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<document id="5185F3A7479C8A1519254A42C2FA56EF" ID-DOI="10.11646/zootaxa.5514.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13849314" ID-ZooBank="F2D3EAD5-33F3-4D4C-A257-D942E1AEB6CA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727440832015" checkinUser="plazi" docAuthor="Macgowan, Iain &amp; Tschirnhaus, Michael Von" docDate="2024" docId="03F4013A8B5C3769FF03F8D9FACD2882" docLanguage="en" docName="zootaxa.5514.1.1.pdf" docOrigin="Zootaxa 5514 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5514.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Silba akloa Macgowan &amp; Tschirnhaus, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="5" masterDocId="FFCD79428B5E376DFF94FFA3FFEF290B" masterDocTitle="Records of Lonchaeidae (Diptera, Cyclorrhapha) from Togo and other African countries, with descriptions of new species" masterLastPageNumber="27" masterPageNumber="1" pageNumber="3" updateTime="1727442337463" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DAD55436A32ECEE81D1A2C0FE1A55F9C">Records of Lonchaeidae (Diptera, Cyclorrhapha) from Togo and other African countries, with descriptions of new species</mods:title>
</mods:titleInfo>
<mods:name id="733053D0259325FE8CD282648E0A9C19" type="personal">
<mods:role id="5448BE165C7BBBD744CDE6401CBFC00B">
<mods:roleTerm id="EBB1AC9FAFB41BCB73EB49625A776BDF">Author</mods:roleTerm>
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<mods:namePart id="2FFE8E08CCFE284D46313E46EF1E902D">Macgowan, Iain</mods:namePart>
<mods:affiliation id="6715775FFCEFB4B68A9F1423C943C35E">Research Associate, National Museums of Scotland, Collection Centre, 242 West Granton Road, Edinburgh, Scotland. EH 5 1 JA</mods:affiliation>
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<mods:namePart id="4DFB7B22ABDECED32295616DB2930C82">Tschirnhaus, Michael Von</mods:namePart>
<mods:affiliation id="B97C8327F7FE4CB9806CC137D910590A">Fakultät Biologie, Universität Bielefeld, Postfach 100 131, 33501 Bielefeld, Germany</mods:affiliation>
</mods:name>
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<mods:title id="A413E31C9197CF1FEF53495024214E32">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4B0B5541634B2DA3CE96570BBAEC2F76">
<mods:date id="970E2D11961511BC39BC3B167CAA5B78">2024</mods:date>
<mods:detail id="564520E674F4D424DDD6760B8BB1ACF7" type="pubDate">
<mods:number id="4081A9E65765423FF928CF35F05B06C0">2024-09-25</mods:number>
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<mods:number id="C9237C1292B7B7A533B8A054A34EFD6A">5514</mods:number>
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<mods:number id="9A37573AF6E94DDDBF132A59A106917E">1</mods:number>
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<mods:url id="495D0A109FACCABE5DE98036B0DE2625">http://dx.doi.org/10.11646/zootaxa.5514.1.1</mods:url>
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<mods:classification id="FB540EC5C79767A1D7D77746DF038851">journal article</mods:classification>
<mods:identifier id="C85DFA91BAE53881AFD81993AE119F11" type="DOI">10.11646/zootaxa.5514.1.1</mods:identifier>
<mods:identifier id="ACFCB02DBAC2C4A0A52BE2EAB5CBDC81" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F4013A8B5C3769FF03F8D9FACD2882" ID-DOI="http://doi.org/10.5281/zenodo.13849392" ID-Zenodo-Dep="13849392" LSID="urn:lsid:plazi:treatment:03F4013A8B5C3769FF03F8D9FACD2882" httpUri="http://treatment.plazi.org/id/03F4013A8B5C3769FF03F8D9FACD2882" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<subSubSection id="C347E3A78B5C376FFF03F8D9FDDF2E9F" box="[151,560,1914,1941]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BE2B02C8B5C376FFF03F8D9FDDF2E9F" blockId="2.[151,560,1914,1941]" box="[151,560,1914,1941]" pageId="2" pageNumber="3">
<heading id="D0AA07408B5C376FFF03F8D9FDDF2E9F" bold="true" box="[151,560,1914,1941]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B9296C3E8B5C376FFF03F8D9FDDF2E9F" bold="true" box="[151,560,1914,1941]" pageId="2" pageNumber="3">
<taxonomicName id="4C5DCBAF8B5C376FFF03F8D9FEF02E9E" authorityName="Macgowan &amp; Tschirnhaus" authorityYear="2024" box="[151,287,1914,1941]" class="Insecta" family="Lonchaeidae" genus="Silba" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="akloa" status="sp. nov.">
<emphasis id="B9296C3E8B5C376FFF03F8D9FEF02E9E" bold="true" box="[151,287,1914,1941]" italics="true" pageId="2" pageNumber="3">Silba akloa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21AD1458B5C376FFEB3F8D8FE6B2E9E" box="[295,388,1915,1941]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1366ACA98B5C376FFE07F8D9FDC62E9F" box="[403,553,1914,1940]" captionStart="FIGURES 3–5" captionStartId="3.[152,270,1157,1181]" captionTargetBox="[155,1431,386,1129]" captionTargetId="figure-117@3.[151,1436,381,1134]" captionTargetPageId="3" captionText="FIGURES 3–5. Silba akola sp. nov. 3. Epandrium and associated features, lateral view. 4. Same, ventral view, Prg = pregonite, Pog = postgonite, Ph = phallus, E = epandrium, Sur = surstylus, Pren = prensisetae, Cer = cerci. 5. Phallus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849318" httpUri="https://zenodo.org/record/13849318/files/figure.png" pageId="2" pageNumber="3">Figures 3–5</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C347E3A78B5C376EFF03F81FFE702E99" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
<paragraph id="8BE2B02C8B5C376EFF03F81FFF34286E" blockId="2.[151,1436,1980,2043]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Description:
<typeStatus id="54E60E8E8B5C376FFEA5F81FFE762EDD" box="[305,409,1980,2006]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
male:
<emphasis id="B9296C3E8B5C376FFE63F81FFDDB2EDD" box="[503,564,1980,2006]" italics="true" pageId="2" pageNumber="3">Head</emphasis>
: Eye bare. Frons, sub-shining black, dulled by greyish pollinosity and microsculpture. Ratio of frons width at point just above lunule to eye width 0.4:1 Orbital plate bare apart from the orbital seta, sub-shining black, dulled by microsculpture. Frontal and interfrontal setulae short, at most 0.2x length of orbital seta, longer setulae present just on margin above lunule. Lunule, ground colour black, covered in greyish pollinosity, parafacial slightly grey pollinose as is the face. Antenna; postpedicel black, long, reaching down below mouth margin, length to depth ratio approximately 2.7:1. Arista yellowish basally, width of plumosity at maximum approximately 0.75x depth of postpedicel. Anterior genal setulae in a single row of 5–6 along mouth margin. Palpus black.
</paragraph>
<caption id="DF22E0A48B5D376EFF0CFB26FABA2DC9" ID-DOI="http://doi.org/10.5281/zenodo.13849318" ID-Zenodo-Dep="13849318" httpUri="https://zenodo.org/record/13849318/files/figure.png" pageId="3" pageNumber="4" startId="3.[152,270,1157,1181]" targetBox="[155,1431,386,1129]" targetPageId="3" targetType="figure">
<paragraph id="8BE2B02C8B5D376EFF0CFB26FABA2DC9" blockId="3.[151,1437,1157,1218]" pageId="3" pageNumber="4">
<emphasis id="B9296C3E8B5D376EFF0CFB26FEAF2D95" bold="true" box="[152,320,1157,1182]" pageId="3" pageNumber="4">FIGURES 3–5.</emphasis>
<taxonomicName id="4C5DCBAF8B5D376EFED2FB25FE582D95" authorityName="Macgowan &amp; Tschirnhaus" authorityYear="2024" box="[326,439,1158,1182]" class="Insecta" family="Lonchaeidae" genus="Silba" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="akola" status="sp. nov.">
<emphasis id="B9296C3E8B5D376EFED2FB25FE582D95" box="[326,439,1158,1182]" italics="true" pageId="3" pageNumber="4">Silba akola</emphasis>
</taxonomicName>
<emphasis id="B9296C3E8B5D376EFE29FB25FDE22D95" bold="true" box="[445,525,1158,1182]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="A21AD1458B5D376EFE29FB25FDE22D95" box="[445,525,1158,1182]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
3. Epandrium and associated features, lateral view. 4. Same, ventral view, Prg = pregonite, Pog = postgonite, Ph = phallus, E = epandrium, Sur = surstylus, Pren = prensisetae, Cer = cerci. 5. Phallus, lateral view.
</paragraph>
</caption>
<paragraph id="8BE2B02C8B5D376EFF53FAB7FBA32F21" blockId="3.[151,1437,1300,2011]" pageId="3" pageNumber="4">
<emphasis id="B9296C3E8B5D376EFF53FAB7FEF72C25" box="[199,280,1300,1326]" italics="true" pageId="3" pageNumber="4">Thorax</emphasis>
: Mesonotum sub-shining blue-black, with a covering of short setulae, these approximately 0.3x length of the orbital seta. Anepisternum with a vertical row of three posterior and three anterior setae, with a scattering of setulae anteriorly. One seta on proepimeron and one on proepisternum. Katepisternum with two setae near the dorsal margin, the posterior slightly stronger and positioned slightly more dorsal than the anterior, a scattering of short setulae located along the anterior margin of this sclerite otherwise the surface is bare. Scutellum with a covering of greyish pollinosity, on margin with two setulae between apical and lateral setae on each side, one broken setula evident between apical setae. Calypter whitish with a rather dense white fringe of uniform length. Wing slightly fumose with pale brown veins, wing length
<quantity id="4CA51DC98B5D376EFD17F9B3FD3A2F21" box="[643,725,1552,1578]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="4" unit="mm" value="3.6">3.6mm</quantity>
. Haltere dark. Legs entirely dark.
</paragraph>
<paragraph id="8BE2B02C8B5D376EFF53F997FF1F2E65" blockId="3.[151,1437,1300,2011]" pageId="3" pageNumber="4">
<emphasis id="B9296C3E8B5D376EFF53F997FE922F45" box="[199,381,1588,1614]" italics="true" pageId="3" pageNumber="4">Male terminalia</emphasis>
: In lateral view surstylus extending posteriorly as a large rounded lobe beyond the shell of the epandrium, posterior prensisetae visible at the base of the cerci. Cercus 0.4x the height of the epandrium, bearing long, strong setae along posterior and ventral margins. Epandrium approximately 1.2x higher than wide with long setae along posterior and ventral margins. In ventral view inner surface of surstylus covered in microsculpture, five large peg-like prensisetae located on posterior margin of surstylus, the posterior two located in a transverse alignment on a setulose posterior projection. Ventral margin of surstylus bearing 4–5 very strong setae posteriorly, these becoming weaker anteriorly. Postgonites large and setulose, bifurcate apically. Phallus relatively simple, distiphallus a long slightly curved tube, partly separated from the deeper basiphallus which has a slightly scaled surface.
</paragraph>
<paragraph id="8BE2B02C8B5D376EFF53F8DBFE702E99" blockId="3.[151,1437,1300,2011]" box="[199,415,1912,1938]" pageId="3" pageNumber="4">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection id="C347E3A78B5D3769FF53F83FFBE429F2" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BE2B02C8B5D3769FF53F83FFBE429F2" blockId="3.[151,1437,1300,2011]" lastBlockId="4.[151,1437,151,393]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="B9296C3E8B5D376EFF53F83FFED52EBD" bold="true" box="[199,314,1948,1974]" pageId="3" pageNumber="4">Diagnosis</emphasis>
. This species with the posterior prensisetae situated on a setulose posterior projection of the surstylus is closely related to
<taxonomicName id="4C5DCBAF8B5D376EFE13F863FCEE2ED0" authority="MacGowan, 2005" authorityName="MacGowan" authorityYear="2005" box="[391,769,1984,2011]" class="Insecta" family="Lonchaeidae" genus="Silba" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="figurata">
<emphasis id="B9296C3E8B5D376EFE13F863FDC92ED1" box="[391,550,1984,2010]" italics="true" pageId="3" pageNumber="4">Silba figurata</emphasis>
<bibRefCitation id="EFCCCDDD8B5D376EFDA5F863FCEE2ED0" author="MacGowan, I." box="[561,769,1984,2011]" pageId="3" pageNumber="4" pagination="1 - 23" refId="ref13076" refString="MacGowan, I. (2005) New species of Lonchaeidae (Diptera: Schizophora) from central and southern Africa. Zootaxa, 967 (1), 1 - 23. https: // doi. org / 10.11646 / zootaxa. 967.1.1" type="journal article" year="2005">MacGowan, 2005</bibRefCitation>
</taxonomicName>
. However, in ventral view the posterior prensisetae of
<taxonomicName id="4C5DCBAF8B5D3769FA13F863FF1E29BA" authorityName="MacGowan" authorityYear="2005" class="Insecta" family="Lonchaeidae" genus="Silba" kingdom="Animalia" lastPageId="4" lastPageNumber="5" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="figurata">
<emphasis id="B9296C3E8B5D3769FA13F863FF1E29BA" italics="true" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">S. figurata</emphasis>
</taxonomicName>
are in a posterior—anterior row whilst in
<taxonomicName id="4C5DCBAF8B5A3769FD49FF34FCD629BA" authorityName="Macgowan &amp; Tschirnhaus" authorityYear="2024" box="[733,825,151,177]" class="Insecta" family="Lonchaeidae" genus="Silba" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="akola">
<emphasis id="B9296C3E8B5A3769FD49FF34FCD629BA" box="[733,825,151,177]" italics="true" pageId="4" pageNumber="5">S. akola</emphasis>
</taxonomicName>
they are transversely arranged. In addition, the inner surface of the surstylus of
<taxonomicName id="4C5DCBAF8B5A3769FE56FF18FDF229DE" authorityName="Macgowan &amp; Tschirnhaus" authorityYear="2024" box="[450,541,187,213]" class="Insecta" family="Lonchaeidae" genus="Silba" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="akloa">
<emphasis id="B9296C3E8B5A3769FE56FF18FDF229DE" box="[450,541,187,213]" italics="true" pageId="4" pageNumber="5">S. akloa</emphasis>
</taxonomicName>
is more extensively covered in micro-sculpture and the phallus, with its deeper basiphallus, is only partly separated from the distiphallus and without spicules.
</paragraph>
</subSubSection>
<subSubSection id="C347E3A78B5A3769FF53FEA0FE25286E" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BE2B02C8B5A3769FF53FEA0FE25286E" blockId="4.[151,1437,151,393]" pageId="4" pageNumber="5">
<emphasis id="B9296C3E8B5A3769FF53FEA0FE812817" bold="true" box="[199,366,258,285]" pageId="4" pageNumber="5">Type material</emphasis>
.
<materialsCitation id="3B35BA718B5A3769FEE8FEA0FE25286E" collectionCode="SDEI" collectorName="M. von Tschirnhaus. Specimen" country="Togo" elevation="8" latitude="7.512" location="Domi creek" longLatPrecision="78" longitude="0.61" municipality="Akloa" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" stateProvince="Plateaux" typeStatus="holotype">
<typeStatus id="54E60E8E8B5A3769FEE8FEA0FE062816" box="[380,489,259,285]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D5B7BA58B5A3769FE67FEA0FDF62816" box="[499,537,259,285]" count="1" pageId="4" pageNumber="5" type="male">1♂</specimenCount>
.
<emphasis id="B9296C3E8B5A3769FDB1FEA1FD972817" bold="true" box="[549,632,258,284]" pageId="4" pageNumber="5">
<collectingCountry id="F34AF0BC8B5A3769FDB1FEA1FD972817" box="[549,632,258,284]" name="Togo" pageId="4" pageNumber="5">TOGO</collectingCountry>
</emphasis>
:
<collectingRegion id="49997ECE8B5A3769FD13FEA0FC802816" box="[647,879,259,285]" country="Togo" name="Plateaux" pageId="4" pageNumber="5">Région des Plateaux</collectingRegion>
,
<collectingMunicipality id="6B862A568B5A3769FCEDFEA0FC2F2816" box="[889,960,259,285]" pageId="4" pageNumber="5">Akloa</collectingMunicipality>
, at
<location id="8E82E6F78B5A3769FC73FEA0FB842816" LSID="urn:lsid:plazi:treatment:03F4013A8B5C3769FF03F8D9FACD2882:8E82E6F78B5A3769FC73FEA0FB842816" box="[999,1131,259,285]" country="Togo" latitude="7.512" longLatPrecision="78" longitude="0.61" municipality="Akloa" name="Domi creek" pageId="4" pageNumber="5" stateProvince="Plateaux">Domi creek</location>
coming from the Cascades d’Akloa.
<geoCoordinate id="EE69D6EB8B5A3769FF6BFE85FEBD284A" box="[255,338,294,321]" degrees="7.512" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="55" value="7.512">7.512N</geoCoordinate>
<geoCoordinate id="EE69D6EB8B5A3769FEC3FE84FE46284A" box="[343,425,295,321]" degrees="0.610" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="55" value="0.61">0.610E</geoCoordinate>
, 22.4.200
<quantity id="4CA51DC98B5A3769FD8EFE84FD9E284A" box="[538,625,295,321]" metricMagnitude="0" metricUnit="m" metricValue="8.28" pageId="4" pageNumber="5" unit="m" value="8.28">
<elevation id="0070571F8B5A3769FD8EFE84FD9E284A" box="[538,625,295,321]" metricMagnitude="0" metricUnit="m" metricValue="8.28" pageId="4" pageNumber="5" unit="m" value="8.28">8. 280m</elevation>
</quantity>
, cocoa / avocado plantation, leg.
<collectorName id="26A8D5FA8B5A3769FC4CFE84FAC3284A" box="[984,1324,295,321]" pageId="4" pageNumber="5">M. von Tschirnhaus. Specimen</collectorName>
deposited in
<collectionCode id="ED4C28E98B5A3769FF20FEE8FF19286E" box="[180,246,331,357]" pageId="4" pageNumber="5">SDEI</collectionCode>
; SDEI-Dip-01000.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C347E3A78B5A3769FF53FECCFACD2882" box="[199,1314,367,393]" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BE2B02C8B5A3769FF53FECCFACD2882" blockId="4.[151,1437,151,393]" box="[199,1314,367,393]" pageId="4" pageNumber="5">
<emphasis id="B9296C3E8B5A3769FF53FECCFEA72882" bold="true" box="[199,328,367,393]" pageId="4" pageNumber="5">Etymology</emphasis>
. The specific epithet refers to the town of Akloa near which the
<typeStatus id="54E60E8E8B5A3769FB8DFECCFB942882" box="[1049,1147,367,393]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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