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<mods:title id="A3C1861D2CB50DB05385B5A6085ADF57">Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar</mods:title>
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<treatment id="6C38C875FFBA5B58FDE4FBD3E3ABFECF" ID-DOI="http://doi.org/10.5281/zenodo.7872298" ID-GBIF-Taxon="209579351" ID-Zenodo-Dep="7872298" LSID="urn:lsid:plazi:treatment:6C38C875FFBA5B58FDE4FBD3E3ABFECF" httpUri="http://treatment.plazi.org/id/6C38C875FFBA5B58FDE4FBD3E3ABFECF" lastPageId="14" lastPageNumber="130" pageId="13" pageNumber="130">
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<paragraph id="E42E7963FFBA5B5BFDE4FBD3E186FBB6" blockId="13.[574,1014,1061,1087]" box="[574,1014,1061,1087]" pageId="13" pageNumber="130">
<heading id="BF66CE0FFFBA5B5BFDE4FBD3E186FBB6" box="[574,1014,1061,1087]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="130" reason="2">
Subgenus
<taxonomicName id="239102E0FFBA5B5BFD6DFBD3E186FBB6" authority="Dejean, 1833" authorityName="Dejean" authorityYear="1833" box="[695,1014,1061,1087]" class="Insecta" family="Staphylinidae" genus="Microsaurus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="D6E5A571FFBA5B5BFD6DFBD3E122FBB6" bold="true" box="[695,850,1061,1087]" italics="true" pageId="13" pageNumber="130">Microsaurus</emphasis>
Dejean, 1833
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFBA5B5BFF67FB91E75EFAB5" pageId="13" pageNumber="130" type="discussion">
<paragraph id="E42E7963FFBA5B5BFF67FB91E35FFB08" blockId="13.[189,303,1127,1153]" box="[189,303,1127,1153]" pageId="13" pageNumber="130">
<heading id="BF66CE0FFFBA5B5BFF67FB91E35FFB08" bold="true" box="[189,303,1127,1153]" fontSize="11" level="3" pageId="13" pageNumber="130" reason="3">
<emphasis id="D6E5A571FFBA5B5BFF67FB91E35FFB08" bold="true" box="[189,303,1127,1153]" pageId="13" pageNumber="130">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFBA5B5BFF67FB63E75EFAB5" blockId="13.[188,1400,1173,1340]" pageId="13" pageNumber="130">
This diverse subgenus is in great need of generic revision as it is rendered polyphyletic by several genera and subgenera of
<taxonomicName id="239102E0FFBA5B5BFE42FB4EE072FB5B" authorityName="Kraatz" authorityYear="1857" box="[408,514,1208,1234]" pageId="13" pageNumber="126" rank="tribe" tribe="Quediini">Quediini</taxonomicName>
, including
<taxonomicName id="239102E0FFBA5B5BFD57FB4EE17FFB5B" box="[653,783,1208,1234]" class="Insecta" family="Staphylinidae" genus="Malaisdius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="D6E5A571FFBA5B5BFD57FB4EE17FFB5B" box="[653,783,1208,1234]" italics="true" pageId="13" pageNumber="130">Malaisdius</emphasis>
</taxonomicName>
(see further comments above under “
<taxonomicName id="239102E0FFBA5B5BFB38FB4FE707FB5B" authorityName="Dejean" authorityYear="1833" box="[1250,1399,1209,1234]" pageId="13" pageNumber="144" rank="subGenus" subGenus="Microsaurus">
<emphasis id="D6E5A571FFBA5B5BFB38FB4FE707FB5B" box="[1250,1399,1209,1234]" italics="true" pageId="13" pageNumber="130">Microsaurus</emphasis>
</taxonomicName>
lineage”). Currently, it is not possible to give a meaningful diagnosis of
<taxonomicName id="239102E0FFBA5B5BFBFDFB2DE74BFB7F" authorityName="Stephens" authorityYear="1829" box="[1063,1339,1243,1270]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="subGenus" subGenus="Microsaurus">
<emphasis id="D6E5A571FFBA5B5BFBFDFB2DE6F9FB7C" box="[1063,1161,1243,1269]" italics="true" pageId="13" pageNumber="130">Quedius</emphasis>
(
<emphasis id="D6E5A571FFBA5B5BFB40FB2AE75FFB7C" box="[1178,1327,1244,1269]" italics="true" pageId="13" pageNumber="130">Microsaurus</emphasis>
)
</taxonomicName>
. The species groups listed below are redefined from
<bibRefCitation id="80000492FFBA5B5BFD38FB09E1DBFA90" author="Smetana A." box="[738,939,1279,1305]" pageId="13" pageNumber="130" refId="ref13920" refString="Smetana A. 2017 a. Quediine Subtribes of Staphylinini (Coleoptera, Staphylinidae, Staphylininae) of Mainland China. J. Farkac, Prague." type="book" year="2017">Smetana (2017a)</bibRefCitation>
, likely monophyletic and correspond to clades in
<bibRefCitation id="80000492FFBA5B5BFEF6FAD4E06DFAB5" author="Brunke A. J. &amp; Hansen A. K. &amp; Salnitska M. &amp; Kypke J. L. &amp; Predeus A. V. &amp; Escalona H. E. &amp; Chapados J. T. &amp; Eyres J. &amp; Richter R. &amp; Smetana A. &amp; Slipinski S. A. &amp; Zwick A. &amp; Hajek J. &amp; Leschen R. A. B. &amp; Solodovnikov A. &amp; Dettman J. R." box="[300,541,1314,1340]" pageId="13" pageNumber="130" pagination="396 - 421" refId="ref13357" refString="Brunke A. J., Hansen A. K., Salnitska M., Kypke J. L., Predeus A. V., Escalona H. E., Chapados J. T., Eyres J., Richter R., Smetana A., Slipinski S. A., Zwick A., Hajek J., Leschen R. A. B., Solodovnikov A. &amp; Dettman J. R. 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle megaradiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46: 396 - 421. https: // doi. org / 10.1111 / syen. 12468" type="journal article" year="2021">
Brunke
<emphasis id="D6E5A571FFBA5B5BFE53FAD5E3B4FAB5" box="[393,452,1314,1340]" italics="true" pageId="13" pageNumber="130">et al.</emphasis>
(2021)
</bibRefCitation>
. They will be treated in much greater detail in a future publication.
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFBA5B58FF67FA93E3ABFECF" lastPageId="14" lastPageNumber="131" pageId="13" pageNumber="130" type="multiple">
<paragraph id="E42E7963FFBA5B5BFF67FA93E034FA08" blockId="13.[189,580,1381,1409]" box="[189,580,1381,1409]" pageId="13" pageNumber="130">
<emphasis id="D6E5A571FFBA5B5BFF67FA93E034FA08" bold="true" box="[189,580,1381,1409]" italics="true" pageId="13" pageNumber="130">Apicicornis group, sensu nov.</emphasis>
</paragraph>
<paragraph id="E42E7963FFBA5B58FF67FA6FE3ABFECF" blockId="13.[188,1401,1433,1988]" lastBlockId="14.[189,1398,265,327]" lastPageId="14" lastPageNumber="131" pageId="13" pageNumber="130">
<bibRefCitation id="80000492FFBA5B5BFF67FA6FE3F4FA3A" author="Smetana A." box="[189,388,1433,1459]" pageId="13" pageNumber="130" refId="ref13920" refString="Smetana A. 2017 a. Quediine Subtribes of Staphylinini (Coleoptera, Staphylinidae, Staphylininae) of Mainland China. J. Farkac, Prague." type="book" year="2017">Smetana (2017a)</bibRefCitation>
(and additional references listed therein) have recognized the Beesoni and Apicicornis species groups of
<taxonomicName id="239102E0FFBA5B5BFE44FA4AE0C5FA5E" authorityName="Stephens" authorityYear="1829" box="[414,693,1468,1495]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="subGenus" subGenus="Microsaurus">
<emphasis id="D6E5A571FFBA5B5BFE44FA4AE070FA5F" box="[414,512,1468,1494]" italics="true" pageId="13" pageNumber="130">Quedius</emphasis>
(
<emphasis id="D6E5A571FFBA5B5BFDCCFA4BE0DBFA5F" box="[534,683,1469,1494]" italics="true" pageId="13" pageNumber="130">Microsaurus</emphasis>
)
</taxonomicName>
as separate yet similar taxa that are commonly collected and abundant components of the East Palaearctic and Oriental fauna. Recently, it was shown that the Beesoni group rendered the Apicicornis group paraphyletic but they together formed a monophyletic Clade D within the
<taxonomicName id="239102E0FFBA5B5BFE18F9D0E027F9B6" authorityName="Dejean" authorityYear="1833" box="[450,599,1574,1599]" pageId="13" pageNumber="144" rank="subGenus" subGenus="Microsaurus">
<emphasis id="D6E5A571FFBA5B5BFE18F9D0E027F9B6" box="[450,599,1574,1599]" italics="true" pageId="13" pageNumber="130">Microsaurus</emphasis>
</taxonomicName>
lineage, with the exception of
<taxonomicName id="239102E0FFBA5B5BFC0FF9D0E707F9C9" authority="Smetana, 2007" authorityName="Smetana" authorityYear="2007" box="[981,1399,1574,1600]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="species" species="masatakai">
<emphasis id="D6E5A571FFBA5B5BFC0FF9D0E6CAF9C9" box="[981,1210,1574,1600]" italics="true" pageId="13" pageNumber="130">Quedius masatakai</emphasis>
<bibRefCitation id="80000492FFBA5B5BFB1FF9D0E707F9C9" author="Smetana A." box="[1221,1399,1574,1600]" pageId="13" pageNumber="130" pagination="68 - 75" refId="ref13669" refString="Smetana A. 2007. Three species of the subgenus Microsaurus Dejean, 1833 of the genus Quedius Stephens, 1829 (Coleoptera, Staphylinidae, Staphylinini, Quediina) from Northern Laos. Elytra 35: 68 - 75." type="journal article" year="2007">Smetana, 2007</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="239102E0FFBA5B5BFF2CF9BFE02CF9EA" authority="Smetana, 2007" authorityName="Smetana" authorityYear="2007" box="[246,604,1609,1635]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="species" species="masasatoi">
<emphasis id="D6E5A571FFBA5B5BFF2CF9BFE3E8F9EA" box="[246,408,1609,1635]" italics="true" pageId="13" pageNumber="130">Q. masasatoi</emphasis>
<bibRefCitation id="80000492FFBA5B5BFE7CF9BFE02CF9EA" author="Smetana A." box="[422,604,1609,1635]" pageId="13" pageNumber="130" pagination="68 - 75" refId="ref13669" refString="Smetana A. 2007. Three species of the subgenus Microsaurus Dejean, 1833 of the genus Quedius Stephens, 1829 (Coleoptera, Staphylinidae, Staphylinini, Quediina) from Northern Laos. Elytra 35: 68 - 75." type="journal article" year="2007">Smetana, 2007</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="80000492FFBA5B5BFDA8F9BFE119F9EA" author="Brunke A. J. &amp; Hansen A. K. &amp; Salnitska M. &amp; Kypke J. L. &amp; Predeus A. V. &amp; Escalona H. E. &amp; Chapados J. T. &amp; Eyres J. &amp; Richter R. &amp; Smetana A. &amp; Slipinski S. A. &amp; Zwick A. &amp; Hajek J. &amp; Leschen R. A. B. &amp; Solodovnikov A. &amp; Dettman J. R." box="[626,873,1609,1635]" pageId="13" pageNumber="130" pagination="396 - 421" refId="ref13357" refString="Brunke A. J., Hansen A. K., Salnitska M., Kypke J. L., Predeus A. V., Escalona H. E., Chapados J. T., Eyres J., Richter R., Smetana A., Slipinski S. A., Zwick A., Hajek J., Leschen R. A. B., Solodovnikov A. &amp; Dettman J. R. 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle megaradiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46: 396 - 421. https: // doi. org / 10.1111 / syen. 12468" type="journal article" year="2021">
Brunke
<emphasis id="D6E5A571FFBA5B5BFD03F9BCE16BF9EA" box="[729,795,1609,1635]" italics="true" pageId="13" pageNumber="130">et al.</emphasis>
2021
</bibRefCitation>
). Here the former two groups, minus the Masasatoi group (see below), are combined to form a newly defined Apicicornis species group that can be recognized globally within the lineage of
<taxonomicName id="239102E0FFBA5B5BFD06F966E101F920" authorityName="Dejean" authorityYear="1833" box="[732,881,1680,1705]" pageId="13" pageNumber="144" rank="subGenus" subGenus="Microsaurus">
<emphasis id="D6E5A571FFBA5B5BFD06F966E101F920" box="[732,881,1680,1705]" italics="true" pageId="13" pageNumber="130">Microsaurus</emphasis>
</taxonomicName>
by the following combination of character states: antennomeres 610 simple (not serrate or bipectinate) but inflated (i.e., not flattened,
<figureCitation id="7CAA65E6FFBA5B5BFAD6F944E718F944" box="[1292,1384,1714,1741]" captionStart="Fig" captionStartId="22.[189,228,1840,1864]" captionTargetBox="[192,1395,440,1798]" captionTargetId="figure-53@22.[189,1398,436,1802]" captionTargetPageId="22" captionText="Fig. 5. A. Malaisdius ruficeps (Scheerpeltz, 1965), dorsal abdomen. B. M. smetanai gen. et sp. nov., dorsal abdomen. CE. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965. C. Antenna. D. Pronotum (white arrow: marginal punctures removed from marginal bead; black arrow: anterior angles with marginal bead entirely visible in dorsal view). E. Elytra and basal abdomen. F. Q. (M.) masasatoi Smetana, 2007, pronotum (arrows indicating punctures of the dorsal row). Abbreviation: Pf = posterior frontal puncture. Scale bars: AB, DE = 1 mm; C, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7872192" httpUri="https://zenodo.org/record/7872192/files/figure.png" pageId="13" pageNumber="130">Fig. 5C</figureCitation>
), antennomere 10 at most weakly transverse, many species with all antennomeres elongate; eyes large but with distinct temple, eye strongly protruding from head outline and with posterior frontal puncture situated at about the hind margin of eye (
<figureCitation id="7CAA65E6FFBA5B5BFD6AF8EAE160F8BE" box="[688,784,1820,1847]" captionStart="Fig" captionStartId="22.[189,228,1840,1864]" captionTargetBox="[192,1395,440,1798]" captionTargetId="figure-53@22.[189,1398,436,1802]" captionTargetPageId="22" captionText="Fig. 5. A. Malaisdius ruficeps (Scheerpeltz, 1965), dorsal abdomen. B. M. smetanai gen. et sp. nov., dorsal abdomen. CE. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965. C. Antenna. D. Pronotum (white arrow: marginal punctures removed from marginal bead; black arrow: anterior angles with marginal bead entirely visible in dorsal view). E. Elytra and basal abdomen. F. Q. (M.) masasatoi Smetana, 2007, pronotum (arrows indicating punctures of the dorsal row). Abbreviation: Pf = posterior frontal puncture. Scale bars: AB, DE = 1 mm; C, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7872192" httpUri="https://zenodo.org/record/7872192/files/figure.png" pageId="13" pageNumber="130">Fig. 5D</figureCitation>
); pronotum at least slightly explanate, with at least some marginal punctures removed from the marginal bead (
<figureCitation id="7CAA65E6FFBA5B5BFC52F8C9E197F8D3" box="[904,999,1855,1882]" captionStart="Fig" captionStartId="22.[189,228,1840,1864]" captionTargetBox="[192,1395,440,1798]" captionTargetId="figure-53@22.[189,1398,436,1802]" captionTargetPageId="22" captionText="Fig. 5. A. Malaisdius ruficeps (Scheerpeltz, 1965), dorsal abdomen. B. M. smetanai gen. et sp. nov., dorsal abdomen. CE. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965. C. Antenna. D. Pronotum (white arrow: marginal punctures removed from marginal bead; black arrow: anterior angles with marginal bead entirely visible in dorsal view). E. Elytra and basal abdomen. F. Q. (M.) masasatoi Smetana, 2007, pronotum (arrows indicating punctures of the dorsal row). Abbreviation: Pf = posterior frontal puncture. Scale bars: AB, DE = 1 mm; C, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7872192" httpUri="https://zenodo.org/record/7872192/files/figure.png" pageId="13" pageNumber="130">Fig. 5D</figureCitation>
), anterior angles broadly rounded and with marginal bead entirely visible as seen in dorsal view (
<figureCitation id="7CAA65E6FFBA5B5BFC70F894E679F8F4" box="[938,1033,1890,1917]" captionStart="Fig" captionStartId="22.[189,228,1840,1864]" captionTargetBox="[192,1395,440,1798]" captionTargetId="figure-53@22.[189,1398,436,1802]" captionTargetPageId="22" captionText="Fig. 5. A. Malaisdius ruficeps (Scheerpeltz, 1965), dorsal abdomen. B. M. smetanai gen. et sp. nov., dorsal abdomen. CE. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965. C. Antenna. D. Pronotum (white arrow: marginal punctures removed from marginal bead; black arrow: anterior angles with marginal bead entirely visible in dorsal view). E. Elytra and basal abdomen. F. Q. (M.) masasatoi Smetana, 2007, pronotum (arrows indicating punctures of the dorsal row). Abbreviation: Pf = posterior frontal puncture. Scale bars: AB, DE = 1 mm; C, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7872192" httpUri="https://zenodo.org/record/7872192/files/figure.png" pageId="13" pageNumber="130">Fig. 5D</figureCitation>
); scutellum impunctate; elytral punctures uniform in size, setose and evenly distributed (
<figureCitation id="7CAA65E6FFBA5B5BFCBEF870E1CFF829" box="[868,959,1925,1952]" captionStart="Fig" captionStartId="22.[189,228,1840,1864]" captionTargetBox="[192,1395,440,1798]" captionTargetId="figure-53@22.[189,1398,436,1802]" captionTargetPageId="22" captionText="Fig. 5. A. Malaisdius ruficeps (Scheerpeltz, 1965), dorsal abdomen. B. M. smetanai gen. et sp. nov., dorsal abdomen. CE. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965. C. Antenna. D. Pronotum (white arrow: marginal punctures removed from marginal bead; black arrow: anterior angles with marginal bead entirely visible in dorsal view). E. Elytra and basal abdomen. F. Q. (M.) masasatoi Smetana, 2007, pronotum (arrows indicating punctures of the dorsal row). Abbreviation: Pf = posterior frontal puncture. Scale bars: AB, DE = 1 mm; C, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7872192" httpUri="https://zenodo.org/record/7872192/files/figure.png" pageId="13" pageNumber="130">Fig. 5E</figureCitation>
), in most species (not
<taxonomicName id="239102E0FFBA5B5BFB11F870E707F829" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[1227,1399,1926,1952]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="species" species="rutilipennis">
<emphasis id="D6E5A571FFBA5B5BFB11F870E707F829" box="[1227,1399,1926,1952]" italics="true" pageId="13" pageNumber="130">Q. rutilipennis</emphasis>
</taxonomicName>
and several others) fine and dense. The Apicicornis group as defined here should be treated as a separate genus but this is outside of the scope of this paper and a more thorough morphological study is needed for a formal description.
</paragraph>
</subSubSection>
</treatment>
</document>