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<mods:title id="EAB9A835AAE94C8B26FA51A13BDB2AD2">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<treatment id="03D7FF1CFF68FF49FF48FBB7326DFE41" ID-DOI="http://doi.org/10.5281/zenodo.3683198" ID-GBIF-Taxon="162246760" ID-Zenodo-Dep="3683198" LSID="urn:lsid:plazi:treatment:03D7FF1CFF68FF49FF48FBB7326DFE41" httpUri="http://treatment.plazi.org/id/03D7FF1CFF68FF49FF48FBB7326DFE41" lastPageId="164" lastPageNumber="162" pageId="163" pageNumber="162">
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<heading id="D089F966FF68FF4EFF48FBB73398FF01" bold="true" box="[192,740,217,241]" fontSize="10" level="2" pageId="163" pageNumber="162" reason="3">
<taxonomicName id="4C7E3589FF68FF4EFF48FBB73398FF01" authority="(Linnaeus, 1767)" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[192,740,217,241]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="B90A9218FF68FF4EFF48FBB73398FF01" bold="true" box="[192,740,217,241]" pageId="163" pageNumber="162">
<emphasis id="B90A9218FF68FF4EFF48FBB73087FF01" bold="true" box="[192,507,217,241]" italics="true" pageId="163" pageNumber="162">Cyclocephala amazona</emphasis>
(
<bibRefCitation id="EFEF33FBFF68FF4EFD84FBB733A0FF01" author="Linnaeus" box="[524,732,217,241]" firstAuthor="Linnaeus" pageId="163" pageNumber="162" pagination="533 - 1327" refId="ref212835" refString="Linnaeus, C. 1767. Systema naturae. Editio duodecima reformata. Tom. I. Pars II. Laur. Salvii; Stockholm. p. 533 - 1327." type="book chapter" year="1767">Linnaeus, 1767</bibRefCitation>
)
</emphasis>
</taxonomicName>
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</paragraph>
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<paragraph id="8BC14E0AFF68FF4EFF60FB9133C1FEE6" blockId="163.[232,701,255,278]" box="[232,701,255,278]" pageId="163" pageNumber="162">
<treatmentCitationGroup id="AB6E6924FF68FF4EFF60FB9133C1FEE6" box="[232,701,255,278]" pageId="163" pageNumber="162">
<treatmentCitation id="0ADF681BFF68FF4EFF60FB9133C1FEE6" author="Linnaeus" box="[232,701,255,278]" page="551" pageId="163" pageNumber="162" year="1767">
<taxonomicName id="4C7E3589FF68FF4EFF60FB9133C1FEE6" ID-CoL="6Y7SM" authority="Linnaeus 1767: 551" authorityName="Linnaeus" authorityPageNumber="551" authorityYear="1767" box="[232,701,255,278]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazonus">
<emphasis id="B90A9218FF68FF4EFF60FB91309DFEE5" box="[232,481,255,277]" italics="true" pageId="163" pageNumber="162">Scarabaeus amazonus</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFE6FFA6E33C1FEE6" author="Linnaeus" box="[487,701,256,278]" firstAuthor="Linnaeus" pageId="163" pageNumber="162" pagination="533 - 1327" refId="ref212835" refString="Linnaeus, C. 1767. Systema naturae. Editio duodecima reformata. Tom. I. Pars II. Laur. Salvii; Stockholm. p. 533 - 1327." type="book chapter" year="1767">Linnaeus 1767: 551</bibRefCitation>
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</paragraph>
<paragraph id="8BC14E0AFF68FF4EFF48FA483063FE8E" blockId="163.[192,1440,294,382]" pageId="163" pageNumber="162">
<emphasis id="B90A9218FF68FF4EFF48FA483074FECE" bold="true" box="[192,264,294,318]" pageId="163" pageNumber="162">Note.</emphasis>
Two presently recognized subspecies of
<taxonomicName id="4C7E3589FF68FF4EFD65FA48357EFECE" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[749,1026,294,318]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="B90A9218FF68FF4EFD65FA48357EFECE" box="[749,1026,294,318]" italics="true" pageId="163" pageNumber="162">Cyclocephala amazona</emphasis>
</taxonomicName>
have been described, one of which occurs in the research area. The other one is
<taxonomicName id="4C7E3589FF68FF4EFD52FA2835ECFEAE" authority="Hohne, 1923" authorityName="Hohne" authorityYear="1923" box="[730,1168,326,350]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="amazona" subSpecies="boliviensis">
<emphasis id="B90A9218FF68FF4EFD52FA28328CFEAE" box="[730,1008,326,350]" italics="true" pageId="163" pageNumber="162">C. amazona boliviensis</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFC7EFA2835ECFEAE" author="Hohne, W." box="[1014,1168,326,350]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="163" pageNumber="162" pagination="252 - 255" part="1923" refId="ref208127" refString="Hohne, W. 1923 a. Neue Dynastiden (Col.). Deutsche Entomologische Zeitschrift 1923 (3): 252 - 255." title="Neue Dynastiden (Col.)" type="journal article" year="1923">Höhne, 1923</bibRefCitation>
</taxonomicName>
(recorded from
<collectingCountry id="F3690E9AFF68FF4EFAC5FA2834E7FEAE" box="[1357,1435,326,350]" name="Brazil" pageId="163" pageNumber="162">Brazil</collectingCountry>
,
<collectingCountry id="F3690E9AFF68FF4EFF48FA08306BFE8E" box="[192,279,358,382]" name="Bolivia" pageId="163" pageNumber="162">Bolivia</collectingCountry>
)
</paragraph>
<paragraph id="8BC14E0AFF68FF4EFF48FAD8328BFE3E" blockId="163.[192,1015,438,462]" box="[192,1015,438,462]" pageId="163" pageNumber="162">
Subspecies
<taxonomicName id="4C7E3589FF68FF4EFEC5FAD8328BFE3E" authority="amazona (Linnaeus, 1767)" authorityName="amazona (Linnaeus" authorityYear="1767" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[333,1015,438,462]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="amazona" subSpecies="amazona">
<emphasis id="B90A9218FF68FF4EFEC5FAD8328BFE3E" bold="true" box="[333,1015,438,462]" pageId="163" pageNumber="162">
<emphasis id="B90A9218FF68FF4EFEC5FAD83272FE3D" bold="true" box="[333,782,438,462]" italics="true" pageId="163" pageNumber="162">Cyclocephala amazona amazona</emphasis>
(
<bibRefCitation id="EFEF33FBFF68FF4EFC96FAD83292FE3E" author="Linnaeus" box="[798,1006,438,462]" firstAuthor="Linnaeus" pageId="163" pageNumber="162" pagination="533 - 1327" refId="ref212835" refString="Linnaeus, C. 1767. Systema naturae. Editio duodecima reformata. Tom. I. Pars II. Laur. Salvii; Stockholm. p. 533 - 1327." type="book chapter" year="1767">Linnaeus, 1767</bibRefCitation>
)
</emphasis>
</taxonomicName>
</paragraph>
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<treatmentCitationGroup id="AB6E6924FF68FF4EFF60FAB2323EFE03" box="[232,834,476,499]" pageId="163" pageNumber="162">
<taxonomicName id="4C7E3589FF68FF4EFF60FAB2323EFE03" ID-CoL="6Y7SM" authority="Linnaeus 1767: 551" authorityName="Linnaeus" authorityPageNumber="551" authorityYear="1767" box="[232,834,476,499]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazonus">
<treatmentCitation id="0ADF681BFF68FF4EFF60FAB233C1FE02" author="Linnaeus" box="[232,701,476,499]" page="551" pageId="163" pageNumber="162" year="1767">
<emphasis id="B90A9218FF68FF4EFF60FAB2309DFE02" box="[232,481,476,498]" italics="true" pageId="163" pageNumber="162">Scarabaeus amazonus</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFE6FFAB333C1FE02" author="Linnaeus" box="[487,701,476,499]" firstAuthor="Linnaeus" pageId="163" pageNumber="162" pagination="533 - 1327" refId="ref212835" refString="Linnaeus, C. 1767. Systema naturae. Editio duodecima reformata. Tom. I. Pars II. Laur. Salvii; Stockholm. p. 533 - 1327." type="book chapter" year="1767">Linnaeus 1767: 551</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF68FF4EFD43FAB33246FE03" box="[715,826,477,499]" name="Suriname" pageId="163" pageNumber="162">Suriname</collectingCountry>
)
</taxonomicName>
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</paragraph>
<paragraph id="8BC14E0AFF68FF4EFF60FA94326AFDFF" blockId="163.[232,969,476,585]" box="[232,790,505,527]" pageId="163" pageNumber="162">
<treatmentCitationGroup id="AB6E6924FF68FF4EFF60FA94326AFDFF" box="[232,790,505,527]" pageId="163" pageNumber="162">
=
<taxonomicName id="4C7E3589FF68FF4EFF75FA97326AFDFF" authority="Fabricius 1781: 39" authorityName="Fabricius" authorityPageNumber="39" authorityYear="1781" box="[253,790,505,527]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="signata">
<treatmentCitation id="0ADF681BFF68FF4EFF75FA9733D8FDFF" author="Fabricius, J. C." box="[253,676,505,527]" page="39" pageId="163" pageNumber="162" year="1781">
<emphasis id="B90A9218FF68FF4EFF75FA9730AFFDFF" box="[253,467,505,527]" italics="true" pageId="163" pageNumber="162">Melolontha signata</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFE52FA9733D8FDFF" author="Fabricius, J. C." bookContentInfo="552 p." box="[474,676,505,527]" journalOrPublisher="Tom. I. Carol. Ernest. Bohnii; Hamburg and Kiel, Germany" pageId="163" pageNumber="162" refId="ref201797" refString="Fabricius, J. C. 1781. Species insectorum exhibentes eorum differentias specificas, synonyma aucto- rum, loca natalia, metamorphosin, adjectis observationibus, descriptionibus. Tom. I. Carol. Ernest. Bohnii; Hamburg and Kiel, Germany. 552 p." title="Species insectorum exhibentes eorum differentias specificas, synonyma aucto- rum, loca natalia, metamorphosin, adjectis observationibus, descriptionibus" type="book" year="1781">Fabricius 1781: 39</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF68FF4EFD3AFA973273FDFF" box="[690,783,505,527]" name="Jamaica" pageId="163" pageNumber="162">Jamaica</collectingCountry>
)
</taxonomicName>
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</paragraph>
<paragraph id="8BC14E0AFF68FF4EFF60F9793212FDDC" blockId="163.[232,969,476,585]" box="[232,878,534,556]" pageId="163" pageNumber="162">
<treatmentCitationGroup id="AB6E6924FF68FF4EFF60F9793212FDDC" box="[232,878,534,556]" pageId="163" pageNumber="162">
=
<treatmentCitation id="0ADF681BFF68FF4EFF75F97833D3FDDB" author="Fabricius, J. C." box="[253,687,534,556]" page="132" pageId="163" pageNumber="162" year="1798">
<taxonomicName id="4C7E3589FF68FF4EFF75F97833D3FDDB" authority="Fabricius 1798: 132" authorityName="Fabricius" authorityPageNumber="132" authorityYear="1798" box="[253,687,534,556]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B90A9218FF68FF4EFF75F97830ACFDDB" box="[253,464,534,555]" italics="true" pageId="163" pageNumber="162">Melolontha pallens</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFE5FF97833D3FDDB" author="Fabricius, J. C." bookContentInfo="572 p." box="[471,687,534,556]" journalOrPublisher="Proft et Storch; Copenhagen" pageId="163" pageNumber="162" refId="ref201973" refString="Fabricius, J. C. 1798. Supplementum entomologiae systematicae. Proft et Storch; Copenhagen. 572 p." title="Supplementum entomologiae systematicae" type="book" year="1798">Fabricius 1798: 132</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF68FF4EFD34F978321AFDDC" box="[700,870,534,556]" name="French Guiana" pageId="163" pageNumber="162">French Guiana</collectingCountry>
)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC14E0AFF68FF4EFF60F95A32B5FDB9" blockId="163.[232,969,476,585]" box="[232,969,563,585]" pageId="163" pageNumber="162">
<treatmentCitationGroup id="AB6E6924FF68FF4EFF60F95A32B5FDB9" box="[232,969,563,585]" pageId="163" pageNumber="162">
=
<treatmentCitation id="0ADF681BFF68FF4EFF75F95D339DFDB8" author="Bates, H. W." box="[253,737,563,585]" page="300" pageId="163" pageNumber="162" year="1888">
<taxonomicName id="4C7E3589FF68FF4EFF75F95D339DFDB8" authority="H. W. Bates 1888: 300" authorityName="H. W. Bates" authorityPageNumber="300" authorityYear="1888" box="[253,737,563,585]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="detecta">
<emphasis id="B90A9218FF68FF4EFF75F95D309FFDB8" box="[253,483,563,584]" italics="true" pageId="163" pageNumber="162">Cyclocephala detecta</emphasis>
H. W.
<bibRefCitation id="EFEF33FBFF68FF4EFDA7F95A339DFDB8" author="Bates, H. W." bookContentInfo="432 p." box="[559,737,563,585]" journalOrPublisher="Frederick Ducane Godman and Osbert Salvin; London" pageId="163" pageNumber="162" pagination="161 - 336" refId="ref193515" refString="Bates, H. W. 1888. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. p. 161 - 336. In: Biologia Centrali-Americana. Frederick Ducane Godman and Osbert Salvin; London. 432 p." title="Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana" year="1888">Bates 1888: 300</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF68FF4EFD67F95D323EFDB9" box="[751,834,563,585]" name="Mexico" pageId="163" pageNumber="162">Mexico</collectingCountry>
,
<collectingCountry id="F3690E9AFF68FF4EFCC4F95D32BDFDB9" box="[844,961,563,585]" name="Nicaragua" pageId="163" pageNumber="162">Nicaragua</collectingCountry>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FF68FF49FF48F937326DFE41" lastPageId="164" lastPageNumber="163" pageId="163" pageNumber="162" type="distribution">
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<emphasis id="B90A9218FF68FF4EFF48F9373090FD81" bold="true" box="[192,492,601,625]" pageId="163" pageNumber="162">
Distribution.
<collectingCountry id="F3690E9AFF68FF4EFEF1F937309AFD81" box="[377,486,601,625]" name="Guyana" pageId="163" pageNumber="162">Guyana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFE7AF937339FFD82" author="Endrodi, S." box="[498,739,601,626]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="163" pageNumber="162" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 147</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF68FF4EFD66F9373219FD81" author="Endrodi, S." bookContentInfo="800 p." box="[750,869,601,625]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="163" pageNumber="162" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 66</bibRefCitation>
, 132 (as
<taxonomicName id="4C7E3589FF68FF4EFC46F9343582FD81" authority="amazonica" box="[974,1278,602,625]" class="Insecta" family="Scarabaeidae" genus="Lachnosterna" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="amazonica" subSpecies="amazonica">
<emphasis id="B90A9218FF68FF4EFC46F9343582FD81" box="[974,1278,602,625]" italics="true" pageId="163" pageNumber="162">C. amazonica amazonica</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFA9CF9373039FD61" author="Dechambre, R. - P." journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="163" pageNumber="162" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 160</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF68FF4EFEDAF914330BFD62" author="Moore" box="[338,631,633,658]" etAl="et al." firstAuthor="Moore" pageId="163" pageNumber="162" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 142</bibRefCitation>
.
<emphasis id="B90A9218FF68FF4EFD0BF917326DFD61" bold="true" box="[643,785,633,657]" pageId="163" pageNumber="162">
<collectingCountry id="F3690E9AFF68FF4EFD0BF9173277FD61" box="[643,779,633,657]" name="Suriname" pageId="163" pageNumber="162">Suriname</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFC91F9173577FD61" author="Linnaeus" box="[793,1035,633,657]" firstAuthor="Linnaeus" pageId="163" pageNumber="162" pagination="533 - 1327" refId="ref212835" refString="Linnaeus, C. 1767. Systema naturae. Editio duodecima reformata. Tom. I. Pars II. Laur. Salvii; Stockholm. p. 533 - 1327." type="book chapter" year="1767">Linnaeus 1767: 551</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF68FF4EFBB7F917342DFD61" authorityName="Linnaeus" authorityYear="1767" box="[1087,1361,633,657]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazonus">
<emphasis id="B90A9218FF68FF4EFBB7F917342DFD61" box="[1087,1361,633,657]" italics="true" pageId="163" pageNumber="162">Scarabaeus amazonus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFAE1F91430ADFD41" author="Uyttenboogaart" firstAuthor="Uyttenboogaart" pageId="163" pageNumber="162" pagination="112 - 121" refId="ref227872" refString="Uyttenboogaart, D. L. 1902. List of beetles collected in the months of August and September 1900 in Surinam (Dutch Guyana) and on the island of Barbadoes. Tijdschrift voor Entomologie 45 (1 - 2): 112 - 121." type="journal article" year="1902">Uyttenboogaart 1902: 116</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF68FF4EFD8DF9F433A4FD42" authority="Drury" authorityName="Drury" box="[517,728,666,690]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B90A9218FF68FF4EFD8DF9F433FEFD41" box="[517,642,666,689]" italics="true" pageId="163" pageNumber="162">C. signata</emphasis>
Drury
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFD65F9F732A1FD42" author="Endrodi, S." box="[749,989,665,690]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="163" pageNumber="162" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 147</bibRefCitation>
(designation
<typeStatus id="54C5F0A8FF68FF4EFB0CF9F43477FD42" box="[1156,1291,666,690]" pageId="163" pageNumber="162" type="neotype">NEOTYPE</typeStatus>
<taxonomicName id="4C7E3589FF68FF4EFA9BF9F73047FD21" authorityName="Linnaeus" authorityYear="1767" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazonus">
<emphasis id="B90A9218FF68FF4EFA9BF9F73047FD21" italics="true" pageId="163" pageNumber="162">Scarabaeus amazonus</emphasis>
</taxonomicName>
), 1985a: 66, 132 (as
<taxonomicName id="4C7E3589FF68FF4EFDA2F9D43229FD21" authority="amazonica" box="[554,853,698,721]" class="Insecta" family="Scarabaeidae" genus="Lachnosterna" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="amazonica" subSpecies="amazonica">
<emphasis id="B90A9218FF68FF4EFDA2F9D43229FD21" box="[554,853,698,721]" italics="true" pageId="163" pageNumber="162">C. amazonica amazonica</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFCE2F9D73507FD21" author="Dechambre, R. - P." box="[874,1147,697,722]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="163" pageNumber="162" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 160</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF68FF4EFB0DF9D434E0FD22" author="Moore" box="[1157,1436,697,722]" etAl="et al." firstAuthor="Moore" pageId="163" pageNumber="162" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 142</bibRefCitation>
; MHCN; NZCS; RMNH.
<emphasis id="B90A9218FF68FF4EFE6DF9B733C0FD01" bold="true" box="[485,700,729,753]" pageId="163" pageNumber="162">
<collectingCountry id="F3690E9AFF68FF4EFE6DF9B733CAFD01" box="[485,694,729,753]" name="French Guiana" pageId="163" pageNumber="162">French Guiana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFD4BF9B732CFFD02" author="Fabricius, J. C." bookContentInfo="572 p." box="[707,947,729,754]" journalOrPublisher="Proft et Storch; Copenhagen" pageId="163" pageNumber="162" refId="ref201973" refString="Fabricius, J. C. 1798. Supplementum entomologiae systematicae. Proft et Storch; Copenhagen. 572 p." title="Supplementum entomologiae systematicae" type="book" year="1798">Fabricius 1798: 132</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF68FF4EFC6DF9B735B2FD01" authorityName="Fabricius" authorityYear="1798" box="[997,1230,729,753]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B90A9218FF68FF4EFC6DF9B735B2FD01" box="[997,1230,729,753]" italics="true" pageId="163" pageNumber="162">Melolontha pallens</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFB6DF9B73193FCE2" author="Endrodi, S." journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="163" pageNumber="162" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 147</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF68FF4EFF75F9973004FCE1" author="Endrodi, S." bookContentInfo="800 p." box="[253,376,761,785]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="163" pageNumber="162" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 66</bibRefCitation>
, 132 (as
<taxonomicName id="4C7E3589FF68FF4EFE62F9943263FCE1" authority="amazonica" box="[490,799,762,785]" class="Insecta" family="Scarabaeidae" genus="Lachnosterna" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="amazonica" subSpecies="amazonica">
<emphasis id="B90A9218FF68FF4EFE62F9943263FCE1" box="[490,799,762,785]" italics="true" pageId="163" pageNumber="162">C. amazonica amazonica</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFCB1F9973529FCE1" author="Dechambre, R. - P." box="[825,1109,761,786]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="163" pageNumber="162" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 160</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF68FF4EFBECF9943432FCE2" author="Duranton, M." box="[1124,1358,762,786]" journalOrPublisher="Coleopteres, numero special" pageId="163" pageNumber="162" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton 2011: 11</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF68FF4EFA0DF9943038FCC1" authorityName="Moser" authorityYear="1918" class="Insecta" family="Scarabaeidae" genus="Lachnosterna" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis id="B90A9218FF68FF4EFA0DF9943038FCC1" italics="true" pageId="163" pageNumber="162">C. amazonica</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFED2F8773357FCC1" author="Ponchel" box="[346,555,793,818]" firstAuthor="Ponchel" pageId="163" pageNumber="162" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011: 60</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF68FF4EFDBEF8773260FCC2" author="Dupuis, F." box="[566,796,793,818]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="163" pageNumber="162" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b: 109</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF68FF4EFCAFF8743538FCC2" author="Moore" box="[807,1092,793,818]" etAl="et al." firstAuthor="Moore" pageId="163" pageNumber="162" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 142</bibRefCitation>
; MHCN; NZCS.
<emphasis id="B90A9218FF68FF4EFA85F87734DCFCC1" bold="true" box="[1293,1440,793,817]" pageId="163" pageNumber="162">“Guianas”:</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFF48F8543003FCA2" author="Arrow, G. J." bookContentInfo="124 p." box="[192,383,825,850]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="163" pageNumber="162" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 8</bibRefCitation>
, 14 (as
<taxonomicName id="4C7E3589FF68FF4EFE57F854330DFCA1" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[479,625,826,849]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="B90A9218FF68FF4EFE57F854330DFCA1" box="[479,625,826,849]" italics="true" pageId="163" pageNumber="162">C. amazona</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF68FF4EFD2DF8543263FCA1" authorityName="Fabricius" authorityYear="1798" box="[677,799,825,849]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B90A9218FF68FF4EFD2DF8543263FCA1" box="[677,799,825,849]" italics="true" pageId="163" pageNumber="162">C. pallens</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFCB0F857352EFCA1" author="Blackwelder, R. E." box="[824,1106,825,850]" journalOrPublisher="United States National Museum Bulletin" pageId="163" pageNumber="162" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 251</bibRefCitation>
, 252 (as
<taxonomicName id="4C7E3589FF68FF4EFB48F854342EFCA1" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1216,1362,826,849]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="B90A9218FF68FF4EFB48F854342EFCA1" box="[1216,1362,826,849]" italics="true" pageId="163" pageNumber="162">C. amazona</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF68FF4EFA0DF854306AFC81" authorityName="Fabricius" authorityYear="1798" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B90A9218FF68FF4EFA0DF854306AFC81" italics="true" pageId="163" pageNumber="162">C. pallens</emphasis>
</taxonomicName>
).
<emphasis id="B90A9218FF68FF4EFEA3F83730B5FC81" bold="true" box="[299,457,857,881]" pageId="163" pageNumber="162">
<collectingCountry id="F3690E9AFF68FF4EFEA3F8373002FC81" box="[299,382,857,881]" name="Brazil" pageId="163" pageNumber="162">Brazil</collectingCountry>
(NA):
</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFE46F83733CAFC82" author="Endrodi, S." box="[462,694,857,882]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="163" pageNumber="162" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 147</bibRefCitation>
(NA), 1985a: 66, 132 (as
<taxonomicName id="4C7E3589FF68FF4EFC51F834347DFC81" authority="amazonica" box="[985,1281,858,881]" class="Insecta" family="Scarabaeidae" genus="Lachnosterna" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="amazonica" subSpecies="amazonica">
<emphasis id="B90A9218FF68FF4EFC51F834347DFC81" box="[985,1281,858,881]" italics="true" pageId="163" pageNumber="162">C. amazonica amazonica</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFA9EF837303AFC61" author="Dechambre, R. - P." journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="163" pageNumber="162" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 160</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF68FF4EFEDBF8143306FC62" author="Moore" box="[339,634,889,914]" etAl="et al." firstAuthor="Moore" pageId="163" pageNumber="162" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 142</bibRefCitation>
.
<emphasis id="B90A9218FF68FF4EFD0EF8173260FC61" bold="true" box="[646,796,889,913]" pageId="163" pageNumber="162">
<collectingCountry id="F3690E9AFF68FF4EFD0EF817326AFC61" box="[646,790,889,913]" name="Venezuela" pageId="163" pageNumber="162">Venezuela</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFCACF814328CFC61" author="Arrow, G. J." bookContentInfo="124 p." box="[804,1008,889,914]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="163" pageNumber="162" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 16</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF68FF4EFBAEF81435D8FC61" box="[1062,1188,890,913]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B90A9218FF68FF4EFBAEF81435D8FC61" box="[1062,1188,890,913]" italics="true" pageId="163" pageNumber="162">C. signata</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFB35F8173190FC41" author="Blackwelder, R. E." journalOrPublisher="United States National Museum Bulletin" pageId="163" pageNumber="162" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 252</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF68FF4EFEABF8F430E3FC41" authorityName="Fabricius" authorityYear="1798" box="[291,415,921,945]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B90A9218FF68FF4EFEABF8F430E3FC41" box="[291,415,921,945]" italics="true" pageId="163" pageNumber="162">C. pallens</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF68FF4EFE5CF8F4332FFC41" box="[468,595,922,945]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B90A9218FF68FF4EFE5CF8F4332FFC41" box="[468,595,922,945]" italics="true" pageId="163" pageNumber="162">C. signata</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFDE4F8F7321EFC42" author="Endrodi, S." box="[620,866,921,946]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="163" pageNumber="162" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 147</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF68FF4EFCF8F8F73296FC41" author="Endrodi, S." bookContentInfo="800 p." box="[880,1002,921,945]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="163" pageNumber="162" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 66</bibRefCitation>
, 132 (as
<taxonomicName id="4C7E3589FF68FF4EFBD3F8F434ECFC41" authority="amazonica" box="[1115,1424,922,945]" class="Insecta" family="Scarabaeidae" genus="Lachnosterna" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="amazonica" subSpecies="amazonica">
<emphasis id="B90A9218FF68FF4EFBD3F8F434ECFC41" box="[1115,1424,922,945]" italics="true" pageId="163" pageNumber="162">C. amazonica amazonica</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFF48F8D730A6FC21" author="Dechambre, R. - P." box="[192,474,953,978]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="163" pageNumber="162" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 160</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF68FF4EFE60F8D43272FC22" author="Moore" box="[488,782,953,978]" etAl="et al." firstAuthor="Moore" pageId="163" pageNumber="162" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 142</bibRefCitation>
.
<emphasis id="B90A9218FF68FF4EFC92F8D7320FFC21" bold="true" box="[794,883,953,977]" pageId="163" pageNumber="162">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFCF3F8D73523FC22" author="Fabricius, J. C." bookContentInfo="552 p." box="[891,1119,953,978]" journalOrPublisher="Tom. I. Carol. Ernest. Bohnii; Hamburg and Kiel, Germany" pageId="163" pageNumber="162" refId="ref201797" refString="Fabricius, J. C. 1781. Species insectorum exhibentes eorum differentias specificas, synonyma aucto- rum, loca natalia, metamorphosin, adjectis observationibus, descriptionibus. Tom. I. Carol. Ernest. Bohnii; Hamburg and Kiel, Germany. 552 p." title="Species insectorum exhibentes eorum differentias specificas, synonyma aucto- rum, loca natalia, metamorphosin, adjectis observationibus, descriptionibus" type="book" year="1781">Fabricius 1781: 39</bibRefCitation>
(Jamaica - as
<taxonomicName id="4C7E3589FF68FF4EFA9EF8D73067FC01" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B90A9218FF68FF4EFA9EF8D73067FC01" italics="true" pageId="163" pageNumber="162">Melolontha signata</emphasis>
</taxonomicName>
); H. W.
<bibRefCitation id="EFEF33FBFF68FF4EFE0CF8B43336FC02" author="Bates, H. W." bookContentInfo="432 p." box="[388,586,986,1010]" journalOrPublisher="Frederick Ducane Godman and Osbert Salvin; London" pageId="163" pageNumber="162" pagination="161 - 336" refId="ref193515" refString="Bates, H. W. 1888. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. p. 161 - 336. In: Biologia Centrali-Americana. Frederick Ducane Godman and Osbert Salvin; London. 432 p." title="Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana" year="1888">Bates 1888: 300</bibRefCitation>
(Mexico, Nicaragua - as
<taxonomicName id="4C7E3589FF68FF4EFC09F8B43285FC01" authorityName="H. W. Bates" authorityYear="1888" box="[897,1017,985,1009]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="detecta">
<emphasis id="B90A9218FF68FF4EFC09F8B43285FC01" box="[897,1017,985,1009]" italics="true" pageId="163" pageNumber="162">C. detecta</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFB9AF8B4359EFC02" author="Arrow, G. J." bookContentInfo="124 p." box="[1042,1250,985,1010]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="163" pageNumber="162" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 10</bibRefCitation>
, 16 (Honduras, Mexico, Panama - as
<taxonomicName id="4C7E3589FF68FF4EFE33F894334CFBE1" authorityName="H. W. Bates" authorityYear="1888" box="[443,560,1017,1041]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="detecta">
<emphasis id="B90A9218FF68FF4EFE33F894334CFBE1" box="[443,560,1017,1041]" italics="true" pageId="163" pageNumber="162">C. detecta</emphasis>
</taxonomicName>
; Colombia, Grenada, Nicaragua, Panama, Peru, Trinidad and Tobago - as
<taxonomicName id="4C7E3589FF68FF4EFF48FF743040FBC1" box="[192,316,1050,1073]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B90A9218FF68FF4EFF48FF743040FBC1" box="[192,316,1050,1073]" italics="true" pageId="163" pageNumber="162">C. signata</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFEDBFF773316FBC1" author="Blackwelder, R. E." box="[339,618,1049,1074]" journalOrPublisher="United States National Museum Bulletin" pageId="163" pageNumber="162" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 251</bibRefCitation>
, 252 (Honduras, Mexico, Nicaragua, Panama - as
<taxonomicName id="4C7E3589FF68FF4EFB41FF743443FBC1" authorityName="H. W. Bates" authorityYear="1888" box="[1225,1343,1049,1073]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="detecta">
<emphasis id="B90A9218FF68FF4EFB41FF743443FBC1" box="[1225,1343,1049,1073]" italics="true" pageId="163" pageNumber="162">C. detecta</emphasis>
</taxonomicName>
; Belize, Colombia, Cuba, Grenada, Jamaica, Nicaragua, Panama, Peru, Trinidad and Tobago - as
<taxonomicName id="4C7E3589FF68FF4EFA9AFF5434ECFBA1" box="[1298,1424,1082,1105]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B90A9218FF68FF4EFA9AFF5434ECFBA1" box="[1298,1424,1082,1105]" italics="true" pageId="163" pageNumber="162">C. signata</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFF48FF3730D0FB82" author="Endrodi, S." box="[192,428,1113,1138]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="163" pageNumber="162" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 147</bibRefCitation>
(Chile, Costa Rica, Colombia, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru - as
<taxonomicName id="4C7E3589FF68FF4EFD25FF1432D2FB61" authority="amazona" authorityName="amazona (Linnaeus" authorityYear="1767" box="[685,942,1146,1169]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="amazona" subSpecies="amazona">
<emphasis id="B90A9218FF68FF4EFD25FF1432D2FB61" box="[685,942,1146,1169]" italics="true" pageId="163" pageNumber="162">C. amazona amazona</emphasis>
</taxonomicName>
; Antilles - as
<taxonomicName id="4C7E3589FF68FF4EFBC5FF143446FB61" box="[1101,1338,1146,1169]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="amazona" subSpecies="signata">
<emphasis id="B90A9218FF68FF4EFBC5FF143446FB61" box="[1101,1338,1146,1169]" italics="true" pageId="163" pageNumber="162">C. amazona signata</emphasis>
</taxonomicName>
), 1985a: 66, 132 (Central America, Ecuador, Paraguay, Peru - as
<taxonomicName id="4C7E3589FF68FF4EFCFDFFF435DBFB41" authority="amazonica" box="[885,1191,1178,1201]" class="Insecta" family="Scarabaeidae" genus="Lachnosterna" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="amazonica" subSpecies="amazonica">
<emphasis id="B90A9218FF68FF4EFCFDFFF435DBFB41" box="[885,1191,1178,1201]" italics="true" pageId="163" pageNumber="162">C. amazonica amazonica</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF68FF4EFB36FFF73190FB21" author="Dechambre, R. - P." journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="163" pageNumber="162" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 160</bibRefCitation>
(Barbados, Bolivia, Central America, Chile, Colombia, Cuba, Ecuador, Grenada, Mexico, Paraguay, Peru, Trinidad and Tobago);
<bibRefCitation id="EFEF33FBFF68FF4EFDAFFFB73264FB02" author="Lachaume" box="[551,792,1241,1266]" firstAuthor="Lachaume" pageId="163" pageNumber="162" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume 1992: 13</bibRefCitation>
(Central AmericaParaguay, West Indies);
<bibRefCitation id="EFEF33FBFF68FF4EFAB0FFB73054FAE2" author="Ratcliffe" firstAuthor="Ratcliffe" pageId="163" pageNumber="162" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 83</bibRefCitation>
(Costa RicaParaguay/Chile);
<bibRefCitation id="EFEF33FBFF68FF4EFD2DFF9432BBFAE2" author="Moore" box="[677,967,1273,1298]" etAl="et al." firstAuthor="Moore" pageId="163" pageNumber="162" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 142</bibRefCitation>
(Bolivia, Chile, Colombia, Costa Rica, Ecuador, Panama, Paraguay, Peru, Trinidad and Tobago, West Indies).
</paragraph>
<paragraph id="8BC14E0AFF68FF4EFF48FE2F344FF9E9" blockId="163.[192,1440,1345,1561]" pageId="163" pageNumber="162">
<emphasis id="B90A9218FF68FF4EFF48FE2F3062FAA9" bold="true" box="[192,286,1345,1369]" pageId="163" pageNumber="162">Note 1.</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFEACFE2F330EFAA9" author="Uyttenboogaart" box="[292,626,1345,1370]" firstAuthor="Uyttenboogaart" pageId="163" pageNumber="162" pagination="112 - 121" refId="ref227872" refString="Uyttenboogaart, D. L. 1902. List of beetles collected in the months of August and September 1900 in Surinam (Dutch Guyana) and on the island of Barbadoes. Tijdschrift voor Entomologie 45 (1 - 2): 112 - 121." type="journal article" year="1902">Uyttenboogaart (1902: 116)</bibRefCitation>
incorrectly named Drury as the author of the
<taxonomicName id="4C7E3589FF68FF4EFB16FE2F34DCFAA9" box="[1182,1440,1345,1369]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B90A9218FF68FF4EFB16FE2F34DCFAA9" box="[1182,1440,1345,1369]" italics="true" pageId="163" pageNumber="162">Cyclocephala signata</emphasis>
</taxonomicName>
he recorded for
<collectingCountry id="F3690E9AFF68FF4EFEF5FE0F3085FA89" box="[381,505,1377,1401]" name="Suriname" pageId="163" pageNumber="162">Suriname</collectingCountry>
. In Uyttenboogaarts time, two species could be called
<taxonomicName id="4C7E3589FF68FF4EFB04FE0C3474FA89" box="[1164,1288,1378,1401]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B90A9218FF68FF4EFB04FE0C3474FA89" box="[1164,1288,1378,1401]" italics="true" pageId="163" pageNumber="162">C. signata</emphasis>
</taxonomicName>
. These were
<taxonomicName id="4C7E3589FF68FF4EFF48FEEF331EFA6A" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[192,610,1409,1434]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B90A9218FF68FF4EFF48FEEF30D9FA69" box="[192,421,1409,1433]" italics="true" pageId="163" pageNumber="162">Melolontha signata</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFE22FEEF331EFA6A" author="Fabricius, J. C." bookContentInfo="552 p." box="[426,610,1409,1434]" journalOrPublisher="Tom. I. Carol. Ernest. Bohnii; Hamburg and Kiel, Germany" pageId="163" pageNumber="162" refId="ref201797" refString="Fabricius, J. C. 1781. Species insectorum exhibentes eorum differentias specificas, synonyma aucto- rum, loca natalia, metamorphosin, adjectis observationibus, descriptionibus. Tom. I. Carol. Ernest. Bohnii; Hamburg and Kiel, Germany. 552 p." title="Species insectorum exhibentes eorum differentias specificas, synonyma aucto- rum, loca natalia, metamorphosin, adjectis observationibus, descriptionibus" type="book" year="1781">Fabricius, 1781</bibRefCitation>
</taxonomicName>
(p. 39), which is now a synonym of
<taxonomicName id="4C7E3589FF68FF4EFC70FEEC35F8FA69" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1016,1156,1410,1433]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="B90A9218FF68FF4EFC70FEEC35F8FA69" box="[1016,1156,1410,1433]" italics="true" pageId="163" pageNumber="162">C. amazona</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7E3589FF68FF4EFB4BFEEC3187FA4A" authority="Olivier, 1789" authorityName="Olivier" authorityYear="1789" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B90A9218FF68FF4EFB4BFEEC343DFA69" box="[1219,1345,1410,1433]" italics="true" pageId="163" pageNumber="162">M. signata</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFACEFEEF3187FA4A" author="Olivier" firstAuthor="Olivier" pageId="163" pageNumber="162" refId="ref218493" refString="Olivier, G. A. 1789. Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome premier. Baudouin; Paris. [Note. Each genus is separately paginated]." type="book" year="1789">Olivier, 1789</bibRefCitation>
</taxonomicName>
(p. 28), which is now a synonym of
<taxonomicName id="4C7E3589FF68FF4EFD26FECC32B0FA49" authority="(Illiger, 1806)" baseAuthorityName="Illiger" baseAuthorityYear="1806" box="[686,972,1441,1465]" class="Insecta" family="Aphodiidae" genus="Didactylia" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="notata">
<emphasis id="B90A9218FF68FF4EFD26FECC3263FA49" box="[686,799,1442,1465]" italics="true" pageId="163" pageNumber="162">C. notata</emphasis>
(Illiger, 1806)
</taxonomicName>
(p. 235). According to
<bibRefCitation id="EFEF33FBFF68FF4EFB57FECF306CFA2A" author="Uyttenboogaart" firstAuthor="Uyttenboogaart" pageId="163" pageNumber="162" pagination="112 - 121" refId="ref227872" refString="Uyttenboogaart, D. L. 1902. List of beetles collected in the months of August and September 1900 in Surinam (Dutch Guyana) and on the island of Barbadoes. Tijdschrift voor Entomologie 45 (1 - 2): 112 - 121." type="journal article" year="1902">Uyttenboogaart (1902)</bibRefCitation>
, the species he found was very common and flew into lighted rooms at night. This species is the present
<taxonomicName id="4C7E3589FF68FF4EFEADFE8C30CAFA09" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[293,438,1506,1529]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="B90A9218FF68FF4EFEADFE8C30CAFA09" box="[293,438,1506,1529]" italics="true" pageId="163" pageNumber="162">C. amazona</emphasis>
</taxonomicName>
, as confirmed by our examination of the original specimens of Uyttenboogaart in RMNH.
<taxonomicName id="4C7E3589FF68FF4EFEAFFD6F3362F9E9" baseAuthorityName="Illiger" baseAuthorityYear="1806" box="[295,542,1537,1561]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="notata">
<emphasis id="B90A9218FF68FF4EFEAFFD6F3362F9E9" box="[295,542,1537,1561]" italics="true" pageId="163" pageNumber="162">Cyclocephala notata</emphasis>
</taxonomicName>
is a species from the Antilles and does not occur in the Guianas.
</paragraph>
<paragraph id="8BC14E0AFF68FF4EFF48FD473401F992" blockId="163.[192,1440,1577,1634]" pageId="163" pageNumber="162">
<emphasis id="B90A9218FF68FF4EFF48FD473067F9B1" bold="true" box="[192,283,1577,1601]" pageId="163" pageNumber="162">Note 2.</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFEA8FD443341F9B2" author="Moore" box="[288,573,1577,1602]" etAl="et al." firstAuthor="Moore" pageId="163" pageNumber="162" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. (2018b: 107)</bibRefCitation>
conclude that the
<typeStatus id="54C5F0A8FF68FF4EFC9AFD443213F9B2" box="[786,879,1578,1602]" pageId="163" pageNumber="162" type="neotype">neotype</typeStatus>
of
<taxonomicName id="4C7E3589FF68FF4EFC07FD4735EBF9B1" authorityName="Linnaeus" authorityYear="1767" box="[911,1175,1577,1601]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazonus">
<emphasis id="B90A9218FF68FF4EFC07FD4735EBF9B1" box="[911,1175,1577,1601]" italics="true" pageId="163" pageNumber="162">Scarabaeus amazonus</emphasis>
</taxonomicName>
designated in
<bibRefCitation id="EFEF33FBFF68FF4EFAB7FD47302BF992" author="Endrodi, S." journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="163" pageNumber="162" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a: 147)</bibRefCitation>
is invalid because it violates several requirements of Article 75 of the
<bibRefCitation id="EFEF33FBFF68FF4EFB3AFD243405F992" author="ICZN" box="[1202,1401,1609,1634]" firstAuthor="ICZN" pageId="163" pageNumber="162" refId="ref209183" refString="ICZN [International Commission on Zoological Nomenclature]. 1964. International code of zoological nomenclature adopted by the XV International Congress of Zoology. Second edition. International Trust for Zoological Nomenclature; London. 176 p." type="book" year="1964">ICZN (1964: 81)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC14E0AFF68FF4EFF48FD1F30AAF919" blockId="163.[192,1440,1649,1770]" pageId="163" pageNumber="162">
<emphasis id="B90A9218FF68FF4EFF48FD1F305CF979" bold="true" box="[192,288,1649,1673]" pageId="163" pageNumber="162">Note 3.</emphasis>
According to
<bibRefCitation id="EFEF33FBFF68FF4EFE43FD1F33D3F97A" author="Ratcliffe" box="[459,687,1649,1674]" firstAuthor="Ratcliffe" pageId="163" pageNumber="162" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe (2003: 83</bibRefCitation>
, 165),
<bibRefCitation id="EFEF33FBFF68FF4EFC89FD1F350CF97A" author="Ratcliffe and Cave" box="[769,1136,1649,1674]" firstAuthor="Ratcliffe" pageId="163" pageNumber="162" pagination="1 - 424" refId="ref222249" refString="Ratcliffe, B. C., and R. D. Cave. 2006. The dynastine scarab beetles of Honduras, Nicaragua, and El Salvador (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 21: 1 - 424." type="journal article" year="2006">Ratcliffe and Cave (2006: 132</bibRefCitation>
, 167) and
<bibRefCitation id="EFEF33FBFF68FF4EFB79FD1F3039F95A" author="Ratcliffe" etAl="et al." firstAuthor="Ratcliffe" pageId="163" pageNumber="162" pagination="1 - 661" refId="ref222393" refString="Ratcliffe, B. C., R. D. Cave, and E. B. Cano. 2013. The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 27: 1 - 661." type="journal article" year="2013">Ratcliffe et al. (2013: 201</bibRefCitation>
, 595), the purported specimens of
<taxonomicName id="4C7E3589FF68FF4EFD77FDFF3566F959" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[767,1050,1681,1705]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="B90A9218FF68FF4EFD77FDFF3566F959" box="[767,1050,1681,1705]" italics="true" pageId="163" pageNumber="162">Cyclocephala amazona</emphasis>
</taxonomicName>
from
<collectingCountry id="F3690E9AFF68FF4EFBE2FDFF347CF959" box="[1130,1280,1681,1705]" name="El Salvador" pageId="163" pageNumber="162">El Salvador</collectingCountry>
,
<collectingCountry id="F3690E9AFF68FF4EFA98FDFF34E0F959" box="[1296,1436,1681,1705]" name="Guatemala" pageId="163" pageNumber="162">Guatemala</collectingCountry>
,
<collectingCountry id="F3690E9AFF68FF4EFF48FDDF3041F939" box="[192,317,1713,1737]" name="Honduras" pageId="163" pageNumber="162">Honduras</collectingCountry>
,
<collectingCountry id="F3690E9AFF68FF4EFEC2FDDF30DCF939" box="[330,416,1713,1737]" name="Mexico" pageId="163" pageNumber="162">Mexico</collectingCountry>
and
<collectingCountry id="F3690E9AFF68FF4EFE57FDDF3323F939" box="[479,607,1713,1737]" name="Nicaragua" pageId="163" pageNumber="162">Nicaragua</collectingCountry>
in
<bibRefCitation id="EFEF33FBFF68FF4EFD01FDDF32F1F93A" author="Endrodi, S." box="[649,909,1713,1738]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="163" pageNumber="162" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a: 147)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFF68FF4EFC44FDDF3589F939" author="Dechambre, R. - P." box="[972,1269,1713,1738]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="163" pageNumber="162" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre (1979c: 160)</bibRefCitation>
are in fact
<taxonomicName id="4C7E3589FF68FF4EFA0DFDDC30AEF919" authority="Casey, 1915" authorityName="Casey" authorityYear="1915" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="multiplex">
<emphasis id="B90A9218FF68FF4EFA0DFDDC3048F919" italics="true" pageId="163" pageNumber="162">C. multiplex</emphasis>
<bibRefCitation id="EFEF33FBFF68FF4EFEB3FDBC30AEF919" author="Casey, T. L." box="[315,466,1745,1770]" journalOrPublisher="Memoirs on the Coleoptera" pageId="163" pageNumber="162" pagination="1 - 394" part="6" refId="ref196237" refString="Casey, T. L. 1915. A review of the American species of Rutelinae, Dynastinae and Cetoniinae. Memoirs on the Coleoptera 6: 1 - 394." title="A review of the American species of Rutelinae, Dynastinae and Cetoniinae" type="journal article" year="1915">Casey, 1915</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC14E0AFF68FF49FF48FD97326DFE41" blockId="163.[192,1441,1785,1905]" lastBlockId="164.[192,1440,217,433]" lastPageId="164" lastPageNumber="163" pageId="163" pageNumber="162">
<emphasis id="B90A9218FF68FF4EFF48FD973060F8E1" bold="true" box="[192,284,1785,1809]" pageId="163" pageNumber="162">Note 4.</emphasis>
<taxonomicName id="4C7E3589FF68FF4EFEA9FD97334BF8E1" authorityName="Casey" authorityYear="1915" box="[289,567,1785,1809]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="multiplex">
<emphasis id="B90A9218FF68FF4EFEA9FD97334BF8E1" box="[289,567,1785,1809]" italics="true" pageId="163" pageNumber="162">Cyclocephala multiplex</emphasis>
</taxonomicName>
is described in
<bibRefCitation id="EFEF33FBFF68FF4EFD63FD9432C1F8E2" author="Casey, T. L." box="[747,957,1785,1810]" journalOrPublisher="Memoirs on the Coleoptera" pageId="163" pageNumber="162" pagination="1 - 394" part="6" refId="ref196237" refString="Casey, T. L. 1915. A review of the American species of Rutelinae, Dynastinae and Cetoniinae. Memoirs on the Coleoptera 6: 1 - 394." title="A review of the American species of Rutelinae, Dynastinae and Cetoniinae" type="journal article" year="1915">Casey (1915: 139)</bibRefCitation>
based on five specimens from
<collectingCountry id="F3690E9AFF68FF4EFAA8FD9734E0F8E1" box="[1312,1436,1785,1809]" name="Honduras" pageId="163" pageNumber="162">Honduras</collectingCountry>
. In
<bibRefCitation id="EFEF33FBFF68FF4EFF6BFC7430BDF8C2" author="Arrow, G. J." bookContentInfo="124 p." box="[227,449,1817,1842]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="163" pageNumber="162" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow (1937b: 10)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFF68FF4EFE77FC773257F8C1" author="Blackwelder, R. E." box="[511,811,1817,1842]" journalOrPublisher="United States National Museum Bulletin" pageId="163" pageNumber="162" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder (1944: 251)</bibRefCitation>
, this species is considered a synonym of
<taxonomicName id="4C7E3589FF68FF4EFAA9FC7434E4F8C1" authorityName="H. W. Bates" authorityYear="1888" box="[1313,1432,1817,1841]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="detecta">
<emphasis id="B90A9218FF68FF4EFAA9FC7434E4F8C1" box="[1313,1432,1817,1841]" italics="true" pageId="163" pageNumber="162">C. detecta</emphasis>
</taxonomicName>
, which was described in H. W.
<bibRefCitation id="EFEF33FBFF68FF4EFDAEFC543384F8A2" author="Bates, H. W." bookContentInfo="432 p." box="[550,760,1850,1874]" journalOrPublisher="Frederick Ducane Godman and Osbert Salvin; London" pageId="163" pageNumber="162" pagination="161 - 336" refId="ref193515" refString="Bates, H. W. 1888. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. p. 161 - 336. In: Biologia Centrali-Americana. Frederick Ducane Godman and Osbert Salvin; London. 432 p." title="Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana" year="1888">Bates (1888: 300)</bibRefCitation>
based on two specimens from
<collectingCountry id="F3690E9AFF68FF4EFBEAFC5735C4F8A1" box="[1122,1208,1849,1873]" name="Mexico" pageId="163" pageNumber="162">Mexico</collectingCountry>
and an unspecified number of specimens from
<collectingCountry id="F3690E9AFF68FF4EFD8BFC3733FBF881" box="[515,647,1881,1905]" name="Nicaragua" pageId="163" pageNumber="162">Nicaragua</collectingCountry>
.
<taxonomicName id="4C7E3589FF68FF4EFD07FC3732F6F881" authorityName="H. W. Bates" authorityYear="1888" box="[655,906,1881,1905]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="detecta">
<emphasis id="B90A9218FF68FF4EFD07FC3732F6F881" box="[655,906,1881,1905]" italics="true" pageId="163" pageNumber="162">Cyclocephala detecta</emphasis>
</taxonomicName>
is synonymized with
<taxonomicName id="4C7E3589FF68FF49FB03FC343036FF01" authority="in Endrodi (1964: 466)" authorityName="in Endrodi" authorityPageNumber="466" authorityYear="1964" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" lastPageId="164" lastPageNumber="163" order="Coleoptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="B90A9218FF68FF4EFB03FC343466F881" box="[1163,1306,1882,1905]" italics="true" pageId="163" pageNumber="162">C. amazona</emphasis>
in
<bibRefCitation id="EFEF33FBFF68FF49FAB5FC373036FF01" author="Endrodi, S." journalOrPublisher="Folia Entomologica Hungarica (N. S.)" lastPageId="164" lastPageNumber="163" pageId="163" pageNumber="162" pagination="433 - 470" part="17" refId="ref200586" refString="Endrodi, S. 1964. Eine Reihe von neuen Cyclocephala - Arten (Col., Melolonthidae, Dynastinae). Folia Entomologica Hungarica (N. S.) 17 (31): 433 - 470." title="Eine Reihe von neuen Cyclocephala - Arten (Col., Melolonthidae, Dynastinae)" type="journal article" year="1964">Endrödi (1964: 466)</bibRefCitation>
</taxonomicName>
, and both
<taxonomicName id="4C7E3589FF6FFF49FE43FBB4333EFF01" authorityName="H. W. Bates" authorityYear="1888" box="[459,578,217,241]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="detecta">
<emphasis id="B90A9218FF6FFF49FE43FBB4333EFF01" box="[459,578,217,241]" italics="true" pageId="164" pageNumber="163">C. detecta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E3589FF6FFF49FD08FBB4326AFF01" authorityName="Casey" authorityYear="1915" box="[640,790,217,241]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="multiplex">
<emphasis id="B90A9218FF6FFF49FD08FBB4326AFF01" box="[640,790,217,241]" italics="true" pageId="164" pageNumber="163">C. multiplex</emphasis>
</taxonomicName>
are mentioned as synonyms of
<taxonomicName id="4C7E3589FF6FFF49FB10FBB4309BFEE2" authority="amazona in Endrodi (1966 a: 147)" authorityName="in Endrodi" authorityPageNumber="147" authorityYear="1966" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="subSpecies" species="amazona" subSpecies="amazona">
<emphasis id="B90A9218FF6FFF49FB10FBB434DCFF01" box="[1176,1440,218,241]" italics="true" pageId="164" pageNumber="163">C. amazona amazona</emphasis>
in
<bibRefCitation id="EFEF33FBFF6FFF49FF6AFB97309BFEE2" author="Endrodi, S." box="[226,487,249,274]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="164" pageNumber="163" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a: 147)</bibRefCitation>
</taxonomicName>
and of
<taxonomicName id="4C7E3589FF6FFF49FDCFFB943568FEE1" authority="in Endrodi (1985 a: 66)" authorityName="in Endrodi" authorityPageNumber="66" authorityYear="1985" box="[583,1044,249,273]" class="Insecta" family="Scarabaeidae" genus="Lachnosterna" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis id="B90A9218FF6FFF49FDCFFB943393FEE1" box="[583,751,250,273]" italics="true" pageId="164" pageNumber="163">C. amazonica</emphasis>
in
<bibRefCitation id="EFEF33FBFF6FFF49FC92FB973568FEE1" author="Endrodi, S." bookContentInfo="800 p." box="[794,1044,249,273]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="164" pageNumber="163" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">Endrödi (1985a: 66)</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EFEF33FBFF6FFF49FBA9FB973460FEE1" author="Ratcliffe" box="[1057,1308,249,274]" firstAuthor="Ratcliffe" pageId="164" pageNumber="163" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe (2003: 165)</bibRefCitation>
reinstates
<taxonomicName id="4C7E3589FF6FFF49FF48FA74302AFEC1" authorityName="Casey" authorityYear="1915" box="[192,342,281,305]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="multiplex">
<emphasis id="B90A9218FF6FFF49FF48FA74302AFEC1" box="[192,342,281,305]" italics="true" pageId="164" pageNumber="163">C. multiplex</emphasis>
</taxonomicName>
as a valid species and mentions that all supposed specimens of
<taxonomicName id="4C7E3589FF6FFF49FBEDFA74358AFEC1" authorityName="in Endrodi" authorityYear="1964" box="[1125,1270,282,305]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="B90A9218FF6FFF49FBEDFA74358AFEC1" box="[1125,1270,282,305]" italics="true" pageId="164" pageNumber="163">C. amazona</emphasis>
</taxonomicName>
from north of
<collectingCountry id="F3690E9AFF6FFF49FF48FA54303EFEA1" box="[192,322,313,338]" name="Costa Rica" pageId="164" pageNumber="163">Costa Rica</collectingCountry>
were in fact
<taxonomicName id="4C7E3589FF6FFF49FE55FA54330DFEA1" authorityName="Casey" authorityYear="1915" box="[477,625,313,337]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="multiplex">
<emphasis id="B90A9218FF6FFF49FE55FA54330DFEA1" box="[477,625,313,337]" italics="true" pageId="164" pageNumber="163">C. multiplex</emphasis>
</taxonomicName>
. By doing so he ignores
<taxonomicName id="4C7E3589FF6FFF49FC1CFA543575FEA1" authorityName="H. W. Bates" authorityYear="1888" box="[916,1033,313,337]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="detecta">
<emphasis id="B90A9218FF6FFF49FC1CFA543575FEA1" box="[916,1033,313,337]" italics="true" pageId="164" pageNumber="163">C. detecta</emphasis>
</taxonomicName>
, which, based on its
<typeStatus id="54C5F0A8FF6FFF49FA8CFA543445FEA2" box="[1284,1337,314,338]" pageId="164" pageNumber="163">type</typeStatus>
locality, is almost certainly a senior synonym of
<taxonomicName id="4C7E3589FF6FFF49FD2AFA34324BFE81" authorityName="Casey" authorityYear="1915" box="[674,823,345,369]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="multiplex">
<emphasis id="B90A9218FF6FFF49FD2AFA34324BFE81" box="[674,823,345,369]" italics="true" pageId="164" pageNumber="163">C. multiplex</emphasis>
</taxonomicName>
rather than a junior synonym of
<taxonomicName id="4C7E3589FF6FFF49FB43FA343427FE81" authorityName="in Endrodi" authorityYear="1964" box="[1227,1371,346,369]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="amazona">
<emphasis id="B90A9218FF6FFF49FB43FA343427FE81" box="[1227,1371,346,369]" italics="true" pageId="164" pageNumber="163">C. amazona</emphasis>
</taxonomicName>
. This is also mentioned in
<bibRefCitation id="EFEF33FBFF6FFF49FE4AFA143385FE62" author="Moore" box="[450,761,377,402]" etAl="et al." firstAuthor="Moore" pageId="164" pageNumber="163" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. (2018b: 217)</bibRefCitation>
. The
<typeStatus id="54C5F0A8FF6FFF49FCB5FA14320EFE62" box="[829,882,378,402]" pageId="164" pageNumber="163">type</typeStatus>
series of
<taxonomicName id="4C7E3589FF6FFF49FC61FA14351DFE61" authorityName="H. W. Bates" authorityYear="1888" box="[1001,1121,377,401]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="detecta">
<emphasis id="B90A9218FF6FFF49FC61FA14351DFE61" box="[1001,1121,377,401]" italics="true" pageId="164" pageNumber="163">C. detecta</emphasis>
</taxonomicName>
should be compared with those of
<taxonomicName id="4C7E3589FF6FFF49FEAEFAF430C0FE41" authorityName="Casey" authorityYear="1915" box="[294,444,409,433]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="multiplex">
<emphasis id="B90A9218FF6FFF49FEAEFAF430C0FE41" box="[294,444,409,433]" italics="true" pageId="164" pageNumber="163">C. multiplex</emphasis>
</taxonomicName>
to confirm this supposition.
</paragraph>
</subSubSection>
</treatment>
</document>