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<document id="3CABBDC1D37758450975A570BF78A206" ID="10.11646/zootaxa.4268.3.9SLASH11063" ID-CLB-Dataset="33029" ID-DOI="10.11646/zootaxa.4268.3.9" ID-GBIF-Dataset="11fda489-d4f7-4efd-8bbf-dad2d8b6c27f" ID-ISSN="1175-5326" ID-Zenodo-Dep="580559" ID-ZooBank="F41A9A05-FC5A-44A6-A2CF-18F8BECFE22A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495012171270" checkinUser="plazi" docAuthor="Selvakumar, C., Subramanian, K. A., Chandra, Kailash, Sivaramakrishnan, K. G., Jehamalar, E. Eyarin &amp; Sinha, Bikramjit" docDate="2017" docId="2A0887B0EF5EE963FF18FDA7FB3FFD27" docLanguage="en" docName="zootaxa.4268.3.9SLASH11063.pdf" docOrigin="Zootaxa 4268 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Choroterpes (Dilatognathus) nigella Kang &amp; Yang 1994" docType="treatment" docVersion="7" lastPageNumber="447" masterDocId="D631FFC8EF5BE96BFF8FFFE0FFD9FFA2" masterDocTitle="A new species and a new record of the subgenus Dilatognathus Kluge 2012 (Ephemeroptera: Leptophlebiidae: genus Choroterpes Eaton, 1881) from India" masterLastPageNumber="447" masterPageNumber="439" pageNumber="444" updateTime="1698439675173" updateUser="ExternalLinkService">
<mods:mods id="B3BBDC7EFE81406D55C4DA4202C41072" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8FE14CF7CA8F0D6AA222D91052D31A52">
<mods:title id="EBBF424EDF7458B4F8937D36C8D4C95B">A new species and a new record of the subgenus Dilatognathus Kluge 2012 (Ephemeroptera: Leptophlebiidae: genus Choroterpes Eaton, 1881) from India</mods:title>
</mods:titleInfo>
<mods:name id="1888D3E776005E81C9011462ED8EB29F" type="personal">
<mods:role id="0A185F677AF0EB73A0769130F495EB92">
<mods:roleTerm id="959AC020477567635B3303BDC0462163">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6761A7C2B057F7060F6E56943DC55016">Selvakumar, C.</mods:namePart>
</mods:name>
<mods:name id="B8ED53538B004F8E16C7F1A4AA530C27" type="personal">
<mods:role id="3C4C6EB6ADE860A78CD1B0215D9A2EA6">
<mods:roleTerm id="66463855E01B6A6251358981C6DEF909">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D0FDDAD21FA6DEF1C4469006529732E">Subramanian, K. A.</mods:namePart>
</mods:name>
<mods:name id="156698CE374D29387D16F24BA4D7DB4C" type="personal">
<mods:role id="345C4E166152F6BDA8735D49B72E7CBE">
<mods:roleTerm id="202B48A9C572FC4BD82B2623AACF27FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="86DC3B85139F0C649A9D09F8463BD31C">Chandra, Kailash</mods:namePart>
</mods:name>
<mods:name id="05A3DC279DAE7F75DA9BABE767B89FAE" type="personal">
<mods:role id="C829430E371A133497D44959F8479FE5">
<mods:roleTerm id="EB32BDB8E2FE8FBE408117B5D1873A45">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CA53E4C98C498B545D0940E950A4F84B">Sivaramakrishnan, K. G.</mods:namePart>
</mods:name>
<mods:name id="E4B0987BC26926C5C6B6F769B2BCE1DB" type="personal">
<mods:role id="D0E4FC94BAEB05F1F2DBC5A7C1574459">
<mods:roleTerm id="CC96AC76A6E118F2F3E11005C7CFE9B3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0672706C0FF03C2B22ABBDC44D750D8">Jehamalar, E. Eyarin</mods:namePart>
</mods:name>
<mods:name id="B524E18557BD138986CB11144D796E47" type="personal">
<mods:role id="E37B87927CF7B9DBD26F3759212ED8C3">
<mods:roleTerm id="3B96ADF6F2DA778862FD230200F00B5E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5E60A4E5640BF9DC7AD16121F57A934B">Sinha, Bikramjit</mods:namePart>
</mods:name>
<mods:typeOfResource id="465425E2F365EE5D1494F560D2F59547">text</mods:typeOfResource>
<mods:relatedItem id="2A115071B5B3BA6F8F0B679172DC1377" type="host">
<mods:titleInfo id="F72AC832B643D56E446A3227CED5C217">
<mods:title id="4FD7CE31BD3FFD1AB108FDF561C00650">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DA84DE5F2D1EDD74B5737AE6F646F00A">
<mods:date id="DE6CB1A0F29D0E63304D3094D4939CA9">2017</mods:date>
<mods:detail id="9CB083DCB1197AB2A46C54E8132975EB" type="volume">
<mods:number id="919D24E30B365D52E025922FA8D32D71">4268</mods:number>
</mods:detail>
<mods:detail id="CE1CAA40262402F0A2C4A94CC0352A97" type="issue">
<mods:number id="E56A8F264BFC67370D8B07F4B78B5DA6">3</mods:number>
</mods:detail>
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<mods:start id="7DB5CF2EAD9574D60DE1B8E9562F5AD0">439</mods:start>
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</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0B6F7F23A3A625D61F63E5BF08DFF514">journal article</mods:classification>
<mods:identifier id="42484D6E112A7C56D3A59E538A6AFF00" type="CLB-Dataset">33029</mods:identifier>
<mods:identifier id="1844F686984E078F9464AE2A870AA5E1" type="DOI">10.11646/zootaxa.4268.3.9</mods:identifier>
<mods:identifier id="B9EE5E5A96A34E6474F8C90858692C21" type="GBIF-Dataset">11fda489-d4f7-4efd-8bbf-dad2d8b6c27f</mods:identifier>
<mods:identifier id="61DAFF606CBBB42B824D08A07375D027" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AF6634502293D82B643C836FA7AE77B8" type="Zenodo-Dep">580559</mods:identifier>
<mods:identifier id="AD8073A98FE437136D5B719168B0C727" type="ZooBank">F41A9A05-FC5A-44A6-A2CF-18F8BECFE22A</mods:identifier>
</mods:mods>
<treatment id="2A0887B0EF5EE963FF18FDA7FB3FFD27" ID-DOI="http://doi.org/10.5281/zenodo.6029290" ID-GBIF-Taxon="130810176" ID-Zenodo-Dep="6029290" LSID="urn:lsid:plazi:treatment:2A0887B0EF5EE963FF18FDA7FB3FFD27" httpUri="http://treatment.plazi.org/id/2A0887B0EF5EE963FF18FDA7FB3FFD27" lastPageId="8" lastPageNumber="447" pageId="5" pageNumber="444">
<subSubSection id="EABB652DEF5EE96EFF18FDA7FEF7FD21" pageId="5" pageNumber="444" type="nomenclature">
<paragraph id="A21E36A6EF5EE96EFF18FDA7FC8DFDC0" blockId="5.[151,852,583,643]" box="[151,852,583,610]" pageId="5" pageNumber="444">
<heading id="F95681CAEF5EE96EFF18FDA7FC8DFDC0" bold="true" box="[151,852,583,610]" fontSize="11" level="1" pageId="5" pageNumber="444" reason="1">
<taxonomicName id="65A14D25EF5EE96EFF18FDA7FC8DFDC0" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" box="[151,852,583,610]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="nigella" subGenus="Dilatognathus">
<emphasis id="90D5EAB4EF5EE96EFF18FDA7FC8DFDC0" bold="true" box="[151,852,583,610]" pageId="5" pageNumber="444">
<emphasis id="90D5EAB4EF5EE96EFF18FDA7FEF2FDC3" bold="true" box="[151,299,583,609]" italics="true" pageId="5" pageNumber="444">Choroterpes</emphasis>
(
<emphasis id="90D5EAB4EF5EE96EFEB3FDA7FE30FDC3" bold="true" box="[316,489,583,609]" italics="true" pageId="5" pageNumber="444">Dilatognathus</emphasis>
)
<emphasis id="90D5EAB4EF5EE96EFE75FDA7FD94FDC3" bold="true" box="[506,589,583,609]" italics="true" pageId="5" pageNumber="444">nigella</emphasis>
(
<bibRefCitation id="C6304B57EF5EE96EFDD3FDA8FC95FDC0" author="Kang" box="[604,844,584,610]" pageId="5" pageNumber="444" refString="Kang, S. C. &amp; Yang, C. T. (1994) Leptophlebiidae of Taiwan (Ephemeroptera). Journal of Taiwan Museum, 47 (1), 57 - 81." type="journal article" year="1994">Kang &amp; Yang 1994</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A21E36A6EF5EE96EFF18FD8AFEF7FD21" blockId="5.[151,852,583,643]" box="[151,302,618,643]" pageId="5" pageNumber="444">
(
<figureCitation id="3A9A2A23EF5EE96EFF10FD8AFEFFFD21" box="[159,294,618,643]" captionStart-0="FIGURES 21 – 27" captionStart-1="FIGURES 28 – 36" captionStartId-0="6.[151,264,1953,1976]" captionStartId-1="7.[151,264,1953,1976]" captionTargetBox-0="[204,1379,200,1922]" captionTargetBox-1="[222,1367,201,1921]" captionTargetId-0="figure@6.[196,1390,193,1932]" captionTargetId-1="figure@7.[212,1376,193,1932]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 21 – 27. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 21. Dorsal view of male; 22. Labrum; 23. Hypopharynx; 24. Left mandible; 25. Right mandible; 26. Maxillae; 27. Labium." captionText-1="FIGURES 28 – 36. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 28. Foreleg; 29. Midleg; 30. Hindleg; 31, Claw; 32. Gill II; 33. Gill III; 34. Gill IV; 35. Gill V; 36. Gill VII." httpUri-0="https://zenodo.org/record/580563/files/figure.png" httpUri-1="https://zenodo.org/record/580564/files/figure.png" pageId="5" pageNumber="444">Figs. 21–36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EABB652DEF5EE96EFF18FD52FB20FC95" pageId="5" pageNumber="444" type="materials_examined">
<paragraph id="A21E36A6EF5EE96EFF18FD52FB20FC95" blockId="5.[151,1437,690,2020]" pageId="5" pageNumber="444">
<materialsCitation id="12C93CFBEF5EE96EFF18FD52FC56FD4D" ID-GBIF-Occurrence="1503335935" collectingDate="2015-04-21" collectorName="K. A. Subramanian" county="Material" elevation="625" latitude="27.3964" location="Ranga River" longLatPrecision="6" longitude="93.7573" municipality="Lower Subansiri District" pageId="5" pageNumber="444" specimenCount="3" specimenCount-larva="3" stateProvince="Arunachal Pradesh">
<emphasis id="90D5EAB4EF5EE96EFF18FD52FE58FD69" bold="true" box="[151,385,690,715]" pageId="5" pageNumber="444">
<collectingCounty id="4B7F4E2AEF5EE96EFF18FD52FF26FD69" box="[151,255,690,715]" pageId="5" pageNumber="444">Material</collectingCounty>
examined.
</emphasis>
<specimenCount id="B4A7FD2FEF5EE96EFE06FD53FE3EFD69" box="[393,487,690,715]" pageId="5" pageNumber="444" type="larva">3 larvae</specimenCount>
,
<collectingRegion id="6065F844EF5EE96EFE7CFD52FD15FD69" box="[499,716,690,715]" country="India" name="Arunachal Pradesh" pageId="5" pageNumber="444">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="427AACDCEF5EE96EFD57FD53FC2BFD69" box="[728,1010,690,715]" pageId="5" pageNumber="444">Lower Subansiri District</collectingMunicipality>
,
<location id="A77E607DEF5EE96EFC71FD53FB57FD69" LSID="urn:lsid:plazi:treatment:2A0887B0EF5EE963FF18FDA7FB3FFD27:A77E607DEF5EE96EFC71FD53FB57FD69" box="[1022,1166,690,715]" county="Material" latitude="27.3964" longLatPrecision="6" longitude="93.7573" municipality="Lower Subansiri District" name="Ranga River" pageId="5" pageNumber="444" stateProvince="Arunachal Pradesh">Ranga River</location>
,
<geoCoordinate id="C7955061EF5EE96EFB15FD52FACDFD68" box="[1178,1300,690,715]" direction="north" orientation="latitude" pageId="5" pageNumber="444" precision="5" value="27.3964">27.3964 N</geoCoordinate>
,
<geoCoordinate id="C7955061EF5EE96EFAAEFD52FA4EFD68" box="[1313,1431,690,715]" direction="east" orientation="longitude" pageId="5" pageNumber="444" precision="5" value="93.7573">93.7573 E</geoCoordinate>
,
<elevation id="298CD195EF5EE96EFF18FD36FF3DFD4C" box="[151,228,726,751]" metricMagnitude="2" metricUnit="m" metricValue="6.25" pageId="5" pageNumber="444" unit="m" value="625.0">
<quantity id="65599B43EF5EE96EFF18FD36FF3DFD4C" box="[151,228,726,751]" metricMagnitude="2" metricUnit="m" metricValue="6.25" pageId="5" pageNumber="444" unit="m" value="625.0">625 m</quantity>
</elevation>
,
<collectingDate id="C65BE98EEF5EE96EFF7AFD36FEB7FD4D" box="[245,366,726,751]" pageId="5" pageNumber="444" value="2015-04-21">
<date id="D61F1066EF5EE96EFF7AFD36FEB7FD4D" box="[245,366,726,751]" pageId="5" pageNumber="444" value="2015-04-21">21.iv.2015</date>
</collectingDate>
, colls.
<collectorName id="0F545370EF5EE96EFE49FD37FD71FD4D" box="[454,680,726,751]" pageId="5" pageNumber="444">K. A. Subramanian</collectorName>
&amp; Bikramjit Sinha
</materialsCitation>
;
<materialsCitation id="12C93CFBEF5EE96EFC10FD37FAF6FCB0" ID-GBIF-Occurrence="1503335937" collectingDate="2013-03-23" collectorName="Srimoyee Basu" country="India" county="Darjeeling" elevation="554" latitude="27.1696" location="Rishi river" longLatPrecision="6" longitude="88.6351" municipality="Rishikhola" pageId="5" pageNumber="444" specimenCount="10" specimenCount-larva="10" stateProvince="West Bengal">
<specimenCount id="B4A7FD2FEF5EE96EFC10FD37FBD7FD4D" box="[927,1038,726,751]" pageId="5" pageNumber="444" type="larva">10 larvae</specimenCount>
,
<collectingCountry id="DAB67636EF5EE96EFB90FD37FBA8FD4D" box="[1055,1137,727,751]" name="India" pageId="5" pageNumber="444">INDIA</collectingCountry>
,
<collectingRegion id="6065F844EF5EE96EFB0EFD37FACFFD4D" box="[1153,1302,726,751]" country="India" name="West Bengal" pageId="5" pageNumber="444">West Bengal</collectingRegion>
,
<collectingCounty id="4B7F4E2AEF5EE96EFAA9FD36FA45FD4D" box="[1318,1436,726,751]" pageId="5" pageNumber="444">Darjeeling</collectingCounty>
(
<collectingRegion id="6065F844EF5EE96EFF11FD1AFF28FCB1" box="[158,241,762,787]" country="India" name="Sikkim" pageId="5" pageNumber="444">Sikkim</collectingRegion>
border),
<collectingMunicipality id="427AACDCEF5EE96EFED6FD1AFE0CFCB1" box="[345,469,762,787]" pageId="5" pageNumber="444">Rishikhola</collectingMunicipality>
,
<location id="A77E607DEF5EE96EFE6EFD1AFD83FCB1" LSID="urn:lsid:plazi:treatment:2A0887B0EF5EE963FF18FDA7FB3FFD27:A77E607DEF5EE96EFE6EFD1AFD83FCB1" box="[481,602,762,787]" country="India" county="Darjeeling" latitude="27.1696" longLatPrecision="6" longitude="88.6351" municipality="Rishikhola" name="Rishi river" pageId="5" pageNumber="444" stateProvince="West Bengal">Rishi river</location>
,
<geoCoordinate id="C7955061EF5EE96EFDE9FD1AFD06FCB0" box="[614,735,762,787]" direction="north" orientation="latitude" pageId="5" pageNumber="444" precision="5" value="27.1696">27.1696 N</geoCoordinate>
,
<geoCoordinate id="C7955061EF5EE96EFD65FD1AFCB9FCB0" box="[746,864,762,787]" direction="east" orientation="longitude" pageId="5" pageNumber="444" precision="5" value="88.6351">88.6351 E</geoCoordinate>
,
<elevation id="298CD195EF5EE96EFCE3FD1AFC6CFCB0" box="[876,949,762,787]" metricMagnitude="2" metricUnit="m" metricValue="5.54" pageId="5" pageNumber="444" unit="m" value="554.0">
<quantity id="65599B43EF5EE96EFCE3FD1AFC6CFCB0" box="[876,949,762,787]" metricMagnitude="2" metricUnit="m" metricValue="5.54" pageId="5" pageNumber="444" unit="m" value="554.0">554 m</quantity>
</elevation>
,
<collectingDate id="C65BE98EEF5EE96EFC4EFD1AFBE7FCB1" box="[961,1086,762,787]" pageId="5" pageNumber="444" value="2013-03-23">
<date id="D61F1066EF5EE96EFC4EFD1AFBE7FCB1" box="[961,1086,762,787]" pageId="5" pageNumber="444" value="2013-03-23">23.iii.2013</date>
</collectingDate>
, coll.
<collectorName id="0F545370EF5EE96EFB0EFD1AFAF6FCB0" box="[1153,1327,762,787]" pageId="5" pageNumber="444">Srimoyee Basu</collectorName>
</materialsCitation>
;
<materialsCitation id="12C93CFBEF5EE96EFAB4FD1AFB20FC95" ID-GBIF-Occurrence="1503335936" collectingDate="2007-02-03" collectorName="J. Lyngdoh" country="India" county="Jaintia Hills district" location="Malidar village" municipality="Wah Malidar" pageId="5" pageNumber="444" specimenCount="5" specimenCount-larva="5" stateProvince="Meghalaya">
<specimenCount id="B4A7FD2FEF5EE96EFAB4FD1AFA41FCB1" box="[1339,1432,762,787]" pageId="5" pageNumber="444" type="larva">5 larvae</specimenCount>
,
<collectingRegion id="6065F844EF5EE96EFF18FCFEFECFFC95" box="[151,278,798,823]" country="India" name="Meghalaya" pageId="5" pageNumber="444">Meghalaya</collectingRegion>
,
<collectingCounty id="4B7F4E2AEF5EE96EFEAEFCFEFDD9FC95" box="[289,512,798,823]" pageId="5" pageNumber="444">Jaintia Hills district</collectingCounty>
,
<collectingMunicipality id="427AACDCEF5EE96EFD84FCFEFD47FC95" box="[523,670,798,823]" pageId="5" pageNumber="444">Wah Malidar</collectingMunicipality>
,
<location id="A77E607DEF5EE96EFD25FCFEFC80FC95" LSID="urn:lsid:plazi:treatment:2A0887B0EF5EE963FF18FDA7FB3FFD27:A77E607DEF5EE96EFD25FCFEFC80FC95" box="[682,857,798,823]" country="India" county="Jaintia Hills district" municipality="Wah Malidar" name="Malidar village" pageId="5" pageNumber="444" stateProvince="Meghalaya">Malidar village</location>
,
<collectingDate id="C65BE98EEF5EE96EFCEBFCFEFC10FC95" box="[868,969,798,823]" pageId="5" pageNumber="444" value="2007-02-03">
<date id="D61F1066EF5EE96EFCEBFCFEFC10FC95" box="[868,969,798,823]" pageId="5" pageNumber="444" value="2007-02-03">3.ii.2007</date>
</collectingDate>
, colls.
<collectorName id="0F545370EF5EE96EFB97FCFFFB4CFC95" box="[1048,1173,798,823]" pageId="5" pageNumber="444">J. Lyngdoh</collectorName>
&amp; Party.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="EABB652DEF5EE96EFF48FCA2FE56F8DA" pageId="5" pageNumber="444" type="description">
<paragraph id="A21E36A6EF5EE96EFF48FCA2FB53FAB1" blockId="5.[151,1437,690,2020]" pageId="5" pageNumber="444">
<emphasis id="90D5EAB4EF5EE96EFF48FCA2FE69FCF9" bold="true" box="[199,432,834,859]" pageId="5" pageNumber="444">Description. Larva.</emphasis>
Body length
<quantity id="65599B43EF5EE96EFDC4FCA2FD57FCF8" box="[587,654,834,859]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="444" unit="mm" value="5.0">5 mm</quantity>
, cerci
<quantity id="65599B43EF5EE96EFD56FCA2FCC4FCF8" box="[729,797,834,859]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="444" unit="mm" value="5.0">5 mm</quantity>
, median filament
<quantity id="65599B43EF5EE96EFC66FCA2FBF2FCF8" box="[1001,1067,834,859]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="444" unit="mm" value="6.0">6 mm</quantity>
(
<figureCitation id="3A9A2A23EF5EE96EFBB4FCA2FB56FCF9" box="[1083,1167,834,859]" captionStart="FIGURES 21 – 27" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[204,1379,200,1922]" captionTargetId="figure@6.[196,1390,193,1932]" captionTargetPageId="6" captionText="FIGURES 21 – 27. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 21. Dorsal view of male; 22. Labrum; 23. Hypopharynx; 24. Left mandible; 25. Right mandible; 26. Maxillae; 27. Labium." httpUri="https://zenodo.org/record/580563/files/figure.png" pageId="5" pageNumber="444">Fig. 21</figureCitation>
). Labrum wide, median emargination deep, with sharp semicircular impression on dorsal surface; proximal transverse row of setae interrupted medially; instead of distal transverse row, a wide fringe of irregularly situated setae, smaller than setae of the proximal transverse row; most part of dorsal surface with irregularly arranged long and thin setae, somewhat smaller than setae of distal stripe; closely set long setae laterally (
<figureCitation id="3A9A2A23EF5EE96EFC2CFC32FC22FC49" box="[931,1019,978,1003]" captionStart="FIGURES 21 – 27" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[204,1379,200,1922]" captionTargetId="figure@6.[196,1390,193,1932]" captionTargetPageId="6" captionText="FIGURES 21 – 27. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 21. Dorsal view of male; 22. Labrum; 23. Hypopharynx; 24. Left mandible; 25. Right mandible; 26. Maxillae; 27. Labium." httpUri="https://zenodo.org/record/580563/files/figure.png" pageId="5" pageNumber="444">Fig. 22</figureCitation>
). Lingua of hypopharynx with well developed lateral processes, anterior margin shallowly cleft (
<figureCitation id="3A9A2A23EF5EE96EFCD8FC16FC77FBAD" box="[855,942,1014,1039]" captionStart="FIGURES 21 – 27" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[204,1379,200,1922]" captionTargetId="figure@6.[196,1390,193,1932]" captionTargetPageId="6" captionText="FIGURES 21 – 27. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 21. Dorsal view of male; 22. Labrum; 23. Hypopharynx; 24. Left mandible; 25. Right mandible; 26. Maxillae; 27. Labium." httpUri="https://zenodo.org/record/580563/files/figure.png" pageId="5" pageNumber="444">Fig. 23</figureCitation>
). Mandibles with outer margin moderately convex and with closely set long setae medially (
<figureCitation id="3A9A2A23EF5EE96EFD47FBFAFC8AFB91" box="[712,851,1050,1075]" captionStart="FIGURES 21 – 27" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[204,1379,200,1922]" captionTargetId="figure@6.[196,1390,193,1932]" captionTargetPageId="6" captionText="FIGURES 21 – 27. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 21. Dorsal view of male; 22. Labrum; 23. Hypopharynx; 24. Left mandible; 25. Right mandible; 26. Maxillae; 27. Labium." httpUri="https://zenodo.org/record/580563/files/figure.png" pageId="5" pageNumber="444">Figs. 24–25</figureCitation>
). Maxilla with inner-apical angle produced as tusklike process with 3 simple setae at base; palp elongated and bears long filtering setae; 2nd segment in apical part of inner side with a few setae forming a longitudinal row; 3rd segment with a proximal semicircular transverse row of setae on inner side and fields of irregularly situated setae along with ventral and dorsal sides (
<figureCitation id="3A9A2A23EF5EE96EFB29FB6CFB23FB07" box="[1190,1274,1164,1189]" captionStart="FIGURES 21 – 27" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[204,1379,200,1922]" captionTargetId="figure@6.[196,1390,193,1932]" captionTargetPageId="6" captionText="FIGURES 21 – 27. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 21. Dorsal view of male; 22. Labrum; 23. Hypopharynx; 24. Left mandible; 25. Right mandible; 26. Maxillae; 27. Labium." httpUri="https://zenodo.org/record/580563/files/figure.png" pageId="5" pageNumber="444">Fig. 26</figureCitation>
). Labium with paraglossae widened, expanded laterally; palp elongated, with long filtering setae form regular longitudinal rows; segment 2 with double row on outer side and a row on dorsal side; segment 3 with a row on outer side and a row on inner side; dorsal side of 3rd segment also bears smaller irregularly situated setae (
<figureCitation id="3A9A2A23EF5EE96EFBA6FB1BFBA7FAB1" box="[1065,1150,1275,1300]" captionStart="FIGURES 21 – 27" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[204,1379,200,1922]" captionTargetId="figure@6.[196,1390,193,1932]" captionTargetPageId="6" captionText="FIGURES 21 – 27. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 21. Dorsal view of male; 22. Labrum; 23. Hypopharynx; 24. Left mandible; 25. Right mandible; 26. Maxillae; 27. Labium." httpUri="https://zenodo.org/record/580563/files/figure.png" pageId="5" pageNumber="444">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="A21E36A6EF5EE96EFF48FAC0FDAEF991" blockId="5.[151,1437,690,2020]" pageId="5" pageNumber="444">
Legs pale yellowish; each femur with dark brown spots at middle and near apex; fore femur also with a brown spot near base, widest proximally; middle and hind femur widest at middle. Femora with irregularly situated stout setae of various lengths, dorsal side of fore femur with few setae, dorsal side of middle and hind femur with many setae. Stout setae on inner side of fore tibiae situated dense and irregularly placed, pointed and bipectinate. Inner side of middle tibia with few stout setae. Inner side of hind tibia with a sparse row of stout setae; outer side of hind tibia with stout setae of variable length, as on outer side of femur. Outer sides of all tibiae with irregularly situated thin hairs, whose length exceed tibia width (
<figureCitation id="3A9A2A23EF5EE96EFD0CFA17FCD4F9AD" box="[643,781,1527,1552]" captionStart="FIGURES 28 – 36" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[222,1367,201,1921]" captionTargetId="figure@7.[212,1376,193,1932]" captionTargetPageId="7" captionText="FIGURES 28 – 36. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 28. Foreleg; 29. Midleg; 30. Hindleg; 31, Claw; 32. Gill II; 33. Gill III; 34. Gill IV; 35. Gill V; 36. Gill VII." httpUri="https://zenodo.org/record/580564/files/figure.png" pageId="5" pageNumber="444">Figs. 28–30</figureCitation>
). Claws apically hooked, a row of subequal sized denticles on claws (
<figureCitation id="3A9A2A23EF5EE96EFE86F9FBFE85F991" box="[265,348,1563,1588]" captionStart="FIGURES 28 – 36" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[222,1367,201,1921]" captionTargetId="figure@7.[212,1376,193,1932]" captionTargetPageId="7" captionText="FIGURES 28 – 36. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 28. Foreleg; 29. Midleg; 30. Hindleg; 31, Claw; 32. Gill II; 33. Gill III; 34. Gill IV; 35. Gill V; 36. Gill VII." httpUri="https://zenodo.org/record/580564/files/figure.png" pageId="5" pageNumber="444">Fig. 31</figureCitation>
). Hind wing pad present.
</paragraph>
<paragraph id="A21E36A6EF5EE96EFF48F9DFFD4DF8F6" blockId="5.[151,1437,690,2020]" pageId="5" pageNumber="444">
Abdomen without stout setae; posterolateral margins of abdominal terga with pointed denticles, size of denticles increases from tergum 6 to tergum 9. Posterior margins of abdominal terga 1–2 with row of with small, spine-like denticles; terga 3–7 with row of long, thin, spine-like denticles; terga 8–9 with row of short spine-like denticles at middle, long denticles at corner; sterna without denticles. Each segment of abdominal tergum brown with lighter blanks adjacent to anterior margin. Gill I absent; gills II–VII with dorsal and ventral lamellae terminated in 3 processes, with median process longer than laterals; gills II–V equal size, gill VI smaller, gill VII smallest (
<figureCitation id="3A9A2A23EF5EE96EFE89F8F7FE4AF892" box="[262,403,1815,1840]" captionStart="FIGURES 28 – 36" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[222,1367,201,1921]" captionTargetId="figure@7.[212,1376,193,1932]" captionTargetPageId="7" captionText="FIGURES 28 – 36. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 28. Foreleg; 29. Midleg; 30. Hindleg; 31, Claw; 32. Gill II; 33. Gill III; 34. Gill IV; 35. Gill V; 36. Gill VII." httpUri="https://zenodo.org/record/580564/files/figure.png" pageId="5" pageNumber="444">Figs. 32–36</figureCitation>
). Terminal filament longer than cerci with a whorl of setae on alternate segments; setae longer than length of corresponding segment.
</paragraph>
<paragraph id="A21E36A6EF5EE96EFF48F8BFFE56F8DA" blockId="5.[151,1437,690,2020]" box="[199,399,1887,1912]" pageId="5" pageNumber="444">
<emphasis id="90D5EAB4EF5EE96EFF48F8BFFECAF8DA" bold="true" box="[199,275,1887,1912]" pageId="5" pageNumber="444">Adult.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="EABB652DEF5EE96EFF48F863FC8EF83E" box="[199,855,1923,1948]" pageId="5" pageNumber="444" type="distribution">
<paragraph id="A21E36A6EF5EE96EFF48F863FC8EF83E" blockId="5.[151,1437,690,2020]" box="[199,855,1923,1948]" pageId="5" pageNumber="444">
<emphasis id="90D5EAB4EF5EE96EFF48F863FEB9F83E" bold="true" box="[199,352,1923,1948]" pageId="5" pageNumber="444">Distribution.</emphasis>
<collectingCountry id="DAB67636EF5EE96EFEE8F863FE7AF83E" box="[359,419,1923,1948]" name="India" pageId="5" pageNumber="444">India</collectingCountry>
,
<collectingCountry id="DAB67636EF5EE96EFE21F863FDCDF83E" box="[430,532,1923,1948]" name="Thailand" pageId="5" pageNumber="444">Thailand</collectingCountry>
,
<collectingRegion id="6065F844EF5EE96EFDAFF863FDA8F83E" box="[544,625,1923,1948]" country="China" name="Hainan" pageId="5" pageNumber="444">Hainan</collectingRegion>
and
<collectingCountry id="DAB67636EF5EE96EFD28F863FD20F83E" box="[679,761,1923,1948]" name="Taiwan" pageId="5" pageNumber="444">Taiwan</collectingCountry>
Islands.
</paragraph>
</subSubSection>
<subSubSection id="EABB652DEF5EE96EFF48F847FAECF841" pageId="5" pageNumber="444" type="diagnosis">
<paragraph id="A21E36A6EF5EE96EFF48F847FAECF841" blockId="5.[151,1437,690,2020]" pageId="5" pageNumber="444">
<emphasis id="90D5EAB4EF5EE96EFF48F847FE98F862" bold="true" box="[199,321,1959,1984]" pageId="5" pageNumber="444">Diagnosis.</emphasis>
<taxonomicName id="65A14D25EF5EE96EFEC5F847FC85F81D" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" box="[330,860,1959,1984]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="nigella" subGenus="Dilatognathus">
<emphasis id="90D5EAB4EF5EE96EFEC5F847FE0CF862" box="[330,469,1959,1984]" italics="true" pageId="5" pageNumber="444">Choroterpes</emphasis>
(
<emphasis id="90D5EAB4EF5EE96EFE68F848FDDBF81D" box="[487,514,1960,1983]" italics="true" pageId="5" pageNumber="444">D.</emphasis>
)
<emphasis id="90D5EAB4EF5EE96EFD9BF847FDBBF862" box="[532,610,1959,1984]" italics="true" pageId="5" pageNumber="444">nigella</emphasis>
(
<bibRefCitation id="C6304B57EF5EE96EFDFDF847FC8AF81D" author="Kang" box="[626,851,1959,1984]" pageId="5" pageNumber="444" refString="Kang, S. C. &amp; Yang, C. T. (1994) Leptophlebiidae of Taiwan (Ephemeroptera). Journal of Taiwan Museum, 47 (1), 57 - 81." type="journal article" year="1994">Kang &amp; Yang 1994</bibRefCitation>
)
</taxonomicName>
can be distinguished from all other species by the labrum with deep median emargination and sharp semicircular impression on the dorsal surface (
<figureCitation id="3A9A2A23EF5EE96EFB5BF82BFAF1F841" box="[1236,1320,1995,2020]" captionStart="FIGURES 21 – 27" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[204,1379,200,1922]" captionTargetId="figure@6.[196,1390,193,1932]" captionTargetPageId="6" captionText="FIGURES 21 – 27. Larva of Choroterpes (D.) nigella (Kang &amp; Yang 1994). 21. Dorsal view of male; 22. Labrum; 23. Hypopharynx; 24. Left mandible; 25. Right mandible; 26. Maxillae; 27. Labium." httpUri="https://zenodo.org/record/580563/files/figure.png" pageId="5" pageNumber="444">Fig. 22</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="F6DE662EEF5DE96DFF18F841FC12F874" httpUri="https://zenodo.org/record/580563/files/figure.png" pageId="6" pageNumber="445" targetBox="[204,1379,200,1922]" targetPageId="6">
<paragraph id="A21E36A6EF5DE96DFF18F841FC12F874" blockId="6.[151,1436,1953,2006]" pageId="6" pageNumber="445">
<emphasis id="90D5EAB4EF5DE96DFF18F841FE8AF814" bold="true" box="[151,339,1953,1976]" pageId="6" pageNumber="445">FIGURES 21–27</emphasis>
. Larva of
<taxonomicName id="65A14D25EF5DE96DFE44F841FC1AF815" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" box="[459,963,1953,1975]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="nigella" subGenus="Dilatognathus">
<emphasis id="90D5EAB4EF5DE96DFE44F841FD91F815" box="[459,584,1953,1975]" italics="true" pageId="6" pageNumber="445">Choroterpes</emphasis>
(
<emphasis id="90D5EAB4EF5DE96DFDD2F842FDAFF815" box="[605,630,1954,1975]" italics="true" pageId="6" pageNumber="445">D.</emphasis>
)
<emphasis id="90D5EAB4EF5DE96DFD05F841FD09F815" box="[650,720,1953,1975]" italics="true" pageId="6" pageNumber="445">nigella</emphasis>
(Kang &amp; Yang 1994)
</taxonomicName>
. 21. Dorsal view of male; 22. Labrum; 23. Hypopharynx; 24. Left mandible; 25. Right mandible; 26. Maxillae; 27. Labium.
</paragraph>
</caption>
<caption id="F6DE662EEF5CE96CFF18F841FD19F874" httpUri="https://zenodo.org/record/580564/files/figure.png" pageId="7" pageNumber="446" targetBox="[222,1367,201,1921]" targetPageId="7">
<paragraph id="A21E36A6EF5CE96CFF18F841FD19F874" blockId="7.[151,1436,1953,2006]" pageId="7" pageNumber="446">
<emphasis id="90D5EAB4EF5CE96CFF18F841FE95F814" bold="true" box="[151,332,1953,1976]" pageId="7" pageNumber="446">FIGURES 28–36</emphasis>
. Larva of
<taxonomicName id="65A14D25EF5CE96CFE3AF841FC52F815" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" box="[437,907,1953,1975]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="446" phylum="Arthropoda" rank="species" species="nigella" subGenus="Dilatognathus">
<emphasis id="90D5EAB4EF5CE96CFE3AF841FDEAF815" box="[437,563,1953,1975]" italics="true" pageId="7" pageNumber="446">Choroterpes</emphasis>
(
<emphasis id="90D5EAB4EF5CE96CFDCEF842FD83F815" box="[577,602,1954,1975]" italics="true" pageId="7" pageNumber="446">D.</emphasis>
)
<emphasis id="90D5EAB4EF5CE96CFDE7F841FD76F815" box="[616,687,1953,1975]" italics="true" pageId="7" pageNumber="446">nigella</emphasis>
(Kang &amp; Yang 1994)
</taxonomicName>
. 28. Foreleg; 29. Midleg; 30. Hindleg; 31, Claw; 32. Gill II; 33.Gill III; 34. Gill IV; 35. Gill V; 36. Gill VII.
</paragraph>
</caption>
<subSubSection id="EABB652DEF53E963FF48FF77FB3FFD27" pageId="8" pageNumber="447" type="discussion">
<paragraph id="A21E36A6EF53E963FF48FF77FC55FEBF" blockId="8.[151,1436,151,645]" pageId="8" pageNumber="447">
<emphasis id="90D5EAB4EF53E963FF48FF77FEE2FF12" bold="true" box="[199,315,151,176]" pageId="8" pageNumber="447">Remarks.</emphasis>
<taxonomicName id="65A14D25EF53E963FEC9FF77FCBFFF12" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" box="[326,870,151,176]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="447" phylum="Arthropoda" rank="species" species="nigella" subGenus="Dilatognathus">
<emphasis id="90D5EAB4EF53E963FEC9FF77FE08FF12" box="[326,465,151,176]" italics="true" pageId="8" pageNumber="447">Choroterpes</emphasis>
(
<emphasis id="90D5EAB4EF53E963FE6AFF79FDD8FF12" box="[485,513,153,176]" italics="true" pageId="8" pageNumber="447">D.</emphasis>
)
<emphasis id="90D5EAB4EF53E963FD9AFF77FDBAFF12" box="[533,611,151,176]" italics="true" pageId="8" pageNumber="447">nigella</emphasis>
(
<bibRefCitation id="C6304B57EF53E963FDFAFF78FC87FF12" author="Kang" box="[629,862,152,176]" pageId="8" pageNumber="447" refString="Kang, S. C. &amp; Yang, C. T. (1994) Leptophlebiidae of Taiwan (Ephemeroptera). Journal of Taiwan Museum, 47 (1), 57 - 81." type="journal article" year="1994">Kang &amp; Yang 1994</bibRefCitation>
)
</taxonomicName>
was originally described from
<collectingCountry id="DAB67636EF53E963FB57FF77FAF3FF12" box="[1240,1322,151,176]" name="Taiwan" pageId="8" pageNumber="447">Taiwan</collectingCountry>
based on larvae and eggs.
<bibRefCitation id="C6304B57EF53E963FED9FF5CFE34FF76" author="Kluge" box="[342,493,188,213]" pageId="8" pageNumber="447" refString="Kluge, N. J. (2014) Indonesian species of Dilatognathus Kluge 2012 (Ephemeroptera, Leptophlebiidae, Choroterpes s. l.) and species-specific sexual dimorphism in development of maxilla. Zootaxa, 3786 (1), 44 - 56. http: // dx. doi. org / 10.11646 / zootaxa. 3786.1.2" type="journal article" year="2014">Kluge (2014)</bibRefCitation>
examined the specimens from
<collectingCountry id="DAB67636EF53E963FCC0FF5CFC6AFF77" box="[847,947,188,213]" name="Thailand" pageId="8" pageNumber="447">Thailand</collectingCountry>
and
<collectingRegion id="6065F844EF53E963FC64FF5CFBE5FF77" box="[1003,1084,188,213]" country="China" name="Hainan" pageId="8" pageNumber="447">Hainan</collectingRegion>
<collectingCountry id="DAB67636EF53E963FBCBFF5CFB51FF77" box="[1092,1160,188,213]" name="Iceland" pageId="8" pageNumber="447">Island</collectingCountry>
and provided additional characters. The new record of the larvae
<taxonomicName id="65A14D25EF53E963FDE6FF3FFBA1FF5A" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" box="[617,1144,223,248]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="447" phylum="Arthropoda" rank="species" species="nigella" subGenus="Dilatognathus">
<emphasis id="90D5EAB4EF53E963FDE6FF3FFD2DFF5A" box="[617,756,223,248]" italics="true" pageId="8" pageNumber="447">Choroterpes</emphasis>
(
<emphasis id="90D5EAB4EF53E963FC8AFF01FCF8FF5A" box="[773,801,225,248]" italics="true" pageId="8" pageNumber="447">D.</emphasis>
)
<emphasis id="90D5EAB4EF53E963FCBDFF3FFCA6FF5A" box="[818,895,223,248]" italics="true" pageId="8" pageNumber="447">nigella</emphasis>
(
<bibRefCitation id="C6304B57EF53E963FC1FFF00FBA9FF5A" author="Kang" box="[912,1136,224,248]" pageId="8" pageNumber="447" refString="Kang, S. C. &amp; Yang, C. T. (1994) Leptophlebiidae of Taiwan (Ephemeroptera). Journal of Taiwan Museum, 47 (1), 57 - 81." type="journal article" year="1994">Kang &amp; Yang 1994</bibRefCitation>
)
</taxonomicName>
from
<collectingRegion id="6065F844EF53E963FB30FF3FFA41FF5A" box="[1215,1432,223,248]" country="India" name="Arunachal Pradesh" pageId="8" pageNumber="447">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="6065F844EF53E963FF18FEE4FECDFEBF" box="[151,276,260,285]" country="India" name="Meghalaya" pageId="8" pageNumber="447">Meghalaya</collectingRegion>
and
<collectingRegion id="6065F844EF53E963FEC5FEE5FE0EFEBF" box="[330,471,260,285]" country="India" name="West Bengal" pageId="8" pageNumber="447">West Bengal</collectingRegion>
is extension of its distributional range.
</paragraph>
<paragraph id="A21E36A6EF53E963FF48FEC7FB3FFD27" blockId="8.[151,1436,151,645]" pageId="8" pageNumber="447">
<emphasis id="90D5EAB4EF53E963FF48FEC7FE95FEE2" bold="true" box="[199,332,295,320]" pageId="8" pageNumber="447">Discussion.</emphasis>
<bibRefCitation id="C6304B57EF53E963FED7FEC7FDD3FEE2" author="Flowers" box="[344,522,295,320]" pageId="8" pageNumber="447" refString="Flowers, R. W. (2009) Life was a beach: A panbiogeographic analysis of the cosmopolitan mayfly genus Choroterpes (Ephemeroptera: Leptophlebiidae: Atalophlebiinae). Aquatic Insects, 31 (Supplement 1), 585 - 593. https: // doi. org / 10.1080 / 01650420902909317" type="journal article" year="2009">Flowers (2009)</bibRefCitation>
made a detailed analysis of the origin and present distribution of the genus
<taxonomicName id="65A14D25EF53E963FF18FEACFEFBFEC7" box="[151,290,332,357]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="447" phylum="Arthropoda" rank="genus">
<emphasis id="90D5EAB4EF53E963FF18FEACFEFBFEC7" box="[151,290,332,357]" italics="true" pageId="8" pageNumber="447">Choroterpes</emphasis>
</taxonomicName>
but did not include the Oriental subgenera.
<bibRefCitation id="C6304B57EF53E963FCCFFEACFC05FEC6" author="Kluge" box="[832,988,332,357]" pageId="8" pageNumber="447" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera: Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge (2012)</bibRefCitation>
delineated the salient features of the subgenus
<emphasis id="90D5EAB4EF53E963FE84FE8FFE72FE2A" box="[267,427,367,392]" italics="true" pageId="8" pageNumber="447">Dilatognathus</emphasis>
and has dealt with the evolution of
<emphasis id="90D5EAB4EF53E963FCE8FE8FFBDEFE2A" box="[871,1031,367,392]" italics="true" pageId="8" pageNumber="447">Dilatognathus</emphasis>
<typeStatus id="7D1A8804EF53E963FB9CFE90FB9AFE2A" box="[1043,1091,368,392]" pageId="8" pageNumber="447">type</typeStatus>
of nymphal mouthparts in 5 phylogenetically unrelated taxa of
<taxonomicName id="65A14D25EF53E963FDA7FE74FD07FE0F" box="[552,734,404,429]" class="Insecta" family="Leptophlebiidae" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="447" phylum="Arthropoda" rank="family">Leptophlebiidae</taxonomicName>
viz.,
<emphasis id="90D5EAB4EF53E963FCADFE74FB31FE0F" box="[802,1256,404,429]" italics="true" pageId="8" pageNumber="447">
Dilatognathus,
<taxonomicName id="65A14D25EF53E963FC5BFE74FBB8FE0F" box="[980,1121,404,429]" class="Insecta" family="Leptophlebiidae" genus="Notophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="447" phylum="Arthropoda" rank="genus">Notophlebia</taxonomicName>
,
<taxonomicName id="65A14D25EF53E963FBE0FE74FB31FE0F" box="[1135,1256,404,429]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="447" phylum="Arthropoda" rank="genus">Hagenulus</taxonomicName>
</emphasis>
,
<taxonomicName id="65A14D25EF53E963FB76FE74FAB0FE0F" box="[1273,1385,404,429]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="447" phylum="Arthropoda" rank="genus">
<emphasis id="90D5EAB4EF53E963FB76FE74FAB0FE0F" box="[1273,1385,404,429]" italics="true" pageId="8" pageNumber="447">Ulmeritus</emphasis>
</taxonomicName>
and Hermanellognatha (
<bibRefCitation id="C6304B57EF53E963FEFAFE57FE26FE72" author="Kluge" box="[373,511,439,464]" pageId="8" pageNumber="447" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera: Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge 2012</bibRefCitation>
). Both the species described herein have
<emphasis id="90D5EAB4EF53E963FC41FE57FBB7FE72" box="[974,1134,439,464]" italics="true" pageId="8" pageNumber="447">Dilatognathus</emphasis>
<typeStatus id="7D1A8804EF53E963FBF9FE58FB7FFE72" box="[1142,1190,440,464]" pageId="8" pageNumber="447">type</typeStatus>
mouthparts, one with a long maxillary tusk and another without a maxillary tusk. Mouthpart architecture of both the species endorses the phylogenetic approach of
<bibRefCitation id="C6304B57EF53E963FE47FE1FFDBAFDBA" author="Kluge" box="[456,611,511,536]" pageId="8" pageNumber="447" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera: Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge (2012)</bibRefCitation>
and his view of presence or absence of maxillary tusks and different patterns of filter-feeding accessories within the subgenus
<emphasis id="90D5EAB4EF53E963FCBCFDC4FC0AFD9F" box="[819,979,548,573]" italics="true" pageId="8" pageNumber="447">Dilatognathus</emphasis>
. The occurrence of
<emphasis id="90D5EAB4EF53E963FB4DFDC4FAB5FD9F" box="[1218,1388,548,573]" italics="true" pageId="8" pageNumber="447">Dilatognathus-</emphasis>
<typeStatus id="7D1A8804EF53E963FAE3FDC5FA45FD9F" box="[1388,1436,549,573]" pageId="8" pageNumber="447">type</typeStatus>
mouthparts in different genera of the tribes, Choroterpini,
<taxonomicName id="65A14D25EF53E963FCC4FDA7FC1EFDC2" box="[843,967,583,608]" class="Insecta" family="Leptophlebiidae" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="447" phylum="Arthropoda" rank="tribe" tribe="Hagenulini">Hagenulini</taxonomicName>
and Iscini, is ultimately an instance of convergent evolution to filter feeding in rock bottomed, silt encrusted streams with moderate flow.
</paragraph>
</subSubSection>
</treatment>
</document>
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