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<document ID-CLB-Dataset="55558" ID-DOI="http://dx.doi.org/10.3897/alpento.6.80349" ID-GBIF-Dataset="6a6dbf8e-2456-4ac3-b685-4509c0dbff9e" ID-Pensoft-Pub="2535-0889-6-13" ID-Pensoft-UUID="7266BBB11F3C52A397FD698561E728F0" ID-ZooBank="CAAB885C7DB84C4B9D0153B07656840B" ModsDocID="2535-0889-6-13" checkinTime="1652359712937" checkinUser="pensoft" docAuthor="Reyes-Hernandez, Jose L., Hansen, Aslak Kappel &amp; Solodovnikov, Alexey" docDate="2022" docId="24C4CA6F71AD58349C4DE6144886BE14" docLanguage="en" docName="AlpEntomol 6: 13-18" docOrigin="Alpine Entomology 6" docPubDate="2022-05-11" docSource="http://dx.doi.org/10.3897/alpento.6.80349" docTitle="Ikaros navarretei Reyes-Hernández &amp; Hansen &amp; Solodovnikov 2022, sp. nov." docType="treatment" docUuid="2C1E9003-82DD-4CF6-900D-16451E23B9E4" docUuidSource="ZooBank" docVersion="5" id="7266BBB11F3C52A397FD698561E728F0" lastPageNumber="13" masterDocId="7266BBB11F3C52A397FD698561E728F0" masterDocTitle="Ikaros navarretei (Coleoptera, Staphylinidae, Staphylininae), a new apterous rove beetle species from high elevations in Colombia" masterLastPageNumber="18" masterPageNumber="13" pageNumber="13" updateTime="1732867490599" updateUser="ExternalLinkService">
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<mods:title id="B81930486F5CDCD40010919C70A7DDCD">Ikaros navarretei (Coleoptera, Staphylinidae, Staphylininae), a new apterous rove beetle species from high elevations in Colombia</mods:title>
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<mods:namePart id="4CCCF844D202682FA45DB143B58537AC">Reyes-Hernandez, Jose L.</mods:namePart>
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<mods:affiliation id="D416D56EB3F45C146C31DE8E10FD3D74">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, Copenhagen, 2100, Denmark</mods:affiliation>
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<mods:namePart id="619FE1535364C183E373E793916766F0">Hansen, Aslak Kappel</mods:namePart>
<mods:nameIdentifier id="1AFDCD56EDA683EF9AFC7AEF5682B7AF" type="ORCID">https://orcid.org/0000-0003-2089-7233</mods:nameIdentifier>
<mods:affiliation id="90F8D94F2CCE20DBCD35702C81B201A3">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, Copenhagen, 2100, Denmark</mods:affiliation>
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<mods:namePart id="B6786AA352CC53EE4DBEF4D096DA52C1">Solodovnikov, Alexey</mods:namePart>
<mods:nameIdentifier id="FCFFFF63281E40F5AEFCDB0C1D171F3B" type="ORCID">https://orcid.org/0000-0003-2031-849X</mods:nameIdentifier>
<mods:affiliation id="7B0A740C9DD158C5F314FEAA1BB574AC">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, Copenhagen, 2100, Denmark</mods:affiliation>
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<mods:title id="249F5CAC7DEE5CFB5A0D7D63EF71A003">Alpine Entomology</mods:title>
</mods:titleInfo>
<mods:part id="DEECE6AE2BC37BC49FE8DE4E21AB1BCF">
<mods:date id="F7FDBDCEA1C5356CA02286D0EA454927">2022</mods:date>
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<mods:number id="E660B8CDEAB8CD6197B8B63739E91A3B">2022-05-11</mods:number>
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<mods:number id="B21052F19AA0E50C0D7711B1E8AC2400">6</mods:number>
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<mods:classification id="CEC7BD11F173E65917F49C6326B6412A">journal article</mods:classification>
<mods:identifier id="2CA93CF7BD09983EEA199B7DDF777F3B" type="DOI">http://dx.doi.org/10.3897/alpento.6.80349</mods:identifier>
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<treatment id="24C4CA6F71AD58349C4DE6144886BE14" ID-GBIF-Taxon="195302499" LSID="urn:lsid:zoobank.org:act:2C1E9003-82DD-4CF6-900D-16451E23B9E4" httpUri="http://treatment.plazi.org/id/24C4CA6F71AD58349C4DE6144886BE14" lastPageNumber="13" pageId="0" pageNumber="13" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Staphylininae">
<subSubSection id="824A9F84690383108B9F90C098C55260" pageId="0" pageNumber="13" type="nomenclature">
<paragraph id="D06983AA7D6E1426732BC45D060EFB2F" pageId="0" pageNumber="13">
<taxonomicName id="E31BB4112981DBDE2365AD1A2B94AA4C" ID-CoL="9GD48" LSID="http://zoobank.org/2C1E9003-82DD-4CF6-900D-16451E23B9E4" authority="Reyes-Hernández &amp; Hansen &amp; Solodovnikov, 2022" authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei" status="sp. nov.">Ikaros navarretei</taxonomicName>
<taxonomicNameLabel id="F1D9E0F2703BE36103FCCD244F3210C8" pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="AFF74D38F7E85FAA193F9C4A7A1B221A" pageId="0" pageNumber="13" type="type locality">
<paragraph id="8AAFEF578257037EECD2D0CDC95C4734" pageId="0" pageNumber="13">Type locality.</paragraph>
<paragraph id="F7DB49423BC50F52DDB7C2A06EFEB273" pageId="0" pageNumber="13">
Colombia, Cauca, Silvia,
<geoCoordinate id="5E5CDE90468061A57C0C57F8EB6A0C4E" degrees="2.5889" direction="north" orientation="latitude" precision="5" value="2.5889">2.5889</geoCoordinate>
,
<geoCoordinate id="5611583C0F3846A1108300413CF2BE1C" degrees="76.24886" direction="west" orientation="longitude" precision="1" value="-76.24886">-76.24886</geoCoordinate>
, 3400 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="43A08805F8ABDD74AC6C450209E0CFD3" pageId="0" pageNumber="13" type="generic placement">
<paragraph id="BBEF1F7A405DCACA38C788E2F4AF8A37" pageId="0" pageNumber="13">Generic placement.</paragraph>
<paragraph id="D17F28461C1187221D19C2BB89B377BE" pageId="0" pageNumber="13">
Our specimen fully agrees with the generic diagnosis and description provided by
<bibRefCitation id="F999DAC6E6E0424DEE33370B57545863" DOI="https://doi.org/10.5852/ejt.2021.744.1303" author="Chatzimanolis, S" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="13" pagination="67 - 82" refId="B1" refString="Chatzimanolis, S, Brunke, AJ, 2021. A new apterous rove beetle genus (Coleoptera: Staphylinidae) from the Northern Andes with an assessment of its phylogenetic position. European Journal of Taxonomy 744: 67 - 82, DOI: https://doi.org/10.5852/ejt.2021.744.1303" title="A new apterous rove beetle genus (Coleoptera: Staphylinidae) from the Northern Andes with an assessment of its phylogenetic position." url="https://doi.org/10.5852/ejt.2021.744.1303" volume="744" year="2021">Chatzimanolis and Brunke (2021)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="44C7FE27690C243A085B26D3CD75DC89" pageId="0" pageNumber="13" type="diagnosis">
<paragraph id="8300B69F66BD547FF0AAE8D8BC1B997A" pageId="0" pageNumber="13">Diagnosis.</paragraph>
<paragraph id="14FF78E5CC9CCEFA1B4F319A2E7667A6" pageId="0" pageNumber="13">
<taxonomicName id="60D0B4399ECB99D9D62DC41EC0BD2A78" authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis id="7910B4703AA5C7E37FA271A4CE418009" italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
sp. nov. can be distinguished from all other species in this genus by the combination of the presence of an arch-like carina on terga III-V; the meshed to isodiametric microsculpture on the dorsal surface of the head, thorax, and mesoscutellum; antennomeres with crown-like macrosetae shorter than the length of antennomere.
</paragraph>
</subSubSection>
<subSubSection id="82C28E291109A7688D1822AB971633C2" pageId="0" pageNumber="13" type="etymology">
<paragraph id="BC969BC52F02A1D9B0200236636C16F6" pageId="0" pageNumber="13">Etymology.</paragraph>
<paragraph id="FFB405B352210AD41F4BCF8D87E0356B" pageId="0" pageNumber="13">
The species epithet is in recognition (patronymic) of
<normalizedToken id="76A01F751F1C9601D7468C127A0878B6" originalValue="José">Jose</normalizedToken>
Luis Navarrete Heredia for his contribution to the knowledge of the family
<taxonomicName id="8F29714BF6197D5A5FB129195DC0A1AC" class="Insecta" family="Staphylinidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
and training of many coleopterologists in Latin America.
</paragraph>
</subSubSection>
<subSubSection id="923A3E4489FDD54791F9427CF87B1D67" pageId="0" pageNumber="13" type="type material">
<paragraph id="993721E334E6F8FB767B946FE1C431FC" pageId="0" pageNumber="13">Type material.</paragraph>
<paragraph id="62C4DF06F9DA862E8EC18709F55AD18D" pageId="0" pageNumber="13">
<emphasis id="E0F09F7D573EE48FD4C999B0C71D6443" bold="true" pageId="0" pageNumber="13">
<emphasis id="7BE711248C138B80C66EB48A0AA7DB2F" italics="true" pageId="0" pageNumber="13">Holotype</emphasis>
.
</emphasis>
Male, point mounted, with genitalia in a separate microvial, with labels as follows: &quot;COLOMB. [COLOMBIA], 20 km E Silvia, Cauca, [~
<geoCoordinate id="E09ED407C86914385E906A18CFD99ED2" degrees="2.5889" direction="north" orientation="latitude" precision="5" value="2.5889">2.5889</geoCoordinate>
,
<geoCoordinate id="5D8903F43710DC2F7DBDAAA5793E4A3D" degrees="76.24886" direction="west" orientation="longitude" precision="1" value="-76.24886">-76.24886</geoCoordinate>
, 3400 m a.s.l.] VII.16.1970 [16 June 1970], II, 000' J.M. Campbell / Xantopygina sic? gen. det. Newton 1998 / HOLOTYPE
<taxonomicName id="115B5CF8C71E85B84AAE7AB8B3C86677" authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis id="0FB1124AA52AD542C941C2A1DA1B1F78" italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
<normalizedToken id="27A0C8D8B57F69FAD2811AA68683423D" originalValue="Reyes-Hernández">Reyes-Hernandez</normalizedToken>
, Hansen and Solodovnikov, des.
<normalizedToken id="16BBF70F386BD0CD1A7DFC6386554D4B" originalValue="Reyes-Hernández">Reyes-Hernandez</normalizedToken>
, Hansen and Solodovnikov 2022&quot; CNC.
</paragraph>
</subSubSection>
<subSubSection id="BDDDCCA0ACFD742D85E68279E0B72A65" pageId="0" pageNumber="13" type="description">
<paragraph id="F1F0FCC3A7EB2C066F4643081AC2C1F5" pageId="0" pageNumber="13">Description.</paragraph>
<paragraph id="5012507B2B4F0159D8306F3435E85AF7" pageId="0" pageNumber="13">
Habitus as in Fig.
<figureCitation id="D4F7DAFC1765CF750A7F9C9DCF8FE05D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal view of Ikaros navarretei sp. nov. A. Habitus; B. Close up of head punctation and meshed to isodiametric microsculpture; C. Close up of pronotum punctation and meshed to isodiametric microsculpture; D. Close up of mesoscutellum and elytra punctation and microsculpture; E. Close up of abdominal tergite III with arch-like carina highlighted by white dashed line." figureDoi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13">1A</figureCitation>
. Total body length 10.06 mm. Forebody length 5.25 mm long.
</paragraph>
<paragraph id="8D1958EDF454AB4326BF2A1166B4AD86" pageId="0" pageNumber="13">Coloration of body reddish-brown, with abdomen having undertones of metallic green-brown.</paragraph>
<paragraph id="C4F76B395DA86BAF67F7B251D2F25390" pageId="0" pageNumber="13">
Head subrectangular, slightly wider than long, HW/HL ratio = 1.1. Epicranium with numerous large punctures, except impunctate center; punctures not contiguous, the distance between punctures typically equals the width of 1-2 punctures; with transversely meshed to isodiametric microsculpture (Fig.
<figureCitation id="6646759A1B0F64B704561F393989733B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal view of Ikaros navarretei sp. nov. A. Habitus; B. Close up of head punctation and meshed to isodiametric microsculpture; C. Close up of pronotum punctation and meshed to isodiametric microsculpture; D. Close up of mesoscutellum and elytra punctation and microsculpture; E. Close up of abdominal tergite III with arch-like carina highlighted by white dashed line." figureDoi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13">1B</figureCitation>
). Labial palpus with palpomere 3 (apical) widest before the apex. Antennomeres with crown-like macrosetae shorter than length of antennomere.
</paragraph>
<caption id="ED699EECA69815E94C1F6376786547BF" doi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13" start="Figure 1" startId="F1">
<paragraph id="44E18CD4AF28B5FD90364784BCE40752" pageId="0" pageNumber="13">
<emphasis id="FEFE45D75A9C52BA71B10F22175561CE" bold="true" pageId="0" pageNumber="13">Figure 1.</emphasis>
Dorsal view of
<taxonomicName id="2959FB8272FF7F6AB81926DB7391323E" authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis id="3887836A2BD1DB00669C031F7B509E44" italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="4A1A923151F48A7FD4A3EC821EEAA19E" bold="true" pageId="0" pageNumber="13">A.</emphasis>
Habitus;
<emphasis id="39F49032B148D63B486099708DAB6E50" bold="true" pageId="0" pageNumber="13">B.</emphasis>
Close up of head punctation and meshed to isodiametric microsculpture;
<emphasis id="BEFB5B3FF4AF83CB798D9D60E10C4315" bold="true" pageId="0" pageNumber="13">C.</emphasis>
Close up of pronotum punctation and meshed to isodiametric microsculpture;
<emphasis id="DE0474B6533C528251922430AB5BFBF6" bold="true" pageId="0" pageNumber="13">D.</emphasis>
Close up of mesoscutellum and elytra punctation and microsculpture;
<emphasis id="700E7B336BFD31BF44741501539E7181" bold="true" pageId="0" pageNumber="13">E.</emphasis>
Close up of abdominal tergite III with arch-like carina highlighted by white dashed line.
</paragraph>
</caption>
<paragraph id="9C4808F67F8CBC88BD108008CF4A0008" pageId="0" pageNumber="13">
Pronotum longer than wide, PW/PL ratio = 0.9; surface of pronotum with a median impunctate area as wide as 3-5 punctures; with multiple rows of irregular punctures in addition to rows flanking impunctate center; with meshed to isodiametric microsculpture (Fig.
<figureCitation id="B1D01EDD2485020F39CDC76A56732560" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal view of Ikaros navarretei sp. nov. A. Habitus; B. Close up of head punctation and meshed to isodiametric microsculpture; C. Close up of pronotum punctation and meshed to isodiametric microsculpture; D. Close up of mesoscutellum and elytra punctation and microsculpture; E. Close up of abdominal tergite III with arch-like carina highlighted by white dashed line." figureDoi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13">1C</figureCitation>
).
</paragraph>
<paragraph id="3175476DC93F877564AEA4CAF06054AA" pageId="0" pageNumber="13">
Elytra shorter than pronotum, EL/PL ratio = 0.83. Elytra with large, deep punctures, the distance between punctures equals to width of 0.5-2 punctures; only with isodiametric microsculpture at mesoscutellum, disc of elytra with micropunctuation (Fig.
<figureCitation id="D707CBB78674E75B24F4FF5EBBCEDBC5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal view of Ikaros navarretei sp. nov. A. Habitus; B. Close up of head punctation and meshed to isodiametric microsculpture; C. Close up of pronotum punctation and meshed to isodiametric microsculpture; D. Close up of mesoscutellum and elytra punctation and microsculpture; E. Close up of abdominal tergite III with arch-like carina highlighted by white dashed line." figureDoi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13">1D</figureCitation>
).
</paragraph>
<paragraph id="18FB0A0ADF95B41883A760B54347BA67" pageId="0" pageNumber="13">
Abdominal terga III-V with arch-like carina (Fig.
<figureCitation id="E46F3CC40F00499B3FCCB88E03629702" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal view of Ikaros navarretei sp. nov. A. Habitus; B. Close up of head punctation and meshed to isodiametric microsculpture; C. Close up of pronotum punctation and meshed to isodiametric microsculpture; D. Close up of mesoscutellum and elytra punctation and microsculpture; E. Close up of abdominal tergite III with arch-like carina highlighted by white dashed line." figureDoi="10.3897/alpento.6.80349.figure1" httpUri="https://binary.pensoft.net/fig/682340" pageId="0" pageNumber="13">1E</figureCitation>
), arch-like carina on terga IV and V nearly straight. Male secondary sexual structures with very shallow emargination on sternum VII; with deep, broad V-shaped emargination on sternum VIII; borders of emargination on sterna VII and VIII appearing
<normalizedToken id="55645511BEFF38C8380FEC47AFC5BE64" originalValue="shaved">'shaved'</normalizedToken>
(with no setae); lateral tergal sclerites IX subcylindrical and the same length as sternum IX; tergum X subtruncate medio-apically; sternum IX with basal portion symmetrical, about 0.57
<normalizedToken id="F0175C10AED22FF58197E0B1CE1538F3" originalValue="×">x</normalizedToken>
as long as distal portion and subtruncate apically.
</paragraph>
<paragraph id="52DEC5BB655A0C5A1A120A6885CE91FC" pageId="0" pageNumber="13">
Aedeagus as in Fig.
<figureCitation id="B606610314BE2AD13345919E196B93CF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aedeagus of Ikaros navarretei sp. nov. A. Aedeagus in parameral view; B. Paramere in antiparameral view with peg setae; C. Aedeagus in lateral view. Scale bar: 1 mm." figureDoi="10.3897/alpento.6.80349.figure2" httpUri="https://binary.pensoft.net/fig/682341" pageId="0" pageNumber="13">2</figureCitation>
; in parameral view apex of paramere nearly reaching apex of median lobe (Fig.
<figureCitation id="9DCB78157986577053C8E3423CF1D721" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aedeagus of Ikaros navarretei sp. nov. A. Aedeagus in parameral view; B. Paramere in antiparameral view with peg setae; C. Aedeagus in lateral view. Scale bar: 1 mm." figureDoi="10.3897/alpento.6.80349.figure2" httpUri="https://binary.pensoft.net/fig/682341" pageId="0" pageNumber="13">2A</figureCitation>
); paramere broadest at the middle of its length, converging to rounded tip (violin-like shape), characteristic arrangement of peg setae in two sublateral rows containing 4-5 setae with an additional single setae closer to apex (Fig.
<figureCitation id="6B242C9093D518FD95E4971346D4DBF4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aedeagus of Ikaros navarretei sp. nov. A. Aedeagus in parameral view; B. Paramere in antiparameral view with peg setae; C. Aedeagus in lateral view. Scale bar: 1 mm." figureDoi="10.3897/alpento.6.80349.figure2" httpUri="https://binary.pensoft.net/fig/682341" pageId="0" pageNumber="13">2B</figureCitation>
); in lateral view paramere narrower apically; median lobe in dorsal view narrowing to small, rounded apex; in lateral view, median lobe becoming narrower near apex, with one subapical tooth (Fig.
<figureCitation id="9A5C9825EE383FCEB7C10B24B267A55D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aedeagus of Ikaros navarretei sp. nov. A. Aedeagus in parameral view; B. Paramere in antiparameral view with peg setae; C. Aedeagus in lateral view. Scale bar: 1 mm." figureDoi="10.3897/alpento.6.80349.figure2" httpUri="https://binary.pensoft.net/fig/682341" pageId="0" pageNumber="13">2C</figureCitation>
).
</paragraph>
<caption id="8C961B686AFAE3E824E12691BECCCFD3" doi="10.3897/alpento.6.80349.figure2" httpUri="https://binary.pensoft.net/fig/682341" pageId="0" pageNumber="13" start="Figure 2" startId="F2">
<paragraph id="2E322D639EF8C6554D35E82CBF761F84" pageId="0" pageNumber="13">
<emphasis id="AD2614BD523E60C3773276947B6A7D88" bold="true" pageId="0" pageNumber="13">Figure 2.</emphasis>
Aedeagus of
<taxonomicName id="D7A4A62A64CC01A2C0A661FF4871253F" authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis id="B094DFDEF7664AA9E13F183FAD5D551E" italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="413B37877FEE038CA8BF69064C8A8720" bold="true" pageId="0" pageNumber="13">A.</emphasis>
Aedeagus in parameral view;
<emphasis id="7B335AB17553A5CA97FA65B511E29ABF" bold="true" pageId="0" pageNumber="13">B.</emphasis>
Paramere in antiparameral view with peg setae;
<emphasis id="E985682583B0AF1AF3B7ADB9FC74992D" bold="true" pageId="0" pageNumber="13">C.</emphasis>
Aedeagus in lateral view. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="FF49EAC1BB5A6A8CEC13367EC7B58E4A" pageId="0" pageNumber="13" type="distribution">
<paragraph id="BAA03CF5E5F0037C7D491EAF57ACCC86" pageId="0" pageNumber="13">Distribution and habitat.</paragraph>
<paragraph id="7937345FBBE6520D637B5FBAFCC5551A" pageId="0" pageNumber="13">
Known only from the type locality near Silvia in the Cauca Department, Colombia (Fig.
<figureCitation id="5D6BCFE79F589F3C584D61394BCC4D49" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of Ikaros. Elevation colored from green (low) across brown (middle) to white (high). Black square (■) Ikaros apterus. Yellow star (★) Ikaros navarretei sp. nov .. Black circle (●) Ikaros paramo. Black triangle (▲) Ikaros polygonos." figureDoi="10.3897/alpento.6.80349.figure3" httpUri="https://binary.pensoft.net/fig/682342" pageId="0" pageNumber="13">3</figureCitation>
).
</paragraph>
<caption id="9A3823F1AF3C6B6975E40141009EAE6A" doi="10.3897/alpento.6.80349.figure3" httpUri="https://binary.pensoft.net/fig/682342" pageId="0" pageNumber="13" start="Figure 3" startId="F3">
<paragraph id="0B12A3A2E33AC28187D55B89A8C3B1E2" pageId="0" pageNumber="13">
<emphasis id="5D5ADFA38200A1A6FFC6796AF30D79B1" bold="true" pageId="0" pageNumber="13">Figure 3.</emphasis>
Distribution of
<taxonomicName id="910ADCFFE2BC2A60EBFD9915B89F9CD1" genus="Ikaros." lsidName="Ikaros." pageId="0" pageNumber="13" rank="genus">
<emphasis id="7FE62F282C838673C1E54E8FDADD5DA9" italics="true" pageId="0" pageNumber="13">Ikaros.</emphasis>
</taxonomicName>
Elevation colored from green (low) across brown (middle) to white (high). Black square (■)
<taxonomicName id="A0154C7A9597DA3D42E593CEEC0C6052" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros apterus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="apterus">
<emphasis id="1F61DDFFE614A29FD9338930B21A501D" italics="true" pageId="0" pageNumber="13">Ikaros apterus</emphasis>
</taxonomicName>
. Yellow star (★)
<taxonomicName id="2C75A2A84E10B454D07C936C224B575A" authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis id="90F1C102C7661DA32C43BBAA5A32F527" italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
sp. nov.. Black circle (●)
<taxonomicName id="9CEC02F63491F3C2835038A3AAD4911D" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros paramo" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="paramo">
<emphasis id="785F0E4FA6E82ED3C30775CB42FF6EB7" italics="true" pageId="0" pageNumber="13">Ikaros paramo</emphasis>
</taxonomicName>
. Black triangle (▲)
<taxonomicName id="29D080670A054EF2752014D7CBE6896C" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros polygonos" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="polygonos">
<emphasis id="D8774C9CFD12AF90AF8AEFFB9506A0C5" italics="true" pageId="0" pageNumber="13">Ikaros polygonos</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E545B9B784BDF155833C0A3E2822A34D" pageId="0" pageNumber="13" type="notes">
<paragraph id="CFF523FD7DB5D6C4BB74D5F358784BC0" pageId="0" pageNumber="13">Note.</paragraph>
<paragraph id="26C1E46D752E22379DB5B91B22DF428F" pageId="0" pageNumber="13">
In
<taxonomicName id="25E22986FAFD4987835CD95026FEF940" authorityName="Reyes-Hernández &amp; Hansen &amp; Solodovnikov" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros navarretei" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis id="EDA8E52E37BF69FACC2EE70529EFA51D" italics="true" pageId="0" pageNumber="13">Ikaros navarretei</emphasis>
</taxonomicName>
sp. nov. a structure which we call the arch-like carina on the abdominal terga for the sake of compatibility with other literature on
<taxonomicName id="88CA761254E25F11E4ABAFA4CEC86174" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" class="Insecta" family="Staphylinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
, appears to be somewhat similar to a structure usually called in the
<taxonomicName id="64F3882200629D7731AA828AB911FEC9" authorityName="Latreille" authorityYear="1802" lsidName="" pageId="0" pageNumber="13" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
literature as the posterior basal tergal carina (PBTC). This is a very different condition from other species of
<taxonomicName id="03DFA85C64C6A436C6694E43E88B32A4" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Ikaros" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ikaros" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="39AA020032BFA7FD97311E49999C9EE9" italics="true" pageId="0" pageNumber="13">Ikaros</emphasis>
</taxonomicName>
and, as far as we are aware, all other
<taxonomicName id="8700404E0EB6F2D0FA22F831B6E31591" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" class="Insecta" family="Staphylinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
where the arch-like carina is more of a curved fragment that does not reach the spiracles as in the PBTC.
</paragraph>
</subSubSection>
</treatment>
</document>