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<document id="B99C1942F31FB98971EA0BB39F16BE97" ID-CLB-Dataset="26935" ID-DOI="10.5852/ejt.2019.521" ID-GBIF-Dataset="ed2118ff-0995-455d-a2a8-a25971b4e885" ID-Zenodo-Dep="2656607" ID-ZooBank="40C3F1EE-5EE3-4DA8-ADE0-6D57BAC23C26" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1556725635199" checkinUser="plazi" docAuthor="Wahlberg, Emma" docDate="2019" docId="03A6B675FFF6FFFA00DBD5B0FBDEB7DA" docLanguage="en" docName="ejt-521_wahlberg.pdf.imf" docOrigin="European Journal of Taxonomy 521" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Iteaphila macquarti Zetterstedt 1838" docType="treatment" docVersion="11" lastPageNumber="15" masterDocId="FF9FCE0DFFF8FFF50066D663FFB7B660" masterDocTitle="Revision and morphological analysis of the Ragadidae (Insecta, Diptera)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="15" updateTime="1698724386314" updateUser="ExternalLinkService">
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<mods:title id="04C9B0B9E55BF253B83AAAAF419C9FC9">Revision and morphological analysis of the Ragadidae (Insecta, Diptera)</mods:title>
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<mods:namePart id="382F3F7A743CE9B276D126C0E973D322">Wahlberg, Emma</mods:namePart>
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<mods:date id="10B2127459BAB706BB83A40EF6EE589C">2019</mods:date>
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<treatment id="03A6B675FFF6FFFA00DBD5B0FBDEB7DA" ID-DOI="http://doi.org/10.5281/zenodo.3489120" ID-GBIF-Taxon="156884886" ID-Zenodo-Dep="3489120" LSID="urn:lsid:plazi:treatment:03A6B675FFF6FFFA00DBD5B0FBDEB7DA" httpUri="http://treatment.plazi.org/id/03A6B675FFF6FFFA00DBD5B0FBDEB7DA" lastPageId="15" lastPageNumber="15" pageId="14" pageNumber="15">
<subSubSection id="C31554E8FFF6FFFB00DBD5B0FD75B58D" box="[189,706,979,1006]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BB00763FFF6FFFB00DBD5B0FD75B58D" blockId="14.[189,706,979,1006]" box="[189,706,979,1006]" pageId="14" pageNumber="15">
<heading id="D0F8B00FFFF6FFFB00DBD5B0FD75B58D" bold="true" box="[189,706,979,1006]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B97BDB71FFF6FFFB00DBD5B0FD75B58D" bold="true" box="[189,706,979,1006]" pageId="14" pageNumber="15">
Material examined of
<taxonomicName id="4C0F7CE0FFF6FFFB01B7D5B0FD75B58D" ID-CoL="3QDZ9" authorityName="Zetterstedt" authorityYear="1838" box="[465,706,979,1006]" class="Insecta" family="Empididae" genus="Iteaphila" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="macquarti">
<emphasis id="B97BDB71FFF6FFFB01B7D5B0FD75B58D" bold="true" box="[465,706,979,1006]" italics="true" pageId="14" pageNumber="15">Iteaphila macquarti</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31554E8FFF6FFFB008CD261FDECB28D" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BB00763FFF6FFFB008CD261FED1B27C" blockId="14.[189,1122,1026,1088]" box="[234,358,1026,1052]" pageId="14" pageNumber="15">
<typeStatus id="54B4B9C1FFF6FFFB008CD261FED1B27C" box="[234,358,1026,1052]" pageId="14" pageNumber="15" type="lectotype">
<emphasis id="B97BDB71FFF6FFFB008CD261FED1B27C" bold="true" box="[234,358,1026,1052]" pageId="14" pageNumber="15">Lectotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BB00763FFF6FFFB00DBD245FBD5B220" blockId="14.[189,1122,1026,1088]" box="[189,1122,1061,1088]" pageId="14" pageNumber="15">
<materialsCitation id="3B670D3EFFF6FFFB00DBD245FBD5B220" ID-GBIF-Occurrence="2237540417" box="[189,1122,1061,1088]" collectingDate="4 Jun." collectionCode="MZLU" collectorName="D. Boheman" country="Norway" location="NORWAY" pageId="14" pageNumber="15" specimenCode="MZLU 3028" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<collectingCountry id="F31847F3FFF6FFFB00DBD245FE8FB220" box="[189,312,1062,1088]" name="Norway" pageId="14" pageNumber="15">NORWAY</collectingCountry>
<specimenCount id="9D09CCEAFFF6FFFB0137D245FEC8B220" box="[337,383,1062,1088]" pageId="14" pageNumber="15" type="male">1 ♂</specimenCount>
; “Werdalen, Naes”; [
<collectingDate id="EFF5D84BFFF6FFFB0211D245FD73B220" box="[631,708,1061,1088]" pageId="14" pageNumber="15">
<date id="FFB121A3FFF6FFFB0211D245FD73B220" box="[631,708,1061,1088]" pageId="14" pageNumber="15">4 Jun.</date>
</collectingDate>
];
<collectorName id="26FA62B5FFF6FFFB02BED245FCDBB220" box="[728,876,1062,1088]" pageId="14" pageNumber="15">D. Boheman</collectorName>
leg.;
<specimenCode id="DBA9AF18FFF6FFFB03C9D245FBF8B220" box="[943,1103,1062,1088]" collectionCode="MZLU" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="14" pageNumber="15">MZLU 3028</specimenCode>
:1.
</materialsCitation>
</paragraph>
<paragraph id="8BB00763FFF6FFFB008CD204FE11B2E1" blockId="14.[189,1399,1127,1261]" box="[234,422,1127,1153]" pageId="14" pageNumber="15">
<emphasis id="B97BDB71FFF6FFFB008CD204FE11B2E1" bold="true" box="[234,422,1127,1153]" pageId="14" pageNumber="15">Other material</emphasis>
</paragraph>
<paragraph id="8BB00763FFF6FFFB00DBD2E8FDECB28D" blockId="14.[189,1399,1127,1261]" pageId="14" pageNumber="15">
<materialsCitation id="3B670D3EFFF6FFFB00DBD2E8FAB2B2A9" ID-GBIF-Occurrence="2237540433" collectingDate="2006-06-26" collectingDateMax="2006-07-15" collectingDateMin="2006-06-26" collectionCode="NHRS" collectorName="Swedish Malaise Trap Project" country="Sweden" county="Nuolja" location="Lappland" municipality="Abisko" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F31847F3FFF6FFFB00DBD2E8FE80B2C5" box="[189,311,1163,1189]" name="Sweden" pageId="14" pageNumber="15">SWEDEN</collectingCountry>
<specimenCount id="9D09CCEAFFF6FFFB013DD2E8FE39B2C6" box="[347,398,1163,1190]" pageId="14" pageNumber="15" type="male">1 ♂</specimenCount>
;
<location id="8ED051B8FFF6FFFB01C7D2E8FDA4B2C5" LSID="urn:lsid:plazi:treatment:03A6B675FFF6FFFA00DBD5B0FBDEB7DA:8ED051B8FFF6FFFB01C7D2E8FDA4B2C5" box="[417,531,1163,1189]" country="Sweden" county="Nuolja" municipality="Abisko" name="Lappland" pageId="14" pageNumber="15">Lappland</location>
,
<location id="8ED051B8FFF6FFFB0242D2E8FDC2B2C5" LSID="urn:lsid:plazi:treatment:03A6B675FFF6FFFA00DBD5B0FBDEB7DA:8ED051B8FFF6FFFB0242D2E8FDC2B2C5" box="[548,629,1163,1189]" country="Sweden" county="Nuolja" municipality="Abisko" name="Kiruna" pageId="14" pageNumber="15">Kiruna</location>
kommun,
<collectingMunicipality id="6BD49D19FFF6FFFB029BD2E8FCE5B2C5" box="[765,850,1163,1189]" pageId="14" pageNumber="15">Abisko</collectingMunicipality>
nationalpark,
<collectingCounty id="62D17FEFFFF6FFFB0460D2E8FBEDB2C5" box="[1030,1114,1163,1189]" pageId="14" pageNumber="15">Nuolja</collectingCounty>
;
<collectingDate id="EFF5D84BFFF6FFFB040CD2E8FAC5B2C5" box="[1130,1394,1163,1190]" pageId="14" pageNumber="15" value="2006-06-26" valueMax="2006-07-15" valueMin="2006-06-26">
<date id="FFB121A3FFF6FFFB040CD2E8FAC5B2C5" box="[1130,1394,1163,1190]" pageId="14" pageNumber="15" value="2006-06-26" valueMax="2006-07-15" valueMin="2006-06-26">26 Jun.15 Jul. 2006</date>
</collectingDate>
;
<collectorName id="26FA62B5FFF6FFFB00DBD2CCFDACB2A9" box="[189,539,1199,1225]" pageId="14" pageNumber="15">Swedish Malaise Trap Project</collectorName>
leg.;
<collectingMethod id="524E7F74FFF6FFFB023BD2CCFD45B2A9" box="[605,754,1199,1225]" pageId="14" pageNumber="15">Malaise trap</collectingMethod>
, bare mountain; DNA voucher AD7E; NHRS
</materialsCitation>
<materialsCitation id="3B670D3EFFF6FFFB0578D2CCFDE0B28D" ID-GBIF-Occurrence="2237540427" collectingDate="2006-06-26" collectingDateMax="2006-07-15" collectingDateMin="2006-06-26" collectionCode="NHRS" collectorName="Swedish Malaise Trap Project" country="Sweden" county="Nuolja" location="Lappland" municipality="Abisko" pageId="14" pageNumber="15" specimenCount="23" specimenCount-female="23">
<specimenCount id="9D09CCEAFFF6FFFB0578D2CCFAD8B2A9" box="[1310,1391,1199,1225]" pageId="14" pageNumber="15" type="female">23 ♀♀</specimenCount>
; same data as for preceding; NHRS
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31554E8FFF6FFFB00DBD377FCE7B038" pageId="14" pageNumber="15" type="description">
<paragraph id="8BB00763FFF6FFFB00DBD377FEF9B34E" blockId="14.[189,334,1300,1326]" box="[189,334,1300,1326]" pageId="14" pageNumber="15">
<heading id="D0F8B00FFFF6FFFB00DBD377FEF9B34E" bold="true" box="[189,334,1300,1326]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B97BDB71FFF6FFFB00DBD377FEF9B34E" bold="true" box="[189,334,1300,1326]" pageId="14" pageNumber="15">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB00763FFF6FFFB00DBD327FCE7B038" blockId="14.[189,1399,1347,1624]" pageId="14" pageNumber="15">
Eyes bare; males holoptic. Scape with setae; postpedicel tapering, at least 3 times as long as wide; stylus cylindrical (
<figureCitation id="13341BE6FFF6FFFB012AD304FE1DB3E2" box="[332,426,1383,1410]" captionStart="Fig" captionStartId="7.[189,232,1856,1882]" captionTargetBox="[189,1395,324,1808]" captionTargetId="figure@7.[189,1399,324,1816]" captionTargetPageId="7" captionText="Fig. 6. Habitus, lateral view. A. Ragas unica Walker, 1837, head and fore leg (NHRS). B. Iteaphila macquarti Zetterstedt, 1838, antenna (NHRS). C. Iteaphila macquarti, head and fore leg (NHRS). D. Ragas unica, mouthparts (NHRS). E. Iteaphila macquarti, mouthparts (NHRS). F. Ragas unica, 1 ♂, genitalia, lateral (NHRS). G. Ragas unica, 1 ♂, right epandrial lobe, dorsal (NHRS). H. Iteaphila macquarti, 1 ♂, genitalia, lateral (NHRS)." httpUri="https://zenodo.org/record/2656619/files/figure.png" pageId="14" pageNumber="15">Fig. 6B</figureCitation>
). Labrum straight in both sexes (6E); epipharyngal blades and ventroapical comb absent. Postgena bare or with setae, not spine like (
<figureCitation id="13341BE6FFF6FFFB0370D3E8FCC4B3C5" box="[790,883,1418,1445]" captionStart="Fig" captionStartId="7.[189,232,1856,1882]" captionTargetBox="[189,1395,324,1808]" captionTargetId="figure@7.[189,1399,324,1816]" captionTargetPageId="7" captionText="Fig. 6. Habitus, lateral view. A. Ragas unica Walker, 1837, head and fore leg (NHRS). B. Iteaphila macquarti Zetterstedt, 1838, antenna (NHRS). C. Iteaphila macquarti, head and fore leg (NHRS). D. Ragas unica, mouthparts (NHRS). E. Iteaphila macquarti, mouthparts (NHRS). F. Ragas unica, 1 ♂, genitalia, lateral (NHRS). G. Ragas unica, 1 ♂, right epandrial lobe, dorsal (NHRS). H. Iteaphila macquarti, 1 ♂, genitalia, lateral (NHRS)." httpUri="https://zenodo.org/record/2656619/files/figure.png" pageId="14" pageNumber="15">Fig. 6C</figureCitation>
). Prosternum separated from proepisternum and forming an isolated sclerite; laterotergite bare. Stout spine-like setae present on anterior face of fore coxa and inner ventral margin of fore trochanter absent (
<figureCitation id="13341BE6FFF6FFFB0338D3B1FC0CB38D" box="[862,955,1490,1517]" captionStart="Fig" captionStartId="7.[189,232,1856,1882]" captionTargetBox="[189,1395,324,1808]" captionTargetId="figure@7.[189,1399,324,1816]" captionTargetPageId="7" captionText="Fig. 6. Habitus, lateral view. A. Ragas unica Walker, 1837, head and fore leg (NHRS). B. Iteaphila macquarti Zetterstedt, 1838, antenna (NHRS). C. Iteaphila macquarti, head and fore leg (NHRS). D. Ragas unica, mouthparts (NHRS). E. Iteaphila macquarti, mouthparts (NHRS). F. Ragas unica, 1 ♂, genitalia, lateral (NHRS). G. Ragas unica, 1 ♂, right epandrial lobe, dorsal (NHRS). H. Iteaphila macquarti, 1 ♂, genitalia, lateral (NHRS)." httpUri="https://zenodo.org/record/2656619/files/figure.png" pageId="14" pageNumber="15">Fig. 6C</figureCitation>
); tubercle lacking on fore coxa; male fore tarsal claws present. Costal bristle absent; subcosta reaching costa; R
<subScript id="178B0526FFF6FFFB0456D064FBFCB075" attach="left" box="[1072,1099,1543,1557]" fontSize="6" pageId="14" pageNumber="15">4+5</subScript>
branched; CuA recurved (
<figureCitation id="13341BE6FFF6FFFB00A0D07AFE95B053" box="[198,290,1561,1587]" captionStart="Fig" captionStartId="10.[189,232,1815,1841]" captionTargetBox="[194,1398,718,1768]" captionTargetId="figure@10.[189,1399,718,1774]" captionTargetPageId="10" captionText="Fig. 9. Right wing. A. Ragas unica Walker, 1837, drawing with terminology on characters (left) and photo (right) (NHRS). B. Dipsomyia spinifera Bezzi, 1909 (MTD). C. Hydropeza longipennae (Miller, 1923) (dark colour is from underlying board to which the wing is glued) (NZAC 04021412). D. Zanclotus dioktes Wilder, 1982 (USNM). E. Iteaphila macquarti Zetterstedt, 1838 (NHRS). F. Anthepiscopus oedalinus (Zetterstedt, 1838) (NHRS-GULI000058764). G. Hormopeza obliterata Zetterstedt, 1838 (NHRS-000061621)." httpUri="https://zenodo.org/record/2656625/files/figure.png" pageId="14" pageNumber="15">Fig. 9E</figureCitation>
). Females: tergite 10 fused to cercus (
<figureCitation id="13341BE6FFF6FFFB0285D07AFCF6B053" box="[739,833,1561,1587]" captionStart="Fig" captionStartId="9.[189,232,1919,1945]" captionTargetBox="[200,1398,1402,1868]" captionTargetId="figure@9.[189,1399,1397,1877]" captionTargetPageId="9" captionText="Fig. 8. AB. Hormopeza obliterata Zetterstedt, 1838. A. Antennae and mouthparts, ♀ (NHRS- 000061621). B. Apical part of tarsus, ♂ (NHRS-000061670). CD. Terminalia, ♀. C. Iteaphila macquarti Zetterstedt, 1838, dorsolateral (NHRS). D. Hormopeza obliterata, dorsal (NHRS-000061621)." httpUri="https://zenodo.org/record/2656623/files/figure.png" pageId="14" pageNumber="15">Fig. 8C</figureCitation>
). Males: postgonites form fused processes from hypandrium; the apex of the phallus is simple (
<figureCitation id="13341BE6FFF6FFFB0282D05EFCF4B038" box="[740,835,1597,1624]" captionStart="Fig" captionStartId="7.[189,232,1856,1882]" captionTargetBox="[189,1395,324,1808]" captionTargetId="figure@7.[189,1399,324,1816]" captionTargetPageId="7" captionText="Fig. 6. Habitus, lateral view. A. Ragas unica Walker, 1837, head and fore leg (NHRS). B. Iteaphila macquarti Zetterstedt, 1838, antenna (NHRS). C. Iteaphila macquarti, head and fore leg (NHRS). D. Ragas unica, mouthparts (NHRS). E. Iteaphila macquarti, mouthparts (NHRS). F. Ragas unica, 1 ♂, genitalia, lateral (NHRS). G. Ragas unica, 1 ♂, right epandrial lobe, dorsal (NHRS). H. Iteaphila macquarti, 1 ♂, genitalia, lateral (NHRS)." httpUri="https://zenodo.org/record/2656619/files/figure.png" pageId="14" pageNumber="15">Fig. 6H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31554E8FFF6FFFB00DBD01DFAF8B08B" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BB00763FFF6FFFB00DBD01DFEE1B0F8" blockId="14.[189,342,1662,1688]" box="[189,342,1662,1688]" pageId="14" pageNumber="15">
<heading id="D0F8B00FFFF6FFFB00DBD01DFEE1B0F8" bold="true" box="[189,342,1662,1688]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B97BDB71FFF6FFFB00DBD01DFEE1B0F8" bold="true" box="[189,342,1662,1688]" pageId="14" pageNumber="15">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BB00763FFF6FFFB00DBD0CDFAF8B08B" blockId="14.[189,1399,1710,1771]" pageId="14" pageNumber="15">
The genus
<taxonomicName id="4C0F7CE0FFF6FFFB0159D0CDFE1EB0A8" authorityName="Zetterstedt" authorityYear="1838" box="[319,425,1710,1736]" class="Insecta" family="Empididae" genus="Iteaphila" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B97BDB71FFF6FFFB0159D0CDFE1EB0A8" box="[319,425,1710,1736]" italics="true" pageId="14" pageNumber="15">Iteaphila</emphasis>
</taxonomicName>
is the most widely spread group within the Ragadidae, with representatives in the Nearctic, Palaearctic and Oriental regions (
<bibRefCitation id="EF9E7A92FFF6FFFB02D0D0B2FC41B08B" author="Shamshev I. V. &amp; Sinclair B. J." box="[694,1014,1745,1771]" firstAuthor="Shamshev" journalOrPublisher="European Journal of Entomology" pageId="14" pageNumber="15" pagination="441 - 450" part="106" refId="ref6415" refString="Shamshev I. V. &amp; Sinclair B. J. 2009. Revision of the Iteaphila setosa group (Diptera: Empididae). European Journal of Entomology 106: 441 - 450. https: // doi. org / 10.14411 / eje. 2009.055" title="Revision of the Iteaphila setosa group (Diptera: Empididae)" type="journal article" year="2009">Shamshev &amp; Sinclair 2009</bibRefCitation>
;
<bibRefCitation id="EF9E7A92FFF6FFFB0464D0B2FAF4B08B" author="Sinclair B. J. &amp; Shamshev I. V." box="[1026,1347,1745,1771]" firstAuthor="Sinclair" journalOrPublisher="Zootaxa" pageId="14" pageNumber="15" pagination="1 - 61" part="3561" refId="ref6687" refString="Sinclair B. J. &amp; Shamshev I. V. 2012. World revision of Iteaphila macquarti group (Diptera: Empididae). Zootaxa 3561: 1 - 61." title="World revision of Iteaphila macquarti group (Diptera: Empididae)" type="journal article" year="2012">Sinclair &amp; Shamshev 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31554E8FFF6FFFA00DBD170FBDEB7DA" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BB00763FFF6FFFB00DBD170FEF3B14D" blockId="14.[189,324,1811,1837]" box="[189,324,1811,1837]" pageId="14" pageNumber="15">
<heading id="D0F8B00FFFF6FFFB00DBD170FEF3B14D" bold="true" box="[189,324,1811,1837]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B97BDB71FFF6FFFB00DBD170FEF3B14D" bold="true" box="[189,324,1811,1837]" pageId="14" pageNumber="15">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="8BB00763FFF6FFFA00DBD121FBDEB7DA" blockId="14.[189,1399,1858,2027]" lastBlockId="15.[189,1399,267,442]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<bibRefCitation id="EF9E7A92FFF6FFFB00DBD121FE42B13C" author="Sinclair B. J. &amp; Saigusa T." box="[189,501,1858,1884]" firstAuthor="Sinclair" journalOrPublisher="Entomological Science" pageId="14" pageNumber="15" pagination="507 - 522" part="4" refId="ref6654" refString="Sinclair B. J. &amp; Saigusa T. 2001. Revision of the world species of Ragas Walker (Diptera: Empidoidea). Entomological Science 4: 507 - 522." title="Revision of the world species of Ragas Walker (Diptera: Empidoidea)" type="journal article" year="2001">Sinclair &amp; Saigusa (2001)</bibRefCitation>
transferred
<taxonomicName id="4C0F7CE0FFF6FFFB02E1D120FC70B13C" authority="(Bezzi 1924)" authorityName="Bezzi" authorityYear="1924" baseAuthorityName="Bezzi" baseAuthorityYear="1924" box="[647,967,1858,1884]" class="Insecta" family="Empididae" genus="Ragas" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B97BDB71FFF6FFFB02E1D120FC94B13C" box="[647,803,1859,1884]" italics="true" pageId="14" pageNumber="15">Ragas setosa</emphasis>
(Bezzi 1924)
</taxonomicName>
to the genus
<taxonomicName id="4C0F7CE0FFF6FFFB040AD121FB61B13C" authorityName="Zetterstedt" authorityYear="1838" box="[1132,1238,1858,1884]" class="Insecta" family="Empididae" genus="Iteaphila" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B97BDB71FFF6FFFB040AD121FB61B13C" box="[1132,1238,1858,1884]" italics="true" pageId="14" pageNumber="15">Iteaphila</emphasis>
</taxonomicName>
based on the presence of a straight labrum, the absence of spine like setae on fore coxa and postgena, the palpus projection and the shape of the male terminalia.
<bibRefCitation id="EF9E7A92FFF6FFFB0372D1EAFBC7B1C4" author="Shamshev I. V. &amp; Sinclair B. J." box="[788,1136,1929,1956]" firstAuthor="Shamshev" journalOrPublisher="European Journal of Entomology" pageId="14" pageNumber="15" pagination="441 - 450" part="106" refId="ref6415" refString="Shamshev I. V. &amp; Sinclair B. J. 2009. Revision of the Iteaphila setosa group (Diptera: Empididae). European Journal of Entomology 106: 441 - 450. https: // doi. org / 10.14411 / eje. 2009.055" title="Revision of the Iteaphila setosa group (Diptera: Empididae)" type="journal article" year="2009">Shamshev &amp; Sinclair (2009)</bibRefCitation>
assigned this species together with several other from the southern Europe, the northern Africa and the Middle East to the
<taxonomicName id="4C0F7CE0FFF6FFFB00A1D1B2FE32B18B" baseAuthorityName="Bezzi" baseAuthorityYear="1925" box="[199,389,2001,2027]" class="Insecta" family="Empididae" genus="Iteaphila" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B97BDB71FFF6FFFB00A1D1B2FE32B18B" box="[199,389,2001,2027]" italics="true" pageId="14" pageNumber="15">Iteaphila setosa</emphasis>
</taxonomicName>
-group. This group, however, possesses species with numerous peculiar characters such as both ventral and dorsal surstyli, costal bristle, antennal shape and modification in male hind femur, and its affinity to other species in
<taxonomicName id="4C0F7CE0FFF7FFFA02D3D753FCA8B72A" authorityName="Zetterstedt" authorityYear="1838" box="[693,799,304,330]" class="Insecta" family="Empididae" genus="Iteaphila" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B97BDB71FFF7FFFA02D3D753FCA8B72A" box="[693,799,304,330]" italics="true" pageId="15" pageNumber="16">Iteaphila</emphasis>
</taxonomicName>
is unclear. For the analysis herein, the characters present in the
<typeStatus id="54B4B9C1FFF7FFFA010CD735FE2AB710" box="[362,413,342,368]" pageId="15" pageNumber="16">type</typeStatus>
species were used and the above listed characters treated as uniquely or possibly secondarily derived in the group. Further examination of morphological characters such as mouthparts, as well as DNA data is needed to clarify the phylogenetic position of this group.
</paragraph>
</subSubSection>
</treatment>
</document>