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<mods:title id="436D4DC1AE0D436A316C4D8CC0876FF2">A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215</mods:title>
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<paragraph id="CE177823FFA7F95B5BB4FF23F6E2FF06" blockId="55.[151,701,151,212]" box="[151,701,151,176]" pageId="55" pageNumber="56">
<heading id="955FCF4FFFA7F95B5BB4FF23F6E2FF06" bold="true" box="[151,701,151,176]" fontSize="10" level="2" pageId="55" pageNumber="56" reason="2">
<taxonomicName id="09A803A0FFA7F95B5BB4FF23F6E2FF06" ID-CoL="83b9343c-7d0f-4eae-9715-ebc1fff51168" authority="Van Straelen, 1936" authorityName="Van Straelen" authorityYear="1936" box="[151,701,151,176]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="inflatus">
<emphasis id="FCDCA431FFA7F95B5BB4FF23F6E2FF06" bold="true" box="[151,701,151,176]" pageId="55" pageNumber="56">
<emphasis id="FCDCA431FFA7F95B5BB4FF23F587FF06" bold="true" box="[151,472,151,176]" italics="true" pageId="55" pageNumber="56">Cenomanocarcinus inflatus</emphasis>
<bibRefCitation id="AA3905D2FFA7F95B5AFCFF2CF6E2FF06" author="Van Straelen, V." box="[479,701,151,176]" pageId="55" pageNumber="56" pagination="1 - 50" refId="ref147628" refString="Van Straelen, V. (1936) Crustaces decapodes nouveaux ou peu connus de l'epoque cretacique. Bulletin du Musee royal d'Histoire naturelle de Belgique, 12 (45), 1 - 50, pls. 1 - 4." type="journal article" year="1936">Van Straelen, 1936</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="86B22BA8FFA7F95B5BB4FF08F570FF62" box="[151,303,188,212]" pageId="55" pageNumber="56" type="description">
<paragraph id="CE177823FFA7F95B5BB4FF08F570FF62" blockId="55.[151,701,151,212]" box="[151,303,188,212]" pageId="55" pageNumber="56">
(
<figureCitation id="569364A6FFA7F95B5BBCFF08F578FF62" box="[159,295,188,212]" captionStart="FIGURE 18" captionStartId="52.[151,250,1789,1812]" captionTargetBox="[151,1436,193,1768]" captionTargetId="figure-154@52.[151,1436,193,1768]" captionTargetPageId="52" captionText="FIGURE 18. A, D. Cenomanocarcinus inflatus Van Straelen, 1936 (Cenomanocarcinidae), MNHN J08587 (lectotype), photograph of MAB k. 2968 (cast), upper Cenomanian, Butte de Gazonfier, Le Mans (northwestern France); dorsal and left lateral views (direction of photography in D indicated by arrow in A); B, C, E, Cenomanocarcinus vanstraeleni Stenzel, 1945, MAB k. 2909 (B, C; indeterminate sex), upper Albian-lower Cenomanian (Pawpaw Formation), Tarrant County, Texas (U.S.A); dorsal and left lateral views (direction of photography in C indicated by arrow in B); MAB k. 2885 (E; indeterminate sex), dorsal view of carapace; F, Cenomanocarcinus beardi Schweitzer, Feldmann, Fam, Hessin, Hetrick, Nyborg &amp; Ross, 2003a, MAB k. 2880 (indeterminate sex), Cenomanian (Woodbine Formation), Wagner Park, Texas (U.S.A), dorsal view of carapace. ebs, epibranchial spine; pbs, posterobranchial spine. Scale bars: 10mm." figureDoi="http://doi.org/10.5281/zenodo.5248664" httpUri="https://zenodo.org/record/5248664/files/figure.png" pageId="55" pageNumber="56">Fig. 18A, D</figureCitation>
