163 lines
20 KiB
XML
163 lines
20 KiB
XML
<document id="9DC8BF79BEF0B479AE195DEFF5B50CDF" ID-DOI="10.11646/zootaxa.3215.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5248640" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629765289367" checkinUser="felipe" docAuthor="Van Bakel, Barry W. M., Guinot, Danièle, Artal, Pedro, Fraaije, René H. B. & Jagt, John W. M." docDate="2012" docId="4601C935FFA8F9555BB4FA32F115FDA1" docLanguage="en" docName="zt03215p216.pdf" docOrigin="Zootaxa 3215 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3215.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Campylostoma matutiforme Bell 1858" docType="treatment" docVersion="7" lastPageNumber="58" masterDocId="BA38B14DFF90F96C5B23FFB4F45FFFB6" masterDocTitle="A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215" masterLastPageNumber="216" masterPageNumber="1" pageNumber="57" updateTime="1699109082209" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2EA9D479842154F60694D8E965A0251D">A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215</mods:title>
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<mods:namePart id="E276CC9A3CD58C586CF863221EBFF035">Van Bakel, Barry W. M.</mods:namePart>
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<mods:namePart id="E31930F9D0824D871AC4EE94793E3D59">Guinot, Danièle</mods:namePart>
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<mods:namePart id="055718D8B8AFBECA46F8B96E62C89052">Artal, Pedro</mods:namePart>
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<mods:namePart id="5654DB1CC35E1C59136CFB7B228B9101">Fraaije, René H. B.</mods:namePart>
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<mods:namePart id="9A1EEB155DA3D7F329FE8D0B73795212">Jagt, John W. M.</mods:namePart>
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<treatment id="4601C935FFA8F9555BB4FA32F115FDA1" ID-DOI="http://doi.org/10.5281/zenodo.5250372" ID-Zenodo-Dep="5250372" LSID="urn:lsid:plazi:treatment:4601C935FFA8F9555BB4FA32F115FDA1" httpUri="http://treatment.plazi.org/id/4601C935FFA8F9555BB4FA32F115FDA1" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
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<subSubSection id="86B22BA8FFA8F9545BB4FA32F60BFA29" box="[151,596,1414,1439]" pageId="56" pageNumber="57" type="nomenclature">
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<paragraph id="CE177823FFA8F9545BB4FA32F60BFA29" blockId="56.[151,596,1414,1474]" box="[151,596,1414,1439]" pageId="56" pageNumber="57">
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<heading id="955FCF4FFFA8F9545BB4FA32F60BFA29" bold="true" box="[151,596,1414,1439]" fontSize="10" level="2" pageId="56" pageNumber="57" reason="2">
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<taxonomicName id="09A803A0FFA8F9545BB4FA32F60BFA29" ID-CoL="a236c064-6dde-418b-afa6-fc4db9f6b6f6" authority="Bell, 1858" authorityName="Bell" authorityYear="1858" box="[151,596,1414,1439]" class="Malacostraca" family="Cenomanocarcinidae" genus="Campylostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="matutiforme">
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<emphasis id="FCDCA431FFA8F9545BB4FA32F60BFA29" bold="true" box="[151,596,1414,1439]" pageId="56" pageNumber="57">
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<emphasis id="FCDCA431FFA8F9545BB4FA32F587FA29" bold="true" box="[151,472,1414,1439]" italics="true" pageId="56" pageNumber="57">Campylostoma matutiforme</emphasis>
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<bibRefCitation id="AA3905D2FFA8F9545AFCFA32F60BFA29" author="Bell, T." box="[479,596,1414,1439]" pageId="56" pageNumber="57" pagination="1 - 11" refId="ref118308" refString="Bell, T. (1858) A monograph of the fossil malacostracous Crustacea of Great Britain. Part I. Crustacea of the London Clay. Monographs of the Palaeontographical Society (London), 10 (40) (1856), viii + 44 pp., pls. 1 - 11." type="book chapter" year="1858">Bell, 1858</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="86B22BA8FFA8F9545BB4FA1EF574FA74" box="[151,299,1450,1474]" pageId="56" pageNumber="57" type="description">
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<paragraph id="CE177823FFA8F9545BB4FA1EF574FA74" blockId="56.[151,596,1414,1474]" box="[151,299,1450,1474]" pageId="56" pageNumber="57">
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(
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<figureCitation id="569364A6FFA8F9545BBCFA1EF57DFA74" box="[159,290,1450,1474]" captionStart="FIGURE 16" captionStartId="50.[151,250,1843,1866]" captionTargetBox="[151,1436,195,1820]" captionTargetId="figure-123@50.[151,1436,195,1820]" captionTargetPageId="50" captionText="FIGURE 16. Campylostoma matutiforme Bell, 1858 (Cenomanocarcinidae); A, D–F, NHM In. 35231 (male), lower Eocene (Ypresian), Sheppey (Kent, southern England); ventral, posterior, left lateral, and abdominal views; B, C, NHM In. 32654/ 32655 (indeterminate sex), lower Eocene (Ypresian), Herne Bay (Kent, southern England); frontal and ventral views latter showing pterygostomes and mxp3. a1, a2, a3, a4, a5, a6, abdominal somites 1, 2, 3, 4, 5, 6; cx1, cx2, cx3, cx4, cx5, P1, P2, P3, P4, P5 coxas; lb, linea brachyura; P1, P2, P3, P4, pereiopods 1, 2, 3, 4; pt, pterygostome; sl, subantennary lobe of pterygostome. Scale bars: A-E: 5mm; F: 2mm." figureDoi="http://doi.org/10.5281/zenodo.5248744" httpUri="https://zenodo.org/record/5248744/files/figure.png" pageId="56" pageNumber="57">Fig. 16A–F</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="86B22BA8FFA8F9555BB4FA45F75EFF62" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="materials_examined">
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<paragraph id="CE177823FFA8F9545BB4FA45F149F9C0" blockId="56.[151,1436,1521,2014]" pageId="56" pageNumber="57">
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<emphasis id="FCDCA431FFA8F9545BB4FA45F5DFF9BC" bold="true" box="[151,384,1521,1546]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Material examined.</emphasis>
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<materialsCitation id="7EC0727EFFA8F9545AA5FA46F14DF9C0" collectionCode="NHM" country="United Kingdom" county="Isle of Sheppey" location="Partial" municipality="lower Eocene" pageId="56" pageNumber="57" specimenCount="1" stateProvince="Kent" typeStatus="syntype">
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<location id="CB772EF8FFA8F9545AA5FA46F590F9BC" LSID="urn:lsid:plazi:treatment:4601C935FFA8F9555BB4FA32F115FDA1:CB772EF8FFA8F9545AA5FA46F590F9BC" box="[390,463,1522,1546]" country="United Kingdom" county="Isle of Sheppey" municipality="lower Eocene" name="Partial" pageId="56" pageNumber="57" stateProvince="Kent">Partial</location>
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carapace with mxp3,
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<location id="CB772EF8FFA8F95459E0FA46F709F9BC" LSID="urn:lsid:plazi:treatment:4601C935FFA8F9555BB4FA32F115FDA1:CB772EF8FFA8F95459E0FA46F709F9BC" box="[707,854,1522,1546]" country="United Kingdom" county="Isle of Sheppey" municipality="lower Eocene" name="London Clay" pageId="56" pageNumber="57" stateProvince="Kent">London Clay</location>
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,
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<location id="CB772EF8FFA8F9545842FA46F798F9BC" LSID="urn:lsid:plazi:treatment:4601C935FFA8F9555BB4FA32F115FDA1:CB772EF8FFA8F9545842FA46F798F9BC" box="[865,967,1522,1546]" country="United Kingdom" county="Isle of Sheppey" municipality="lower Eocene" name="Ypresian" pageId="56" pageNumber="57" stateProvince="Kent">Ypresian</location>
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,
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||
<location id="CB772EF8FFA8F95458F2FA46F033F9BF" LSID="urn:lsid:plazi:treatment:4601C935FFA8F9555BB4FA32F115FDA1:CB772EF8FFA8F95458F2FA46F033F9BF" box="[977,1132,1522,1546]" country="United Kingdom" county="Isle of Sheppey" municipality="lower Eocene" name="lower Eocene" pageId="56" pageNumber="57" stateProvince="Kent">lower Eocene</location>
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,
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<location id="CB772EF8FFA8F9545F54FA46F0AFF9BC" LSID="urn:lsid:plazi:treatment:4601C935FFA8F9555BB4FA32F115FDA1:CB772EF8FFA8F9545F54FA46F0AFF9BC" box="[1143,1264,1522,1546]" country="United Kingdom" county="Isle of Sheppey" municipality="lower Eocene" name="Herne Bay" pageId="56" pageNumber="57" stateProvince="Kent">Herne Bay</location>
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,
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<collectingRegion id="0C6CB6C1FFA8F9545FD8FA46F16BF9BF" box="[1275,1332,1522,1545]" country="United Kingdom" name="Kent" pageId="56" pageNumber="57">Kent</collectingRegion>
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,
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<collectingCountry id="B6BF38B3FFA8F9545E1DFA46F1C4F9BC" box="[1342,1435,1522,1546]" name="United Kingdom" pageId="56" pageNumber="57">England</collectingCountry>
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(
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<collectionCode id="A8B9E0E6FFA8F9545BBDF9A2F4BEF998" box="[158,225,1558,1582]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="56" pageNumber="57" type="Herbarium">NHM</collectionCode>
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In. 32654/32655);
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<typeStatus id="1113C681FFA8F9545AE1F9A3F643F999" box="[450,540,1559,1583]" pageId="56" pageNumber="57" type="syntype">syntype</typeStatus>
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, carapace with sternite 4, P1 coxae and mxp3, London Clay, Ypresian,
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<collectingMunicipality id="2E73E259FFA8F9545E7FF9A2F4B3F9E4" pageId="56" pageNumber="57">lower Eocene</collectingMunicipality>
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,
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<collectingCounty id="277600AFFFA8F9545BDBF98EF5F7F9E4" box="[248,424,1594,1618]" pageId="56" pageNumber="57">Isle of Sheppey</collectingCounty>
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,
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<collectingCountry id="B6BF38B3FFA8F9545A97F98EF64EF9E4" box="[436,529,1594,1618]" name="United Kingdom" pageId="56" pageNumber="57">England</collectingCountry>
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(
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<collectionCode id="A8B9E0E6FFA8F9545903F98EF63CF9E4" box="[544,611,1594,1618]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="56" pageNumber="57" type="Herbarium">NHM</collectionCode>
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In. 7314); partial carapace with sternite 4, abdomen and bases of pereiopods, same provenance (
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<collectionCode id="A8B9E0E6FFA8F9545A89F9EAF5B2F9C0" box="[426,493,1630,1654]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="56" pageNumber="57" type="Herbarium">NHM</collectionCode>
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In. 35231); 2 carapaces, same provenance (
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<collectionCode id="A8B9E0E6FFA8F95458F4F9EAF045F9C0" box="[983,1050,1630,1654]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="56" pageNumber="57" type="Herbarium">NHM</collectionCode>
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In. 39424, In. 59083)
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</materialsCitation>
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||
.
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</paragraph>
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<paragraph id="CE177823FFA8F9555BE4F936F75EFF62" blockId="56.[151,1436,1521,2014]" lastBlockId="57.[151,1436,151,536]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
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<emphasis id="FCDCA431FFA8F9545BE4F936F592F92C" bold="true" box="[199,461,1665,1690]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Emended description.</emphasis>
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Pleural suture distinct, running just below anterolateral, posterolateral margins. Pterygostome large, anteriorly tumid, with blunt granular crest, extended anteriorly into prominent subantennary lobe; buccal cavity narrow, margins weakly concave, rimmed. Mxp3: coxae large, flabelliform, inserted in front of P1; exopod relatively short, wedge shaped with narrow distal end; endopod elongated, slender, basis-ischium, merus equally long, merus strongly grooved. Sternites 3‒6 preserved; sternite 3 small, incompletely preserved, separated from sternite 4 by oblique lateral grooves; sternite 4 trapezoidal, lateral sides weakly raised, lateral margins slightly concave, nearly vertical groove at level of P1 gynglyme separating large raised episternite 4; episternite 5 raised, oblique, at least one prominence present for abdominal holding; sutures 4/5, 5/6 similarly shaped, only lateral, crescent shaped, medial part vertical, demarcating flat, undivided medial floor of sternum. Abdomen curvature in prolongation with carapace, all somites free, somite 1 short, wide, somite 3 narrowest, somites 3‒5 with raised medial ridge, which is strong, acute in somite 4, somite 6 longer, telson unknown. P1‒P
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<quantity id="0950D5C6FFA9F9555F19FF23F039FF19" box="[1082,1126,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="57" pageNumber="58" unit="in" value="3.0">3 in</quantity>
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the same level, P4 weakly tilted, slightly reduced, P5 subdorsal, strongly reduced.
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</paragraph>
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</subSubSection>
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<subSubSection id="86B22BA8FFA9F9555BE4FF54F115FDA1" pageId="57" pageNumber="58" type="discussion">
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<paragraph id="CE177823FFA9F9555BE4FF54F115FDA1" blockId="57.[151,1436,151,536]" pageId="57" pageNumber="58">
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<emphasis id="FCDCA431FFA9F9555BE4FF54F564FF4E" bold="true" box="[199,315,224,248]" inLineHeading="true" pageId="57" pageNumber="58" reason="1">Remarks.</emphasis>
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Study of the material examined necessitates an amendment of the description; the carapace was described by
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<bibRefCitation id="AA3905D2FFA9F9555A0DFEB0F584FEAA" author="Bell, T." box="[302,475,260,284]" pageId="57" pageNumber="58" pagination="1 - 11" refId="ref118308" refString="Bell, T. (1858) A monograph of the fossil malacostracous Crustacea of Great Britain. Part I. Crustacea of the London Clay. Monographs of the Palaeontographical Society (London), 10 (40) (1856), viii + 44 pp., pls. 1 - 11." type="book chapter" year="1858">Bell (1858: 23)</bibRefCitation>
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and
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<bibRefCitation id="AA3905D2FFA9F9555930FEB0F684FEAA" author="Carter, J." box="[531,731,260,284]" pageId="57" pageNumber="58" pagination="15 - 44" refId="ref121658" refString="Carter, J. (1898) A contribution to the palaeontology of the decapod Crustacea of England. Quarterly Review of the Geological Society of London, 54, 15 - 44, pls. 1 - 2." type="journal article" year="1898">Carter (1898: 30)</bibRefCitation>
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||
.
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<taxonomicName id="09A803A0FFA9F95559C5FEB0F040FEAA" authorityName="Bell" authorityYear="1858" box="[742,1055,260,284]" class="Malacostraca" family="Cenomanocarcinidae" genus="Campylostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="matutiforme">
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<emphasis id="FCDCA431FFA9F95559C5FEB0F040FEAA" box="[742,1055,260,284]" italics="true" pageId="57" pageNumber="58">Campylostoma matutiforme</emphasis>
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</taxonomicName>
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(see
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<bibRefCitation id="AA3905D2FFA9F9555F7FFEB0F0A7FEAA" author="Bell, T." box="[1116,1272,260,284]" pageId="57" pageNumber="58" pagination="1 - 11" refId="ref118308" refString="Bell, T. (1858) A monograph of the fossil malacostracous Crustacea of Great Britain. Part I. Crustacea of the London Clay. Monographs of the Palaeontographical Society (London), 10 (40) (1856), viii + 44 pp., pls. 1 - 11." type="book chapter" year="1858">Bell 1858: 23</bibRefCitation>
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, pl. 3, figs. 8– 10) was deemed a podotreme crab, possibly a cenomanocarcinid, by
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<bibRefCitation id="AA3905D2FFA9F9555880FE93F0ECFE89" author="Guinot, D. & Vega, F. J. & Van Bakel, B. W. M." box="[931,1203,295,320]" pageId="57" pageNumber="58" pagination="681 - 719" refId="ref130419" refString="Guinot, D., Vega, F. J. & Van Bakel, B. W. M. (2008) Cenomanocarcinidae n. fam., a new Cretaceous podotreme family (Crustacea, Brachyura, Raninoidia), with comments on related families. Geodiversitas, 30, 681 - 719." type="journal article" year="2008">
|
||
Guinot
|
||
<emphasis id="FCDCA431FFA9F95558DAFE9DF074FEF6" box="[1017,1067,296,320]" italics="true" pageId="57" pageNumber="58">et al</emphasis>
|
||
. (2008: 21)
|
||
</bibRefCitation>
|
||
. The re-examination of several specimens in the NHM collections (
|
||
<figureCitation id="569364A6FFA9F95559B8FEF8F742FED2" box="[667,797,332,356]" captionStart="FIGURE 16" captionStartId="50.[151,250,1843,1866]" captionTargetBox="[151,1436,195,1820]" captionTargetId="figure-123@50.[151,1436,195,1820]" captionTargetPageId="50" captionText="FIGURE 16. Campylostoma matutiforme Bell, 1858 (Cenomanocarcinidae); A, D–F, NHM In. 35231 (male), lower Eocene (Ypresian), Sheppey (Kent, southern England); ventral, posterior, left lateral, and abdominal views; B, C, NHM In. 32654/ 32655 (indeterminate sex), lower Eocene (Ypresian), Herne Bay (Kent, southern England); frontal and ventral views latter showing pterygostomes and mxp3. a1, a2, a3, a4, a5, a6, abdominal somites 1, 2, 3, 4, 5, 6; cx1, cx2, cx3, cx4, cx5, P1, P2, P3, P4, P5 coxas; lb, linea brachyura; P1, P2, P3, P4, pereiopods 1, 2, 3, 4; pt, pterygostome; sl, subantennary lobe of pterygostome. Scale bars: A-E: 5mm; F: 2mm." figureDoi="http://doi.org/10.5281/zenodo.5248744" httpUri="https://zenodo.org/record/5248744/files/figure.png" pageId="57" pageNumber="58">Fig. 16A–F</figureCitation>
|
||
) confirm this placement. The much elongated mxp3 with large, flabelliform coxae, sculpture of the pterygostome with pronounced, pointed subantennary lobes, crownshaped sternite 3, subtrapezoidal sternite 4, match the cenomanocarcinid configuration. The carapace does not show the strong branchial crests that are typical of
|
||
<taxonomicName id="09A803A0FFA9F95559C4FE0CF7A5FE79" box="[743,1018,440,464]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="FCDCA431FFA9F95559C4FE0CF79FFE66" box="[743,960,440,464]" italics="true" pageId="57" pageNumber="58">Cenomanocarcinus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, and the orbits are much wider. The presence of an abdominal holding system on episternite 5 is confirmed in
|
||
<specimenCount id="D8AEB3AAFFA9F95558DBFE68F0C8FE42" box="[1016,1175,476,500]" pageId="57" pageNumber="58" type="generic">one specimen</specimenCount>
|
||
, where a single, sharp tubercle is visible; because of poor preservation, it is unclear if a more posteriorly situated peg was present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |