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<document id="E6FBFBDD9F4A5FD347DDBC18D9141A30" ID-CLB-Dataset="21951" ID-DOI="10.5852/ejt.2017.370" ID-GBIF-Dataset="635f947d-f5a3-4e05-abc8-5bccbe01f637" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838462" ID-ZooBank="1163140F-FC96-4B65-A03A-37E9BE9D9218" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590087831364" checkinUser="carolina" docAuthor="Hosoishi, Shingo &amp; Ogata, Kazuo" docDate="2017" docId="99519B48565F110FFD81FEDCFBA9FB7B" docLanguage="en" docName="ejt-2017-370.pdf.imf" docOrigin="European Journal of Taxonomy 370" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Crematogaster khmerensis Hosoishi &amp; Ogata 2017, sp. nov." docType="treatment" docUuid="D2CC1731-9989-4708-9513-36E1458CE012" docUuidSource="ZooBank" docVersion="8" lastPageNumber="11" masterDocId="6568E33056571105FFC6FFD7FFACFFE5" masterDocTitle="Phylogenetic analysis and systematic position of two new species of the ant genus Crematogaster (Hymenoptera, Formicidae) from Southeast Asia" masterLastPageNumber="17" masterPageNumber="1" pageNumber="9" updateTime="1698840711414" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2413B850E326DA53DBFCF5F7FEDAE9B5">Phylogenetic analysis and systematic position of two new species of the ant genus Crematogaster (Hymenoptera, Formicidae) from Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name id="FB908E7CCABC1845A225E1BD42A32E86" type="personal">
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<mods:roleTerm id="A3F307C3B116E3704404059221E31250">Author</mods:roleTerm>
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<mods:namePart id="A2BB0822429EBDDCCDAD4BB7ADFC58CE">Hosoishi, Shingo</mods:namePart>
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<mods:affiliation id="7FB185BF0ACB787134BFC37989B32800">Institute of Tropical Agriculture, Kyushu University, 6 - 10 - 1 Hakozaki, Higashi-ku, Fukuoka, 812 - 8581 Japan. &amp; urn: lsid: zoobank. org: author: 75 C 51 C 3 B- 5 C 72 - 4229 - 8599 - 6 AE 254086690 &amp; Corresponding author: hosoishi @ gmail. com</mods:affiliation>
<mods:nameIdentifier id="33D9FD968A204174E6DCF73F6B500360" type="email">hosoishi@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A5F1CF8ED69AFB4FE77EBADDE405775C">Ogata, Kazuo</mods:namePart>
<mods:nameIdentifier id="A0E509C323A7D20A62AA55F1AFCD4BB6" type="LSID">2A3EC860-EE95-4D5F-AAEC-E87838815AFF</mods:nameIdentifier>
<mods:affiliation id="98756A174DE0C57F4EC6025079E30BEC">Institute of Tropical Agriculture, Kyushu University, 6 - 10 - 1 Hakozaki, Higashi-ku, Fukuoka, 812 - 8581 Japan. &amp; Email: kogata @ agr. kyushu-u. ac. jp &amp; urn: lsid: zoobank. org: author: 2 A 3 EC 860 - EE 95 - 4 D 5 F-AAEC-E 87838815 AFF</mods:affiliation>
<mods:nameIdentifier id="B055327F8109F36287FFB30B319B5217" type="email">kogata@agr.kyushu</mods:nameIdentifier>
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<mods:typeOfResource id="9B87BECD386F2C4801F420BD923D582A">text</mods:typeOfResource>
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<mods:title id="0321F9CA476BB4F14EECC9B718AA602E">European Journal of Taxonomy</mods:title>
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<mods:part id="83DAF41FC7BB173FB7FA9716DEABBBA8">
<mods:date id="D967072D774CC9E660C222FD208CA31B">2017</mods:date>
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<mods:number id="20EBD1F5C07BF84EE8523B9E07121A18">2017-11-24</mods:number>
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<treatment id="99519B48565F110FFD81FEDCFBA9FB7B" ID-DOI="http://doi.org/10.5281/zenodo.3852184" ID-GBIF-Taxon="164246450" ID-Zenodo-Dep="3852184" LSID="urn:lsid:zoobank.org:act:D2CC1731-9989-4708-9513-36E1458CE012" httpUri="http://treatment.plazi.org/id/99519B48565F110FFD81FEDCFBA9FB7B" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="59E279D5565F110DFD81FEDCFC41FEC3" box="[583,1005,267,294]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="11472A5E565F110DFD81FEDCFC41FEC3" blockId="8.[189,1208,267,409]" box="[583,1005,267,294]" pageId="8" pageNumber="9">
<heading id="4A0F9D32565F110DFD81FEDCFC41FEC3" box="[583,1005,267,294]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="D6F851DD565F110DFD81FEDCFC27FEC3" authority="Hosoishi &amp; Ogata, 2017" authorityName="Hosoishi &amp; Ogata" authorityYear="2017" box="[583,907,267,294]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="khmerensis" status="sp. nov.">
<emphasis id="238CF64C565F110DFD81FEDCFC27FEC3" bold="true" box="[583,907,267,294]" italics="true" pageId="8" pageNumber="9">Crematogaster khmerensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38BF4B37565F110DFC54FEDBFC41FEC3" box="[914,1005,268,294]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="59E279D5565F110DFEBDFEE6FE0AFD85" pageId="8" pageNumber="9" type="description">
<paragraph id="11472A5E565F110DFEBDFEE6FE9AFE7C" blockId="8.[189,1208,267,409]" pageId="8" pageNumber="9">
<uri id="6569265C565F110DFEBDFEE6FB14FEA9" box="[379,1208,305,332]" pageId="8" pageNumber="9">
urn:lsid:zoobank.org:act:
<uuid id="655E108B565F110DFD5CFEE6FB14FEA9" box="[666,1208,305,332]" pageId="8" pageNumber="9">D2CC1731-9989-4708-9513-36E1458CE012</uuid>
</uri>
<figureCitation id="89C336DB565F110DFD15FE8DFC8DFE92" box="[723,801,346,375]" captionStart="Fig" captionStartId="9.[189,232,1896,1922]" captionTargetBox="[223,1364,266,1856]" captionTargetId="figure@9.[223,1364,265,1856]" captionTargetPageId="9" captionText="Fig. 4. AD. Crematogaster khmerensis sp. nov., worker. A. Body in lateral view. B. Full-face view of head. C. Dorsal view of mesosoma. D. Petiole and postpetiole in dorsal view. — EH. — Crematogaster pfeifferi sp. nov., worker. E. Body in lateral view. F. Full-face view of head. G. Dorsal view of mesosoma. H. Petiole and postpetiole in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3838470" httpUri="https://zenodo.org/record/3838470/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
AD
<emphasis id="238CF64C565F110DFF7BFEA8FE9AFE7C" bold="true" box="[189,310,383,409]" pageId="8" pageNumber="9">Diagnosis</emphasis>
</paragraph>
<paragraph id="11472A5E565F110DFF7BFE67FE0AFD85" blockId="8.[189,1399,432,608]" pageId="8" pageNumber="9">
<taxonomicName id="D6F851DD565F110DFF7BFE67FE50FE2F" authority="Hosoishi &amp; Ogata, 2017" authorityName="Hosoishi &amp; Ogata" authorityYear="2017" box="[189,508,432,458]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="khmerensis" status="sp. nov.">
<emphasis id="238CF64C565F110DFF7BFE67FE50FE2F" box="[189,508,432,458]" italics="true" pageId="8" pageNumber="9">Crematogaster khmerensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38BF4B37565F110DFDCFFE67FDC5FE2F" box="[521,617,432,458]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the subgenus
<emphasis id="238CF64C565F110DFC6AFE67FAADFE2F" box="[940,1281,432,458]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="D6F851DD565F110DFC6AFE67FBF7FE2F" authorityName="Lund" authorityYear="1831" box="[940,1115,432,458]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" sensu="stricto">Crematogaster</taxonomicName>
<taxonomicNameLabel id="38BF4B37565F110DFBAEFE66FAADFE2F" box="[1128,1281,433,458]" pageId="8" pageNumber="9" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
(
<bibRefCitation id="756957AF565F110DFAD0FE67FEA7FE15" author="Blaimer B. B." pageId="8" pageNumber="9" pagination="47 - 67" refId="ref6997" refString="Blaimer B. B. 2012 b. A subgeneric revision of Crematogaster and discussion of regional species-groups (Hymenoptera: Formicidae). Zootaxa 3482: 47 - 67." type="journal article" year="2012">Blaimer 2012b</bibRefCitation>
). This small-sized species of
<taxonomicName id="D6F851DD565F110DFDB5FE01FC8EFE15" authorityName="Lund" authorityYear="1831" box="[627,802,470,496]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="238CF64C565F110DFDB5FE01FC8EFE15" box="[627,802,470,496]" italics="true" pageId="8" pageNumber="9">Crematogaster</emphasis>
</taxonomicName>
ants (HW 0.60.66; WL 0.630.69) is similar to
<taxonomicName id="D6F851DD565F110DFF7BFE2CFE96FDF0" authority="Hosoishi &amp; Ogata, 2017" authorityName="Hosoishi &amp; Ogata" authorityYear="2017" box="[189,314,507,533]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pfeifferi" status="sp. nov.">
<emphasis id="238CF64C565F110DFF7BFE2CFF7DFDF0" box="[189,209,507,533]" italics="true" pageId="8" pageNumber="9">C</emphasis>
.
<emphasis id="238CF64C565F110DFF1BFE2CFE96FDF0" box="[221,314,507,533]" italics="true" pageId="8" pageNumber="9">pfeifferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38BF4B37565F110DFEF9FE2CFE36FDF0" box="[319,410,507,533]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, but can be distinguished by the propodeal spiracles touching metapleural gland bulla, short propodeal spines (PSL 0.080.09), petiole broader anteriorly than posteriorly and undeveloped subpetiolar process.
</paragraph>
</subSubSection>
<subSubSection id="59E279D5565F110DFF7BFD5DFB68FD31" box="[189,1220,650,725]" pageId="8" pageNumber="9" type="etymology">
<paragraph id="11472A5E565F110DFF7BFD5DFEE8FD41" blockId="8.[189,324,650,676]" box="[189,324,650,676]" pageId="8" pageNumber="9">
<heading id="4A0F9D32565F110DFF7BFD5DFEE8FD41" bold="true" box="[189,324,650,676]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="238CF64C565F110DFF7BFD5DFEE8FD41" bold="true" box="[189,324,650,676]" pageId="8" pageNumber="9">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="11472A5E565F110DFF7BFD6DFB68FD31" blockId="8.[189,1220,698,725]" box="[189,1220,698,725]" pageId="8" pageNumber="9">
The specific name refers to the former kingdom name of
<collectingCountry id="69EF6ACE565F110DFC9DFD6DFC7BFD31" box="[859,983,698,724]" name="Cambodia" pageId="8" pageNumber="9">Cambodia</collectingCountry>
, the Khmer Empire.
</paragraph>
</subSubSection>
<subSubSection id="59E279D5565F110DFF7BFD29FEB2FBAD" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="11472A5E565F110DFF7BFD29FE2DFCFD" blockId="8.[189,385,766,792]" box="[189,385,766,792]" pageId="8" pageNumber="9">
<heading id="4A0F9D32565F110DFF7BFD29FE2DFCFD" bold="true" box="[189,385,766,792]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="238CF64C565F110DFF7BFD29FE2DFCFD" bold="true" box="[189,385,766,792]" pageId="8" pageNumber="9">Type specimens</emphasis>
</heading>
</paragraph>
<materialsCitation id="A1902003565F110DFF2CFCF8FD38FC71" ID-GBIF-Occurrence="2621728301" collectingDate="2010-11-24" collectionCode="THNHM" collectorName="S. Hosoishi" country="Cambodia" location="Kampong Thom Province" pageId="8" pageNumber="9" specimenCode="KUMANT040" specimenCount="1" stateProvince="Kampong Thom Province" typeStatus="holotype">
<paragraph id="11472A5E565F110DFF2CFCF8FEF0FCAC" blockId="8.[189,1398,815,916]" box="[234,348,815,841]" pageId="8" pageNumber="9">
<emphasis id="238CF64C565F110DFF2CFCF8FEF0FCAC" bold="true" box="[234,348,815,841]" pageId="8" pageNumber="9">
<typeStatus id="CE4394FC565F110DFF2CFCF8FEF0FCAC" box="[234,348,815,841]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="11472A5E565F110DFF7BFC83FD38FC71" blockId="8.[189,1398,815,916]" pageId="8" pageNumber="9">
<collectingCountry id="69EF6ACE565F110DFF7BFC83FEF1FC8B" box="[189,349,852,878]" name="Cambodia" pageId="8" pageNumber="9">CAMBODIA</collectingCountry>
:
<specimenCount id="07FEE1D7565F110DFEADFC83FE6CFC8B" box="[363,448,852,878]" pageId="8" pageNumber="9" type="worker">worker</specimenCount>
,
<collectingRegion id="D33CE4BC565F110DFE0DFC82FD50FC8B" box="[459,764,852,879]" country="Cambodia" name="Kampong Thom" pageId="8" pageNumber="9">Kampong Thom Province</collectingRegion>
,
<date id="65460C9E565F110DFCC1FC83FC0BFC8B" box="[775,935,852,879]" pageId="8" pageNumber="9" value="2010-11-24">
<collectingDate id="7502F576565F110DFCC1FC83FC0BFC8B" box="[775,935,852,879]" pageId="8" pageNumber="9" value="2010-11-24">24 Nov. 2010</collectingDate>
</date>
,
<collectorName id="BC0D4F88565F110DFC77FC83FB9BFC8B" box="[945,1079,852,878]" pageId="8" pageNumber="9">S. Hosoishi</collectorName>
leg., (TUS arboreal SF3-1) (
<specimenCode id="415E8225565F110DFF03FCADFED4FC71" box="[197,376,890,916]" pageId="8" pageNumber="9">KUMANT040</specimenCode>
: deposited at
<collectionCode id="77E9B29B565F110DFDD1FCADFD24FC71" box="[535,648,890,916]" pageId="8" pageNumber="9">THNHM</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="11472A5E565F110DFF2CFC69FECBFC3D" blockId="8.[189,1399,958,1096]" box="[234,359,958,984]" pageId="8" pageNumber="9">
<emphasis id="238CF64C565F110DFF2CFC69FECBFC3D" bold="true" box="[234,359,958,984]" pageId="8" pageNumber="9">
<typeStatus id="CE4394FC565F110DFF2CFC69FECBFC3D" box="[234,359,958,984]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="11472A5E565F110DFF7BFC34FEB2FBAD" blockId="8.[189,1399,958,1096]" pageId="8" pageNumber="9">
<collectingCountry id="69EF6ACE565F110DFF7BFC34FEF1FC18" box="[189,349,995,1021]" name="Cambodia" pageId="8" pageNumber="9">CAMBODIA</collectingCountry>
:
<specimenCount id="07FEE1D7565F110DFEAAFC34FE4EFC18" box="[364,482,995,1021]" pageId="8" pageNumber="9" type="worker">3 workers</specimenCount>
, same locality data as for
<typeStatus id="CE4394FC565F110DFCC9FC34FCDAFC18" box="[783,886,995,1021]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
(TUS arboreal SF3-3, TUS arboreal SF3-6) (KUMANT041: deposited at
<collectionCode id="77E9B29B565F110DFDDEFBDEFDD8FBC6" box="[536,628,1033,1059]" pageId="8" pageNumber="9">BMNH</collectionCode>
; KUMANT042: deposited at
<collectionCode id="77E9B29B565F110DFC17FBDEFB8DFBC6" box="[977,1057,1033,1059]" pageId="8" pageNumber="9">CASC</collectionCode>
; KUMANT043: deposited at
<collectionCode id="77E9B29B565F110DFF7BFBF9FEBEFBAD" box="[189,274,1070,1096]" country="Japan" httpUri="http://grbio.org/cool/txbq-e9hf" name="Kyushu University Entomology Collection" pageId="8" pageNumber="9">KUEC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="59E279D5565F110FFF7BFBA5FD4BFBC2" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="11472A5E565F110DFF7BFBA5FDA4FB6E" blockId="8.[189,520,1137,1164]" box="[189,520,1137,1164]" pageId="8" pageNumber="9">
<heading id="4A0F9D32565F110DFF7BFBA5FDA4FB6E" bold="true" box="[189,520,1137,1164]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="238CF64C565F110DFF7BFBA5FDA4FB6E" bold="true" box="[189,520,1137,1164]" pageId="8" pageNumber="9">Measurements and indices</emphasis>
</heading>
</paragraph>
<paragraph id="11472A5E565F110DFF7BFB74FE93FACB" blockId="8.[189,1398,1186,1326]" pageId="8" pageNumber="9">
HW 0.600.66; HL 0.600.67; CI 97102; SL 0.430.50; SI 6976; EL 0.140.15; PW 0.350.40; WL 0.630.69; PSL 0.080.09; PtL 0.220.23; PtW 0.200.25; PtH 0.120.14; PpL 0.120.14; PpW 0.200.23; PtHI 5264; PtWI 87114; PpWI 154167; WI 92100 (
<typeStatus id="CE4394FC565F110DFC08FB3AFB9BFAE2" box="[974,1079,1261,1287]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
and three
<typeStatus id="CE4394FC565F110DFB6BFB39FABEFAED" box="[1197,1298,1262,1288]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
workers measured).
</paragraph>
<paragraph id="11472A5E565F110DFF7BFA81FEE2FA95" blockId="8.[189,334,1366,1392]" box="[189,334,1366,1392]" pageId="8" pageNumber="9">
<heading id="4A0F9D32565F110DFF7BFA81FEE2FA95" bold="true" box="[189,334,1366,1392]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="238CF64C565F110DFF7BFA81FEE2FA95" bold="true" box="[189,334,1366,1392]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="11472A5E565F110DFF2CFA5FFEE0FA47" blockId="8.[189,612,1416,1479]" box="[234,332,1416,1442]" pageId="8" pageNumber="9">
<emphasis id="238CF64C565F110DFF2CFA5FFEE0FA47" bold="true" box="[234,332,1416,1442]" pageId="8" pageNumber="9">Worker</emphasis>
</paragraph>
<paragraph id="11472A5E565F110DFF7BFA7AFDC8FA22" blockId="8.[189,612,1416,1479]" box="[189,612,1453,1479]" pageId="8" pageNumber="9">Workers presumably monomorphic.</paragraph>
<paragraph id="11472A5E565F110DFF7BFA26FB3BF923" blockId="8.[189,1399,1521,1735]" pageId="8" pageNumber="9">Head subquadrate in full-face view, with rounded posterior corners and subparallel sides. Occipital carinae developed. Mandible with four teeth, apical tooth large. Anterior margin of clypeus convex; anterolateral margins of clypeus protruded anteriorly; posterior margin of clypeus rounded between frontal lobes. Frontal carinae almost parallel. Antennae 11-segmented; antennal club 3-segmented. Scape reaching posterior corner of head. Basal flagellar segment (antennal segment III) as broad as long. Compound eyes slightly projecting beyond lateral margins of head in full face view.</paragraph>
<paragraph id="11472A5E565F110DFF7BF927FB80F80E" blockId="8.[189,1399,1776,2027]" pageId="8" pageNumber="9">Pronotal shoulders angulate, but without distinct rugulae laterally. Mesonotal dorsum convex in lateral view. Pronotum and mesonotum not forming same dorsal outline in lateral view. Mesothoracic spiracle forms large pit concealed by posterior pronotal lobe. Metapleural gland opening slit-shaped. Propodeal spiracle large and circular, situated at posterolateral corners, touching metapleural gland bulla. Metanotal groove straight in dorsal view, deep and forming concave region between mesonotum and propodeum. In dorsal view, feeble longitudinal rugulae connecting between mesonotum and propodeum; the boundary distinct. Propodeal spines developed and directed vertically in lateral view.</paragraph>
<caption id="45877AD6565E110CFF7BF8BFFD0DF809" ID-DOI="http://doi.org/10.5281/zenodo.3838470" ID-Zenodo-Dep="3838470" httpUri="https://zenodo.org/record/3838470/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1896,1922]" targetBox="[223,1364,266,1856]" targetPageId="9">
<paragraph id="11472A5E565E110CFF7BF8BFFD0DF809" blockId="9.[189,1399,1896,2028]" pageId="9" pageNumber="10">
<emphasis id="238CF64C565E110CFF7BF8BFFEE0F867" bold="true" box="[189,332,1896,1922]" pageId="9" pageNumber="10">Fig. 4. AD</emphasis>
.
<taxonomicName id="D6F851DD565E110CFE9DF8BEFD39F867" authority="Hosoishi &amp; Ogata, 2017" authorityName="Hosoishi &amp; Ogata" authorityYear="2017" box="[347,661,1896,1923]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="khmerensis" status="sp. nov.">
<emphasis id="238CF64C565E110CFE9DF8BEFD39F867" box="[347,661,1896,1923]" italics="true" pageId="9" pageNumber="10">Crematogaster khmerensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38BF4B37565E110CFD5AF8BEFD56F866" box="[668,762,1897,1923]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, worker.
<emphasis id="238CF64C565E110CFCA1F8BFFCD0F867" bold="true" box="[871,892,1896,1922]" pageId="9" pageNumber="10">A</emphasis>
. Body in lateral view.
<emphasis id="238CF64C565E110CFB4FF8BEFB31F866" bold="true" box="[1161,1181,1897,1923]" pageId="9" pageNumber="10">B</emphasis>
. Full-face view of head.
<emphasis id="238CF64C565E110CFEC7F85CFEBAF840" bold="true" box="[257,278,1931,1957]" pageId="9" pageNumber="10">C</emphasis>
. Dorsal view of mesosoma.
<emphasis id="238CF64C565E110CFD9BF85BFDDEF843" bold="true" box="[605,626,1932,1958]" pageId="9" pageNumber="10">D</emphasis>
. Petiole and postpetiole in dorsal view. —
<emphasis id="238CF64C565E110CFB98F85BFB3BF843" bold="true" box="[1118,1175,1932,1958]" pageId="9" pageNumber="10">EH</emphasis>
. —
<taxonomicName id="D6F851DD565E110CFB0EF85BFEB6F82C" authority="Hosoishi &amp; Ogata, 2017" authorityName="Hosoishi &amp; Ogata" authorityYear="2017" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pfeifferi" status="sp. nov.">
<emphasis id="238CF64C565E110CFB0EF85BFEB6F82C" italics="true" pageId="9" pageNumber="10">Crematogaster pfeifferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38BF4B37565E110CFED9F878FED6F82C" box="[287,378,1967,1993]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, worker.
<emphasis id="238CF64C565E110CFE24F878FE5AF82C" bold="true" box="[482,502,1967,1993]" pageId="9" pageNumber="10">E</emphasis>
. Body in lateral view. F. Full-face view of head.
<emphasis id="238CF64C565E110CFBE6F878FB9BF82C" bold="true" box="[1056,1079,1967,1993]" pageId="9" pageNumber="10">G</emphasis>
. Dorsal view of mesosoma.
<emphasis id="238CF64C565E110CFF7BF805FF78F809" bold="true" box="[189,212,2002,2028]" pageId="9" pageNumber="10">H</emphasis>
. Petiole and postpetiole in dorsal view.
</paragraph>
</caption>
<paragraph id="11472A5E565D110FFF7BFEDCFC20FE7D" blockId="10.[189,1398,267,408]" pageId="10" pageNumber="11">Petiole scoop shaped, but flattened broader anteriorly, longer than broad; spiracle situated anteriorly midway between dorsal and ventral margin of petiole in lateral view, directed laterally. Subpetiolar process developed as a short lamellate denticle. Postpetiole without longitudinal median sulcus, but bilobed behind; spiracle situated anteriorly on lateral surface.</paragraph>
<paragraph id="11472A5E565D110FFF7BFE15FCA6FD7E" blockId="10.[189,1399,450,667]" pageId="10" pageNumber="11">Dorsal surface of head mostly smooth or weakly shagreened. Clypeus mostly smooth and shining. Pronotal collar areolately sculptured. Promesonotum weakly punctate. Lateral surface of pronotum smooth and shining. Mesopleuron mostly smooth and shining, but sculptured on surrounding area. Anterodorsal surface of propodeum weakly punctate; posteior half smooth and shining. Lateral surface of propodeum smooth and shining. Dorsal and lateral surfaces of petiole smooth and shining. Dorsal and lateral surfaces of postpetiole smooth and shining.</paragraph>
<paragraph id="11472A5E565D110FFF7BFD11FC9EFC9D" blockId="10.[189,1398,710,889]" pageId="10" pageNumber="11">Standing pilosity sparse. Dorsal face of head with appressed setae; one pair of erect setae on frontal lobes. Clypeus with one pair of erect setae. Anterior clypeal margin with two pairs of long setae mixed with short setae laterally. Scape with decumbent setae. Mesontum with appressed setae sparsely. Petiole with one pair of suberect setae posteriorly. Postpetiole with one pair of suberect setae posteriorly. Fourth abdominal tergite with short appressed setae sparsely.</paragraph>
<paragraph id="11472A5E565D110FFF7BFC74FE36FC58" blockId="10.[189,410,931,957]" box="[189,410,931,957]" pageId="10" pageNumber="11">Body color brown.</paragraph>
<paragraph id="11472A5E565D110FFF2CFC30FE1AFBE4" blockId="10.[189,743,999,1064]" box="[234,438,999,1025]" pageId="10" pageNumber="11">
<emphasis id="238CF64C565D110FFF2CFC30FE1AFBE4" bold="true" box="[234,438,999,1025]" pageId="10" pageNumber="11">Queen and male</emphasis>
</paragraph>
<paragraph id="11472A5E565D110FFF7BFBDAFD4BFBC2" blockId="10.[189,743,999,1064]" box="[189,743,1037,1064]" pageId="10" pageNumber="11">Queens and males of this species are unknown.</paragraph>
</subSubSection>
<subSubSection id="59E279D5565D110FFF7BFB85FBA9FB7B" box="[189,1029,1106,1182]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="11472A5E565D110FFF7BFB85FEFAFB89" blockId="10.[189,342,1106,1132]" box="[189,342,1106,1132]" pageId="10" pageNumber="11">
<heading id="4A0F9D32565D110FFF7BFB85FEFAFB89" bold="true" box="[189,342,1106,1132]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="238CF64C565D110FFF7BFB85FEFAFB89" bold="true" box="[189,342,1106,1132]" pageId="10" pageNumber="11">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="11472A5E565D110FFF7BFB54FBA9FB7B" blockId="10.[189,1029,1155,1182]" box="[189,1029,1155,1182]" pageId="10" pageNumber="11">
This species is known only from the
<typeStatus id="CE4394FC565D110FFDA8FB53FD0DFB7B" box="[622,673,1156,1182]" pageId="10" pageNumber="11">type</typeStatus>
locality in
<collectingCountry id="69EF6ACE565D110FFCE3FB54FC32FB78" box="[805,926,1155,1181]" name="Cambodia" pageId="10" pageNumber="11">Cambodia</collectingCountry>
(
<figureCitation id="89C336DB565D110FFC68FB54FC59FB7B" box="[942,1013,1155,1182]" captionStart="Fig" captionStartId="11.[189,232,1895,1921]" captionTargetBox="[189,1399,850,1855]" captionTargetId="figure@11.[189,1399,850,1855]" captionTargetPageId="11" captionText="Fig. 5. Phylogeny and distribution of the Crematogaster borneensis-group, C. khmerensisi sp. nov., C. fruhstorferi, C. ssAUS5, C. mjobergi and the C. tetracantha-group. Closed circle indicates C. khmerensis sp. nov., closed square indicates C. pfeifferi sp. nov. It is noted that C. pfeifferi sp. nov. is not represented in the phylogeny." figureDoi="http://doi.org/10.5281/zenodo.3838472" httpUri="https://zenodo.org/record/3838472/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>