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<document id="09502452985940EE5E053C4239511C18" ID-CLB-Dataset="29126" ID-DOI="10.11646/zootaxa.4497.4.3" ID-GBIF-Dataset="f09380a8-4d31-4e79-9b72-74ee26438553" ID-ISSN="1175-5326" ID-Zenodo-Dep="1456311" ID-ZooBank="7DFD16E7-D628-48DA-AAD6-4AB1E4D679E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539238820092" checkinUser="plazi" docAuthor="Reid, C. A. M. &amp; Beatson, M." docDate="2018" docId="2A2D13766B6BFF9FFF1AFAA907795A9B" docLanguage="en" docName="zootaxa.4497.4.3.pdf" docOrigin="Zootaxa 4497 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cheiloxena tuberosa Reid 1992" docType="treatment" docVersion="9" lastPageNumber="529" masterDocId="D6146B0E6B71FF83FF8DFFF604615833" masterDocTitle="Revision of the Australian leaf beetle genus Cheiloxena Baly, 1860 (Coleoptera: Chrysomelidae: Spilopyrinae)" masterLastPageNumber="534" masterPageNumber="501" pageNumber="527" updateTime="1698574941947" updateUser="ExternalLinkService">
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<mods:title id="9D09F48C07F1B7AB64744888FF409088">Revision of the Australian leaf beetle genus Cheiloxena Baly, 1860 (Coleoptera: Chrysomelidae: Spilopyrinae)</mods:title>
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<mods:namePart id="FD6020B18D3EC3B403390C791705A276">Reid, C. A. M.</mods:namePart>
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<treatment id="2A2D13766B6BFF9FFF1AFAA907795A9B" ID-DOI="http://doi.org/10.5281/zenodo.5958946" ID-GBIF-Taxon="148673167" ID-Zenodo-Dep="5958946" LSID="urn:lsid:plazi:treatment:2A2D13766B6BFF9FFF1AFAA907795A9B" httpUri="http://treatment.plazi.org/id/2A2D13766B6BFF9FFF1AFAA907795A9B" lastPageId="28" lastPageNumber="529" pageId="26" pageNumber="527">
<subSubSection id="EA9EF1EB6B6BFF99FF1AFAA9062C5DA8" pageId="26" pageNumber="527" type="nomenclature">
<paragraph id="A23BA2606B6BFF99FF1AFAA9067D5D49" blockId="26.[151,589,1375,1435]" box="[151,540,1375,1402]" pageId="26" pageNumber="527">
<heading id="F973150C6B6BFF99FF1AFAA9067D5D49" bold="true" box="[151,540,1375,1402]" fontSize="11" level="1" pageId="26" pageNumber="527" reason="1">
<taxonomicName id="6584D9E36B6BFF99FF1AFAA9067D5D49" authority="Reid, 1992" authorityName="Reid" authorityYear="1992" box="[151,540,1375,1402]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6BFF99FF1AFAA9067D5D49" bold="true" box="[151,540,1375,1402]" pageId="26" pageNumber="527">
<emphasis id="90F07E726B6BFF99FF1AFAA905F05D4A" bold="true" box="[151,401,1375,1401]" italics="true" pageId="26" pageNumber="527">Cheiloxena tuberosa</emphasis>
<bibRefCitation id="C615DF916B6BFF99FE15FA96067D5D49" box="[408,540,1376,1402]" pageId="26" pageNumber="527" refId="ref20270" refString="Reid, C. A. M. (1992) Revision of the genus Cheiloxena Baly (Coleoptera: Chrysomelidae: Eumolpinae). Memoirs of the Museum of Fictoria, 53 (1), 101 - 114. https: // doi. org / 10.24199 / j. mmv. 1992.53.04" type="journal article">Reid, 1992</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A23BA2606B6BFF99FF1AFA74062C5DA8" blockId="26.[151,589,1375,1435]" box="[151,589,1410,1435]" pageId="26" pageNumber="527">
(
<figureCitation id="3ABFBEE56B6BFF99FF12FA7404945DA8" box="[159,245,1410,1435]" captionStart="FIGURES 710" captionStartId="7.[151,264,1975,1998]" captionTargetBox="[194,1366,311,1853]" captionTargetId="figure@7.[152,1435,139,1955]" captionTargetPageId="7" captionText="FIGURES 710. Habitus of Cheiloxena species: 7, C. insignis Baly (Launching Place, Victoria); 8, C. insignis (Lake Mountain, Victoria); 9, C. monga Reid &amp; Beatson (holotype); 10, C. tuberosa Reid; Not to scale." httpUri="https://zenodo.org/record/1456317/files/figure.png" pageId="26" pageNumber="527">Figs 10</figureCitation>
,
<figureCitation id="3ABFBEE56B6BFF99FE8CFA75057E5DA8" box="[257,287,1411,1435]" captionStart="FIGURES 1321" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[230,1395,232,1777]" captionTargetId="figure@13.[152,1435,141,1956]" captionTargetPageId="13" captionText="FIGURES 1321. Lateral of Cheiloxena species: 13, C. aitori Reid &amp; Beatson, male (holotype); 14, C. blackburni Reid; 15, C. conani Reid &amp; Beatson; 16, C. frenchae Blackburn, (holotype); 17, C. insignis Blackburn; 18, C. monga Reid &amp; Beatson (holotype); 19, C. tuberosa Reid; 20, C. westwoodii Baly (northern form, Dorrigo); 21, C. westwoodii Baly (southern form, Victoria). Not to scale." httpUri="https://zenodo.org/record/1456321/files/figure.png" pageId="26" pageNumber="527">19</figureCitation>
,
<figureCitation id="3ABFBEE56B6BFF99FEA6FA75052B5DA8" box="[299,330,1411,1435]" captionStart="FIGURES 2230" captionStartId="14.[151,264,1700,1723]" captionTargetBox="[212,1381,282,1587]" captionTargetId="figure@14.[150,1436,193,1679]" captionTargetPageId="14" captionText="FIGURES 2230. Anterior view, Cheiloxena species: 22, C. aitori Reid &amp; Beatson, male (holotype); 23, C. blackburni Reid; 24, C. conani Reid &amp; Beatson; 25, C. frenchae Blackburn; 26, C. insignis Baly; 27, C. monga Reid &amp; Beatson; 28, C. tuberosa Reid; 29, C. westwoodii Baly (northern form, Dorrigo); 30, C. westwoodii Baly (southern form, Victoria)." httpUri="https://zenodo.org/record/1456323/files/figure.png" pageId="26" pageNumber="527">28</figureCitation>
,
<figureCitation id="3ABFBEE56B6BFF99FED8FA7505FC5DA8" box="[341,413,1411,1435]" captionStart="FIGURES 3842" captionStartId="17.[151,264,1957,1980]" captionTargetBox="[281,1374,340,1843]" captionTargetId="figure@17.[152,1435,193,2009]" captionTargetPageId="17" captionText="FIGURES 3842. Cheiloxena species: 38, C. tuberosa Reid, male antenna; 39, C. insignis Blackburn, male antenna; 40, C. insignis, metafemur; 41, C. tuberosa, pro- (left) and meta- (right) tarsi; 42, C. frenchae Blackburn, apex of tarsus." httpUri="https://zenodo.org/record/1456329/files/figure.png" pageId="26" pageNumber="527">38, 41</figureCitation>
,
<figureCitation id="3ABFBEE56B6BFF99FE24FA7505A65DA8" box="[425,455,1411,1435]" captionStart="FIGURES 4351" captionStartId="18.[151,264,1789,1812]" captionTargetId="figure@18.[152,1435,193,1768]" captionTargetPageId="18" captionText="FIGURES 4351. Cheiloxena species, male genitalia lateral and dorsal: 43, C. aitori Reid &amp; Beatson; 44, C. blackburni Reid; 45, C. conani Reid &amp; Beatson; 46, C. frenchae Blackburn; 47, C. insignis Blackburn; 48, C. monga Reid &amp; Beatson; 49, C. tuberosa Reid; 50, C. westwoodii Baly (northern form, Dorrigo); 51, C. westwoodii (southern form, Victoria). All images to scale, scale bar = 2mm." httpUri="https://zenodo.org/record/1457873/files/figure.png" pageId="26" pageNumber="527">49</figureCitation>
,
<figureCitation id="3ABFBEE56B6BFF99FE5EFA7405935DA8" box="[467,498,1410,1435]" captionStart="FIGURES 6268" captionStartId="20.[151,264,1947,1970]" captionTargetBox="[222,1410,255,1799]" captionTargetId="figure@20.[152,1435,111,1926]" captionTargetPageId="20" captionText="FIGURES 6268. Cheiloxena species, female genitalia: 62, C. aitori Reid &amp; Beatson; 63, C. blackburni Reid; 64, C. conani Reid &amp; Beatson; 65, C. frenchae Blackburn; 66, C. insignis Blackburn; 67, C. tuberosa Reid; 68, C. westwoodii Baly (northern form, Dorrigo)." httpUri="https://zenodo.org/record/1456333/files/figure.png" pageId="26" pageNumber="527">67</figureCitation>
,
<figureCitation id="3ABFBEE56B6BFF99FE73FA75067D5DA8" box="[510,540,1411,1435]" captionStart="FIGURES 6976" captionStartId="21.[151,264,1871,1894]" captionText="FIGURES 6976. Cheiloxena species, female genitalia, sternite 8: 69, C. aitori Reid &amp; Beatson; 70, C. blackburni Reid; 71, C. conani Reid &amp; Beatson; 72, C. frenchae Blackburn; 73, C. insignis Blackburn; 74, C. tuberosa Reid; 75, C. westwoodii Baly (northern form, Dorrigo); 76, C. westwoodii (southern form, Victoria)." pageId="26" pageNumber="527">74</figureCitation>
,
<figureCitation id="3ABFBEE56B6BFF99FDAAFA7506255DA8" box="[551,580,1411,1435]" captionStart="FIGURE 77" captionStartId="31.[151,250,1971,1993]" captionTargetId="figure@31.[151,1435,193,1950]" captionTargetPageId="31" captionText="FIGURE 77. Eastern Australia showing distribution of Cheiloxena species." httpUri="https://zenodo.org/record/1457875/files/figure.png" pageId="26" pageNumber="527">77</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EA9EF1EB6B6BFF99FF1AFA3C00585DD3" box="[151,1081,1482,1504]" pageId="26" pageNumber="527" type="reference_group">
<paragraph id="A23BA2606B6BFF99FF1AFA3C00585DD3" blockId="26.[151,1081,1482,1504]" box="[151,1081,1482,1504]" pageId="26" pageNumber="527">
<treatmentCitationGroup id="8294854E6B6BFF99FF1AFA3C00585DD3" box="[151,1081,1482,1504]" pageId="26" pageNumber="527">
<taxonomicName id="6584D9E36B6BFF99FF1AFA3C06705DD3" authority="Reid, 1992: 104" authorityName="Reid" authorityPageNumber="104" authorityYear="1992" box="[151,529,1482,1504]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6BFF99FF1AFA3C05095DD3" box="[151,360,1482,1504]" italics="true" pageId="26" pageNumber="527">Cheiloxena tuberosa</emphasis>
<treatmentCitation id="232584716B6BFF99FEE3FA3C06705DD3" author="" box="[366,529,1482,1504]" page="104" pageId="26" pageNumber="527">
<bibRefCitation id="C615DF916B6BFF99FEE3FA3C05805DD3" box="[366,481,1482,1504]" pageId="26" pageNumber="527" refId="ref20270" refString="Reid, C. A. M. (1992) Revision of the genus Cheiloxena Baly (Coleoptera: Chrysomelidae: Eumolpinae). Memoirs of the Museum of Fictoria, 53 (1), 101 - 114. https: // doi. org / 10.24199 / j. mmv. 1992.53.04" type="journal article">Reid, 1992</bibRefCitation>
: 104
</treatmentCitation>
</taxonomicName>
(type locality: Lamington National Park, Queensland)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="EA9EF1EB6B6BFF98FF1AF9FB07FA58E7" lastPageId="27" lastPageNumber="528" pageId="26" pageNumber="527" type="materials_examined">
<paragraph id="A23BA2606B6BFF99FF1AF9FB00A25EE9" blockId="26.[151,1437,1549,2006]" pageId="26" pageNumber="527">
<emphasis id="90F07E726B6BFF99FF1AF9FB05E45E15" bold="true" box="[151,389,1549,1574]" pageId="26" pageNumber="527">Material examined.</emphasis>
Types: Holotype: Ƌ*/ Lamington NP,
<date id="D63A84A06B6BFF99FCDDF9FB07C45E15" box="[848,933,1549,1574]" pageId="26" pageNumber="527" value="1983-11">xi.1983</date>
M Lowman/ no. 15/ Holotype
<taxonomicName id="6584D9E36B6BFF99FA91F9FB05575E79" authority="Reid" authorityName="Reid" authorityYear="1992" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6BFF99FA91F9FB04995E7A" italics="true" pageId="26" pageNumber="527">Cheiloxena tuberosa</emphasis>
Reid
</taxonomicName>
/ (ANIC); Paratypes (3): New South Wales: ♂/ Dorrigo W Heron/ paratype
<taxonomicName id="6584D9E36B6BFF99FB3CF9C704AA5E5D" authority="Reid" authorityName="Reid" authorityYear="1992" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6BFF99FB3CF9C701FD5E7A" box="[1201,1436,1585,1609]" italics="true" pageId="26" pageNumber="527">Cheiloxena tuberosa</emphasis>
Reid
</taxonomicName>
/ (ANIC); ♀*/ Mt Warning in rainforest,
<date id="D63A84A06B6BFF99FD1BF9A3076F5E5D" box="[662,782,1621,1646]" pageId="26" pageNumber="527" value="1974-11-11">11.xi.1974</date>
G Williams/ paratype
<taxonomicName id="6584D9E36B6BFF99FB86F9A3014C5E5D" authority="Reid" authorityName="Reid" authorityYear="1992" box="[1035,1325,1621,1646]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6BFF99FB86F9A300925E5E" box="[1035,1267,1621,1645]" italics="true" pageId="26" pageNumber="527">Cheiloxena tuberosa</emphasis>
Reid
</taxonomicName>
/ (ANIC); Queensland: Ƌ*/ Mt Glorious St. For., Qld,
<date id="D63A84A06B6BFF99FD0FF98F07635EA1" box="[642,770,1657,1682]" pageId="26" pageNumber="527" value="1985-11" valueMax="1985-12-31" valueMin="1985-11-00">xixii.1985</date>
Y. Bassett coll. ex
<taxonomicName id="6584D9E36B6BFF99FC5AF98F01495EA2" box="[983,1320,1657,1681]" class="Magnoliopsida" family="Malvaceae" genus="Argyrodendron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="26" pageNumber="527" phylum="Tracheophyta" rank="species" species="actinophyllum">
<emphasis id="90F07E726B6BFF99FC5AF98F01495EA2" box="[983,1320,1657,1681]" italics="true" pageId="26" pageNumber="527">Argyrodendron actinophyllum</emphasis>
</taxonomicName>
r/f/ ANIC coleoptera voucher no 87-0134/ CO ANT 10/
<taxonomicName id="6584D9E36B6BFF99FD47F96B000B5E85" authority="Edlin" authorityName="Edlin" box="[714,1130,1693,1718]" class="Magnoliopsida" family="Malvaceae" genus="Argyrodendron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="26" pageNumber="527" phylum="Tracheophyta" rank="species" species="actinophyllum">
<emphasis id="90F07E726B6BFF99FD47F96B00405E86" box="[714,1057,1693,1717]" italics="true" pageId="26" pageNumber="527">Argyrodendron actinophyllum</emphasis>
Edlin
</taxonomicName>
subtropical rainforest Mt Glorious SF IT1 (MT),
<date id="D63A84A06B6BFF99FE2DF93706255EE9" box="[416,580,1729,1754]" pageId="26" pageNumber="527" value="1987-11-19" valueMax="1987-11-26" valueMin="1987-11-19">1926.xi.1987</date>
, Y Bassett/ paratype
<taxonomicName id="6584D9E36B6BFF99FCBFF93700355EE9" authority="Reid" authorityName="Reid" authorityYear="1992" box="[818,1108,1729,1754]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6BFF99FCBFF937007B5EEA" box="[818,1050,1729,1753]" italics="true" pageId="26" pageNumber="527">Cheiloxena tuberosa</emphasis>
Reid
</taxonomicName>
/ (ANIC).
</paragraph>
<paragraph id="A23BA2606B6BFF98FF4AF91007FA58E7" blockId="26.[151,1437,1549,2006]" lastBlockId="27.[151,1437,151,2013]" lastPageId="27" lastPageNumber="528" pageId="26" pageNumber="527">
<materialsCitation id="12ECA83D6B6BFF99FF4AF91000585F11" ID-GBIF-Occurrence="1915499071" collectingDate="2013-11-07" collectionCode="NSW" collectorName="Bielsdown SF &amp; Border Ranges NP" country="United Kingdom" county="Non-types" location="Coramba Rds" municipality="Bielsdown" pageId="26" pageNumber="527" specimenCount="1" stateProvince="New South Wales">
<collectingCounty id="4B5ADAEC6B6BFF99FF4AF910055D5ECD" box="[199,316,1766,1790]" pageId="26" pageNumber="527">Non-types</collectingCounty>
(17):
<emphasis id="90F07E726B6BFF99FE0DF91306315ECD" bold="true" box="[384,592,1765,1790]" pageId="26" pageNumber="527">
<collectingRegion id="60406C826B6BFF99FE0DF91306315ECD" box="[384,592,1765,1790]" country="Australia" name="New South Wales" pageId="26" pageNumber="527">
New
<collectingRegion id="60406C826B6BFF99FE31F91306315ECD" box="[444,592,1765,1790]" inferredBySuffix="true" pageId="26" pageNumber="527">
South
<collectingCountry id="DA93E2F06B6BFF99FD84F91306315ECD" box="[521,592,1765,1790]" name="United Kingdom" pageId="26" pageNumber="527">Wales</collectingCountry>
</collectingRegion>
</collectingRegion>
</emphasis>
: 1Ƌ/
<collectorName id="0F71C7B66B6BFF99FD1DF913074E5ECD" box="[656,815,1765,1790]" pageId="26" pageNumber="527">Bielsdown SF</collectorName>
, jn
<collectingMunicipality id="425F381A6B6BFF99FCDAF91307AC5ECD" box="[855,973,1765,1790]" pageId="26" pageNumber="527">Bielsdown</collectingMunicipality>
/
<location id="A75BF4BB6B6BFF99FC58F91300135ECD" LSID="urn:lsid:plazi:treatment:2A2D13766B6BFF9FFF1AFAA907795A9B:A75BF4BB6B6BFF99FC58F91300135ECD" box="[981,1138,1765,1790]" country="United Kingdom" county="Non-types" municipality="Bielsdown" name="Coramba Rds" pageId="26" pageNumber="527" stateProvince="New South Wales">Coramba Rds</location>
, 30:17S 152:44E dry rf by rd,
<collectingDate id="C67E7D486B6BFF99FF4CF8FF054C5F11" box="[193,301,1801,1826]" pageId="26" pageNumber="527" value="2013-11-07">
<date id="D63A84A06B6BFF99FF4CF8FF054C5F11" box="[193,301,1801,1826]" pageId="26" pageNumber="527" value="2013-11-07">7.xi.2013</date>
</collectingDate>
, Reid &amp; Tan (AMS); 1/ eastern section
<collectorName id="0F71C7B66B6BFF99FC82F8FF078B5F12" box="[783,1002,1801,1826]" pageId="26" pageNumber="527">Border Ranges NP</collectorName>
,
<collectionCode id="C4953AA56B6BFF99FC77F8FC00585F11" box="[1018,1081,1802,1826]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="26" pageNumber="527">NSW</collectionCode>
</materialsCitation>
,
<date id="D63A84A06B6BFF99FBC5F8FF00AB5F11" box="[1096,1226,1801,1826]" pageId="26" pageNumber="527" value="1996-12-14">14.xii.1996</date>
, S. Watkins coll./ (ANIC); ♀/ Dorrigo NP, 2500,
<date id="D63A84A06B6BFF99FD8BF8DB06E15F75" box="[518,640,1837,1862]" pageId="26" pageNumber="527" value="1973-11-18">18.xi.1973</date>
, A&amp;M Walford-Huggins/ WH 7745/ (QMB);
<materialsCitation id="12ECA83D6B6BFF99FB03F8D8048F5FBD" ID-GBIF-Occurrence="1915499066" collectingDate="2009-12-16" collectingDateMax="2009-12-17" collectingDateMin="2009-12-16" collectionCode="DPIO" collectorName="The Glade &amp; Dorrigo NP &amp; R de Keyzer" country="United Kingdom" location="Whian Whian State Forest" pageId="26" pageNumber="527" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="New South Wales">
<specimenCount id="B48269E96B6BFF99FB03F8D800C45F76" box="[1166,1189,1838,1861]" pageId="26" pageNumber="527" type="male"></specimenCount>
,
<specimenCount id="B48269E96B6BFF99FB3EF8D800A55F75" box="[1203,1220,1838,1862]" pageId="26" pageNumber="527" type="female"></specimenCount>
/
<collectorName id="0F71C7B66B6BFF99FB55F8DB012E5F75" box="[1240,1359,1837,1862]" pageId="26" pageNumber="527">The Glade</collectorName>
picnic area,
<collectorName id="0F71C7B66B6BFF99FF58F8A7053B5F5A" box="[213,346,1873,1898]" pageId="26" pageNumber="527">Dorrigo NP</collectorName>
,
<collectingDate id="C67E7D486B6BFF99FEEBF8A706735F59" box="[358,530,1873,1898]" pageId="26" pageNumber="527" value="2009-12-16" valueMax="2009-12-17" valueMin="2009-12-16">
<date id="D63A84A06B6BFF99FEEBF8A706735F59" box="[358,530,1873,1898]" pageId="26" pageNumber="527" value="2009-12-16" valueMax="2009-12-17" valueMin="2009-12-16">1617.xii.2009</date>
</collectingDate>
,
<collectorName id="0F71C7B66B6BFF99FD93F8A406CE5F59" box="[542,687,1873,1898]" pageId="26" pageNumber="527">R de Keyzer</collectorName>
, at MV light (AMS); 1/
<location id="A75BF4BB6B6BFF99FC4BF8A700945F5A" LSID="urn:lsid:plazi:treatment:2A2D13766B6BFF9FFF1AFAA907795A9B:A75BF4BB6B6BFF99FC4BF8A700945F5A" box="[966,1269,1873,1898]" country="United Kingdom" name="Whian Whian State Forest" pageId="26" pageNumber="527" stateProvince="New South Wales">Whian Whian State Forest</location>
,
<collectedFrom id="1198E19D6B6BFF99FA8FF8A401FA5F5A" box="[1282,1435,1873,1897]" pageId="26" pageNumber="527">
on
<taxonomicName id="6584D9E36B6BFF99FAABF8A401FA5F5A" box="[1318,1435,1873,1897]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="26" pageNumber="527" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="90F07E726B6BFF99FAABF8A401FA5F5A" box="[1318,1435,1873,1897]" italics="true" pageId="26" pageNumber="527">E. grandis</emphasis>
</taxonomicName>
</collectedFrom>
(
<collectionCode id="C4953AA56B6BFF99FF12F88004865FBD" box="[159,231,1910,1934]" pageId="26" pageNumber="527">DPIO</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="12ECA83D6B6BFF99FF73F88307515FBD" ID-GBIF-Occurrence="1915499063" box="[254,816,1909,1934]" collectionCode="QMB" collectorName="Deane" country="United Kingdom" location="Queensland" pageId="26" pageNumber="527" specimenCount="1" specimenCount-female="1" stateProvince="Queensland">
<collectingRegion id="60406C826B6BFF99FF73F88305EA5FBD" box="[254,395,1909,1934]" country="Australia" name="Queensland" pageId="26" pageNumber="527">
<emphasis id="90F07E726B6BFF99FF73F88305EA5FBD" bold="true" box="[254,395,1909,1934]" pageId="26" pageNumber="527">Queensland</emphasis>
</collectingRegion>
:
<specimenCount id="B48269E96B6BFF99FE12F88005D15FBD" box="[415,432,1910,1934]" pageId="26" pageNumber="527" type="female"></specimenCount>
/ no further data,
<collectorName id="0F71C7B66B6BFF99FD0AF88006AE5FBE" box="[647,719,1910,1933]" pageId="26" pageNumber="527">Deane</collectorName>
(
<collectionCode id="C4953AA56B6BFF99FD6EF88007495FBD" box="[739,808,1910,1934]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="26" pageNumber="527">QMB</collectionCode>
)
</materialsCitation>
; ♂/ Lamington NP,
<collectingMethod id="7BC5DA776B6BFF99FBAFF88000885FBE" box="[1058,1257,1909,1934]" pageId="26" pageNumber="527">canopy light trap</collectingMethod>
2,
<date id="D63A84A06B6BFF99FA99F883011F5FBD" box="[1300,1406,1909,1934]" pageId="26" pageNumber="527" value="1995-01-13">13.i.1995</date>
R Kitching (AMS); ♂ 2♀/ [Lamington] NP
<date id="D63A84A06B6BFF99FDFDF86F06A45F81" box="[624,709,1945,1970]" pageId="26" pageNumber="527" value="1920-11">xi.1920</date>
H Hacker (QMB);
<materialsCitation id="12ECA83D6B6BFF99FC2EF86C01F65F81" ID-GBIF-Occurrence="1915499070" box="[931,1431,1945,1970]" collectingDate="1970-12-05" collectionCode="QMB" collectorName="FT Fricke" country="United Kingdom" location="Queensland" pageId="26" pageNumber="527" specimenCount="1" specimenCount-female="1" stateProvince="Queensland">
<specimenCount id="B48269E96B6BFF99FC2EF86C07D55F81" box="[931,948,1946,1970]" pageId="26" pageNumber="527" type="female"></specimenCount>
/ ditto except
<collectingDate id="C67E7D486B6BFF99FBD9F86F00A75F81" box="[1108,1222,1945,1970]" pageId="26" pageNumber="527" value="1970-12-05">
<date id="D63A84A06B6BFF99FBD9F86F00A75F81" box="[1108,1222,1945,1970]" pageId="26" pageNumber="527" value="1970-12-05">5.xii.1970</date>
</collectingDate>
<collectorName id="0F71C7B66B6BFF99FB40F86C015F5F81" box="[1229,1342,1945,1970]" pageId="26" pageNumber="527">FT Fricke</collectorName>
(AMS)
</materialsCitation>
; ♀/ Mt Glorious 22k NW Brisbane, rainforest,
<quantity id="657C0F856B6BFF99FD3CF84B06955FE5" box="[689,756,1981,2006]" metricMagnitude="2" metricUnit="m" metricValue="6.35" pageId="26" pageNumber="527" unit="m" value="635.0">635m</quantity>
,
<date id="D63A84A06B6BFF99FC8FF84B07E35FE5" box="[770,898,1981,2006]" pageId="26" pageNumber="527" value="1983-12-13">13.xii.1983</date>
A Hiller (QMB); ♀/ Mt Glorious,
<date id="D63A84A06B6BFF99FA90F84B01E65FE5" box="[1309,1415,1981,2006]" pageId="26" pageNumber="527" value="1983-01-21">21.i.1983</date>
J Sedlacek (QMB);
<materialsCitation id="12ECA83D6B6AFF98FEE7FF6E06885883" ID-GBIF-Occurrence="1915499068" box="[362,745,151,176]" collectingDate="1978-01-17" collectionCode="QMB" collectorName="FT Fricke" country="United Kingdom" location="Queensland" pageId="27" pageNumber="528" specimenCount="1" specimenCount-female="1" stateProvince="Queensland">
<specimenCount id="B48269E96B6AFF98FEE7FF6E051A5883" box="[362,379,152,176]" pageId="27" pageNumber="528" type="female"></specimenCount>
/ ditto except
<date id="D63A84A06B6AFF98FDADFF6106EB5883" box="[544,650,151,176]" pageId="27" pageNumber="528" value="1978-01-17">
<collectingDate id="C67E7D486B6AFF98FDADFF6106EB5883" box="[544,650,151,176]" pageId="27" pageNumber="528" value="1978-01-17">17.i.1978</collectingDate>
</date>
(
<collectionCode id="C4953AA56B6AFF98FD11FF6E06805883" box="[668,737,152,176]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="27" pageNumber="528">QMB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="12ECA83D6B6AFF98FD75FF6E01405883" ID-GBIF-Occurrence="1915499067" box="[760,1313,151,176]" collectingDate="1972-02-15" collectionCode="QMB" collectorName="Monteith" country="United Kingdom" location="Queensland" pageId="27" pageNumber="528" specimenCount="1" specimenCount-male="1" stateProvince="GB">
<specimenCount id="B48269E96B6AFF98FD75FF6E076D5883" box="[760,780,152,176]" pageId="27" pageNumber="528" type="male"></specimenCount>
/ ditto except
<collectingDate id="C67E7D486B6AFF98FC22FF6100405883" box="[943,1057,151,176]" pageId="27" pageNumber="528" value="1972-02-15">
<date id="D63A84A06B6AFF98FC22FF6100405883" box="[943,1057,151,176]" pageId="27" pageNumber="528" value="1972-02-15">15.ii.1972</date>
</collectingDate>
<collectingRegion id="60406C826B6AFF98FBA7FF6E00305883" box="[1066,1105,152,176]" country="Guinea-Bissau" name="Gabu" pageId="27" pageNumber="528">GB</collectingRegion>
<collectorName id="0F71C7B66B6AFF98FBD6FF6100A35883" box="[1115,1218,151,176]" pageId="27" pageNumber="528">Monteith</collectorName>
(
<collectionCode id="C4953AA56B6AFF98FB59FF6E01785883" box="[1236,1305,152,176]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="27" pageNumber="528">QMB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="12ECA83D6B6AFF98FABDFF6E07F658E7" ID-GBIF-Occurrence="1915499058" collectingDate="1987-11-21" collectingDateMax="1987-11-22" collectingDateMin="1987-11-21" collectionCode="QMB" collectorName="Goomburra &amp; Monteith" country="United Kingdom" location="Upper Dalrymple Ck" pageId="27" pageNumber="528" specimenCount="1" specimenCount-female="1" stateProvince="GB">
<specimenCount id="B48269E96B6AFF98FABDFF6E01205883" box="[1328,1345,152,176]" pageId="27" pageNumber="528" type="female"></specimenCount>
/
<location id="A75BF4BB6B6AFF98FADBFF6E055958E6" LSID="urn:lsid:plazi:treatment:2A2D13766B6BFF9FFF1AFAA907795A9B:A75BF4BB6B6AFF98FADBFF6E055958E6" country="United Kingdom" name="Upper Dalrymple Ck" pageId="27" pageNumber="528" stateProvince="GB">Upper Dalrymple Ck</location>
, via
<collectorName id="0F71C7B66B6AFF98FEE3FF4A059358E6" box="[366,498,188,213]" pageId="27" pageNumber="528">Goomburra</collectorName>
,
<collectingDate id="C67E7D486B6AFF98FE70FF4A06FF58E6" box="[509,670,188,213]" pageId="27" pageNumber="528" value="1987-11-21" valueMax="1987-11-22" valueMin="1987-11-21">
<date id="D63A84A06B6AFF98FE70FF4A06FF58E6" box="[509,670,188,213]" pageId="27" pageNumber="528" value="1987-11-21" valueMax="1987-11-22" valueMin="1987-11-21">2122.xi.1987</date>
</collectingDate>
<collectingRegion id="60406C826B6AFF98FD28FF4A06AD58E7" box="[677,716,188,212]" country="Guinea-Bissau" name="Gabu" pageId="27" pageNumber="528">GB</collectingRegion>
<collectorName id="0F71C7B66B6AFF98FD5EFF4A075B58E6" box="[723,826,188,213]" pageId="27" pageNumber="528">Monteith</collectorName>
(
<collectionCode id="C4953AA56B6AFF98FCC4FF4A07EE58E7" box="[841,911,188,212]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="27" pageNumber="528">QMB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EA9EF1EB6B6AFF98FF4AFF290083592E" pageId="27" pageNumber="528" type="diagnosis">
<paragraph id="A23BA2606B6AFF98FF4AFF290083592E" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="528">
<emphasis id="90F07E726B6AFF98FF4AFF29052058CB" bold="true" box="[199,321,223,248]" pageId="27" pageNumber="528">Diagnosis.</emphasis>
<taxonomicName id="6584D9E36B6AFF98FEC6FF16065758CB" authorityName="Reid" authorityYear="1992" box="[331,566,224,248]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6AFF98FEC6FF16065758CB" box="[331,566,224,248]" italics="true" pageId="27" pageNumber="528">Cheiloxena tuberosa</emphasis>
</taxonomicName>
is distinguished by: anterior margin of clypeus deeply excavate; pronotum without dorsal ridges but with 12 lateral tubercles; each elytron with four large conical tubercles.
</paragraph>
</subSubSection>
<subSubSection id="EA9EF1EB6B6AFF9FFF4AFED1014A5957" lastPageId="28" lastPageNumber="529" pageId="27" pageNumber="528" type="description">
<paragraph id="A23BA2606B6AFF98FF4AFED105655AFE" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="528">
<emphasis id="90F07E726B6AFF98FF4AFED105385973" bold="true" box="[199,345,295,320]" pageId="27" pageNumber="528">Description.</emphasis>
Length: male
<quantity id="657C0F856B6AFF98FD8AFED106C0590C" box="[519,673,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.15" metricValueMin="0.95" pageId="27" pageNumber="528" unit="mm" value="10.5" valueMax="11.5" valueMin="9.5">9.511.5 mm</quantity>
, female
<quantity id="657C0F856B6AFF98FC85FED107C5590C" box="[776,932,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.15" metricValueMin="0.85" pageId="27" pageNumber="528" unit="mm" value="10.0" valueMax="11.5" valueMin="8.5">8.511.5 mm</quantity>
; body convex in profile, length about 2.7x height; colour entirely black, except apex last maxillary palp and labial palpi reddish-brown, with ashy grey or golden scale-like setae; scale-like setae adpressed except semi-erect on largest elytral tubercles, variable in proportion (length
<date id="D63A84A06B6AFF98FEFBFE6205DD599F" box="[374,444,404,428]" pageId="27" pageNumber="528" value="2010-10-03">310x</date>
width), thinner and sparser on depressed areas of pronotum and elytra, and apical ventrites, distributed as follows: almost evenly on head but golden behind eyes and white anteriorly; variegated on pronotum with band of dense golden setae on anterior ridge and posterior margin and white setae in five irregular broad stripes following elevated areas, almost glabrous between; scutellum clothed with elongate golden setae; elytra with irregular mosaic of adpressed white setae except tubercles with elongate semi-erect golden setae and golden adpressed setae posterior to tubercles, ventral surfaces with dense white adpressed setae except thin and sparse on apical ventrite, setae of appendages thin but dense, except antennomeres 811 minutely setose; surface sculpture: head pronotum, elytra and venter dull, punctate and densely microsculptured, except shining apices of tubercles.
