L = 0.5 mm; E, G, J, M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872196" httpUri="https://zenodo.org/record/7872196/files/figure.png" pageId="10" pageNumber="127">Fig. 6A</figureCitation>
|
||
);
|
||
<collectingCountry id="9C8639F3FFBD5B5CFC1DF9B6E649F9D3" box="[967,1081,1600,1626]" name="Myanmar" pageId="10" pageNumber="127">Myanmar</collectingCountry>
|
||
......................................... ........................................................................................
|
||
<taxonomicName id="239102E0FFBD5B5CFCA2F995E69DF9F7" ID-CoL="4R3V3" authority="(Scheerpeltz, 1965)" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1965" box="[888,1261,1635,1662]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="species" species="ruficeps" status="comb. nov.">
|
||
<emphasis id="D6E5A571FFBD5B5CFCA2F995E671F9F4" bold="true" box="[888,1025,1635,1661]" italics="true" pageId="10" pageNumber="127">M. ruficeps</emphasis>
|
||
(
|
||
<bibRefCitation id="80000492FFBD5B5CFBCAF995E695F9F7" author="Scheerpeltz O." box="[1040,1253,1635,1662]" pageId="10" pageNumber="127" pagination="93 - 371" refId="ref13560" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Arkiv for Zoologi 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CDD6180AFFBD5B5CFB2FF995E707F9F7" box="[1269,1399,1635,1662]" pageId="10" pageNumber="127" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="E42E7963FFBD5B5CFF1BF971E707F883" blockId="10.[189,1399,1529,1802]" pageId="10" pageNumber="127">
|
||
<keyLead id="5F60DA56FFBD5B5CFF1BF971E707F883" pageId="10" pageNumber="127">
|
||
– Body more evenly colored, yellow-orange, with darker areas on the elytra (
|
||
<figureCitation id="7CAA65E6FFBD5B5CFBA1F970E6A9F929" box="[1147,1241,1670,1696]" captionStart="Fig" captionStartId="16.[189,228,1872,1896]" captionTargetBox="[190,1396,472,1833]" captionTargetId="figure-78@16.[189,1398,471,1834]" captionTargetPageId="16" captionText="Fig. 2. A–E. Dorsal habitus. A. Malaisdius ruficeps (Scheerpeltz, 1965), ♂, paratype (NHW). B. Malaisdius smetanai gen. et. sp. nov., ♂, holotype (NME). C. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965, ♂, holotype (NHRS). D. Q. (M.) piceolineatus Scheerpeltz, 1965, ♂ holotype (NHRS). E. Q. (M.) impressithorax Scheerpeltz, 1965, ♀ holotype (NHRS). F. Forebody of Q. (M.) impressithorax Scheerpeltz, 1965. Abbreviation: Pf = posterior frontal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872182" httpUri="https://zenodo.org/record/7872182/files/figure.png" pageId="10" pageNumber="127">Fig. 2B</figureCitation>
|
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); hind femur with apical dark band (
|
||
<figureCitation id="7CAA65E6FFBD5B5CFDDAF95FE02EF94D" box="[512,606,1705,1732]" captionStart="Fig" captionStartId="16.[189,228,1872,1896]" captionTargetBox="[190,1396,472,1833]" captionTargetId="figure-78@16.[189,1398,471,1834]" captionTargetPageId="16" captionText="Fig. 2. A–E. Dorsal habitus. A. Malaisdius ruficeps (Scheerpeltz, 1965), ♂, paratype (NHW). B. Malaisdius smetanai gen. et. sp. nov., ♂, holotype (NME). C. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965, ♂, holotype (NHRS). D. Q. (M.) piceolineatus Scheerpeltz, 1965, ♂ holotype (NHRS). E. Q. (M.) impressithorax Scheerpeltz, 1965, ♀ holotype (NHRS). F. Forebody of Q. (M.) impressithorax Scheerpeltz, 1965. Abbreviation: Pf = posterior frontal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872182" httpUri="https://zenodo.org/record/7872182/files/figure.png" pageId="10" pageNumber="127">Fig. 2B</figureCitation>
|
||
); elytral punctures in loose rows (
|
||
<figureCitation id="7CAA65E6FFBD5B5CFC23F95FE625F94D" box="[1017,1109,1705,1732]" captionStart="Fig" captionStartId="21.[189,228,1904,1928]" captionTargetBox="[191,1395,503,1860]" captionTargetId="figure-97@21.[189,1398,499,1864]" captionTargetPageId="21" captionText="Fig. 4. A–B. Indoquedius dispersepunctatus(Scheerpeltz, 1965). A. Elytra. B. Dorsal abdomen. C–D. I. recticollis (Scheerpeltz, 1965). C. Elytra. D. Dorsal abdomen. E–F. Malaisdius smetanai gen. et sp. nov. E. Head. F. Elytra. Abbreviations: Af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supraantennal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872188" httpUri="https://zenodo.org/record/7872188/files/figure.png" pageId="10" pageNumber="127">Fig. 4F</figureCitation>
|
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); paramere in basal half distinctly narrower than median lobe, apex hooked (
|
||
<figureCitation id="7CAA65E6FFBD5B5CFC8AF93BE1D9F961" box="[848,937,1741,1768]" captionStart="Fig" captionStartId="24.[189,228,1872,1896]" captionTargetBox="[229,1350,707,1815]" captionTargetId="figure-144@24.[223,1364,704,1834]" captionTargetPageId="24" captionText="Fig. 6. A, F, H–I, K–L.Aedeagus, in situ. A, F, H, K. Ventral view (inset showing paired teeth). I, L. Lateral view. B–D. Median lobe. B. Ventral view. C–D. Lateral view. E, G, J, M. Underside of paramere. A–E. Malaisdius ruficeps (Scheerpeltz, 1965). F–G. M. smetanai gen. et sp. nov. H–J. Quedius (Microsaurus) piceolineatus Scheerpeltz, 1965. K–M. Q. (M.) masasatoi Smetana, 2007. Scale bars: A–D, F, H–I, K–L = 0.5 mm; E, G, J, M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872196" httpUri="https://zenodo.org/record/7872196/files/figure.png" pageId="10" pageNumber="127">Fig. 6F</figureCitation>
|
||
);
|
||
<collectingCountry id="9C8639F3FFBD5B5CFC65F93BE675F96E" box="[959,1029,1741,1767]" name="Nepal" pageId="10" pageNumber="127">Nepal</collectingCountry>
|
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................................................ ................................................................................................................
|
||
<taxonomicName id="239102E0FFBD5B5CFBF3F906E6CFF883" authorityName="Brunke, des Brunke" authorityYear="2023" box="[1065,1215,1776,1802]" class="Insecta" family="Staphylinidae" genus="Malaisdius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="species" species="smetanai" status="gen. et sp. nov.">
|
||
<emphasis id="D6E5A571FFBD5B5CFBF3F906E6CFF883" bold="true" box="[1065,1215,1776,1802]" italics="true" pageId="10" pageNumber="127">M. smetanai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CDD6180AFFBD5B5CFB1DF906E707F883" box="[1223,1399,1776,1802]" pageId="10" pageNumber="127" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |