279 lines
37 KiB
XML
279 lines
37 KiB
XML
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<mods:title id="BAEE89BF53D50121FE4A623407EA2A02">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<treatment id="03D7FF1CFF1DFF3BFF48FAC434DCFBFE" ID-DOI="http://doi.org/10.5281/zenodo.3682627" ID-GBIF-Taxon="162246959" ID-Zenodo-Dep="3682627" LSID="urn:lsid:plazi:treatment:03D7FF1CFF1DFF3BFF48FAC434DCFBFE" httpUri="http://treatment.plazi.org/id/03D7FF1CFF1DFF3BFF48FAC434DCFBFE" lastPageNumber="213" pageId="214" pageNumber="213">
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<subSubSection id="C3641D81FF1DFF3BFF48FAC43332FE31" box="[192,590,425,449]" pageId="214" pageNumber="213" type="nomenclature">
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<paragraph id="8BC14E0AFF1DFF3BFF48FAC43332FE31" blockId="214.[192,590,425,449]" box="[192,590,425,449]" pageId="214" pageNumber="213">
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<taxonomicName id="4C7E3589FF1DFF3BFF48FAC43332FE31" authority="Erichson, 1848" authorityName="Erichson" authorityYear="1848" box="[192,590,425,449]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas">
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<emphasis id="B90A9218FF1DFF3BFF48FAC43332FE31" bold="true" box="[192,590,425,449]" pageId="214" pageNumber="213">
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<emphasis id="B90A9218FF1DFF3BFF48FAC43001FE31" bold="true" box="[192,381,426,449]" italics="true" pageId="214" pageNumber="213">Tomarus gyas</emphasis>
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<bibRefCitation id="EFEF33FBFF1DFF3BFE0DFAC73332FE31" author="Erichson, W. F." box="[389,590,425,449]" editor="R. Schomburgk" journalOrPublisher="J. J. Weber; Leipzig, Germany" pageId="214" pageNumber="213" pagination="553 - 1260" refId="ref201522" refString="Erichson, W. F. 1848. Insecten. p. 553 - 617. In: R. Schomburgk. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil. J. J. Weber; Leipzig, Germany. p. 553 - 1260." title="Insecten" type="book chapter" volumeTitle="Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil" year="1848">Erichson, 1848</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3641D81FF1DFF3BFF60FABE33B0FE16" box="[232,716,464,486]" pageId="214" pageNumber="213" type="reference_group">
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<paragraph id="8BC14E0AFF1DFF3BFF60FABE33B0FE16" blockId="214.[232,716,464,486]" box="[232,716,464,486]" pageId="214" pageNumber="213">
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<treatmentCitationGroup id="AB6E6924FF1DFF3BFF60FABE33B0FE16" box="[232,716,464,486]" pageId="214" pageNumber="213">
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<taxonomicName id="4C7E3589FF1DFF3BFF60FABE33B0FE16" authority="Erichson 1848: 561" authorityName="Erichson" authorityPageNumber="561" authorityYear="1848" box="[232,716,464,486]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas">
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<treatmentCitation id="0ADF681BFF1DFF3BFF60FABE331CFE16" author="Erichson, W. F." box="[232,608,464,486]" page="561" pageId="214" pageNumber="213" year="1848">
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||
<emphasis id="B90A9218FF1DFF3BFF60FABE30FFFE15" box="[232,387,464,485]" italics="true" pageId="214" pageNumber="213">Tomarus gyas</emphasis>
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||
<bibRefCitation id="EFEF33FBFF1DFF3BFE03FABE331CFE16" author="Erichson, W. F." box="[395,608,464,486]" editor="R. Schomburgk" journalOrPublisher="J. J. Weber; Leipzig, Germany" pageId="214" pageNumber="213" pagination="553 - 1260" refId="ref201522" refString="Erichson, W. F. 1848. Insecten. p. 553 - 617. In: R. Schomburgk. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil. J. J. Weber; Leipzig, Germany. p. 553 - 1260." title="Insecten" type="book chapter" volumeTitle="Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil" year="1848">Erichson 1848: 561</bibRefCitation>
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</treatmentCitation>
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(
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||
<collectingCountry id="F3690E9AFF1DFF3BFDE6FABE33BAFE16" box="[622,710,464,486]" name="Guyana" pageId="214" pageNumber="213">Guyana</collectingCountry>
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||
)
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||
</taxonomicName>
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||
</treatmentCitationGroup>
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||
</paragraph>
|
||
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||
<subSubSection id="C3641D81FF1DFF3BFF48FA9834DCFBFE" pageId="214" pageNumber="213" type="distribution">
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||
<paragraph id="8BC14E0AFF1DFF3BFF48FA9834DCFBFE" blockId="214.[192,1440,502,1038]" pageId="214" pageNumber="213">
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||
<emphasis id="B90A9218FF1DFF3BFF48FA983096FDFE" bold="true" box="[192,490,502,526]" pageId="214" pageNumber="213">
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||
Distribution.
|
||
<collectingCountry id="F3690E9AFF1DFF3BFEFFFA983098FDFE" box="[375,484,502,526]" name="Guyana" pageId="214" pageNumber="213">Guyana</collectingCountry>
|
||
:
|
||
</emphasis>
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||
<bibRefCitation id="EFEF33FBFF1DFF3BFE67FA9833A4FDFE" author="Erichson, W. F." box="[495,728,502,526]" editor="R. Schomburgk" journalOrPublisher="J. J. Weber; Leipzig, Germany" pageId="214" pageNumber="213" pagination="553 - 1260" refId="ref201522" refString="Erichson, W. F. 1848. Insecten. p. 553 - 617. In: R. Schomburgk. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil. J. J. Weber; Leipzig, Germany. p. 553 - 1260." title="Insecten" type="book chapter" volumeTitle="Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil" year="1848">Erichson 1848: 561</bibRefCitation>
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; RMNH.
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||
<emphasis id="B90A9218FF1DFF3BFCCFFA9832A9FDFE" bold="true" box="[839,981,502,526]" pageId="214" pageNumber="213">
|
||
<collectingCountry id="F3690E9AFF1DFF3BFCCFFA9832B3FDFE" box="[839,975,502,526]" name="Suriname" pageId="214" pageNumber="213">Suriname</collectingCountry>
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||
:
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||
</emphasis>
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFC52FA9835CAFDFE" author="Endrodi, S." box="[986,1206,502,526]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="214" pageNumber="213" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 73</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFB6DFA993056FDDE" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFB6DFA99343FFDFE" box="[1253,1347,503,526]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFAD9FA99319BFDDE" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
|
||
<emphasis id="B90A9218FF1DFF3BFF7DF9793056FDDE" box="[245,298,535,558]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
|
||
</taxonomicName>
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||
), 1985a: 263 (as
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||
<taxonomicName id="4C7E3589FF1DFF3BFE65F979327AFDDE" box="[493,774,534,558]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFE65F9793335FDDE" box="[493,585,535,558]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFDDFF97933BFFDDE" box="[599,707,535,558]" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
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||
<emphasis id="B90A9218FF1DFF3BFD59F979327AFDDE" box="[721,774,535,558]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
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||
</taxonomicName>
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||
);
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||
<bibRefCitation id="EFEF33FBFF1DFF3BFC93F9783554FDDE" author="Dechambre, R. - P." box="[795,1064,534,558]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="214" pageNumber="213" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFBDDF9793597FDDE" box="[1109,1259,535,558]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="B90A9218FF1DFF3BFBDDF9793597FDDE" box="[1109,1259,535,558]" italics="true" pageId="214" pageNumber="213">Ligyrus gyas</emphasis>
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||
</taxonomicName>
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||
);
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||
<bibRefCitation id="EFEF33FBFF1DFF3BFA89F9783013FDBE" author="Escalona, H. E. & L. J. Joly" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="214" pageNumber="213" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly 2006: 122</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFE29F95933BEFDBE" box="[417,706,566,590]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFE29F9593083FDBE" box="[417,511,567,590]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFD86F9593300FDBE" box="[526,636,567,590]" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
|
||
<emphasis id="B90A9218FF1DFF3BFD04F95933BEFDBE" box="[652,706,567,590]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFD52F958353DFDBE" author="Lopez-Garcia" box="[730,1089,566,590]" etAl="et al." firstAuthor="Lopez-Garcia" pageId="214" pageNumber="213" pagination="491 - 509" refId="ref213049" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garcia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">López-García et al. 2016: 503</bibRefCitation>
|
||
.
|
||
<emphasis id="B90A9218FF1DFF3BFBC3F958345EFDBE" bold="true" box="[1099,1314,566,590]" pageId="214" pageNumber="213">
|
||
<collectingCountry id="F3690E9AFF1DFF3BFBC3F9583461FDBE" box="[1099,1309,566,590]" name="French Guiana" pageId="214" pageNumber="213">French Guiana</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFAA1F959305AFD9E" author="Duranton, M." journalOrPublisher="Coleopteres, numero special" pageId="214" pageNumber="213" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton 2011: 20</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFEDFF939308DFD9E" box="[343,497,599,622]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="B90A9218FF1DFF3BFEDFF939308DFD9E" box="[343,497,599,622]" italics="true" pageId="214" pageNumber="213">Ligyrus gyas</emphasis>
|
||
</taxonomicName>
|
||
) (?);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFDA4F938326EFD9E" author="Dupuis, F." box="[556,786,598,622]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="214" pageNumber="213" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France " Le Coleopteriste "]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b: 107</bibRefCitation>
|
||
, 110.
|
||
<emphasis id="B90A9218FF1DFF3BFCDFF9383284FD9E" bold="true" box="[855,1016,598,622]" pageId="214" pageNumber="213">
|
||
<collectingCountry id="F3690E9AFF1DFF3BFCDFF93832D7FD9E" box="[855,939,598,622]" name="Brazil" pageId="214" pageNumber="213">Brazil</collectingCountry>
|
||
(NA):
|
||
</emphasis>
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFC76F938346DFD9E" author="Blackwelder, R. E." box="[1022,1297,598,622]" journalOrPublisher="United States National Museum Bulletin" pageId="214" pageNumber="213" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFACAF939318AFD7E" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="B90A9218FF1DFF3BFACAF939318AFD7E" italics="true" pageId="214" pageNumber="213">Ligyrus gyas</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFE98F918308FFD7E" author="Endrodi, S." box="[272,499,630,654]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="214" pageNumber="213" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 73</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFDA2F919322CFD7E" box="[554,848,630,654]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFDA2F91933F4FD7E" box="[554,648,631,654]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFD12F9193274FD7E" box="[666,776,631,654]" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
|
||
<emphasis id="B90A9218FF1DFF3BFC92F919322CFD7E" box="[794,848,631,654]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
|
||
</taxonomicName>
|
||
) (NA), 1985a: 263 (as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFBE2F91934F3FD7E" box="[1130,1423,630,654]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFBE2F91935B4FD7E" box="[1130,1224,631,654]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFB51F919343BFD7E" box="[1241,1351,631,654]" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
|
||
<emphasis id="B90A9218FF1DFF3BFAD1F91934F3FD7E" box="[1369,1423,631,654]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFF48F9F830A6FD5E" author="Dechambre, R. - P." box="[192,474,662,686]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="214" pageNumber="213" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFD9AF9F933CCFD5E" box="[530,688,663,686]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="B90A9218FF1DFF3BFD9AF9F933CCFD5E" box="[530,688,663,686]" italics="true" pageId="214" pageNumber="213">Ligyrus gyas</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFD42F9F8354CFD5E" author="Escalona, H. E. & L. J. Joly" box="[714,1072,662,686]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="214" pageNumber="213" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly 2006: 122</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFBE0F9F934F3FD5E" box="[1128,1423,662,686]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFBE0F9F935BAFD5E" box="[1128,1222,663,686]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFB50F9F9343AFD5E" box="[1240,1350,663,686]" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
|
||
<emphasis id="B90A9218FF1DFF3BFAD1F9F934F3FD5E" box="[1369,1423,663,686]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFF48F9D8335FFD3E" author="Lopez-Garcia" box="[192,547,694,718]" etAl="et al." firstAuthor="Lopez-Garcia" pageId="214" pageNumber="213" pagination="491 - 509" refId="ref213049" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garcia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">López-García et al. 2016: 503</bibRefCitation>
|
||
.
|
||
<emphasis id="B90A9218FF1DFF3BFDA4F9D8327BFD3E" bold="true" box="[556,775,694,718]" pageId="214" pageNumber="213">
|
||
<collectingCountry id="F3690E9AFF1DFF3BFDA4F9D833C6FD3E" box="[556,698,694,718]" name="Venezuela" pageId="214" pageNumber="213">Venezuela</collectingCountry>
|
||
(EO):
|
||
</emphasis>
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFC84F9D83295FD3E" author="Endrodi, S." box="[780,1001,694,718]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="214" pageNumber="213" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 73</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFB91F9D93444FD3E" box="[1049,1336,694,718]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFB91F9D9350BFD3E" box="[1049,1143,695,718]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFB0EF9D93588FD3E" box="[1158,1268,695,718]" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
|
||
<emphasis id="B90A9218FF1DFF3BFA8AF9D93444FD3E" box="[1282,1336,695,718]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
|
||
</taxonomicName>
|
||
), 1985a: 263 (as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFE95F9B93341FD1E" box="[285,573,726,750]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFE95F9B93007FD1E" box="[285,379,727,750]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFE02F9B93084FD1E" box="[394,504,727,750]" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
|
||
<emphasis id="B90A9218FF1DFF3BFD8FF9B93341FD1E" box="[519,573,727,750]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFDDCF9B8321BFD1E" author="Dechambre, R. - P." box="[596,871,726,750]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="214" pageNumber="213" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFC10F9B9354EFD1E" box="[920,1074,727,750]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="B90A9218FF1DFF3BFC10F9B9354EFD1E" box="[920,1074,727,750]" italics="true" pageId="214" pageNumber="213">Ligyrus gyas</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFBC1F9B834DCFD1E" author="Escalona, H. E. & L. J. Joly" box="[1097,1440,726,750]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="214" pageNumber="213" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly 2006: 122</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFF61F9993374FCFE" box="[233,520,758,782]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFF61F999303BFCFE" box="[233,327,759,782]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFEDDF99930BFFCFE" box="[341,451,759,782]" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
|
||
<emphasis id="B90A9218FF1DFF3BFE5AF9993374FCFE" box="[466,520,759,782]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
|
||
</taxonomicName>
|
||
) (EO);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFDD5F99832C2FCFE" author="Lopez-Garcia" box="[605,958,758,782]" etAl="et al." firstAuthor="Lopez-Garcia" pageId="214" pageNumber="213" pagination="491 - 509" refId="ref213049" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garcia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">López-García et al. 2016: 503</bibRefCitation>
|
||
.
|
||
<emphasis id="B90A9218FF1DFF3BFC4FF99835D7FCFE" bold="true" box="[967,1195,758,782]" pageId="214" pageNumber="213">“Amazon Basin”:</emphasis>
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFB38F999340AFCFE" author="Arrow, G. J." bookContentInfo="124 p." box="[1200,1398,758,783]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="214" pageNumber="213" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 38</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFF48F8793027FCDE" box="[192,347,791,814]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="B90A9218FF1DFF3BFF48F8793027FCDE" box="[192,347,791,814]" italics="true" pageId="214" pageNumber="213">Ligyrus gyas</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<emphasis id="B90A9218FF1DFF3BFEFBF87830B0FCDE" bold="true" box="[371,460,790,814]" pageId="214" pageNumber="213">Other:</emphasis>
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFE5BF87833CFFCDE" author="Endrodi, S." box="[467,691,790,814]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="214" pageNumber="213" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 73</bibRefCitation>
|
||
(Argentina, Bolivia, Colombia, Ecuador, Guatemala, Mexico, Peru - as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFEB6F859331AFCBE" box="[318,614,822,846]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFEB6F85930E0FCBE" box="[318,412,823,846]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFE27F8593361FCBE" box="[431,541,823,846]" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
|
||
<emphasis id="B90A9218FF1DFF3BFDB8F859331AFCBE" box="[560,614,823,846]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
|
||
</taxonomicName>
|
||
), 1985a: 263 (Argentina, Bolivia, Colombia, Ecuador, Guatemala, Mexico, Peru - as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFE04F83933D8FC9E" box="[396,676,854,878]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFE04F839309BFC9E" box="[396,487,855,878]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFE7DF839331DFC9E" box="[501,609,855,878]" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
|
||
<emphasis id="B90A9218FF1DFF3BFDE7F83933D8FC9E" box="[623,676,855,878]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFD31F83832BFFC9E" author="Dechambre, R. - P." box="[697,963,854,878]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="214" pageNumber="213" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
|
||
(Argentina, Bolivia, Colombia, Ecuador, Guatemala, Mexico, Peru - as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFDA5F81933B4FC7E" box="[557,712,887,910]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="B90A9218FF1DFF3BFDA5F81933B4FC7E" box="[557,712,887,910]" italics="true" pageId="214" pageNumber="213">Ligyrus gyas</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFD57F81832B0FC7E" author="Lachaume" box="[735,972,886,910]" firstAuthor="Lachaume" pageId="214" pageNumber="213" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume 1992: 33</bibRefCitation>
|
||
(Argentina–Mexico - as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFB7DF81934F3FC7E" box="[1269,1423,887,910]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="B90A9218FF1DFF3BFB7DF81934F3FC7E" box="[1269,1423,887,910]" italics="true" pageId="214" pageNumber="213">Ligyrus gyas</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFF48F8F830D6FC5E" author="Ratcliffe" box="[192,426,918,942]" firstAuthor="Ratcliffe" pageId="214" pageNumber="213" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 277</bibRefCitation>
|
||
(Argentina–Mexico);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFD30F8F83561FC5E" author="Escalona, H. E. & L. J. Joly" box="[696,1053,918,942]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="214" pageNumber="213" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly 2006: 122</bibRefCitation>
|
||
(Argentina, Bolivia, Colombia, Costa Rica, Ecuador, Guatemala, Mexico, Panama, Peru - as
|
||
<taxonomicName id="4C7E3589FF1DFF3BFC1FF8D935C8FC3E" box="[919,1204,950,974]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="gyas" subGenus="Tomarus">
|
||
<emphasis id="B90A9218FF1DFF3BFC1FF8D93288FC3E" box="[919,1012,951,974]" italics="true" pageId="214" pageNumber="213">Ligyrus</emphasis>
|
||
(
|
||
<emphasis id="B90A9218FF1DFF3BFB8AF8D9350CFC3E" box="[1026,1136,951,974]" italics="true" pageId="214" pageNumber="213">Tomarus</emphasis>
|
||
)
|
||
<emphasis id="B90A9218FF1DFF3BFBF6F8D935C8FC3E" box="[1150,1204,951,974]" italics="true" pageId="214" pageNumber="213">gyas</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFB42F8D83416FC3E" author="Ponchel" box="[1226,1386,950,974]" firstAuthor="Ponchel" pageId="214" pageNumber="213" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France " Le Coleopteriste "]." type="journal article" year="2011">Ponchel 2011</bibRefCitation>
|
||
(not mentioned);
|
||
<bibRefCitation id="EFEF33FBFF1DFF3BFEDFF8B833CFFC1E" author="Lopez-Garcia" box="[343,691,982,1006]" etAl="et al." firstAuthor="Lopez-Garcia" pageId="214" pageNumber="213" pagination="451 - 492" refId="ref212996" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, and G. Amat-Garcia. 2015. The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution. Zootaxa 4048 (4): 451 - 492." type="journal article" year="2015">López-García et al. 2015: 484</bibRefCitation>
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(Argentina–Mexico, Colombia), 2016: 503 (Argentina, Bolivia, Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Peru).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |