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<document id="8B43B160A33BAF41AB9E8141DD82E028" ID-CLB-Dataset="4064" ID-DOI="10.11646/zootaxa.511.1.1" ID-GBIF-Dataset="d9f10a58-d4f7-4a74-900a-d5f2e041d181" ID-ISSN="1175­5334" ID-Zenodo-Dep="5030215" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624553966208" checkinUser="felipe" docAuthor="Holzenthal, Ralph W. &amp; Contents, Trond Andersen Table Of" docDate="2004" docId="D07687C5FF89FFBB27320AF35BC673B1" docLanguage="en" docName="zt00511.pdf" docOrigin="Zootaxa 511 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.511.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Triaenodes clauseni Holzenthal &amp; Contents 2004, new species" docType="treatment" docVersion="7" lastPageNumber="19" masterDocId="2C4FFFBDFF98FFA9263A0877597B7747" masterDocTitle="The caddisfly genus Triaenodes in the Neotropics (Trichoptera: Leptoceridae)" masterLastPageNumber="80" masterPageNumber="1" pageNumber="18" updateTime="1699020624569" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2F0AF2CF70ECA81864071A4D07F921F1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FDAE793A4D5953FC13B8D736BB7D8314">
<mods:title id="F51C6325097F16E7CA1EC6EAB1753712">The caddisfly genus Triaenodes in the Neotropics (Trichoptera: Leptoceridae)</mods:title>
</mods:titleInfo>
<mods:name id="D1AB81F871886FDC12D8F32D7C8AF6CB" type="personal">
<mods:role id="4F2CB1951B1597A3EF26A77E94416A45">
<mods:roleTerm id="DF197A069E01E0FF6A4AEB14EEC4F9F4">Author</mods:roleTerm>
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<mods:namePart id="1494431304E0DD4FF2C73C6BDA27B13A">Holzenthal, Ralph W.</mods:namePart>
<mods:affiliation id="B7161BBC1C042C566BDA6E990F947424">Department of Entomology, University of Minnesota, 1980 Folwell Ave., Room 219, St. Paul, Minnesota, 55108, U. S. A. (holze 001 @ umn. edu) Museum of Zoology, University of Bergen, Muséplass 3, N- 5007 Bergen, Norway (trond. andersen @ zmb. uib. no)</mods:affiliation>
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<mods:namePart id="4FECD5F039F072C6C0D266D08E453714">Contents, Trond Andersen Table Of</mods:namePart>
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<mods:typeOfResource id="82C31D9CDF8B2A07D2F938055BED1D66">text</mods:typeOfResource>
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<mods:title id="63CF894228DFD12C118DAF8D1F1E5DEF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F1B143ECF922ECBEA7B83F0A79AD373F">
<mods:date id="1F6BA8DDC98F5E99A75F0FFA26C3A452">2004</mods:date>
<mods:detail id="646D5A6154DC71FA079EED351B785974" type="pubDate">
<mods:number id="FADC6A153A6AEED211E5C607B13EEEC5">2004-05-14</mods:number>
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<mods:detail id="633DE594EA9C798024577617005CEE40" type="volume">
<mods:number id="131C935BACE6576229733520B6514E6B">511</mods:number>
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<mods:number id="80BFEFC17743FA5A3368AA549720D495">1</mods:number>
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<mods:start id="274BAF342AC86DEB76114DFDA7974BBD">1</mods:start>
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<mods:url id="44BA3073F839AA611A15FC94E15E3FE9">https://biotaxa.org/Zootaxa/article/view/zootaxa.511.1.1</mods:url>
</mods:location>
<mods:classification id="C615547959558C6DFBE67C3D90266E97">journal article</mods:classification>
<mods:identifier id="F8FA955D8C0FB01524A5EC6960049C63" type="CLB-Dataset">4064</mods:identifier>
<mods:identifier id="669880544EE265C0D9AF7269FC32B9ED" type="DOI">10.11646/zootaxa.511.1.1</mods:identifier>
<mods:identifier id="D8C74222C301D08E9A7C76B3A2946AAF" type="GBIF-Dataset">d9f10a58-d4f7-4a74-900a-d5f2e041d181</mods:identifier>
<mods:identifier id="5D9827B8916A3E0A765C61426D957394" type="ISSN">1175­5334</mods:identifier>
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<treatment id="D07687C5FF89FFBB27320AF35BC673B1" ID-DOI="http://doi.org/10.5281/zenodo.5567233" ID-GBIF-Taxon="188631173" ID-Zenodo-Dep="5567233" LSID="urn:lsid:plazi:treatment:D07687C5FF89FFBB27320AF35BC673B1" httpUri="http://treatment.plazi.org/id/D07687C5FF89FFBB27320AF35BC673B1" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection id="10C56558FF89FFB827320AF35BE275D9" box="[264,665,644,670]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="586036D3FF89FFB827320AF35BE275D9" blockId="17.[264,665,644,705]" box="[264,665,644,670]" pageId="17" pageNumber="18">
<heading id="032881BFFF89FFB827320AF35BE275D9" bold="true" box="[264,665,644,670]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="6AABEAC1FF89FFB827320AF35BE275D9" bold="true" box="[264,665,644,670]" pageId="17" pageNumber="18">
<taxonomicName id="9FDF4D50FF89FFB827320AF3588175D9" authority="Holzenthal &amp; Contents, 2004" authorityName="Holzenthal &amp; Contents" authorityYear="2004" box="[264,506,644,670]" class="Insecta" family="Leptoceridae" genus="Triaenodes" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="clauseni" status="sp. nov.">
<emphasis id="6AABEAC1FF89FFB827320AF3588175D9" bold="true" box="[264,506,644,670]" italics="true" pageId="17" pageNumber="18">Triaenodes clauseni</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="719857BAFF89FFB824330AF35BE275D9" box="[521,665,644,670]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="10C56558FF89FFBB27320AD05D0C7619" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="586036D3FF89FFB827320AD058037586" blockId="17.[264,665,644,705]" box="[264,376,679,705]" pageId="17" pageNumber="18">
<figureCitation id="C0E42A56FF89FFB827320AD058357586" box="[264,334,679,705]" captionStart="FIGURE 9" captionStartId="50.[264,368,1387,1411]" captionTargetBox="[298,1283,317,1369]" captionTargetId="figure-33@50.[260,1327,300,1386]" captionTargetPageId="50" captionText="FIGURE 9. Triaenodes clauseni, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—phallus, lateral." figureDoi="http://doi.org/10.5281/zenodo.5030239" httpUri="https://zenodo.org/record/5030239/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
,
<figureCitation id="C0E42A56FF89FFB827610AD058037586" box="[347,376,679,705]" captionStart="FIGURE 32" captionStartId="73.[264,368,905,929]" captionTargetBox="[472,1109,326,880]" captionTargetId="figure-27@73.[470,1116,326,881]" captionTargetPageId="73" captionText="FIGURE 32. Triaenodes clauseni, new species. Female genitalia: A—lateral." figureDoi="http://doi.org/10.5281/zenodo.5030297" httpUri="https://zenodo.org/record/5030297/files/figure.png" pageId="17" pageNumber="18">32</figureCitation>
</paragraph>
<paragraph id="586036D3FF89FFB827320A805D88747E" blockId="17.[264,1326,759,1905]" pageId="17" pageNumber="18">
Like the former species, this species is similar to
<taxonomicName id="9FDF4D50FF89FFB8255F0A805A9A7457" authorityName="Holzenthal &amp; Contents" authorityYear="2004" box="[869,993,759,785]" class="Insecta" family="Leptoceridae" genus="Triaenodes" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kilambe">
<emphasis id="6AABEAC1FF89FFB8255F0A805A087457" box="[869,883,759,784]" italics="true" pageId="17" pageNumber="18">T</emphasis>
.
<emphasis id="6AABEAC1FF89FFB825BE0A805A9A7457" box="[900,993,759,784]" italics="true" pageId="17" pageNumber="18">kilambe</emphasis>
</taxonomicName>
, but it differs from both
<taxonomicName id="9FDF4D50FF89FFB8232F0A805813747F" authorityName="Holzenthal &amp; Contents" authorityYear="2004" class="Insecta" family="Leptoceridae" genus="Triaenodes" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="chirripo">
<emphasis id="6AABEAC1FF89FFB8232F0A805C587457" box="[1301,1315,759,784]" italics="true" pageId="17" pageNumber="18">T</emphasis>
.
<emphasis id="6AABEAC1FF89FFB827320B685813747F" box="[264,360,799,824]" italics="true" pageId="17" pageNumber="18">chirripo</emphasis>
</taxonomicName>
and
<taxonomicName id="9FDF4D50FF89FFB8279B0B685B60747F" authorityName="Holzenthal &amp; Contents" authorityYear="2004" box="[417,539,799,825]" class="Insecta" family="Leptoceridae" genus="Triaenodes" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kilambe">
<emphasis id="6AABEAC1FF89FFB8279B0B6858D4747F" box="[417,431,799,824]" italics="true" pageId="17" pageNumber="18">T</emphasis>
.
<emphasis id="6AABEAC1FF89FFB827840B685B60747F" box="[446,539,799,824]" italics="true" pageId="17" pageNumber="18">kilambe</emphasis>
</taxonomicName>
in the shape of the dorsomesal process and apex of tergum X.
</paragraph>
<paragraph id="586036D3FF89FFB827020B3058217316" blockId="17.[264,1326,759,1905]" pageId="17" pageNumber="18">
Adult (male: n=7, female: n=2). Forewing length 5.0–5.8,
<quantity id="9F279B36FF89FFB825DE0B305D467426" box="[996,1085,839,865]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="17" pageNumber="18" unit="mm" value="5.4">5.4 mm</quantity>
(male),
<quantity id="9F279B36FF89FFB8229A0B305C567426" box="[1184,1325,839,865]" metricMagnitude="-3" metricUnit="m" metricValue="5.95" metricValueMax="6.2" metricValueMin="5.7" pageId="17" pageNumber="18" unit="mm" value="5.95" valueMax="6.2" valueMin="5.7">5.7–6.2 mm</quantity>
(female); hind wing length 4.2–4.8,
<quantity id="9F279B36FF89FFB824890B185A7574CE" box="[691,782,879,905]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="17" pageNumber="18" unit="mm" value="4.5">4.5 mm</quantity>
(male), 4.8–5.0 mm (female). Eye 0.35–0.40,
<quantity id="9F279B36FF89FFB827320BE0580A74F6" box="[264,369,919,945]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="17" pageNumber="18" unit="mm" value="0.38">0.38 mm</quantity>
wide (male),
<quantity id="9F279B36FF89FFB824220BE05BBF74F6" box="[536,708,919,945]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" metricValueMax="4.1" metricValueMin="3.7" pageId="17" pageNumber="18" unit="mm" value="0.39" valueMax="0.41" valueMin="0.37">0.37–0.41 mm</quantity>
wide (female). Antennal scape 0.44–0.48,
<quantity id="9F279B36FF89FFB822F90BE05C5674F6" box="[1219,1325,919,945]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="17" pageNumber="18" unit="mm" value="0.46">0.46 mm</quantity>
long (male),
<quantity id="9F279B36FF89FFB827A50BC85B31749E" box="[415,586,959,985]" metricMagnitude="-4" metricUnit="m" metricValue="4.1499999999999995" metricValueMax="4.2" metricValueMin="4.1" pageId="17" pageNumber="18" unit="mm" value="0.415" valueMax="0.42" valueMin="0.41">0.41–0.42 mm</quantity>
long (female). Maxillary palp segment lengths (in mm): 0.26– 0.31, 0.28; 0.37–0.42, 0.39; 0.44–0.52, 0.48; 0.24–0.31, 0.27; 0.53–0.58, 0.56 (male); 0.31–0.33, 0.36–0.39, 0.49–0.54, segments IV and V missing (female). Forewing golden brown.
</paragraph>
<paragraph id="586036D3FF89FFB827020C2858397196" blockId="17.[264,1326,759,1905]" pageId="17" pageNumber="18">Male genitalia. Abdominal segment IX anterior margin slightly rounded, slightly produced anteroventrally; pleural region very lightly sclerotized, setose; tergum IX, in dorsal view, with posteromesal margin rounded, with pair of small dorsomesal papillae; sternum IX in lateral view with posterior margin slightly sinuate. Preanal appendage setose, at least twice as long as wide, broad basally, apex subacute. Dorsomesal process of tergum X bifid, originating medially, short, less than half length of tergum X, with apical setae; tergum X, in dorsal view, with lateral margins subtriangular, apex strongly excavate medially; in lateral view, parallel sided, apex rounded, slightly excavate subapicodorsally, apicolateral margins with microtrichia. Inferior appendage subquadrangular basally, setose, with short, tapering, apicoventral projection; recurved process gradually downturned, narrow, tapering throughout length, but constricted subapically, apex attenuate; apicomesal lobe heavily setose, elongate, narrow, parallel sided. Phallobase small, triangular, endothecal membranes prominent, highly convoluted, with pair of parameres, parameres subequal, long, gently curved, tapering, apices slightly curved; phalicata troughlike, straight, widest medially, tapering apically, apex rounded, phallotremal sclerite not evident.</paragraph>
<paragraph id="586036D3FF89FFBB27020EA85D0C7619" blockId="17.[264,1326,759,1905]" lastBlockId="18.[264,1325,284,1270]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
Female genitalia. Abdominal segment VIII with sternum setose posteriorly, pleural region highly membranous. Posterior margin tergum IX triangular in dorsal view, with pair of small papillose lobes. Valves heavily setose, elliptical. Tergum X setose, subquadrate in lateral view, ventrolateral margin rounded, apex acute; in dorsal view constricted medially, apex rounded. Appendage of segment X setose, narrow basally, elliptical. Structures of internal genitalia as in
<figureCitation id="C0E42A56FF8AFFBB242909335BF17619" box="[531,650,324,350]" captionStart="FIGURE 32" captionStartId="73.[264,368,905,929]" captionTargetBox="[472,1109,326,880]" captionTargetId="figure-27@73.[470,1116,326,881]" captionTargetPageId="73" captionText="FIGURE 32. Triaenodes clauseni, new species. Female genitalia: A—lateral." figureDoi="http://doi.org/10.5281/zenodo.5030297" httpUri="https://zenodo.org/record/5030297/files/figure.png" pageId="18" pageNumber="19">Figs. 32A</figureCitation>
; vaginal chamber in ventral view circular.