</paragraph>
<paragraph id="A23BA2606B6AFF98FF4AFD2107615C64" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="528">
Head (
<figureCitation id="3ABFBEE56B6AFF98FE9FFD21050B5AC3" box="[274,362,727,752]" captionStart="FIGURES 710" captionStartId="7.[151,264,1975,1998]" captionTargetBox="[194,1366,311,1853]" captionTargetId="figure@7.[152,1435,139,1955]" captionTargetPageId="7" captionText="FIGURES 710. Habitus of Cheiloxena species: 7, C. insignis Baly (Launching Place, Victoria); 8, C. insignis (Lake Mountain, Victoria); 9, C. monga Reid &amp; Beatson (holotype); 10, C. tuberosa Reid; Not to scale." httpUri="https://zenodo.org/record/1456317/files/figure.png" pageId="27" pageNumber="528">Figs 10</figureCitation>
,
<figureCitation id="3ABFBEE56B6AFF98FEFAFD2E05F45AC3" box="[375,405,728,752]" captionStart="FIGURES 1321" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[230,1395,232,1777]" captionTargetId="figure@13.[152,1435,141,1956]" captionTargetPageId="13" captionText="FIGURES 1321. Lateral of Cheiloxena species: 13, C. aitori Reid &amp; Beatson, male (holotype); 14, C. blackburni Reid; 15, C. conani Reid &amp; Beatson; 16, C. frenchae Blackburn, (holotype); 17, C. insignis Blackburn; 18, C. monga Reid &amp; Beatson (holotype); 19, C. tuberosa Reid; 20, C. westwoodii Baly (northern form, Dorrigo); 21, C. westwoodii Baly (southern form, Victoria). Not to scale." httpUri="https://zenodo.org/record/1456321/files/figure.png" pageId="27" pageNumber="528">19</figureCitation>
,
<figureCitation id="3ABFBEE56B6AFF98FE2FFD2E05A05AC3" box="[418,449,728,752]" captionStart="FIGURES 2230" captionStartId="14.[151,264,1700,1723]" captionTargetBox="[212,1381,282,1587]" captionTargetId="figure@14.[150,1436,193,1679]" captionTargetPageId="14" captionText="FIGURES 2230. Anterior view, Cheiloxena species: 22, C. aitori Reid &amp; Beatson, male (holotype); 23, C. blackburni Reid; 24, C. conani Reid &amp; Beatson; 25, C. frenchae Blackburn; 26, C. insignis Baly; 27, C. monga Reid &amp; Beatson; 28, C. tuberosa Reid; 29, C. westwoodii Baly (northern form, Dorrigo); 30, C. westwoodii Baly (southern form, Victoria)." httpUri="https://zenodo.org/record/1456323/files/figure.png" pageId="27" pageNumber="528">28</figureCitation>
,
<figureCitation id="3ABFBEE56B6AFF98FE40FD2E058C5AC3" box="[461,493,728,752]" captionStart="FIGURES 3842" captionStartId="17.[151,264,1957,1980]" captionTargetBox="[281,1374,340,1843]" captionTargetId="figure@17.[152,1435,193,2009]" captionTargetPageId="17" captionText="FIGURES 3842. Cheiloxena species: 38, C. tuberosa Reid, male antenna; 39, C. insignis Blackburn, male antenna; 40, C. insignis, metafemur; 41, C. tuberosa, pro- (left) and meta- (right) tarsi; 42, C. frenchae Blackburn, apex of tarsus." httpUri="https://zenodo.org/record/1456329/files/figure.png" pageId="27" pageNumber="528">38</figureCitation>
): distinctly narrower than pronotum in both sexes; densely and slightly confluently punctured, punctures small, intervals dull and microsculptured; narrow impunctate smooth area around antennal cavities; flat between eyes, slightly concave between antennae; eyes small, slightly laterally prominent, separated by about 4 eye widths (male) or about 4.5 eye widths (female); gena at shortest point about 0.6x eye length (male) or about 0.4x eye length (female); genal lobe 0.75x shortest length of gena; antennae 33.5x socket diameters apart; antennae about 0.65x body length (male), or about 0.55x body length (female); antennomeres 17 dull with shining apices, 811 duller; antennomeres 1 and 311 elongate, 2 transverse 0.50.7x length of 1, 0.250.3x length of 3, relative lengths of antennomeres, with 3 longest and 11 next, and middle segments generally similar in length: male: 2, &lt;1, &lt;4=5=6, &lt;9=10, &lt;8, &lt;7, &lt;11, &lt;3; female: 2, &lt;1, &lt;6, &lt;4=5=8=9=10, &lt;7, &lt;11, &lt;3; clypeus with 23 pairs of long subapical setae; clypeal anterior margin deeply concave, usually less so in female; male apical maxillary palpomere broadly ovoid with truncate apex.
</paragraph>
<paragraph id="A23BA2606B6AFF98FF4AFB9200EC5F7E" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="528">
Thorax (
<figureCitation id="3ABFBEE56B6AFF98FEABFB92051C5C4F" box="[294,381,1124,1149]" captionStart="FIGURES 710" captionStartId="7.[151,264,1975,1998]" captionTargetBox="[194,1366,311,1853]" captionTargetId="figure@7.[152,1435,139,1955]" captionTargetPageId="7" captionText="FIGURES 710. Habitus of Cheiloxena species: 7, C. insignis Baly (Launching Place, Victoria); 8, C. insignis (Lake Mountain, Victoria); 9, C. monga Reid &amp; Beatson (holotype); 10, C. tuberosa Reid; Not to scale." httpUri="https://zenodo.org/record/1456317/files/figure.png" pageId="27" pageNumber="528">Figs 10</figureCitation>
,
<figureCitation id="3ABFBEE56B6AFF98FE05FB9205C75C4F" box="[392,422,1124,1148]" captionStart="FIGURES 1321" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[230,1395,232,1777]" captionTargetId="figure@13.[152,1435,141,1956]" captionTargetPageId="13" captionText="FIGURES 1321. Lateral of Cheiloxena species: 13, C. aitori Reid &amp; Beatson, male (holotype); 14, C. blackburni Reid; 15, C. conani Reid &amp; Beatson; 16, C. frenchae Blackburn, (holotype); 17, C. insignis Blackburn; 18, C. monga Reid &amp; Beatson (holotype); 19, C. tuberosa Reid; 20, C. westwoodii Baly (northern form, Dorrigo); 21, C. westwoodii Baly (southern form, Victoria). Not to scale." httpUri="https://zenodo.org/record/1456321/files/figure.png" pageId="27" pageNumber="528">19</figureCitation>
,
<figureCitation id="3ABFBEE56B6AFF98FE3FFB9205B15C4F" box="[434,464,1124,1148]" captionStart="FIGURES 2230" captionStartId="14.[151,264,1700,1723]" captionTargetBox="[212,1381,282,1587]" captionTargetId="figure@14.[150,1436,193,1679]" captionTargetPageId="14" captionText="FIGURES 2230. Anterior view, Cheiloxena species: 22, C. aitori Reid &amp; Beatson, male (holotype); 23, C. blackburni Reid; 24, C. conani Reid &amp; Beatson; 25, C. frenchae Blackburn; 26, C. insignis Baly; 27, C. monga Reid &amp; Beatson; 28, C. tuberosa Reid; 29, C. westwoodii Baly (northern form, Dorrigo); 30, C. westwoodii Baly (southern form, Victoria)." httpUri="https://zenodo.org/record/1456323/files/figure.png" pageId="27" pageNumber="528">28</figureCitation>
,
<figureCitation id="3ABFBEE56B6AFF98FE51FB92059D5C4F" box="[476,508,1124,1148]" captionStart="FIGURES 3842" captionStartId="17.[151,264,1957,1980]" captionTargetBox="[281,1374,340,1843]" captionTargetId="figure@17.[152,1435,193,2009]" captionTargetPageId="17" captionText="FIGURES 3842. Cheiloxena species: 38, C. tuberosa Reid, male antenna; 39, C. insignis Blackburn, male antenna; 40, C. insignis, metafemur; 41, C. tuberosa, pro- (left) and meta- (right) tarsi; 42, C. frenchae Blackburn, apex of tarsus." httpUri="https://zenodo.org/record/1456329/files/figure.png" pageId="27" pageNumber="528">41</figureCitation>
): sides almost vertical, no clear distinction between dorsal and ventral (hypomeral) areas of pronotum; pronotal puncturation similar to head, except depressed parts of disc sparsely punctured; interspaces dull; pronotum slightly longer than wide, width 0.95x length, greatest width at anterior third and sides weakly sinuate; anterior angles prominent but tips blunt, 45°; anterior margin produced, truncate at middle; lateral sides with 12 small tubercles in apical half; basal margin weakly convex; pronotum with transverse preapical low ridge of shining tubercles, notched at middle; disc unevenly surfaced, with concave sides and middle and small postmedian swelling; hypomeron and prosternum densely punctured and pubescent, as pronotum, except hypomeral lobe rugose and glabrous; scutellum punctured and setose, as pronotum; elytra irregularly scaled and glabrous, the scales generally sparse except for small white patch near elytral apex on lateral margin; elytron conspicuously tuberculate, with variably sized shiny tipped tubercles arranged in approximately 4 irregular longitudinal rows, as follows: first (innermost) with 3 similar sized massive conical tubercles; second with 2 small conical tubercles at elytral base, one massive conical tubercle beside apical tubercle of inner row, and small tubercle on apical elytral slope; third starting on inner surface of humerus, with elongate ridge at base and 3 small tubercles posteriorly; fourth starting on outer surface of humerus, with 69 small tubercles; elytral disc shallowly arcuately depressed in basal half, posterior to humerus, and concave at sides of middle; elytral punctures large, 1.5
<date id="D63A84A06B6AFF98FF1AF97604D25EAB" box="[151,179,1664,1688]" pageId="27" pageNumber="528">2x</date>
pronotal puncture diameters, and sparse, separated by 11.5x diameters, interspaces dull, microsculptured; discal elytral punctures with two small shining tubercles laterally on rim; elytral apex narrowly truncate; epipleuron finely and shallowly punctured, with scale-like pubescence; mesoventrite punctured as prosternum; metepisternum and metaventrite dull and microsculptured, closely punctured and scaled, except midline impunctate and shining; metatibiae with shallow irregular lateral longitudinal grooves, short and robust; bases of tarsomeres 13 depressed; protarsomeres 13 slightly broader at base in male than female.
</paragraph>
<paragraph id="A23BA2606B6AFF9FFF4AF8A1059E5973" blockId="27.[151,1437,151,2013]" lastBlockId="28.[151,1437,151,680]" lastPageId="28" lastPageNumber="529" pageId="27" pageNumber="528">
Abdomen (
<figureCitation id="3ABFBEE56B6AFF98FEC9F8A105FB5F43" box="[324,410,1879,1904]" captionStart="FIGURES 4351" captionStartId="18.[151,264,1789,1812]" captionTargetId="figure@18.[152,1435,193,1768]" captionTargetPageId="18" captionText="FIGURES 4351. Cheiloxena species, male genitalia lateral and dorsal: 43, C. aitori Reid &amp; Beatson; 44, C. blackburni Reid; 45, C. conani Reid &amp; Beatson; 46, C. frenchae Blackburn; 47, C. insignis Blackburn; 48, C. monga Reid &amp; Beatson; 49, C. tuberosa Reid; 50, C. westwoodii Baly (northern form, Dorrigo); 51, C. westwoodii (southern form, Victoria). All images to scale, scale bar = 2mm." httpUri="https://zenodo.org/record/1457873/files/figure.png" pageId="27" pageNumber="528">Figs 49</figureCitation>
,
<figureCitation id="3ABFBEE56B6AFF98FE2BF8A105A55F43" box="[422,452,1879,1904]" captionStart="FIGURES 5261" captionStartId="19.[151,264,1911,1934]" captionTargetBox="[224,1367,255,1760]" captionTargetId="figure@19.[152,1435,131,1947]" captionTargetPageId="19" captionText="FIGURES 5261. Cheiloxena species, apex of penis: 52, C. aitori Reid &amp; Beatson; 53, C. blackburni Reid; 54, C. conani Reid &amp; Beatson; 55, C. frenchae Blackburn; 56, C. insignis Blackburn (Launching Place, Victoria); 57, C. insignis Blackburn (Lake Mountain, Victoria); 58, C. monga Reid &amp; Beatson; 59, C. tuberosa Reid; 60, C. westwoodii Baly (northern form, Dorrigo); 61, C. westwoodii (southern form, Victoria). All images to same scale." httpUri="https://zenodo.org/record/1456331/files/figure.png" pageId="27" pageNumber="528">59</figureCitation>
,
<figureCitation id="3ABFBEE56B6AFF98FE42F8A1058F5F43" box="[463,494,1879,1904]" captionStart="FIGURES 6268" captionStartId="20.[151,264,1947,1970]" captionTargetBox="[222,1410,255,1799]" captionTargetId="figure@20.[152,1435,111,1926]" captionTargetPageId="20" captionText="FIGURES 6268. Cheiloxena species, female genitalia: 62, C. aitori Reid &amp; Beatson; 63, C. blackburni Reid; 64, C. conani Reid &amp; Beatson; 65, C. frenchae Blackburn; 66, C. insignis Blackburn; 67, C. tuberosa Reid; 68, C. westwoodii Baly (northern form, Dorrigo)." httpUri="https://zenodo.org/record/1456333/files/figure.png" pageId="27" pageNumber="528">67</figureCitation>
,
<figureCitation id="3ABFBEE56B6AFF98FE74F8AE06785F43" box="[505,537,1880,1904]" captionStart="FIGURES 6976" captionStartId="21.[151,264,1871,1894]" captionText="FIGURES 6976. Cheiloxena species, female genitalia, sternite 8: 69, C. aitori Reid &amp; Beatson; 70, C. blackburni Reid; 71, C. conani Reid &amp; Beatson; 72, C. frenchae Blackburn; 73, C. insignis Blackburn; 74, C. tuberosa Reid; 75, C. westwoodii Baly (northern form, Dorrigo); 76, C. westwoodii (southern form, Victoria)." pageId="27" pageNumber="528">74</figureCitation>
): ventrite I shinier than thoracic ventrites but closely and strongly punctured and microreticulate; IIV impunctate and shining at apices, but densely punctured, shining at middle but shallowly microreticulate, dull at sides; ventrite pubescence recumbent, scale-like, but much narrower on ventrite V, erect setae only present on apical margin ventrite V; apex of ventrite V narrowly truncate in both sexes; apex of penis contracted to elongate mucronate tip in dorsal view, short and thick in lateral view, strongly reflexed by about 90°; tegminal keel deep and slightly sinuate in lateral view; female sternite VIII apodeme short and narrow, apical sclerotised area roughly triangular but expanded at sides, longer than wide, apex concave; apex of gonocoxite without conspicuous setae; stylus only slightly elongate; median ventral sclerite elongate; spermatheca falcate, acutely tipped, with coiled duct.
</paragraph>
<paragraph id="A23BA2606B6DFF9FFF4AFEBA014A5957" blockId="28.[151,1437,151,680]" box="[199,1323,332,357]" pageId="28" pageNumber="529">
<emphasis id="90F07E726B6DFF9FFF4AFEBA05705956" bold="true" box="[199,273,332,357]" pageId="28" pageNumber="529">Notes.</emphasis>
The maxillary palpi of
<taxonomicName id="6584D9E36B6DFF9FFD97FEBA06FA5957" authorityName="Reid" authorityYear="1992" box="[538,667,332,356]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6DFF9FFD97FEBA06FA5957" box="[538,667,332,356]" italics="true" pageId="28" pageNumber="529">C. tuberosa</emphasis>
</taxonomicName>
were illustrated by
<bibRefCitation id="C615DF916B6DFF9FFCF7FEBA003F5957" box="[890,1118,332,357]" pageId="28" pageNumber="529" refId="ref20270" refString="Reid, C. A. M. (1992) Revision of the genus Cheiloxena Baly (Coleoptera: Chrysomelidae: Eumolpinae). Memoirs of the Museum of Fictoria, 53 (1), 101 - 114. https: // doi. org / 10.24199 / j. mmv. 1992.53.04" type="journal article">
Reid (1992;
<figureCitation id="3ABFBEE56B6DFF9FFB8BFEBA00375957" box="[1030,1110,332,357]" captionStart="FIGURES 1321" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[230,1395,232,1777]" captionTargetId="figure@13.[152,1435,141,1956]" captionTargetPageId="13" captionText="FIGURES 1321. Lateral of Cheiloxena species: 13, C. aitori Reid &amp; Beatson, male (holotype); 14, C. blackburni Reid; 15, C. conani Reid &amp; Beatson; 16, C. frenchae Blackburn, (holotype); 17, C. insignis Blackburn; 18, C. monga Reid &amp; Beatson (holotype); 19, C. tuberosa Reid; 20, C. westwoodii Baly (northern form, Dorrigo); 21, C. westwoodii Baly (southern form, Victoria). Not to scale." httpUri="https://zenodo.org/record/1456321/files/figure.png" pageId="28" pageNumber="529">Fig. 19</figureCitation>
)
</bibRefCitation>
under
<taxonomicName id="6584D9E36B6DFF9FFB21FEBA01455957" authorityName="Blackburn" authorityYear="1896" box="[1196,1316,332,356]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="90F07E726B6DFF9FFB21FEBA01455957" box="[1196,1316,332,356]" italics="true" pageId="28" pageNumber="529">C. insignis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="EA9EF1EB6B6DFF9FFF4AFE8607795A9B" pageId="28" pageNumber="529" type="reference_group">
<paragraph id="A23BA2606B6DFF9FFF4AFE8607175A2B" blockId="28.[151,1437,151,680]" pageId="28" pageNumber="529">
<taxonomicName id="6584D9E36B6DFF9FFF4AFE8605CE59BB" authorityName="Reid" authorityYear="1992" box="[199,431,368,392]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6DFF9FFF4AFE8605CE59BB" box="[199,431,368,392]" italics="true" pageId="28" pageNumber="529">Cheiloxena tuberosa</emphasis>
</taxonomicName>
is a relatively frequently collected species in the subtropical forests of northeastern
<collectingRegion id="60406C826B6DFF9FFAEAFE860542599E" country="Australia" name="New South Wales" pageId="28" pageNumber="529">
New South
<collectingCountry id="DA93E2F06B6DFF9FFF52FE620542599E" box="[223,291,404,429]" name="United Kingdom" pageId="28" pageNumber="529">Wales</collectingCountry>
</collectingRegion>
and southeastern
<collectingRegion id="60406C826B6DFF9FFE61FE620611599E" box="[492,624,404,429]" country="Australia" name="Queensland" pageId="28" pageNumber="529">Queensland</collectingRegion>
(
<figureCitation id="3ABFBEE56B6DFF9FFD0DFE6206B2599F" box="[640,723,404,429]" captionStart="FIGURE 77" captionStartId="31.[151,250,1971,1993]" captionTargetId="figure@31.[151,1435,193,1950]" captionTargetPageId="31" captionText="FIGURE 77. Eastern Australia showing distribution of Cheiloxena species." httpUri="https://zenodo.org/record/1457875/files/figure.png" pageId="28" pageNumber="529">Fig. 77</figureCitation>
), from Dorrigo to Bellthorpe. The northernmost record is based on a photograph of this distinctive species posted on the internet (
<bibRefCitation id="C615DF916B6DFF9FFC04FE41007A59E3" box="[905,1051,439,464]" pageId="28" pageNumber="529" refId="ref20721" refString="Walter, G. (2016) Weevil at Bellthorpe. Available from: http: // www. bowerbird. org. au / observations / 74731 (accessed 29 May 2018)" type="url">Walter 2016</bibRefCitation>
). The distributions of
<taxonomicName id="6584D9E36B6DFF9FFA95FE4E01FD59E3" authorityName="Reid" authorityYear="1992" box="[1304,1436,440,464]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6DFF9FFA95FE4E01FD59E3" box="[1304,1436,440,464]" italics="true" pageId="28" pageNumber="529">C. tuberosa</emphasis>
</taxonomicName>
and its closest relative and most similar species,
<taxonomicName id="6584D9E36B6DFF9FFD3CFE2A07BF59C7" authority="(Reid 1992)" authorityName="Reid" authorityYear="1992" baseAuthorityName="Reid" baseAuthorityYear="1992" box="[689,990,476,501]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="westwoodii">
<emphasis id="90F07E726B6DFF9FFD3CFE2A072E59C7" box="[689,847,476,500]" italics="true" pageId="28" pageNumber="529">C. westwoodii</emphasis>
(
<bibRefCitation id="C615DF916B6DFF9FFCD0FE2A07B759C7" box="[861,982,476,501]" pageId="28" pageNumber="529" refId="ref20270" refString="Reid, C. A. M. (1992) Revision of the genus Cheiloxena Baly (Coleoptera: Chrysomelidae: Eumolpinae). Memoirs of the Museum of Fictoria, 53 (1), 101 - 114. https: // doi. org / 10.24199 / j. mmv. 1992.53.04" type="journal article">Reid 1992</bibRefCitation>
)
</taxonomicName>
, broadly overlap but the two species are not found together as the latter is restricted to eucalypt woodland.
</paragraph>
<paragraph id="A23BA2606B6DFF9FFF4AFDD207795A9B" blockId="28.[151,1437,151,680]" pageId="28" pageNumber="529">
The host is unknown. It has once been collected in an intercept trap hung on a tree of
<taxonomicName id="6584D9E36B6DFF9FFB7FFDD205475A53" ID-CoL="7NPNY" class="Magnoliopsida" family="Malvaceae" genus="Argyrodendron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="28" pageNumber="529" phylum="Tracheophyta" rank="genus">
<emphasis id="90F07E726B6DFF9FFB7FFDD201FD5A0F" box="[1266,1436,548,572]" italics="true" pageId="28" pageNumber="529">Argyrodendron</emphasis>
(Malvaceae)
</taxonomicName>
, which occurs throughout the range of
<taxonomicName id="6584D9E36B6DFF9FFD51FDBE073F5A53" authorityName="Reid" authorityYear="1992" box="[732,862,584,608]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6DFF9FFD51FDBE073F5A53" box="[732,862,584,608]" italics="true" pageId="28" pageNumber="529">C. tuberosa</emphasis>
</taxonomicName>
, but this seems to be a casual association. A single specimen has been collected on leaves of
<taxonomicName id="6584D9E36B6DFF9FFD01FD9A00605AB7" ID-CoL="3BPZH" box="[652,1025,620,644]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="28" pageNumber="529" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="90F07E726B6DFF9FFD01FD9A07085AB7" box="[652,873,620,644]" italics="true" pageId="28" pageNumber="529">Eucalyptus grandis</emphasis>
(Myrtaceae)
</taxonomicName>
, but without indication of feeding.
<taxonomicName id="6584D9E36B6DFF9FFF1AFD66051F5A9B" authorityName="Reid" authorityYear="1992" box="[151,382,656,680]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="tuberosa">
<emphasis id="90F07E726B6DFF9FFF1AFD66051F5A9B" box="[151,382,656,680]" italics="true" pageId="28" pageNumber="529">Cheiloxena tuberosa</emphasis>
</taxonomicName>
has also been collected at MV light.
</paragraph>
</subSubSection>
</treatment>
</document>