139 lines
17 KiB
XML
139 lines
17 KiB
XML
<document id="DC933E58133112C4B96DB04CBC98A09F" ID-DOI="10.11646/zootaxa.4020.1.7" ID-GBIF-Dataset="82b2f539-5d25-42d6-95c5-e8b16a353e12" ID-ISSN="1175-5326" ID-Zenodo-Dep="237970" ID-ZooBank="BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461269068152" checkinUser="plazi" docAuthor="Bickel, Daniel J." docDate="2015" docId="172D87FDFFB4FF84FF3DA1C605D2FAC3" docLanguage="en" docName="zt04020p182.pdf" docOrigin="Zootaxa 4020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus eboritibia Bickel, sp. nov." docType="treatment" docVersion="8" lastPageNumber="179" masterDocId="EB14FF85FFBDFF8EFFAAA70A0459FFCC" masterDocTitle="The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus" masterLastPageNumber="182" masterPageNumber="169" pageNumber="178" updateTime="1698611248703" updateUser="plazi">
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<mods:mods id="F68050C6CDE86234E183E3C15BBE34CA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CF99D2D5F30D7296FBEDA82F89BE3076">
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<mods:title id="169570424D49CB744B23258682FC0436">The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus</mods:title>
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</mods:titleInfo>
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<mods:name id="4CD0BA2899C28209FEDB47134281FEC0" type="personal">
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<mods:role id="CE52B14968B6CC5EEDDAEBBBD12F9DE8">
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<mods:roleTerm id="B7CC372BE6A901BC56172185543C55F2">Author</mods:roleTerm>
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<mods:namePart id="F516EC86D1DDFA9CAE77B6E1604172A2">Bickel, Daniel J.</mods:namePart>
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<mods:typeOfResource id="AE26AD85DD121697F4AF7A8BA7950D90">text</mods:typeOfResource>
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<mods:title id="A25A3F0719AEB617F2B87BE16B544771">Zootaxa</mods:title>
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<mods:part id="DBE3E09830B3C8CF12A67138BBB81693">
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<mods:date id="AD86691F62AB901EB55DC8409DE17879">2015</mods:date>
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<mods:detail id="8EDBCC1513B0EB64EFE0E3AE826DD6EE" type="volume">
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<mods:number id="5ED2F0FB037FBB555AF93D06F72E5A0A">4020</mods:number>
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</mods:detail>
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<mods:detail id="2F4F137F03804D9D73AA922A82F71740" type="issue">
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<mods:number id="76C48C84EF1F1FB43CD7097DB83A9922">1</mods:number>
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<mods:classification id="08DBED77E6068416688BCABB2FDBD1F1">journal article</mods:classification>
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<mods:identifier id="4E43BA505B176AB84CE13545574C8E1E" type="DOI">10.11646/zootaxa.4020.1.7</mods:identifier>
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<mods:identifier id="3044680D1A3256375A5EEAF2A7AC8C6E" type="GBIF-Dataset">82b2f539-5d25-42d6-95c5-e8b16a353e12</mods:identifier>
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<mods:identifier id="FA50AF4DD8E7F68C7617A91BB800C525" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="CB213C7ACC2A64296480CC8C22C9283F" type="Zenodo-Dep">237970</mods:identifier>
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<mods:identifier id="6A72EF3F49D3544FCD121DDC703EE24E" type="ZooBank">BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE</mods:identifier>
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</mods:mods>
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<treatment id="172D87FDFFB4FF84FF3DA1C605D2FAC3" ID-DOI="http://doi.org/10.5281/zenodo.6121815" ID-GBIF-Taxon="119638524" ID-Zenodo-Dep="6121815" LSID="urn:lsid:plazi:treatment:172D87FDFFB4FF84FF3DA1C605D2FAC3" httpUri="http://treatment.plazi.org/id/172D87FDFFB4FF84FF3DA1C605D2FAC3" lastPageId="10" lastPageNumber="179" pageId="9" pageNumber="178">
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<subSubSection id="D79E6560FFB4FF87FF3DA1C6054EF8CB" pageId="9" pageNumber="178" type="nomenclature">
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<paragraph id="9F3B36EBFFB4FF87FF3DA1C6066FF92A" blockId="9.[151,566,1740,1799]" box="[151,566,1740,1766]" pageId="9" pageNumber="178">
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<heading id="C4738187FFB4FF87FF3DA1C6066FF92A" bold="true" box="[151,566,1740,1766]" fontSize="11" level="1" pageId="9" pageNumber="178" reason="1">
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<emphasis id="ADF0EAF9FFB4FF87FF3DA1C6066FF92A" bold="true" box="[151,566,1740,1766]" pageId="9" pageNumber="178">
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<taxonomicName id="58844D68FFB4FF87FF3DA1C60588F92A" ID-CoL="9HVQV" authority="Bickel" authorityName="Bickel" box="[151,465,1740,1766]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="eboritibia" status="sp. nov.">
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<emphasis id="ADF0EAF9FFB4FF87FF3DA1C60525F92A" bold="true" box="[151,380,1740,1766]" italics="true" pageId="9" pageNumber="178">Systenus eboritibia</emphasis>
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Bickel
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFB4FF87FE72A1C6066FF92A" box="[472,566,1740,1766]" pageId="9" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9F3B36EBFFB4FF87FF3DA1E4054EF8CB" blockId="9.[151,566,1740,1799]" box="[151,279,1774,1799]" pageId="9" pageNumber="178">
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(
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<figureCitation id="07BF2A6EFFB4FF87FF35A1E404BFF8CB" box="[159,230,1774,1799]" captionStart="FIGURE 4" captionStartId="11.[151,250,1770,1792]" captionTargetBox="[160,1436,628,1748]" captionTargetId="figure@11.[151,1436,628,1749]" captionTargetPageId="11" captionText="FIGURE 4. Systenus eboritibia sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. parkeri sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral; e. male postabdomen, right lateral." httpUri="https://zenodo.org/record/237974/files/figure.png" pageId="9" pageNumber="178">Figs 4</figureCitation>
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a, b)
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFB4FF87FF3DA03C0688F8BF" pageId="9" pageNumber="178" type="materials_examined">
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<paragraph id="9F3B36EBFFB4FF87FF3DA03C0688F8BF" blockId="9.[151,1436,1846,2016]" pageId="9" pageNumber="178">
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<emphasis id="ADF0EAF9FFB4FF87FF3DA03C051BF883" bold="true" box="[151,322,1846,1871]" pageId="9" pageNumber="178">
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<typeStatus id="403F8849FFB4FF87FF3DA03C048BF883" box="[151,210,1846,1871]" pageId="9" pageNumber="178">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="403F8849FFB4FF87FEFFA03C05AAF883" box="[341,499,1846,1871]" pageId="9" pageNumber="178" type="holotype">
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<emphasis id="ADF0EAF9FFB4FF87FEFFA03C05AAF883" bold="true" box="[341,499,1846,1871]" pageId="9" pageNumber="178">HOLOTYPE</emphasis>
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</typeStatus>
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♂,
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<collectingCountry id="E793767BFFB4FF87FD82A03D068FF883" box="[552,726,1847,1871]" name="Costa Rica" pageId="9" pageNumber="178">COSTA RICA</collectingCountry>
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:
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<emphasis id="ADF0EAF9FFB4FF87FD4FA03C0728F883" bold="true" box="[741,881,1846,1871]" pageId="9" pageNumber="178">Guanacaste</emphasis>
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:
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<quantity id="587C9B0EFFB4FF87FC2EA03D079AF883" box="[900,963,1846,1871]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="178" unit="km" value="3.0">3 km</quantity>
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SE Rio Naranjo,
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<date id="EB3A102BFFB4FF87FB37A03D0105F883" box="[1181,1372,1847,1871]" pageId="9" pageNumber="178" value="1992-08-01" valueMax="1992-08-10" valueMin="1992-08-01">1—10.VIII.1992</date>
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, F.D. Parker (
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<collectionCode id="F995AE2EFFB4FF87FF44A051051AF8BF" LSID="urn:lsid:biocol.org:col:34802" box="[238,323,1883,1907]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="9" pageNumber="178">LACM</collectionCode>
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, ex.
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<collectionCode id="F995AE2EFFB4FF87FEDDA0510593F8BF" box="[375,458,1883,1907]" pageId="9" pageNumber="178">EMUS</collectionCode>
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). (
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<collectionCode id="F995AE2EFFB4FF87FE4EA051066EF8BF" LSID="urn:lsid:biocol.org:col:34802" box="[484,567,1883,1907]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="9" pageNumber="178">LACM</collectionCode>
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Ent 300002).
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFB4FF84FF6DA0740684FBFB" lastPageId="10" lastPageNumber="179" pageId="9" pageNumber="178" type="description">
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<paragraph id="9F3B36EBFFB4FF84FF6DA0740684FBFB" blockId="9.[151,1436,1846,2016]" lastBlockId="10.[151,1437,151,1295]" lastPageId="10" lastPageNumber="179" pageId="9" pageNumber="178">
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<emphasis id="ADF0EAF9FFB4FF87FF6DA0740508F85B" bold="true" box="[199,337,1918,1943]" pageId="9" pageNumber="178">Description</emphasis>
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.
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<emphasis id="ADF0EAF9FFB4FF87FECAA07405C5F85B" bold="true" box="[352,412,1918,1943]" pageId="9" pageNumber="178">Male</emphasis>
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: body length:
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<quantity id="587C9B0EFFB4FF87FDEAA07506C0F85B" box="[576,665,1919,1943]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="178" unit="mm" value="2.8">2.8 mm</quantity>
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; wing 2.7 x
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<quantity id="587C9B0EFFB4FF87FC8EA0750722F85B" box="[804,891,1919,1943]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="178" unit="mm" value="0.8">0.8 mm</quantity>
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(
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<figureCitation id="07BF2A6EFFB4FF87FC20A0740796F85B" box="[906,975,1918,1943]" captionStart="FIGURE 4" captionStartId="11.[151,250,1770,1792]" captionTargetBox="[160,1436,628,1748]" captionTargetId="figure@11.[151,1436,628,1749]" captionTargetPageId="11" captionText="FIGURE 4. Systenus eboritibia sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. parkeri sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral; e. male postabdomen, right lateral." httpUri="https://zenodo.org/record/237974/files/figure.png" pageId="9" pageNumber="178">Fig. 4</figureCitation>
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a).
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<emphasis id="ADF0EAF9FFB4FF87FC5BA0740074F85B" box="[1009,1069,1918,1943]" italics="true" pageId="9" pageNumber="178">Head</emphasis>
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: head almost circular in anterior view; postcranium dorsally concave; single row of short postorbitals, white with only dorsal setae black, and dorsalmost pair behind ocellar triangle strong; frons, face, dark metallic green with thick brownish pruinosity; clypeus dark blue-green; strong vertical and ocellar setae present; eyes distinctly separated across face; ventral eye facets slightly larger than dorsal facets; palpus yellow with short black setae and strong apical seta; proboscis dark brown, projecting anteriorly, keel-like; antenna dark brown; scape and pedicel short; postpedicel elongate, triangular tapering gradually, covered with short pubescence, about 3 times as long as basal width, and with short apical arista.
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<emphasis id="ADF0EAF9FFB7FF84FE9AA62D05D8FE8C" box="[304,385,295,320]" italics="true" pageId="10" pageNumber="179">Thorax</emphasis>
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: dorsum dark metallic green with bronze reflections and grey pruinosity; pleura green with dense grey pruinosity; posterior third of mesonotum distinctly flattened; thoracic setae black except where noted; 12 pairs of ac with posterior pair bordering mesonotal depression larger and offset laterally; 6 strong dc, decreasing in size anteriorly.
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<emphasis id="ADF0EAF9FFB7FF84FEF4A69F05CAFE60" box="[350,403,405,428]" italics="true" pageId="10" pageNumber="179">Legs</emphasis>
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: coxae mostly dark brown, but distal eighth pale yellow; all trochanters pale yellow; femora mostly dark brown basally but distal quarter of FI and distal eighth of FII and FIII yellow; tibiae and basitarsus I and III with white cuticle with short ivory colored (not black) vestiture (MSSC); TII pale yellow with brownish vestiture; CI and CII with white anterior setae and shorter vestiture, CIII with strong white lateral seta at ½; major leg setae black; I: 3.5; 3.4;.7/ 0.8/ 0.4/ 0.2/ 0.3; leg I bare of major setation; II: 3.9; 4.2; 2.2/ 1.4/ 0.9/ 0.7/ 0.5; TII with strong ad seta and weaker pd seta at ¼, and strong ad seta and small pd seta at 2/3, and with apical ring of ad, pd, av and ventral seta; III: 4.0; 5.2; 1.0/ 1.6/ 1.0/ 0.6/ 0.5; TIII covered with ivory colored short vestiture (MSSC) and with 4 almost evenly spaced black dorsal setae, and with apical ring of dorsal, ad and av seta.
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<emphasis id="ADF0EAF9FFB7FF84FB52A59B0169FD64" box="[1272,1328,657,680]" italics="true" pageId="10" pageNumber="179">Wing</emphasis>
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: hyaline; R2+3 and R4+5 diverging, R2+3 joining costa near 7/8, and R4+5 joining costa subapically; veins M and R4+5 diverging in basal portion of wing with two veins slightly bowed half way between dm-cu crossvein and apex and running subparallel to apex, M joining costa just behind wing apex; anal vein and anal cell present; CuAx ratio: 0.7; lower calypter pale yellow with yellowish marginal setae; halter yellow.