</paragraph>
</subSubSection>
<subSubSection id="10C56558FF8AFFBB2702091C5BC673B1" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="586036D3FF8AFFBB2702091C5B967691" blockId="18.[264,1325,284,1270]" pageId="18" pageNumber="19">
<materialsCitation id="E8B73C8EFF8AFFBB2702091C5B967691" ID-GBIF-Occurrence="3383261351" collectingDate="1987-07-22" collectingDateMax="1987-07-23" collectingDateMin="1987-07-22" collectionCode="UMSP" collectorName="Holzenthal &amp; Morse &amp; Clausen" country="Costa Rica" elevation="600" latitude="10.945" location="Rio Bochinche" longLatPrecision="21" longitude="85.413055" municipality="Cerro Campana" pageId="18" pageNumber="19" specimenCode="UMSP000083714" specimenCount="1" stateProvince="Alajuela" typeStatus="holotype">
<emphasis id="6AABEAC1FF8AFFBB2702091C5A4D76C2" bold="true" box="[312,822,363,389]" pageId="18" pageNumber="19">
<typeStatus id="87648871FF8AFFBB2702091C58D176C2" box="[312,426,363,389]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="20C87643FF8AFFBB27C7091C5BCD76C2" box="[509,694,363,389]" name="Costa Rica" pageId="18" pageNumber="19">COSTA RICA</collectingCountry>
:
<collectingRegion id="9A1BF831FF8AFFBB24F8091C5A4B76C2" box="[706,816,363,389]" country="Costa Rica" name="Alajuela" pageId="18" pageNumber="19">Alajuela</collectingRegion>
:
</emphasis>
<collectingMunicipality id="B804ACA9FF8AFFBB2506091B5A8376C1" box="[828,1016,364,390]" pageId="18" pageNumber="19">Cerro Campana</collectingMunicipality>
,
<location id="5D006008FF8AFFBB2239091B5DCA76C1" LSID="urn:lsid:plazi:treatment:D07687C5FF89FFBB27320AF35BC673B1:5D006008FF8AFFBB2239091B5DCA76C1" box="[1027,1201,364,390]" country="Costa Rica" latitude="10.945" longLatPrecision="21" longitude="85.413055" municipality="Cerro Campana" name="Rio Bochinche" pageId="18" pageNumber="19" stateProvince="Alajuela">Río Bochinche</location>
trib.
<quantity id="9F279B36FF8AFFBB22D5091B5C5176C1" box="[1263,1322,364,390]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="km" value="6.0">6 km</quantity>
(air)
<collectorName id="F52A5305FF8AFFBB277B09E3589376E9" box="[321,488,404,430]" pageId="18" pageNumber="19">NW Dos Rios</collectorName>
,
<geoCoordinate id="3DEB5014FF8AFFBB27C909E35BFE76E9" box="[499,645,404,430]" degrees="10" direction="north" minutes="56" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="42" value="10.945">10°56’42”N</geoCoordinate>
,
<geoCoordinate id="3DEB5014FF8AFFBB24AA09E35A5576E9" box="[656,814,404,430]" degrees="085" direction="east" minutes="24" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="47" value="85.413055">085°24’47”E</geoCoordinate>
,
<quantity id="9F279B36FF8AFFBB250309E35AFE76E9" box="[825,901,404,430]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="m" value="600.0">
<elevation id="D3F2D1E0FF8AFFBB250309E35AFE76E9" box="[825,901,404,430]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="2C611013FF8AFFBB25AB09E35D3E76E9" box="[913,1093,404,430]" pageId="18" pageNumber="19" value="1987-07-22" valueMax="1987-07-23" valueMin="1987-07-22">
<collectingDate id="3C25E9FBFF8AFFBB25AB09E35D3E76E9" box="[913,1093,404,430]" pageId="18" pageNumber="19" value="1987-07-22" valueMax="1987-07-23" valueMin="1987-07-22">22–23.vii.1987</collectingDate>
</date>
,
<collectorName id="F52A5305FF8AFFBB226B09E35DAE76E9" box="[1105,1237,404,430]" pageId="18" pageNumber="19">Holzenthal</collectorName>
,
<collectorName id="F52A5305FF8AFFBB22DA09E35C5076E9" box="[1248,1323,404,430]" pageId="18" pageNumber="19">Morse</collectorName>
&amp;
<collectorName id="F52A5305FF8AFFBB271C09CB58FF7691" box="[294,388,444,470]" pageId="18" pageNumber="19">Clausen</collectorName>
(
<specimenCode id="08799EA8FF8AFFBB27A109CB5B097691" box="[411,626,444,470]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="18" pageNumber="19" type="University or college">UMSP000083714</specimenCode>
) (
<collectionCode id="3ECEAE16FF8AFFBB24B009CB5B9B7691" box="[650,736,444,470]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="18" pageNumber="19" type="University or college">UMSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="586036D3FF8AFFBB270209945AAE7311" blockId="18.[264,1325,284,1270]" pageId="18" pageNumber="19">
<materialsCitation id="E8B73C8EFF8AFFBB270209945B1E7509" ID-GBIF-Occurrence="3383261353" collectingDate="1987-07-22" collectingDateMax="1987-07-23" collectingDateMin="1987-07-22" collectionCode="UMSP" collectorName="Holzenthal &amp; Morse &amp; Clausen" country="Costa Rica" county="Cerro Campana" elevation="600" latitude="10.945" location="Dos Rios" longLatPrecision="21" longitude="85.413055" municipality="Rio Bochinche" pageId="18" pageNumber="19" specimenCount="3" specimenCount-male="3" stateProvince="Alajuela" typeStatus="paratype">
<emphasis id="6AABEAC1FF8AFFBB270209945B8576BA" bold="true" box="[312,766,483,509]" pageId="18" pageNumber="19">
<typeStatus id="87648871FF8AFFBB2702099458C176BA" box="[312,442,483,509]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="20C87643FF8AFFBB27FF09945B0576BA" box="[453,638,483,509]" name="Costa Rica" pageId="18" pageNumber="19">COSTA RICA</collectingCountry>
:
<collectingRegion id="9A1BF831FF8AFFBB24B009945B8376BA" box="[650,760,483,509]" country="Costa Rica" name="Alajuela" pageId="18" pageNumber="19">Alajuela</collectingRegion>
:
</emphasis>
<collectingCounty id="B1014E5FFF8AFFBB253E09935AC476B9" box="[772,959,484,510]" pageId="18" pageNumber="19">Cerro Campana</collectingCounty>
,
<collectingMunicipality id="B804ACA9FF8AFFBB25F109935D0276B9" box="[971,1145,484,510]" pageId="18" pageNumber="19">Río Bochinche</collectingMunicipality>
trib.
<quantity id="9F279B36FF8AFFBB228D09935D8A76B9" box="[1207,1265,484,510]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="km" value="6.0">6 km</quantity>
(air) NW
<location id="5D006008FF8AFFBB27790A7B58CF7561" LSID="urn:lsid:plazi:treatment:D07687C5FF89FFBB27320AF35BC673B1:5D006008FF8AFFBB27790A7B58CF7561" box="[323,436,524,550]" country="Costa Rica" county="Cerro Campana" latitude="10.945" longLatPrecision="21" longitude="85.413055" municipality="Rio Bochinche" name="Dos Rios" pageId="18" pageNumber="19" stateProvince="Alajuela">Dos Rios</location>
,
<geoCoordinate id="3DEB5014FF8AFFBB27F80A7B5B2F7561" box="[450,596,524,550]" degrees="10" direction="north" minutes="56" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="42" value="10.945">10°56’42”N</geoCoordinate>
,
<geoCoordinate id="3DEB5014FF8AFFBB24590A7B5A7A7561" box="[611,769,524,550]" degrees="085" direction="east" minutes="24" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="47" value="85.413055">085°24’47”E</geoCoordinate>
,
<quantity id="9F279B36FF8AFFBB25350A7B5A257561" box="[783,862,524,550]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="m" value="600.0">
<elevation id="D3F2D1E0FF8AFFBB25350A7B5A257561" box="[783,862,524,550]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="2C611013FF8AFFBB25570A7B5D5A7561" box="[877,1057,524,550]" pageId="18" pageNumber="19" value="1987-07-22" valueMax="1987-07-23" valueMin="1987-07-22">
<collectingDate id="3C25E9FBFF8AFFBB25570A7B5D5A7561" box="[877,1057,524,550]" pageId="18" pageNumber="19" value="1987-07-22" valueMax="1987-07-23" valueMin="1987-07-22">22–23.vii.1987</collectingDate>
</date>
,
<collectorName id="F52A5305FF8AFFBB220A0A7B5DCF7561" box="[1072,1204,524,550]" pageId="18" pageNumber="19">Holzenthal</collectorName>
,
<collectorName id="F52A5305FF8AFFBB22F80A7B5C767561" box="[1218,1293,524,550]" pageId="18" pageNumber="19">Morse</collectorName>
&amp;
<collectorName id="F52A5305FF8AFFBB27320A43581D7509" box="[264,358,564,590]" pageId="18" pageNumber="19">Clausen</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB27A20A43588F7509" box="[408,500,564,590]" pageId="18" pageNumber="19" type="male">3 males</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB243D0A435B267509" box="[519,605,564,590]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="18" pageNumber="19" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E8B73C8EFF8AFFBB244E0A435DB57531" ID-GBIF-Occurrence="3383261379" collectingDate="1986-10-01" collectingDateMax="1986-10-04" collectingDateMin="1986-10-01" collectionCode="UMSP" collectorName="Chacon" country="Costa Rica" elevation="980" latitude="10.216111" location="Rio San Lorencito" longLatPrecision="21" longitude="-84.60694" municipality="Reserva Forestal San Ramon" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="paratype">
<collectingMunicipality id="B804ACA9FF8AFFBB244E0A435AA87509" box="[628,979,564,590]" pageId="18" pageNumber="19">Reserva Forestal San Ramón</collectingMunicipality>
,
<location id="5D006008FF8AFFBB25DB0A435DC57509" LSID="urn:lsid:plazi:treatment:D07687C5FF89FFBB27320AF35BC673B1:5D006008FF8AFFBB25DB0A435DC57509" box="[993,1214,564,590]" country="Costa Rica" latitude="10.216111" longLatPrecision="21" longitude="-84.60694" municipality="Reserva Forestal San Ramon" name="Rio San Lorencito" pageId="18" pageNumber="19" stateProvince="Alajuela">Río San Lorencito</location>
&amp; tribs.,
<geoCoordinate id="3DEB5014FF8AFFBB27320A2B58E17531" box="[264,410,604,630]" degrees="10" direction="north" minutes="12" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="58" value="10.216111">10°12’58”N</geoCoordinate>
,
<geoCoordinate id="3DEB5014FF8AFFBB279C0A2B5B307531" box="[422,587,604,630]" degrees="084" direction="west" minutes="36" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="25" value="-84.60694">084°36’25”W</geoCoordinate>
,
<quantity id="9F279B36FF8AFFBB246D0A2B5BD87531" box="[599,675,604,630]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="18" pageNumber="19" unit="m" value="980.0">
<elevation id="D3F2D1E0FF8AFFBB246D0A2B5BD87531" box="[599,675,604,630]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="18" pageNumber="19" unit="m" value="980.0">980 m</elevation>
</quantity>
,
<date id="2C611013FF8AFFBB24950A2B5A4E7531" box="[687,821,604,630]" pageId="18" pageNumber="19" value="1986-10-01" valueMax="1986-10-04" valueMin="1986-10-01">
<collectingDate id="3C25E9FBFF8AFFBB24950A2B5A4E7531" box="[687,821,604,630]" pageId="18" pageNumber="19" value="1986-10-01" valueMax="1986-10-04" valueMin="1986-10-01">1–4.x.1986</collectingDate>
</date>
, I &amp; A
<collectorName id="F52A5305FF8AFFBB25B60A2B5A9D7531" box="[908,998,604,630]" pageId="18" pageNumber="19">Chacón</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB222B0A2B5D1B7531" box="[1041,1120,604,630]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB22550A2B5DBE7531" box="[1135,1221,604,630]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="18" pageNumber="19" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E8B73C8EFF8AFFBB22E00A2B5D6175D9" ID-GBIF-Occurrence="3383261364" collectingDate="1987-03-30" collectingDateMax="1987-04-01" collectingDateMin="1987-03-30" collectionCode="UMSP" collectorName="Holzenthal &amp; Hamilton &amp; Heyn" country="Costa Rica" pageId="18" pageNumber="19" specimenCount="2" specimenCount-male="2" stateProvince="Alajuela" typeStatus="paratype">
<date id="2C611013FF8AFFBB22E00A2B580D75D9" pageId="18" pageNumber="19" value="1987-03-30" valueMax="1987-04-01" valueMin="1987-03-30">
<collectingDate id="3C25E9FBFF8AFFBB22E00A2B580D75D9" pageId="18" pageNumber="19" value="1987-03-30" valueMax="1987-04-01" valueMin="1987-03-30">30.iii.– 1.iv.1987</collectingDate>
</date>
,
<collectorName id="F52A5305FF8AFFBB27B10AF35B7475D9" box="[395,527,644,670]" pageId="18" pageNumber="19">Holzenthal</collectorName>
,
<collectorName id="F52A5305FF8AFFBB24190AF35BE975D9" box="[547,658,644,670]" pageId="18" pageNumber="19">Hamilton</collectorName>
&amp;
<collectorName id="F52A5305FF8AFFBB24FD0AF35A7C75D9" box="[711,775,644,670]" pageId="18" pageNumber="19">Heyn</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB25780AF35ADF75D9" box="[834,932,644,670]" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB25860AF35D6975D9" box="[956,1042,644,670]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="18" pageNumber="19" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E8B73C8EFF8AFFBB22140AF35AE875A9" ID-GBIF-Occurrence="3383261423" collectingDate="1990-08-31" collectionCode="INBIO" collectorName="Huisman &amp; Quesada" country="Costa Rica" elevation="550" latitude="10.96" location="Parque Nacional Guanacaste" longLatPrecision="21" longitude="85.5" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<emphasis id="6AABEAC1FF8AFFBB22140AF35DB175D9" bold="true" box="[1070,1226,644,670]" pageId="18" pageNumber="19">
<collectingRegion id="9A1BF831FF8AFFBB22140AF35DBE75D9" box="[1070,1221,644,670]" country="Costa Rica" name="Guanacaste" pageId="18" pageNumber="19">Guanacaste</collectingRegion>
:
</emphasis>
<location id="5D006008FF8AFFBB22E30AF35B727581" LSID="urn:lsid:plazi:treatment:D07687C5FF89FFBB27320AF35BC673B1:5D006008FF8AFFBB22E30AF35B727581" country="Costa Rica" latitude="10.96" longLatPrecision="21" longitude="85.5" name="Parque Nacional Guanacaste" pageId="18" pageNumber="19" stateProvince="Guanacaste">Parque Nacional Guanacaste</location>
, ca.
<locationDeviation id="49D22FF1FF8AFFBB24710ADB5A107581" box="[587,875,684,710]" location="Est. Maritza" pageId="18" pageNumber="19">
<quantity id="9F279B36FF8AFFBB24710ADB5BD97581" box="[587,674,684,710]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="19" unit="km" value="0.7">0.7 km</quantity>
N Est. Maritza
</locationDeviation>
,
<geoCoordinate id="3DEB5014FF8AFFBB25440ADB5D747581" box="[894,1039,684,710]" degrees="10" direction="north" minutes="57" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="36" value="10.96">10°57’36”N</geoCoordinate>
,
<geoCoordinate id="3DEB5014FF8AFFBB22180ADB5DBB7581" box="[1058,1216,684,710]" degrees="085" direction="east" minutes="30" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="00" value="85.5">085°30’00”E</geoCoordinate>
,
<quantity id="9F279B36FF8AFFBB22E80ADB5C5E7581" box="[1234,1317,684,710]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="19" unit="m" value="550.0">
<elevation id="D3F2D1E0FF8AFFBB22E80ADB5C5E7581" box="[1234,1317,684,710]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="19" unit="m" value="550.0">550 m</elevation>
</quantity>
,
<date id="2C611013FF8AFFBB27320AA358E375A9" box="[264,408,724,750]" pageId="18" pageNumber="19" value="1990-08-31">
<collectingDate id="3C25E9FBFF8AFFBB27320AA358E375A9" box="[264,408,724,750]" pageId="18" pageNumber="19" value="1990-08-31">31.viii.1990</collectingDate>
</date>
,
<collectorName id="F52A5305FF8AFFBB279D0AA35B6A75A9" box="[423,529,724,750]" pageId="18" pageNumber="19">Huisman</collectorName>
&amp;
<collectorName id="F52A5305FF8AFFBB24000AA35BE475A9" box="[570,671,724,750]" pageId="18" pageNumber="19">Quesada</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB24EA0AA35A5A75A9" box="[720,801,724,750]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB25090AA35AF075A9" box="[819,907,724,750]" pageId="18" pageNumber="19">INBIO</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E8B73C8EFF8AFFBB259B0AA358037479" ID-GBIF-Occurrence="3383261400" collectingDate="1997-09-07" collectingDateMax="1997-09-10" collectingDateMin="1997-09-07" collectionCode="NMNH" collectorName="Maes &amp; Hernandez" country="Nicaragua" elevation="220" latitude="14.55" location="Cerro Maz" longLatPrecision="21" longitude="-85.11667" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Jinotega" typeStatus="paratype">
<emphasis id="6AABEAC1FF8AFFBB259B0AA35DA575A9" bold="true" box="[929,1246,724,750]" pageId="18" pageNumber="19">
<collectingCountry id="20C87643FF8AFFBB259B0AA35D2775A9" box="[929,1116,724,750]" name="Nicaragua" pageId="18" pageNumber="19">NICARAGUA</collectingCountry>
:
<collectingRegion id="9A1BF831FF8AFFBB22510AA35DA275A9" box="[1131,1241,724,750]" country="Nicaragua" name="Jinotega" pageId="18" pageNumber="19">Jinotega</collectingRegion>
:
</emphasis>
<location id="5D006008FF8AFFBB22D20AA358447451" LSID="urn:lsid:plazi:treatment:D07687C5FF89FFBB27320AF35BC673B1:5D006008FF8AFFBB22D20AA358447451" country="Nicaragua" latitude="14.55" longLatPrecision="21" longitude="-85.11667" name="Cerro Maz" pageId="18" pageNumber="19" stateProvince="Jinotega">Cerro Maz</location>
,
<geoCoordinate id="3DEB5014FF8AFFBB27750A8B589B7451" box="[335,480,764,790]" degrees="14" direction="north" minutes="33" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="00" value="14.55">14°33’00”N</geoCoordinate>
,
<geoCoordinate id="3DEB5014FF8AFFBB27C80A8B5BED7451" box="[498,662,764,790]" degrees="085" direction="west" minutes="07" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="00" value="-85.11667">085°07’00”W</geoCoordinate>
,
<quantity id="9F279B36FF8AFFBB249C0A8B5B8C7451" box="[678,759,764,790]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="19" unit="m" value="220.0">
<elevation id="D3F2D1E0FF8AFFBB249C0A8B5B8C7451" box="[678,759,764,790]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="19" unit="m" value="220.0">220 m</elevation>
</quantity>
,
<date id="2C611013FF8AFFBB25320A8B5ADE7451" box="[776,933,764,790]" pageId="18" pageNumber="19" value="1997-09-07" valueMax="1997-09-10" valueMin="1997-09-07">
<collectingDate id="3C25E9FBFF8AFFBB25320A8B5ADE7451" box="[776,933,764,790]" pageId="18" pageNumber="19" value="1997-09-07" valueMax="1997-09-10" valueMin="1997-09-07">7–10.ix.1997</collectingDate>
</date>
,
<collectorName id="F52A5305FF8AFFBB258C0A8B5A8E7451" box="[950,1013,764,790]" pageId="18" pageNumber="19">Maes</collectorName>
&amp;
<collectorName id="F52A5305FF8AFFBB221E0A8B5DD87451" box="[1060,1187,764,790]" pageId="18" pageNumber="19">Hernández</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB22ED0A8B5C507451" box="[1239,1323,764,790]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB272A0B53580B7479" box="[272,368,804,830]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E8B73C8EFF8AFFBB27B20B535BE17421" ID-GBIF-Occurrence="3383261449" collectingDate="1996-03" collectionCode="NMNH" collectorName="Maes &amp; Hernandez" country="Nicaragua" elevation="700" latitude="13.733334" location="Cerro Saslaya" longLatPrecision="21" longitude="-85.01667" pageId="18" pageNumber="19" specimenCount="3" specimenCount-male="3" stateProvince="Zelaya" typeStatus="paratype">
<emphasis id="6AABEAC1FF8AFFBB27B20B53589E7479" bold="true" box="[392,485,804,830]" pageId="18" pageNumber="19">
<collectingRegion id="9A1BF831FF8AFFBB27B20B53589A7479" box="[392,481,804,830]" country="Nicaragua" name="Zelaya" pageId="18" pageNumber="19">Zelaya</collectingRegion>
:
</emphasis>
<location id="5D006008FF8AFFBB27CA0B535BE17479" LSID="urn:lsid:plazi:treatment:D07687C5FF89FFBB27320AF35BC673B1:5D006008FF8AFFBB27CA0B535BE17479" box="[496,666,804,830]" country="Nicaragua" latitude="13.733334" longLatPrecision="21" longitude="-85.01667" name="Cerro Saslaya" pageId="18" pageNumber="19" stateProvince="Zelaya">Cerro Saslaya</location>
,
<geoCoordinate id="3DEB5014FF8AFFBB24930B535A417479" box="[681,826,804,830]" degrees="13" direction="north" minutes="44" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="00" value="13.733334">13°44’00”N</geoCoordinate>
,
<geoCoordinate id="3DEB5014FF8AFFBB25700B535A957479" box="[842,1006,804,830]" degrees="085" direction="west" minutes="01" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="00" value="-85.01667">085°01’00”W</geoCoordinate>
,
<quantity id="9F279B36FF8AFFBB25C70B535D357479" box="[1021,1102,804,830]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="19" unit="m" value="700.0">
<elevation id="D3F2D1E0FF8AFFBB25C70B535D357479" box="[1021,1102,804,830]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="19" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="2C611013FF8AFFBB22660B535DC17479" box="[1116,1210,804,830]" pageId="18" pageNumber="19" value="1996-03">
<collectingDate id="3C25E9FBFF8AFFBB22660B535DC17479" box="[1116,1210,804,830]" pageId="18" pageNumber="19" value="1996-03">iii.1996</collectingDate>
</date>
,
<collectorName id="F52A5305FF8AFFBB22F20B535C7C7479" box="[1224,1287,804,830]" pageId="18" pageNumber="19">Maes</collectorName>
&amp;
<collectorName id="F52A5305FF8AFFBB27320B3B58FD7421" box="[264,390,844,870]" pageId="18" pageNumber="19">Hernandez</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB27840B3B5B667421" box="[446,541,844,870]" pageId="18" pageNumber="19" type="male">3 males</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB24080B3B5BE87421" box="[562,659,844,870]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E8B73C8EFF8AFFBB24910B3B5D4674C9" ID-GBIF-Occurrence="3383261412" collectingDate="1979-12-26" collectingDateMax="1980-01-01" collectingDateMin="1979-12-26" collectionCode="NMNH" collectorName="Wolda" country="Panama" latitude="9.0" location="Miramar" longLatPrecision="21" longitude="-82.25" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Bocas del Toro" typeStatus="paratype">
<emphasis id="6AABEAC1FF8AFFBB24910B3B5D767421" bold="true" box="[683,1037,844,870]" pageId="18" pageNumber="19">
<collectingCountry id="20C87643FF8AFFBB24910B3B5A4B7421" box="[683,816,844,870]" name="Panama" pageId="18" pageNumber="19">PANAMA</collectingCountry>
:
<collectingRegion id="9A1BF831FF8AFFBB257B0B3B5D737421" box="[833,1032,844,870]" country="Panama" name="Bocas del Toro" pageId="18" pageNumber="19">Bocas del Toro</collectingRegion>
:
</emphasis>
<location id="5D006008FF8AFFBB22200B3B5DF87421" LSID="urn:lsid:plazi:treatment:D07687C5FF89FFBB27320AF35BC673B1:5D006008FF8AFFBB22200B3B5DF87421" box="[1050,1155,844,870]" country="Panama" latitude="9.0" longLatPrecision="21" longitude="-82.25" name="Miramar" pageId="18" pageNumber="19" stateProvince="Bocas del Toro">Miramar</location>
,
<geoCoordinate id="3DEB5014FF8AFFBB22AE0B3B5C5D7421" box="[1172,1318,844,870]" degrees="09" direction="north" minutes="00" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="00" value="9.0">09°00’00”N</geoCoordinate>
,
<geoCoordinate id="3DEB5014FF8AFFBB27320B0358D774C9" box="[264,428,884,910]" degrees="082" direction="west" minutes="15" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="00" value="-82.25">082°15’00”W</geoCoordinate>
,
<date id="2C611013FF8AFFBB27840B035B3274C9" box="[446,585,884,910]" pageId="18" pageNumber="19" value="1979-12-26">
<collectingDate id="3C25E9FBFF8AFFBB27840B035B3274C9" box="[446,585,884,910]" pageId="18" pageNumber="19" value="1979-12-26">26.xii.1979</collectingDate>
</date>
–
<date id="2C611013FF8AFFBB24610B035BCE74C9" box="[603,693,884,910]" pageId="18" pageNumber="19" value="1980-01-01">
<collectingDate id="3C25E9FBFF8AFFBB24610B035BCE74C9" box="[603,693,884,910]" pageId="18" pageNumber="19" value="1980-01-01">1.i.1980</collectingDate>
</date>
, H
<collectorName id="F52A5305FF8AFFBB24D30B035A4F74C9" box="[745,820,884,910]" pageId="18" pageNumber="19">Wolda</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB25560B035ABB74C9" box="[876,960,884,910]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB25EF0B035D4E74C9" box="[981,1077,884,910]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E8B73C8EFF8AFFBB22770B0358A57499" ID-GBIF-Occurrence="3383261350" collectingDate="1977-08-03" collectingDateMax="1977-09-06" collectingDateMin="1977-08-03" collectionCode="NMNH" collectorName="Wolda" country="Panama" elevation="1050" latitude="8.916667" location="Hornitos" longLatPrecision="21" longitude="-82.26667" municipality="Fortuna Dam Site" pageId="18" pageNumber="19" specimenCount="3" specimenCount-male="3" stateProvince="Chiriqui" typeStatus="paratype">
<emphasis id="6AABEAC1FF8AFFBB22770B035DB874C9" bold="true" box="[1101,1219,884,910]" pageId="18" pageNumber="19">
<collectingRegion id="9A1BF831FF8AFFBB22770B035DC674C9" box="[1101,1213,884,910]" country="Panama" name="Chiriqui" pageId="18" pageNumber="19">Chiriqui</collectingRegion>
:
</emphasis>
<collectingMunicipality id="B804ACA9FF8AFFBB22F50B03580D74F1" pageId="18" pageNumber="19">Fortuna Dam Site</collectingMunicipality>
nr.
<location id="5D006008FF8AFFBB27990BEB5B7674F1" LSID="urn:lsid:plazi:treatment:D07687C5FF89FFBB27320AF35BC673B1:5D006008FF8AFFBB27990BEB5B7674F1" box="[419,525,924,950]" country="Panama" latitude="8.916667" longLatPrecision="21" longitude="-82.26667" municipality="Fortuna Dam Site" name="Hornitos" pageId="18" pageNumber="19" stateProvince="Chiriqui">Hornitos</location>
,
<geoCoordinate id="3DEB5014FF8AFFBB24220BEB5BD174F1" box="[536,682,924,950]" degrees="08" direction="north" minutes="55" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="00" value="8.916667">08°55’00”N</geoCoordinate>
,
<geoCoordinate id="3DEB5014FF8AFFBB248D0BEB5A2074F1" box="[695,859,924,950]" degrees="082" direction="west" minutes="16" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="00" value="-82.26667">082°16’00”W</geoCoordinate>
,
<quantity id="9F279B36FF8AFFBB25520BEB5AB874F1" box="[872,963,924,950]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="19" unit="m" value="1050.0">
<elevation id="D3F2D1E0FF8AFFBB25520BEB5AB874F1" box="[872,963,924,950]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="19" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<date id="2C611013FF8AFFBB25F50BEB5DE874F1" box="[975,1171,924,950]" pageId="18" pageNumber="19" value="1977-08-03" valueMax="1977-09-06" valueMin="1977-08-03">
<collectingDate id="3C25E9FBFF8AFFBB25F50BEB5DE874F1" box="[975,1171,924,950]" pageId="18" pageNumber="19" value="1977-08-03" valueMax="1977-09-06" valueMin="1977-08-03">3.viii.–6.ix.1977</collectingDate>
</date>
, H
<collectorName id="F52A5305FF8AFFBB22860BEB5C7374F1" box="[1212,1288,924,950]" pageId="18" pageNumber="19">Wolda</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB27320BB3581F7499" box="[264,356,964,990]" pageId="18" pageNumber="19" type="male">3 males</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB274C0BB358AD7499" box="[374,470,964,990]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E8B73C8EFF8AFFBB27D60BB35A877499" ID-GBIF-Occurrence="3383261331" box="[492,1020,964,990]" collectingDate="1977-11-09" collectingDateMax="1977-11-15" collectingDateMin="1977-11-09" collectionCode="NMNH" collectorName="Wolda" country="Panama" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="paratype">
<date id="2C611013FF8AFFBB27D60BB35BF27499" box="[492,649,964,990]" pageId="18" pageNumber="19" value="1977-11-09" valueMax="1977-11-15" valueMin="1977-11-09">
<collectingDate id="3C25E9FBFF8AFFBB27D60BB35BF27499" box="[492,649,964,990]" pageId="18" pageNumber="19" value="1977-11-09" valueMax="1977-11-15" valueMin="1977-11-09">9–15.xi.1977</collectingDate>
</date>
, H
<collectorName id="F52A5305FF8AFFBB248C0BB35A797499" box="[694,770,964,990]" pageId="18" pageNumber="19">Wolda</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB25080BB35AF87499" box="[818,899,964,990]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB25AE0BB35A8F7499" box="[916,1012,964,990]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E8B73C8EFF8AFFBB22310BB35B037341" ID-GBIF-Occurrence="3383261319" collectingDate="1978-01-04" collectingDateMax="1978-03-07" collectingDateMin="1978-01-04" collectionCode="NMNH" collectorName="Wolda" country="Panama" pageId="18" pageNumber="19" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="paratype">
<date id="2C611013FF8AFFBB22310BB35DCA7499" box="[1035,1201,964,990]" pageId="18" pageNumber="19" value="1978-01-04" valueMax="1978-03-07" valueMin="1978-01-04">
<collectingDate id="3C25E9FBFF8AFFBB22310BB35DCA7499" box="[1035,1201,964,990]" pageId="18" pageNumber="19" value="1978-01-04" valueMax="1978-03-07" valueMin="1978-01-04">4.i.–7.iii.1978</collectingDate>
</date>
, H
<collectorName id="F52A5305FF8AFFBB22E40BB35C517499" box="[1246,1322,964,990]" pageId="18" pageNumber="19">Wolda</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB270A0B9B58E77341" box="[304,412,1004,1030]" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
,
<specimenCount id="4ED9FD5AFF8AFFBB27910B9B58867341" box="[427,509,1004,1030]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB242A0B9B5B0B7341" box="[528,624,1004,1030]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E8B73C8EFF8AFFBB24B20B9B5D947341" ID-GBIF-Occurrence="3383261458" box="[648,1263,1004,1030]" collectingDate="1978-08-16" collectingDateMax="1978-09-12" collectingDateMin="1978-08-16" collectionCode="NMNH" collectorName="Wolda" country="Panama" pageId="18" pageNumber="19" specimenCount="2" specimenCount-male="2" stateProvince="Chiriqui" typeStatus="paratype">
<date id="2C611013FF8AFFBB24B20B9B5A137341" box="[648,872,1004,1030]" pageId="18" pageNumber="19" value="1978-08-16" valueMax="1978-09-12" valueMin="1978-08-16">
<collectingDate id="3C25E9FBFF8AFFBB24B20B9B5A137341" box="[648,872,1004,1030]" pageId="18" pageNumber="19" value="1978-08-16" valueMax="1978-09-12" valueMin="1978-08-16">16.viii.–12.ix.1978</collectingDate>
</date>
, H
<collectorName id="F52A5305FF8AFFBB25A20B9B5A9F7341" box="[920,996,1004,1030]" pageId="18" pageNumber="19">Wolda</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB222C0B9B5D0F7341" box="[1046,1140,1004,1030]" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB22BD0B9B5D9C7341" box="[1159,1255,1004,1030]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E8B73C8EFF8AFFBB22C40B9B5A067369" ID-GBIF-Occurrence="3383261330" collectingDate="1978-12-20" collectingDateMax="1978-12-26" collectingDateMin="1978-12-20" collectionCode="NMNH" collectorName="Wolda" country="Panama" pageId="18" pageNumber="19" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Chiriqui" typeStatus="paratype">
<date id="2C611013FF8AFFBB22C40B9B58EB7369" pageId="18" pageNumber="19" value="1978-12-20" valueMax="1978-12-26" valueMin="1978-12-20">
<collectingDate id="3C25E9FBFF8AFFBB22C40B9B58EB7369" pageId="18" pageNumber="19" value="1978-12-20" valueMax="1978-12-26" valueMin="1978-12-20">20– 26.xii.1978</collectingDate>
</date>
, H
<collectorName id="F52A5305FF8AFFBB27800C635B7D7369" box="[442,518,1044,1070]" pageId="18" pageNumber="19">Wolda</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB24090C635BE67369" box="[563,669,1044,1070]" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
,
<specimenCount id="4ED9FD5AFF8AFFBB24930C635A7F7369" box="[681,772,1044,1070]" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB252F0C635A0E7369" box="[789,885,1044,1070]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E8B73C8EFF8AFFBB25B00C6358037311" ID-GBIF-Occurrence="3383261408" collectingDate="1979-03-21" collectingDateMax="1979-03-27" collectingDateMin="1979-03-21" collectionCode="NMNH" collectorName="Wolda" country="Panama" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="paratype">
<date id="2C611013FF8AFFBB25B00C635D4C7369" box="[906,1079,1044,1070]" pageId="18" pageNumber="19" value="1979-03-21" valueMax="1979-03-27" valueMin="1979-03-21">
<collectingDate id="3C25E9FBFF8AFFBB25B00C635D4C7369" box="[906,1079,1044,1070]" pageId="18" pageNumber="19" value="1979-03-21" valueMax="1979-03-27" valueMin="1979-03-21">21–27.iii.1979</collectingDate>
</date>
, H
<collectorName id="F52A5305FF8AFFBB22580C635DD57369" box="[1122,1198,1044,1070]" pageId="18" pageNumber="19">Wolda</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB22E10C635C507369" box="[1243,1323,1044,1070]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB272A0C4B580B7311" box="[272,368,1084,1110]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E8B73C8EFF8AFFBB27BF0C4B5AAA7311" ID-GBIF-Occurrence="3383261402" box="[389,977,1084,1110]" collectingDate="1979-06-13" collectingDateMax="1979-07-10" collectingDateMin="1979-06-13" collectionCode="NMNH" collectorName="Wolda" country="Panama" pageId="18" pageNumber="19" specimenCount="2" specimenCount-male="2" stateProvince="Chiriqui" typeStatus="paratype">
<date id="2C611013FF8AFFBB27BF0C4B5B257311" box="[389,606,1084,1110]" pageId="18" pageNumber="19" value="1979-06-13" valueMax="1979-07-10" valueMin="1979-06-13">
<collectingDate id="3C25E9FBFF8AFFBB27BF0C4B5B257311" box="[389,606,1084,1110]" pageId="18" pageNumber="19" value="1979-06-13" valueMax="1979-07-10" valueMin="1979-06-13">13.vi.–10.vii.1979</collectingDate>
</date>
, H
<collectorName id="F52A5305FF8AFFBB24BD0C4B5BA87311" box="[647,723,1084,1110]" pageId="18" pageNumber="19">Wolda</collectorName>
—
<specimenCount id="4ED9FD5AFF8AFFBB24C40C4B5A237311" box="[766,856,1084,1110]" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(
<collectionCode id="3ECEAE16FF8AFFBB25520C4B5AB27311" box="[872,969,1084,1110]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="586036D3FF8AFFBB27020C135A357339" blockId="18.[264,1325,284,1270]" box="[312,846,1124,1150]" pageId="18" pageNumber="19">
Distribution.
<collectingCountry id="20C87643FF8AFFBB27EE0C135B2D7339" box="[468,598,1124,1150]" name="Costa Rica" pageId="18" pageNumber="19">Costa Rica</collectingCountry>
,
<collectingCountry id="20C87643FF8AFFBB24580C135BA67339" box="[610,733,1124,1150]" name="Nicaragua" pageId="18" pageNumber="19">Nicaragua</collectingCountry>
,
<collectingCountry id="20C87643FF8AFFBB24D30C135A317339" box="[745,842,1124,1150]" name="Panama" pageId="18" pageNumber="19">Panama</collectingCountry>
.
</paragraph>
<paragraph id="586036D3FF8AFFBB27020CFB5BC673B1" blockId="18.[264,1325,284,1270]" pageId="18" pageNumber="19">
Etymology. The species is named in honor of Dr. Philip J. Clausen, Curator, University of
<collectingRegion id="9A1BF831FF8AFFBB27630CC358AE7389" box="[345,469,1204,1230]" country="United States of America" name="Minnesota" pageId="18" pageNumber="19">Minnesota</collectingRegion>
Insect Collection, in recognition of his near life­long commitment to the care and well­being of the collection.
</paragraph>
</subSubSection>
</treatment>
</document>
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