)
</paragraph>
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<subSubSection id="86B22BA8FFA7F95B5BB4FEB7F5C0FE31" pageId="55" pageNumber="56" type="materials_examined">
<paragraph id="CE177823FFA7F95B5BB4FEB7F5C0FE31" blockId="55.[151,1436,259,716]" pageId="55" pageNumber="56">
<emphasis id="FCDCA431FFA7F95B5BB4FEB7F5DDFEAA" bold="true" box="[151,386,259,284]" inLineHeading="true" pageId="55" pageNumber="56" reason="1">Material examined.</emphasis>
Dorsal carapace, upper Cenomanian, Butte de Gazonfier, Le Mans,
<collectingCountry id="B6BF38B3FFA7F95B5FACFEB0F084FEAD" box="[1167,1243,260,283]" name="France" pageId="55" pageNumber="56">France</collectingCountry>
(
<collectionCode id="A8B9E0E6FFA7F95B5FC8FEB0F11CFEAA" box="[1259,1347,260,284]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="55" pageNumber="56">MNHN</collectionCode>
<accessionNumber id="D1FBE5C0FFA7F95B5E68FEB0F1C4FEAA" box="[1355,1435,260,284]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J08587" pageId="55" pageNumber="56" type="EnaNcbi">J08587</accessionNumber>
[see
<bibRefCitation id="AA3905D2FFA7F95B5BEFFE9CF5C1FE89" author="Van Straelen, V." box="[204,414,295,319]" pageId="55" pageNumber="56" pagination="1 - 50" refId="ref147628" refString="Van Straelen, V. (1936) Crustaces decapodes nouveaux ou peu connus de l'epoque cretacique. Bulletin du Musee royal d'Histoire naturelle de Belgique, 12 (45), 1 - 50, pls. 1 - 4." type="journal article" year="1936">Van Straelen 1936</bibRefCitation>
: pl. 4, fig. 8];
<collectionCode id="A8B9E0E6FFA7F95B5963FE93F6DFFE89" box="[576,640,295,319]" pageId="55" pageNumber="56">MAB</collectionCode>
k. 2968 represents a cast); dorsal carapace, same provenance (
<collectionCode id="A8B9E0E6FFA7F95B5E67FE93F1C4FE89" box="[1348,1435,295,319]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="55" pageNumber="56">MNHN</collectionCode>
R05504, the original of A. Milne-Edwards
<emphasis id="FCDCA431FFA7F95B59A1FEF9F6C7FED2" box="[642,664,333,356]" italics="true" pageId="55" pageNumber="56">in</emphasis>
<bibRefCitation id="AA3905D2FFA7F95B5983FEF8F729FED2" author="Guillier, A." box="[672,886,332,356]" pageId="55" pageNumber="56" refId="ref129556" refString="Guillier, A. (1886) Geologie du Departement de la Sarthe, 430 pp. Monnoyer, Le Mans." type="book" year="1886">Guillier 1886: 244</bibRefCitation>
); dorsal carapaces, same provenance (
<collectionCode id="A8B9E0E6FFA7F95B5E04FEF8F135FED2" box="[1319,1386,332,356]" pageId="55" pageNumber="56">MHN</collectionCode>
LM 3804,
<collectionCode id="A8B9E0E6FFA7F95B5BFEFEC4F541FE31" box="[221,286,368,391]" pageId="55" pageNumber="56">MHN</collectionCode>
LM 3806).
</paragraph>
</subSubSection>
<subSubSection id="86B22BA8FFA7F95B5BE4FE20F732FD7A" pageId="55" pageNumber="56" type="discussion">
<paragraph id="CE177823FFA7F95B5BE4FE20F732FD7A" blockId="55.[151,1436,259,716]" pageId="55" pageNumber="56">
<emphasis id="FCDCA431FFA7F95B5BE4FE20F564FE1A" bold="true" box="[199,315,404,428]" inLineHeading="true" pageId="55" pageNumber="56" reason="1">Remarks.</emphasis>
Specimen MNHN
<accessionNumber id="D1FBE5C0FFA7F95B593DFE20F630FE1A" box="[542,623,404,428]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J08587" pageId="55" pageNumber="56" type="EnaNcbi">J08587</accessionNumber>
(
<figureCitation id="569364A6FFA7F95B59A1FE20F74EFE1A" box="[642,785,404,428]" captionStart="FIGURE 18" captionStartId="52.[151,250,1789,1812]" captionTargetBox="[151,1436,193,1768]" captionTargetId="figure-154@52.[151,1436,193,1768]" captionTargetPageId="52" captionText="FIGURE 18. A, D. Cenomanocarcinus inflatus Van Straelen, 1936 (Cenomanocarcinidae), MNHN J08587 (lectotype), photograph of MAB k. 2968 (cast), upper Cenomanian, Butte de Gazonfier, Le Mans (northwestern France); dorsal and left lateral views (direction of photography in D indicated by arrow in A); B, C, E, Cenomanocarcinus vanstraeleni Stenzel, 1945, MAB k. 2909 (B, C; indeterminate sex), upper Albian-lower Cenomanian (Pawpaw Formation), Tarrant County, Texas (U.S.A); dorsal and left lateral views (direction of photography in C indicated by arrow in B); MAB k. 2885 (E; indeterminate sex), dorsal view of carapace; F, Cenomanocarcinus beardi Schweitzer, Feldmann, Fam, Hessin, Hetrick, Nyborg &amp; Ross, 2003a, MAB k. 2880 (indeterminate sex), Cenomanian (Woodbine Formation), Wagner Park, Texas (U.S.A), dorsal view of carapace. ebs, epibranchial spine; pbs, posterobranchial spine. Scale bars: 10mm." figureDoi="http://doi.org/10.5281/zenodo.5248664" httpUri="https://zenodo.org/record/5248664/files/figure.png" pageId="55" pageNumber="56">Fig. 18A, D</figureCitation>
) was illustrated by
<bibRefCitation id="AA3905D2FFA7F95B58D8FE20F082FE1A" author="Van Straelen, V." box="[1019,1245,404,428]" pageId="55" pageNumber="56" pagination="1 - 50" refId="ref147628" refString="Van Straelen, V. (1936) Crustaces decapodes nouveaux ou peu connus de l'epoque cretacique. Bulletin du Musee royal d'Histoire naturelle de Belgique, 12 (45), 1 - 50, pls. 1 - 4." type="journal article" year="1936">Van Straelen (1936</bibRefCitation>
: pl. 4, fig. 8) as topotype. The name proposed by A. Milne-Edwards in
<bibRefCitation id="AA3905D2FFA7F95B583AFE03F79BFE79" author="Guillier, A." box="[793,964,439,463]" pageId="55" pageNumber="56" refId="ref129556" refString="Guillier, A. (1886) Geologie du Departement de la Sarthe, 430 pp. Monnoyer, Le Mans." type="book" year="1886">Guillier (1886)</bibRefCitation>
, was a
<emphasis id="FCDCA431FFA7F95B5F3BFE0DF0E9FE66" box="[1048,1206,440,464]" italics="true" pageId="55" pageNumber="56">nomen nudum</emphasis>
because a diagnosis was not given.
<bibRefCitation id="AA3905D2FFA7F95B5A66FE68F613FE42" author="Van Straelen, V." box="[325,588,476,500]" pageId="55" pageNumber="56" pagination="1 - 50" refId="ref147628" refString="Van Straelen, V. (1936) Crustaces decapodes nouveaux ou peu connus de l'epoque cretacique. Bulletin du Musee royal d'Histoire naturelle de Belgique, 12 (45), 1 - 50, pls. 1 - 4." type="journal article" year="1936">Van Straelen (1936: 37</bibRefCitation>
, 38) stated that the type (A. Milne-Edwards original) was probably in the MNHN palaeontological collections. This specimen was designated
<typeStatus id="1113C681FFA7F95B5893FE4BF048FDA1" box="[944,1047,511,535]" pageId="55" pageNumber="56" type="lectotype">lectotype</typeStatus>
by
<bibRefCitation id="AA3905D2FFA7F95B5F6BFDB4F1C8FDA1" author="Breton, G. &amp; Collins, J. S. H." box="[1096,1431,511,535]" pageId="55" pageNumber="56" pagination="17 - 20" refId="ref121011" refString="Breton, G. &amp; Collins, J. S. H. (2007) Decapod fauna from the Cenomanian stratotype. In: Garassino, A., Feldmann, R. M. &amp; Teruzzi, G. (Eds.), 3 rd Symposium on Mesozoic and Cenozoic decapod crustaceans, Museo di Storia Naturale di Milano, May 23 - 25, 2007. Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 35, 17 - 20." type="journal article" year="2007">Breton &amp; Collins (2007: 18)</bibRefCitation>
. MNHN
<accessionNumber id="D1FBE5C0FFA7F95B5BD0FD90F51BFD8A" box="[243,324,548,572]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J08587" pageId="55" pageNumber="56" type="EnaNcbi">J08587</accessionNumber>
is, however, a carapace from the Hébert Collection (Collections de la Sorbonne), whereas the original of A. Milne-Edwards bears registration number MNHN R05504. The latter is considered here the
<typeStatus id="1113C681FFA7F95B5E36FDF3F125FDE9" box="[1301,1402,583,607]" pageId="55" pageNumber="56" type="holotype">holotype</typeStatus>
, of
<taxonomicName id="09A803A0FFA7F95B5BB4FDD8F552FD32" authorityName="Van Straelen" authorityYear="1936" box="[151,269,620,644]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="inflatus">
<emphasis id="FCDCA431FFA7F95B5BB4FDD8F552FD32" box="[151,269,620,644]" italics="true" pageId="55" pageNumber="56">C. inflatus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="AA3905D2FFA7F95B5A6FFDD8F58FFD32" author="ICZN" box="[332,464,620,644]" pageId="55" pageNumber="56" refId="ref131577" refString="ICZN (1999) International Code of Zoological Nomenclature (Fourth edition), xxix + 306 pp. The International Trust for Zoological Nomenclature, London." type="book" year="1999">ICZN 1999</bibRefCitation>
, Art. 73.1.1), and
<bibRefCitation id="AA3905D2FFA7F95B5981FDD8F7DAFD32" author="Van Straelen, V." box="[674,901,620,644]" pageId="55" pageNumber="56" pagination="1 - 50" refId="ref147628" refString="Van Straelen, V. (1936) Crustaces decapodes nouveaux ou peu connus de l'epoque cretacique. Bulletin du Musee royal d'Histoire naturelle de Belgique, 12 (45), 1 - 50, pls. 1 - 4." type="journal article" year="1936">Van Straelen (1936)</bibRefCitation>
as the author of the taxon, because he was the first to present a diagnosis, retaining the name suggested earlier by A. Milne-Edwards in
<bibRefCitation id="AA3905D2FFA7F95B5F56FD3BF17FFD11" author="Guillier, A." box="[1141,1312,655,679]" pageId="55" pageNumber="56" refId="ref129556" refString="Guillier, A. (1886) Geologie du Departement de la Sarthe, 430 pp. Monnoyer, Le Mans." type="book" year="1886">Guillier (1886)</bibRefCitation>
. The
<typeStatus id="1113C681FFA7F95B5E7FFD3BF498FD7A" pageId="55" pageNumber="56" type="lectotype">lectotype</typeStatus>
designation by
<bibRefCitation id="AA3905D2FFA7F95B5A5AFD00F6D4FD7A" author="Breton, G. &amp; Collins, J. S. H." box="[377,651,692,716]" pageId="55" pageNumber="56" pagination="17 - 20" refId="ref121011" refString="Breton, G. &amp; Collins, J. S. H. (2007) Decapod fauna from the Cenomanian stratotype. In: Garassino, A., Feldmann, R. M. &amp; Teruzzi, G. (Eds.), 3 rd Symposium on Mesozoic and Cenozoic decapod crustaceans, Museo di Storia Naturale di Milano, May 23 - 25, 2007. Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 35, 17 - 20." type="journal article" year="2007">Breton &amp; Collins (2007)</bibRefCitation>
is therefore invalid.
</paragraph>
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