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<emphasis id="ADF0EAF9FFB7FF84FC08A4280055FCF7" box="[930,1036,802,827]" italics="true" pageId="10" pageNumber="179">Abdomen</emphasis>
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: black with metallic bronze-green reflections and with dusting of grey pruinosity; posterior margin of tergite 1 with posterior row of long black setae, tergites otherwise covered with short black vestiture; sternites 2–6 membranous or only weakly sclerotised, somewhat recessed; sternite 7 glabrous (to be withdrawn between sternites 5 and 6 at rest) and tergite 7 covered with same black vestiture as preabdomen; hypopygium (
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<figureCitation id="07BF2A6EFFB7FF84FC8CA4B80732FC07" box="[806,875,946,971]" captionStart="FIGURE 4" captionStartId="11.[151,250,1770,1792]" captionTargetBox="[160,1436,628,1748]" captionTargetId="figure@11.[151,1436,628,1749]" captionTargetPageId="11" captionText="FIGURE 4. Systenus eboritibia sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. parkeri sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral; e. male postabdomen, right lateral." httpUri="https://zenodo.org/record/237974/files/figure.png" pageId="10" pageNumber="179">Fig. 4</figureCitation>
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b) dark brown with yellow surstylus and cercus; ventral epandrium prolonged and curved; surstylus basally lobate with some lateral setae and with curved apical dorsal projection (hidden between cerci in
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<figureCitation id="07BF2A6EFFB7FF84FDD3A4F006E4FBDF" box="[633,701,1018,1043]" captionStart="FIGURE 4" captionStartId="11.[151,250,1770,1792]" captionTargetBox="[160,1436,628,1748]" captionTargetId="figure@11.[151,1436,628,1749]" captionTargetPageId="11" captionText="FIGURE 4. Systenus eboritibia sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. parkeri sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral; e. male postabdomen, right lateral." httpUri="https://zenodo.org/record/237974/files/figure.png" pageId="10" pageNumber="179">Fig. 4</figureCitation>
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b); cercus elongate, digitiform and slightly expanded distally, and covered with short yellow setae.
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<emphasis id="ADF0EAF9FFB7FF84FDADA3140607FBFB" bold="true" box="[519,606,1054,1079]" pageId="10" pageNumber="179">Female</emphasis>
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: unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFB7FF84FF6DA3480722FB0B" pageId="10" pageNumber="179" type="discussion">
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<paragraph id="9F3B36EBFFB7FF84FF6DA3480722FB0B" blockId="10.[151,1437,151,1295]" pageId="10" pageNumber="179">
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<emphasis id="ADF0EAF9FFB7FF84FF6DA348056DFB97" bold="true" box="[199,308,1090,1115]" pageId="10" pageNumber="179">Remarks</emphasis>
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.
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<taxonomicName id="58844D68FFB7FF84FEEBA349064EFB97" box="[321,535,1090,1115]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="eboritibia">
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<emphasis id="ADF0EAF9FFB7FF84FEEBA349064EFB97" box="[321,535,1090,1115]" italics="true" pageId="10" pageNumber="179">Systenus eboritibia</emphasis>
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</taxonomicName>
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is known from a single male taken at the Rio Naranjo site,
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<collectingCountry id="E793767BFFB7FF84FB04A3490175FB97" box="[1198,1324,1090,1115]" name="Costa Rica" pageId="10" pageNumber="179">Costa Rica</collectingCountry>
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. It has the diagnostic male characters of a black tapering postpedicel, coxae and femora to femoral knees dark brown, both tibia I and tibia III and basitarsus I and III have white cuticle with short ivory colored (not black) vestiture, and the cercus distally expanded and extending well beyond the surstylus.
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFB7FF84FF6DA3D805D2FAC3" pageId="10" pageNumber="179" type="etymology">
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<paragraph id="9F3B36EBFFB7FF84FF6DA3D805D2FAC3" blockId="10.[151,1437,151,1295]" pageId="10" pageNumber="179">
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<emphasis id="ADF0EAF9FFB7FF84FF6DA3D8051FFB27" bold="true" box="[199,326,1234,1259]" pageId="10" pageNumber="179">Etymology</emphasis>
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. The specific epithet “
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<taxonomicName id="58844D68FFB7FF84FDE3A3D806EEFB27" box="[585,695,1234,1259]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="eboritibia">
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<emphasis id="ADF0EAF9FFB7FF84FDE3A3D806EEFB27" box="[585,695,1234,1259]" italics="true" pageId="10" pageNumber="179">eboritibia</emphasis>
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</taxonomicName>
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” is from the Latin,
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<emphasis id="ADF0EAF9FFB7FF84FC30A3D807BEFB27" box="[922,999,1234,1259]" italics="true" pageId="10" pageNumber="179">eboris,</emphasis>
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meaning ivory, and refers to the ivory colored male tibia III.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |