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<document id="33E77541F8D1E5A81042DDC749B26AAF" ID-CLB-Dataset="4064" ID-DOI="10.11646/zootaxa.511.1.1" ID-GBIF-Dataset="d9f10a58-d4f7-4a74-900a-d5f2e041d181" ID-ISSN="1175­5334" ID-Zenodo-Dep="5030215" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624553966208" checkinUser="felipe" docAuthor="Holzenthal, Ralph W. &amp; Contents, Trond Andersen Table Of" docDate="2004" docId="D07687C5FF96FFA7273209AE5DA07181" docLanguage="en" docName="zt00511.pdf" docOrigin="Zootaxa 511 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.511.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Triaenodes acanthus Holzenthal &amp; Contents 2004, new species" docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="2C4FFFBDFF98FFA9263A0877597B7747" masterDocTitle="The caddisfly genus Triaenodes in the Neotropics (Trichoptera: Leptoceridae)" masterLastPageNumber="80" masterPageNumber="1" pageNumber="15" updateTime="1699020624569" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="26D814744C35181DABCEA569BC8F74B1">The caddisfly genus Triaenodes in the Neotropics (Trichoptera: Leptoceridae)</mods:title>
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<mods:namePart id="2F36599C557A69670517C4FE87E5347E">Holzenthal, Ralph W.</mods:namePart>
<mods:affiliation id="8A0D109945DA12AA1F9C081A6E38FE03">Department of Entomology, University of Minnesota, 1980 Folwell Ave., Room 219, St. Paul, Minnesota, 55108, U. S. A. (holze 001 @ umn. edu) Museum of Zoology, University of Bergen, Muséplass 3, N- 5007 Bergen, Norway (trond. andersen @ zmb. uib. no)</mods:affiliation>
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<mods:namePart id="8421020BE85ABEA1274B1E67E09E22E7">Contents, Trond Andersen Table Of</mods:namePart>
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<mods:title id="2552B203286825C83814CB946EA6B576">Zootaxa</mods:title>
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<mods:date id="241A5B39EC6BD53A957D543C012DE9B0">2004</mods:date>
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<mods:number id="BBACCEE760B35CC8E05171EBDCBDB699">2004-05-14</mods:number>
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<treatment id="D07687C5FF96FFA7273209AE5DA07181" ID-DOI="http://doi.org/10.5281/zenodo.5029851" ID-GBIF-Taxon="188631165" ID-Zenodo-Dep="5029851" LSID="urn:lsid:plazi:treatment:D07687C5FF96FFA7273209AE5DA07181" httpUri="http://treatment.plazi.org/id/D07687C5FF96FFA7273209AE5DA07181" lastPageNumber="15" pageId="14" pageNumber="15">
<subSubSection id="10C56558FF96FFA7273209AE5BD876B4" box="[264,675,473,499]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="586036D3FF96FFA7273209AE5BD876B4" blockId="14.[264,675,473,534]" box="[264,675,473,499]" pageId="14" pageNumber="15">
<heading id="032881BFFF96FFA7273209AE5BD876B4" bold="true" box="[264,675,473,499]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="6AABEAC1FF96FFA7273209AE5BD876B4" bold="true" box="[264,675,473,499]" pageId="14" pageNumber="15">
<taxonomicName id="9FDF4D50FF96FFA7273209AE5B7F76B4" authority="Holzenthal &amp; Contents, 2004" authorityName="Holzenthal &amp; Contents" authorityYear="2004" box="[264,516,473,499]" class="Insecta" family="Leptoceridae" genus="Triaenodes" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="acanthus" status="sp. nov.">
<emphasis id="6AABEAC1FF96FFA7273209AE5B7F76B4" bold="true" box="[264,516,473,499]" italics="true" pageId="14" pageNumber="15">Triaenodes acanthus</emphasis>
</taxonomicName>
,
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</emphasis>
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</paragraph>
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<subSubSection id="10C56558FF96FFA72732098B5C6972E9" pageId="14" pageNumber="15" type="description">
<paragraph id="586036D3FF96FFA72732098B58037551" blockId="14.[264,675,473,534]" box="[264,376,508,534]" pageId="14" pageNumber="15">
<figureCitation id="C0E42A56FF96FFA72732098B58357551" box="[264,334,508,534]" captionStart="FIGURE 6" captionStartId="47.[264,368,1652,1676]" captionTargetBox="[271,1285,322,1628]" captionTargetId="figure-47@47.[260,1327,310,1657]" captionTargetPageId="47" captionText="FIGURE 6. Triaenodes acanthus, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—phallus, lateral; E—apices of parameres, lateral; F—apex of phallus, ventral; G—apex of recurved process of inferior appendage." figureDoi="http://doi.org/10.5281/zenodo.5030229" httpUri="https://zenodo.org/record/5030229/files/figure.png" pageId="14" pageNumber="15">Fig. 6</figureCitation>
,
<figureCitation id="C0E42A56FF96FFA72761098B58037551" box="[347,376,508,534]" captionStart="FIGURE 30" captionStartId="71.[264,368,1406,1430]" captionTargetBox="[264,1323,326,1393]" captionTargetId="figure-29@71.[260,1327,326,1412]" captionTargetPageId="71" captionText="FIGURE 30. Triaenodes anomalus Flint. Female genitalia: A—lateral; B—dorsal; C—ventral." figureDoi="http://doi.org/10.5281/zenodo.5030293" httpUri="https://zenodo.org/record/5030293/files/figure.png" pageId="14" pageNumber="15">30</figureCitation>
</paragraph>
<paragraph id="586036D3FF96FFA727320A3B5B007599" blockId="14.[264,1325,588,1734]" pageId="14" pageNumber="15">The subapicoventral projection on the recurved process of the inferior appendage and the long, striate, denticulate parameres render this new species unique among those related species possessing apicoventral projections of the inferior appendages and digitate dorsomesal processes of tergum X.</paragraph>
<paragraph id="586036D3FF96FFA727020A9B5C797411" blockId="14.[264,1325,588,1734]" pageId="14" pageNumber="15">
Adult (male: n=2). Forewing length
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, hind wing length
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. Eye
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wide. Antennal scape
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long. Maxillary palp segment lengths (in mm): 0.260.31, 0.380.39, 0.530.56, 0.23, 0.52. Forewing light brown, denuded.
</paragraph>
<paragraph id="586036D3FF96FFA727020B135C6972E9" blockId="14.[264,1325,588,1734]" pageId="14" pageNumber="15">Male genitalia. Abdominal segment IX anterior margin nearly straight, slightly produced anteroventrally; pleural region membranous, setose; tergum IX, in dorsal view, with posteromesal margin indistinct, dorsomesal papillae not evident; sternum IX in lateral view with posterior margin sinuate. Preanal appendage setose, about as long as wide, constricted basally, apex subacute. Dorsomesal process of tergum X digitate, originating basally, short, about half length of tergum X, with apical setae; tergum X, in dorsal view, with lateral margins subtriangular, apex slightly excavate medially; in lateral view, parallel sided, apex rounded, slightly excavate subapicoventrally, with lateral flange, apicolateral margins with microtrichia. Inferior appendage subquadrangular basally, setose, with long, tapering, apicoventral projection; recurved process gradually downturned, broad along basal 2/3, with subapicoventral projection, apex sharply attenuate; apicomesal lobe heavily setose, elongate, elliptical. Phallobase small, triangular, endothecal membranes prominent, highly convoluted, with pair of parameres, parameres subequal, long, straight, tapering, with longitudinal striae and subapical denticle; phalicata troughlike, straight, narrow throughout length, apex with deep, wide cleft, phallotremal sclerite large, u­shaped.</paragraph>
</subSubSection>
<subSubSection id="10C56558FF96FFA727020DCB5DA07181" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="586036D3FF96FFA727020DCB5A8772B9" blockId="14.[264,1325,588,1734]" pageId="14" pageNumber="15">
<materialsCitation id="E8B73C8EFF96FFA727020DCB5A8772B9" ID-GBIF-Occurrence="3383261318" collectingDate="1981-05-26" collectionCode="NMNH" collectorName="C &amp; O Flint" country="Mexico" location="Rio Jamapa" pageId="14" pageNumber="15" specimenCode="UMSP000067429" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<emphasis id="6AABEAC1FF96FFA727020DCB5A507291" bold="true" box="[312,811,1468,1494]" pageId="14" pageNumber="15">
<typeStatus id="87648871FF96FFA727020DCB58D17291" box="[312,426,1468,1494]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="20C87643FF96FFA7242C0DCB5BED7291" box="[534,662,1468,1494]" name="Mexico" pageId="14" pageNumber="15">MEXICO</collectingCountry>
:
<collectingRegion id="9A1BF831FF96FFA724950DCB5A5D7291" box="[687,806,1468,1494]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="14" pageNumber="15">Veracruz</collectingRegion>
:
</emphasis>
<location id="5D006008FF96FFA725040DCB5AAD7291" LSID="urn:lsid:plazi:treatment:D07687C5FF96FFA7273209AE5DA07181:5D006008FF96FFA725040DCB5AAD7291" box="[830,982,1468,1494]" country="Mexico" name="Rio Jamapa" pageId="14" pageNumber="15" stateProvince="Veracruz de Ignacio de la Llave">Río Jamapa</location>
,
<quantity id="9F279B36FF96FFA725D40DCB5D427291" box="[1006,1081,1468,1494]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="15" unit="km" value="6.0">6 km</quantity>
.
<collectorName id="F52A5305FF96FFA7226A0DCB5C5D7291" box="[1104,1318,1468,1494]" pageId="14" pageNumber="15">N Coscomatepec</collectorName>
,
<date id="2C611013FF96FFA727320D93580572B9" box="[264,382,1508,1534]" pageId="14" pageNumber="15" value="1981-05-26">
<collectingDate id="3C25E9FBFF96FFA727320D93580572B9" box="[264,382,1508,1534]" pageId="14" pageNumber="15" value="1981-05-26">26.v.1981</collectingDate>
</date>
,
<collectorName id="F52A5305FF96FFA727B00D935B6D72B9" box="[394,534,1508,1534]" pageId="14" pageNumber="15">C &amp; O Flint</collectorName>
<specimenCount id="4ED9FD5AFF96FFA724790D935BE972B9" box="[579,658,1508,1534]" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
(
<specimenCode id="08799EA8FF96FFA7249B0D935A0C72B9" box="[673,887,1508,1534]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="14" pageNumber="15" type="University or college">UMSP000067429</specimenCode>
) (
<collectionCode id="3ECEAE16FF96FFA725B50D935A9472B9" box="[911,1007,1508,1534]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="15" type="Museum">NMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="586036D3FF96FFA727020E7B58067109" blockId="14.[264,1325,588,1734]" pageId="14" pageNumber="15">
<materialsCitation id="E8B73C8EFF96FFA727020E7B58037109" ID-GBIF-Occurrence="3383261403" collectingDate="1965-05-22" collectionCode="NMNH" country="Mexico" county="Fortin" location="Rabago" municipality="Flores" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<emphasis id="6AABEAC1FF96FFA727020E7B5BB37161" bold="true" box="[312,712,1548,1574]" pageId="14" pageNumber="15">
<typeStatus id="87648871FF96FFA727020E7B58D57161" box="[312,430,1548,1574]" pageId="14" pageNumber="15" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="20C87643FF96FFA727870E7B5B467161" box="[445,573,1548,1574]" name="Mexico" pageId="14" pageNumber="15">MEXICO</collectingCountry>
:
<collectingRegion id="9A1BF831FF96FFA724760E7B5BB87161" box="[588,707,1548,1574]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="14" pageNumber="15">Veracruz</collectingRegion>
:
</emphasis>
<collectingCounty id="B1014E5FFF96FFA724EB0E7B5A627161" box="[721,793,1548,1574]" pageId="14" pageNumber="15">Fortin</collectingCounty>
de las
<collectingMunicipality id="B804ACA9FF96FFA7254B0E7B5AC67161" box="[881,957,1548,1574]" pageId="14" pageNumber="15">Flores</collectingMunicipality>
,
<date id="2C611013FF96FFA725F10E7B5D3A7161" box="[971,1089,1548,1574]" pageId="14" pageNumber="15" value="1965-05-22">
<collectingDate id="3C25E9FBFF96FFA725F10E7B5D3A7161" box="[971,1089,1548,1574]" pageId="14" pageNumber="15" value="1965-05-22">22.v.1965</collectingDate>
</date>
,
<location id="5D006008FF96FFA7226A0E7B5DD27161" LSID="urn:lsid:plazi:treatment:D07687C5FF96FFA7273209AE5DA07181:5D006008FF96FFA7226A0E7B5DD27161" box="[1104,1193,1548,1574]" country="Mexico" county="Fortin" municipality="Flores" name="Rabago" pageId="14" pageNumber="15" stateProvince="Veracruz de Ignacio de la Llave">Rabago</location>
<specimenCount id="4ED9FD5AFF96FFA722E30E7B5C517161" box="[1241,1322,1548,1574]" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
(
<collectionCode id="3ECEAE16FF96FFA7272A0E43580B7109" box="[272,368,1588,1614]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="15" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="586036D3FF96FFA727020E2B5B4E7131" blockId="14.[264,1325,588,1734]" box="[312,565,1628,1654]" pageId="14" pageNumber="15">
Distribution.
<collectingCountry id="20C87643FF96FFA727EE0E2B5B4A7131" box="[468,561,1628,1654]" name="Mexico" pageId="14" pageNumber="15">Mexico</collectingCountry>
.
</paragraph>
<paragraph id="586036D3FF96FFA727020EF35DA07181" blockId="14.[264,1325,588,1734]" pageId="14" pageNumber="15">
Etymology. From
<emphasis id="6AABEAC1FF96FFA724350EF35B1571DA" box="[527,622,1668,1693]" italics="true" pageId="14" pageNumber="15">akantha</emphasis>
the Greek word for thorn or prickle, referring to the sharp, thornlike apicoventral projection on the recurved process of the inferior appendage.
</paragraph>
</subSubSection>
</treatment>
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