treatments-xml/data/CA/75/87/CA7587B3FFD2647FF2F9A237FD22719F.xml
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<document id="6CEF3EF45040D616E9489602FDBC581C" ID-DOI="10.11646/zootaxa.5389.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10406827" ID-ZooBank="54C26577-5E99-48E3-9160-F8B02B9A785C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1703001240339" checkinUser="plazi" docAuthor="Morffe, Jans, García, Nayla &amp; Hasegawa, Koichi" docDate="2023" docId="CA7587B3FFD2647FF2F9A237FD22719F" docLanguage="en" docName="zootaxa.5389.2.4.pdf" docOrigin="Zootaxa 5389 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5389.2.4/52511" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Desmicola ryukyuensis Morffe &amp; García &amp; Hasegawa 2023, n. sp." docType="treatment" docVersion="3" lastPageNumber="224" masterDocId="364CFFCBFFD16474F26EA358FF887016" masterDocTitle="Morphological and molecular characterization of Desmicola ryukyuensis n. sp. (Nematoda: Oxyuridomorpha: Thelastomatidae) from the wood-feeding cockroach Panesthia angustipennis yayeyamensis Asahina, 1988 (Blattaria: Blaberidae) in the Ryukyu Archipelago, Japan" masterLastPageNumber="226" masterPageNumber="213" pageNumber="216" updateTime="1703103639246" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="E10725F32F866EBE668D9C539E7B9C82">Morphological and molecular characterization of Desmicola ryukyuensis n. sp. (Nematoda: Oxyuridomorpha: Thelastomatidae) from the wood-feeding cockroach Panesthia angustipennis yayeyamensis Asahina, 1988 (Blattaria: Blaberidae) in the Ryukyu Archipelago, Japan</mods:title>
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<mods:namePart id="3AF7A307ABC6236E297F59F5C359125A">Morffe, Jans</mods:namePart>
<mods:nameIdentifier id="0C73948EF67B377DA70E6AB1949D2D1E" type="ORCID">0000-0002-9968-8129</mods:nameIdentifier>
<mods:affiliation id="8ACBED711D267C5CBBC93FD91D4FC217">Instituto de Ecología y Sistemática, Carretera Varona 11835 e / Oriente y Lindero, La Habana 19, CP 11900, Calabazar, Boyeros, La Habana, Cuba</mods:affiliation>
<mods:nameIdentifier id="2910983F4EA17855A23953E520689792" type="email">koichihasegawa@isc.chubu.ac.jp</mods:nameIdentifier>
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<mods:namePart id="EE493A3EF38140E23FE492C66EFA19B1">García, Nayla</mods:namePart>
<mods:nameIdentifier id="D70746D715E38A911968C74A114AF6C5" type="ORCID">0000-0002-3979-8086</mods:nameIdentifier>
<mods:affiliation id="72492C2DDFA3B87926AD65D561DFDB23">Instituto de Ecología y Sistemática, Carretera Varona 11835 e / Oriente y Lindero, La Habana 19, CP 11900, Calabazar, Boyeros, La Habana, Cuba</mods:affiliation>
<mods:nameIdentifier id="E89B11E62C12FB42F25AF3D8051EB76F" type="email">nayla@ecologia.cu</mods:nameIdentifier>
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<mods:namePart id="4797B337C7D806E41366F2AB21CDDF7C">Hasegawa, Koichi</mods:namePart>
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<mods:nameIdentifier id="E5A53DC4B38E4D59FC22DE910704826D" type="email">koichihasegawa@isc.chubu.ac.jp</mods:nameIdentifier>
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<mods:date id="9F55F4C5AF312B084E846FAB2F599091">2023</mods:date>
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<treatment id="CA7587B3FFD2647FF2F9A237FD22719F" LSID="urn:lsid:plazi:treatment:CA7587B3FFD2647FF2F9A237FD22719F" httpUri="http://treatment.plazi.org/id/CA7587B3FFD2647FF2F9A237FD22719F" lastPageId="11" lastPageNumber="224" pageId="3" pageNumber="216">
<subSubSection id="0AC6652EFFD26477F2F9A237FE70719F" box="[151,504,367,394]" pageId="3" pageNumber="216" type="nomenclature">
<paragraph id="426336A5FFD26477F2F9A237FE70719F" blockId="3.[151,687,367,429]" box="[151,504,367,394]" pageId="3" pageNumber="216">
<heading id="192B81C9FFD26477F2F9A237FE70719F" bold="true" box="[151,504,367,394]" fontSize="11" level="1" pageId="3" pageNumber="216" reason="1">
<emphasis id="70A8EAB7FFD26477F2F9A237FE70719F" bold="true" box="[151,504,367,394]" pageId="3" pageNumber="216">
<taxonomicName id="85DC4D26FFD26477F2F9A237FE27719C" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[151,431,367,394]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="3" pageNumber="216" phylum="Nematoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="70A8EAB7FFD26477F2F9A237FE27719C" bold="true" box="[151,431,367,394]" italics="true" pageId="3" pageNumber="216">Desmicola ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD26477F3D8A237FE70719F" box="[438,504,367,393]" pageId="3" pageNumber="216" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0AC6652EFFD26477F2F9A2CBFD2771BB" box="[151,687,403,429]" pageId="3" pageNumber="216" type="description">
<paragraph id="426336A5FFD26477F2F9A2CBFD2771BB" blockId="3.[151,687,367,429]" box="[151,687,403,429]" pageId="3" pageNumber="216">
<figureCitation id="DAE72A20FFD26477F2F9A2CBFE9971BB" box="[151,273,403,429]" captionStart="FIGURE 1" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[168,1419,181,1914]" captionTargetId="figure-18@5.[168,1419,181,1919]" captionTargetPageId="5" captionText="FIGURE 1. Desmicola ryukyuensis n. sp. Male. A. Oesophageal region, lateral view. B. Cephalic end, optical section. C. Cephalic end (reconstructed from SEM images). D. Tail, lateral view. E. Spicule. F. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10406829" httpUri="https://zenodo.org/record/10406829/files/figure.png" pageId="3" pageNumber="216">Fig. 1 AF</figureCitation>
,
<figureCitation id="DAE72A20FFD26477F372A2CBFE1471BB" box="[284,412,403,429]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2 AH</figureCitation>
,
<figureCitation id="DAE72A20FFD26477F3C9A2CBFDAF71BB" box="[423,551,403,429]" captionStart="FIGURE 3" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[186,1401,181,1914]" captionTargetId="figure-18@7.[186,1401,181,1919]" captionTargetPageId="7" captionText="FIGURE 3. Desmicola ryukyuensis n. sp. Female. A. Oesophageal region, lateral view. B. Cephalic end, optical section. C. Cephalic end (reconstructed from SEM images). D. Tail, lateral view. E. Vagina. F. Genital tract, lateral view. G. Egg. F. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10406833" httpUri="https://zenodo.org/record/10406833/files/figure.png" pageId="3" pageNumber="216">Fig. 3 AH</figureCitation>
,
<figureCitation id="DAE72A20FFD26477F05CA2CBFD2771BB" box="[562,687,403,429]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="3" pageNumber="216">Fig. 4 AK</figureCitation>
</paragraph>
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<subSubSection id="0AC6652EFFD26477F2F9A284FD037274" pageId="3" pageNumber="216" type="materials_examined">
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<emphasis id="70A8EAB7FFD26477F2F9A284FEB271E3" box="[151,314,475,501]" italics="true" pageId="3" pageNumber="216">Type material.</emphasis>
<materialsCitation id="F2B43CF8FFD26477F32CA283FD9C722B" collectingDate="2021-10-24" collectionCode="CZACC" collectorName="H. Nagaya" country="Japan" county="Ryukyu Archipelago" latitude="24.342834" location="Yaeyama District" longLatPrecision="1" longitude="123.912445" municipality="Iriomote Island" pageId="3" pageNumber="216" specimenCode="CZACC 11.7486" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
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:
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<collectingCountry id="3ACB7635FFD26477F3B1A284FDAA71E0" box="[479,546,476,502]" name="Japan" pageId="3" pageNumber="216">Japan</collectingCountry>
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<collectingCounty id="AB024E29FFD26477F040A283FC9271E3" box="[558,794,475,501]" pageId="3" pageNumber="216">Ryukyu Archipelago</collectingCounty>
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<collectingMunicipality id="A207ACDFFFD26477F148A283FC5E71E3" box="[806,982,475,501]" pageId="3" pageNumber="216">Iriomote Island</collectingMunicipality>
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<collectingRegion id="8018F847FFD26477F18DA283FB4F71E3" box="[995,1223,475,501]" country="Japan" name="Okinawa" pageId="3" pageNumber="216">Okinawa Prefecture</collectingRegion>
,
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,
<location id="4703607EFFD26477F2F9A2A7FEC2720C" LSID="urn:lsid:plazi:treatment:CA7587B3FFD2647FF2F9A237FD22719F:4703607EFFD26477F2F9A2A7FEC2720C" box="[151,330,511,538]" country="Japan" county="Ryukyu Archipelago" latitude="24.342834" longLatPrecision="1" longitude="123.912445" municipality="Iriomote Island" name="Taketomi Town" pageId="3" pageNumber="216" stateProvince="Okinawa">Taketomi Town</location>
,
<location id="4703607EFFD26477F33AA2A7FE47720F" LSID="urn:lsid:plazi:treatment:CA7587B3FFD2647FF2F9A237FD22719F:4703607EFFD26477F33AA2A7FE47720F" box="[340,463,511,537]" country="Japan" county="Ryukyu Archipelago" latitude="24.342834" longLatPrecision="1" longitude="123.912445" municipality="Iriomote Island" name="Aira River" pageId="3" pageNumber="216" stateProvince="Okinawa">Aira River</location>
;
<geoCoordinate id="27E85062FFD26477F3B5A2A7FDF1720F" box="[475,633,511,537]" degrees="24" direction="north" minutes="20" orientation="latitude" pageId="3" pageNumber="216" precision="1" seconds="34.2" value="24.342834">24º2034.2”N</geoCoordinate>
,
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; in
<taxonomicName id="85DC4D26FFD26477F137A2A7FA85720F" authorityName="Asahina" authorityYear="1988" box="[857,1293,511,537]" class="Insecta" family="Blaberidae" genus="Panesthia" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="216" phylum="Arthropoda" rank="subSpecies" species="angustipennis" subSpecies="yayeyamensis">
<emphasis id="70A8EAB7FFD26477F137A2A7FA85720F" box="[857,1293,511,537]" italics="true" pageId="3" pageNumber="216">Panesthia angustipennis yayeyamensis</emphasis>
</taxonomicName>
;
<date id="36621065FFD26477F773A2A7FA10720F" box="[1309,1432,511,537]" pageId="3" pageNumber="216" value="2021-10-24">
<collectingDate id="2626E98DFFD26477F773A2A7FA10720F" box="[1309,1432,511,537]" pageId="3" pageNumber="216" value="2021-10-24">24/X/2021</collectingDate>
</date>
;
<collectorName id="EF295373FFD26477F2F9A17CFE867228" box="[151,270,548,574]" pageId="3" pageNumber="216">H. Nagaya</collectorName>
coll.;
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</materialsCitation>
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<materialsCitation id="F2B43CF8FFD26477F073A17CFA8F722B" box="[541,1287,547,574]" collectingDate="2021-10-24" collectionCode="CZACC" collectorName="H. Nagaya" country="Japan" county="Ryukyu Archipelago" latitude="11.7487" location="Yaeyama District" longLatPrecision="7" longitude="-11.7491" municipality="Iriomote Island" pageId="3" pageNumber="216" specimenCount="5" specimenCount-male="5" stateProvince="Okinawa" typeStatus="paratype">
<typeStatus id="9D678807FFD26477F073A17CFD067228" box="[541,654,548,574]" pageId="3" pageNumber="216" type="paratype">Paratypes</typeStatus>
:
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, same data as the latter;
<collectionCode id="24CDAE60FFD26477F1B3A17BFBB2722B" box="[989,1082,547,573]" country="Cuba" httpUri="http://grbio.org/cool/b49z-cz3a" name="Coleccion Zoologia, Academia de Ciencias de Cuba" pageId="3" pageNumber="216">CZACC</collectionCode>
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.
<materialsCitation id="F2B43CF8FFD26477F77EA17BFD0F7274" collectingDate="2021-10-24" collectionCode="CZACC" collectorName="H. Nagaya" country="Japan" county="Ryukyu Archipelago" latitude="11.7492" location="Yaeyama District" longLatPrecision="7" longitude="-11.7505" municipality="Iriomote Island" pageId="3" pageNumber="216" specimenCount="14" specimenCount-female="14" stateProvince="Okinawa" typeStatus="paratype">
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, same data as the latter;
<collectionCode id="24CDAE60FFD26477F333A11FFE327277" box="[349,442,583,609]" country="Cuba" httpUri="http://grbio.org/cool/b49z-cz3a" name="Coleccion Zoologia, Academia de Ciencias de Cuba" pageId="3" pageNumber="216">CZACC</collectionCode>
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.
</paragraph>
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<subSubSection id="0AC6652EFFD2647CF2F9A1C8FA0B761F" lastPageId="8" lastPageNumber="221" pageId="3" pageNumber="216" type="description">
<paragraph id="426336A5FFD26477F2F9A1C8FE2972BF" blockId="3.[151,417,655,681]" box="[151,417,655,681]" pageId="3" pageNumber="216">
<emphasis id="70A8EAB7FFD26477F2F9A1C8FEC972BF" box="[151,321,656,681]" italics="true" pageId="3" pageNumber="216">Measurements.</emphasis>
<tableCitation id="0F5E031EFFD26477F329A1D7FE1372BF" box="[327,411,655,681]" captionStart="TABLE 1" captionStartId="4.[151,239,151,175]" captionTargetPageId="4" captionText="TABLE 1. Morphometrics of Desmicola ryukyuensis n. sp. (Nematoda: Oxyuridomorpha: Thelastomatidae) from Panesthia angustipennis yayeyamensis Asahina, 1988 (Insecta: Blattaria: Blaberidae) from Aira River, Taketomi Town, Yaeyama District, Iriomote Island, Ryukyu Archipelago, Japan. All of the measurements are given in micrometers, unless otherwise indicated, in the form mean ± standard deviation and the range in parentheses. The number of measurements is also given when it is different from the number of specimens (n)." pageId="3" pageNumber="216">Table 1</tableCitation>
.
</paragraph>
<paragraph id="426336A5FFD26477F2F9A180FE9372E7" blockId="3.[151,1437,728,1799]" box="[151,283,728,753]" pageId="3" pageNumber="216">
<emphasis id="70A8EAB7FFD26477F2F9A180FE9372E7" box="[151,283,728,753]" italics="true" pageId="3" pageNumber="216">Description</emphasis>
</paragraph>
<paragraph id="426336A5FFD26477F2F9A1A4FB167711" blockId="3.[151,1437,728,1799]" pageId="3" pageNumber="216">
Male. Body smaller and less robust than that of females; C-shaped in heat-killed specimens. Anterior end truncated. Lateral alae well-developed expanding from
<emphasis id="70A8EAB7FFD26477F0E1A07BFD39732A" box="[655,689,803,828]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
level of 7
<superScript id="B5A99BEDFFD26477F14CA079FCA77339" attach="none" box="[802,815,801,815]" fontSize="6" pageId="3" pageNumber="216">th</superScript>
8
<superScript id="B5A99BEDFFD26477F125A079FCD07339" attach="left" box="[843,856,801,815]" fontSize="6" pageId="3" pageNumber="216">th</superScript>
cuticular annuli posterior to head capsule (
<figureCitation id="DAE72A20FFD26477F752A07AFA1D732A" box="[1340,1429,802,828]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2A</figureCitation>
) to
<emphasis id="70A8EAB7FFD26477F2DDA010FF5D7377" box="[179,213,840,865]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
36 annuli before cloaca (
<figureCitation id="DAE72A20FFD26477F393A01FFDDF7377" box="[509,599,839,865]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2D</figureCitation>
). Head capsule conical, with smooth cuticle (
<figureCitation id="DAE72A20FFD26477F621A01FFB4F7377" box="[1103,1223,839,865]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2A, B</figureCitation>
). Cuticle unarmed, with conspicuous, dilated, wide annuli from base of head capsule to
<emphasis id="70A8EAB7FFD26477F1F1A035FC497390" box="[927,961,877,902]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
level of cloaca (
<figureCitation id="DAE72A20FFD26477F610A034FB747391" box="[1150,1276,876,903]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2A, D</figureCitation>
). Annuli
<emphasis id="70A8EAB7FFD26477F70BA035FA0E7390" box="[1381,1414,877,902]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
9 µm wide next to the head capsule and
<emphasis id="70A8EAB7FFD26477F02AA0CBFDED73BA" box="[580,613,915,940]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
6 µm wide close to cloaca. A series of 79 (mostly 8) modified, ventrally projected annuli present in posterior third of body (
<figureCitation id="DAE72A20FFD26477F0B2A0EFFCBC73C7" box="[732,820,951,977]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2E</figureCitation>
). Ventral projections of modified annuli triangular, the cuticle of their anterior surface presenting a central groove that originates on the tips of the projections (
<figureCitation id="DAE72A20FFD26477F757A084FA0773E1" box="[1337,1423,988,1015]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2F</figureCitation>
). Several paired depressions located at both sides of central groove of each ventral projection (
<figureCitation id="DAE72A20FFD26477F6D8A75AFA84740A" box="[1206,1292,1026,1052]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2F</figureCitation>
). Number of depressions ranging from one pair in shortest projection to four pairs in largest ones (
<figureCitation id="DAE72A20FFD26477F60DA77FFB327457" box="[1123,1210,1063,1089]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2F</figureCitation>
). Mouth hexagonal, with three small triangular lips (
<figureCitation id="DAE72A20FFD26477F06DA714FDD57471" box="[515,605,1100,1127]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2C</figureCitation>
). Four pairs of fused, elongated (
<emphasis id="70A8EAB7FFD26477F1B8A715FC707470" box="[982,1016,1101,1126]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
5 µm in length) sub-median papillae surround the mouth (
<figureCitation id="DAE72A20FFD26477F3EAA72AFE56749A" box="[388,478,1138,1164]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2C</figureCitation>
). Amphids slit-like (
<emphasis id="70A8EAB7FFD26477F0A8A72BFD60749A" box="[710,744,1139,1164]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
2 µm in length), lateral in position (
<figureCitation id="DAE72A20FFD26477F6EBA72AFB56749A" box="[1157,1246,1138,1164]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2C</figureCitation>
). Buccal capsule short, with thickened walls. Oesophagus consisting of muscular, sub-cylindrical corpus, slightly expanded at anterior end and decreasing in diameter towards cylindrical isthmus. Basal bulb rounded, valve-plate well-developed. Cardia small. Intestine simple, sub-rectilinear, its anterior region dilated. Nerve ring encircling corpus at its posterior half,
<emphasis id="70A8EAB7FFD26477F2F9A650FF317537" box="[151,185,1288,1313]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
87% of its length. Excretory pore ventral, located at level of isthmus. Monorchic. Testis ventral, reflexed at
<emphasis id="70A8EAB7FFD26477F2F9A675FF317550" box="[151,185,1325,1350]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
one body-width posterior to basal bulb, distal flexure
<emphasis id="70A8EAB7FFD26477F140A675FCC77550" box="[814,847,1325,1350]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
a body-width long.
<taxonomicName id="85DC4D26FFD26477F655A675FB417550" box="[1083,1225,1324,1350]" class="Insecta" family="Micronoctuidae" genus="Vas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="216" phylum="Arthropoda" rank="species" species="deferens">
<emphasis id="70A8EAB7FFD26477F655A675FB417550" box="[1083,1225,1324,1350]" italics="true" pageId="3" pageNumber="216">Vas deferens</emphasis>
</taxonomicName>
divided into three regions: anterior region filled with rod-like spermatids, median region with large, rounded cells and posterior region with smaller cells, gradually tapering towards its junction with cloaca. Posterior lip of cloaca enlarged, protruded, posteriorly directed. Spicule with distinct, rounded capitulum; shaft straight, thickened at level of first quarter of its length and tapered towards the pointed, ventrally curved tip. Eight copulatory papillae present, two pre-cloacal, two adcloacal and four post-cloacal. Pre-cloacal pair ventromedian, located just anterior to the cloaca, papillae close to each other on top of a conical prominence (
<figureCitation id="DAE72A20FFD26477F0F0A554FC947631" box="[670,796,1548,1575]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2G, H</figureCitation>
). Adcloacal pair of papillae formed by large, conical and prominent papillae, flanking cloaca at lateral sides (
<figureCitation id="DAE72A20FFD26477F0B7A56AFCDD765A" box="[729,853,1586,1612]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2G, H</figureCitation>
). Sensillum of each adcloacal papilla surrounded by
<emphasis id="70A8EAB7FFD26477F2F9A500FF317667" box="[151,185,1624,1649]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
12 small protuberances. First post-cloacal pair consisting of small papillae, situated closely to each other on tip of protruded posterior lip of cloaca (
<figureCitation id="DAE72A20FFD26477F044A524FD2D7681" box="[554,677,1660,1687]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2G, H</figureCitation>
). Second post-cloacal pair consisting of minute papillae, sub-dorsal, located
<emphasis id="70A8EAB7FFD26477F29EA5FBFE9A76AA" box="[240,274,1699,1724]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
at midpoint of caudal filament (
<emphasis id="70A8EAB7FFD26477F0EFA5FBFD2B76AA" box="[641,675,1699,1724]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
40 µm posterior to cloaca) (
<figureCitation id="DAE72A20FFD26477F188A5FAFBCA76AA" box="[998,1090,1698,1724]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2D</figureCitation>
). Tail with the anterior portion conical (
<emphasis id="70A8EAB7FFD26477F296A590FE9276F7" box="[248,282,1736,1761]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
one fifth of the tail length), continuing with filament ending in a sharp tip (
<figureCitation id="DAE72A20FFD26477F60FA59FFB3376F7" box="[1121,1211,1735,1761]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2D</figureCitation>
). Phasmids slit-like, lateral in position, at beginning of caudal filament (
<emphasis id="70A8EAB7FFD26477F0B9A5B5FD717710" box="[727,761,1773,1798]" italics="true" pageId="3" pageNumber="216">ca.</emphasis>
16 µm posterior to cloaca) (
<figureCitation id="DAE72A20FFD26477F659A5B4FB1A7711" box="[1079,1170,1772,1799]" captionStart="FIGURE 2" captionStartId="6.[152,255,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-29@6.[151,1436,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Desmicola ryukyuensis n. sp., SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end, en face view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="3" pageNumber="216">Fig. 2H</figureCitation>
).
</paragraph>
<caption id="16A3662DFFD56470F2F9A3CFFCC67129" pageId="4" pageNumber="217" startId="4.[151,239,151,175]" targetBox="[159,1365,330,1969]" targetIsTable="true" targetPageId="4" targetType="table">
<paragraph id="426336A5FFD56470F2F9A3CFFCC67129" blockId="4.[151,1437,151,319]" pageId="4" pageNumber="217">
<emphasis id="70A8EAB7FFD56470F2F9A3CFFE9970B9" bold="true" box="[151,273,151,175]" pageId="4" pageNumber="217">TABLE 1.</emphasis>
Morphometrics of
<taxonomicName id="85DC4D26FFD56470F39DA3CFFD7970B9" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[499,753,151,175]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="4" pageNumber="217" phylum="Nematoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="70A8EAB7FFD56470F39DA3CFFD7970B9" box="[499,753,151,175]" italics="true" pageId="4" pageNumber="217">Desmicola ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD56470F090A3C0FCB670B9" box="[766,830,152,175]" pageId="4" pageNumber="217" rank="species">n. sp.</taxonomicNameLabel>
(
<taxonomicName id="85DC4D26FFD56470F13CA3CFFC4D70B9" box="[850,965,151,175]" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="225" phylum="Nematoda" rank="phylum">Nematoda</taxonomicName>
: Oxyuridomorpha:
<taxonomicName id="85DC4D26FFD56470F6CCA3CFFADC70B9" authorityName="Travassos" authorityYear="1929" box="[1186,1364,151,175]" class="Chromadorea" family="Thelastomatidae" kingdom="Animalia" order="Rhabditida" pageId="4" pageNumber="217" phylum="Nematoda" rank="family">Thelastomatidae</taxonomicName>
) from
<taxonomicName id="85DC4D26FFD56470F2F9A3E3FD5270C5" authority="Asahina, 1988" authorityName="Asahina" authorityYear="1988" box="[151,730,187,211]" class="Insecta" family="Blaberidae" genus="Panesthia" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="217" phylum="Arthropoda" rank="subSpecies" species="angustipennis" subSpecies="yayeyamensis">
<emphasis id="70A8EAB7FFD56470F2F9A3E3FDB070C5" box="[151,568,187,211]" italics="true" pageId="4" pageNumber="217">Panesthia angustipennis yayeyamensis</emphasis>
Asahina, 1988
</taxonomicName>
(
<taxonomicName id="85DC4D26FFD56470F084A3E3FCB370C5" authorityName="Linnaeus" authorityYear="1758" box="[746,827,187,211]" class="Insecta" kingdom="Animalia" pageId="4" pageNumber="217" phylum="Arthropoda" rank="class">Insecta</taxonomicName>
: Blattaria:
<taxonomicName id="85DC4D26FFD56470F1DFA3E3FBAE70C5" box="[945,1062,187,211]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="217" phylum="Arthropoda" rank="family">Blaberidae</taxonomicName>
) from Aira River, Taketomi Town, Yaeyama District, Iriomote Island, Ryukyu Archipelago, Japan. All of the measurements are given in micrometers, unless otherwise indicated, in the form mean ± standard deviation and the range in parentheses. The number of measurements is also given when it is different from the number of specimens (n).
</paragraph>
</caption>
<paragraph id="426336A5FFD56470F15DA212FAA877A7" pageId="4" pageNumber="217">
<table id="30DCC405FFD59B8BF2F1A212FADD77A7" box="[159,1365,330,1969]" gridcols="4" gridrows="53" pageId="4" pageNumber="217">
<tr id="FCEC34E7FFD59B8BF2F1A212FADD7174" box="[159,1365,330,354]" gridrow="0" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<th id="BF3D5D9BFFD59B8BF15DA212FB987174" box="[819,1040,330,354]" gridcol="2" gridrow="0" pageId="4" pageNumber="217">Males</th>
<th id="BF3D5D9BFFD59B8BF6C4A212FADD7174" box="[1194,1365,330,354]" gridcol="3" gridrow="0" pageId="4" pageNumber="217">Females</th>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A230FADD7196" box="[159,1365,360,384]" gridrow="1" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A230FE4D7196" box="[159,453,360,384]" gridcol="0" gridrow="1" pageId="4" pageNumber="217">
<emphasis id="70A8EAB7FFD56470F35AA230FE2B7196" bold="true" box="[308,419,360,384]" pageId="4" pageNumber="217">Character</emphasis>
</th>
<td id="BF3D5D9BFFD59B8BF015A230FD517196" box="[635,729,360,384]" gridcol="1" gridrow="1" pageId="4" pageNumber="217">Holotype</td>
<td id="BF3D5D9BFFD59B8BF15DA230FB987196" box="[819,1040,360,384]" gridcol="2" gridrow="1" pageId="4" pageNumber="217">Paratypes</td>
<td id="BF3D5D9BFFD59B8BF6C4A230FADD7196" box="[1194,1365,360,384]" gridcol="3" gridrow="1" pageId="4" pageNumber="217">Paratypes</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A2D0FADD71B6" box="[159,1365,392,416]" gridrow="2" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA2D0FB9871B6" box="[819,1040,392,416]" gridcol="2" gridrow="2" pageId="4" pageNumber="217">(n = 5)</td>
<td id="BF3D5D9BFFD59B8BF6C4A2D0FADD71B6" box="[1194,1365,392,416]" gridcol="3" gridrow="2" pageId="4" pageNumber="217">(n = 14)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A2FDFADD71AB" box="[159,1365,421,445]" gridrow="3" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A2FDFE4D71AB" box="[159,453,421,445]" gridcol="0" gridrow="3" pageId="4" pageNumber="217">a</th>
<td id="BF3D5D9BFFD59B8BF015A2FDFD5171AB" box="[635,729,421,445]" gridcol="1" gridrow="3" pageId="4" pageNumber="217">12.0</td>
<td id="BF3D5D9BFFD59B8BF15DA2FDFB9871AB" box="[819,1040,421,445]" gridcol="2" gridrow="3" pageId="4" pageNumber="217">11.5 ± 1.5</td>
<td id="BF3D5D9BFFD59B8BF6C4A2FDFADD71AB" box="[1194,1365,421,445]" gridcol="3" gridrow="3" pageId="4" pageNumber="217">9.9 ± 1.0</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A29DFADD71CB" box="[159,1365,453,477]" gridrow="4" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA29DFB9871CB" box="[819,1040,453,477]" gridcol="2" gridrow="4" pageId="4" pageNumber="217">(8.912.3)</td>
<td id="BF3D5D9BFFD59B8BF6C4A29DFADD71CB" box="[1194,1365,453,477]" gridcol="3" gridrow="4" pageId="4" pageNumber="217">(8.612.0)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A2BDFADD71EB" box="[159,1365,485,509]" gridrow="5" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A2BDFADD71EB" box="[1194,1365,485,509]" gridcol="3" gridrow="5" pageId="4" pageNumber="217">n = 13</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A15BFADD720D" box="[159,1365,515,539]" gridrow="6" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A15BFE4D720D" box="[159,453,515,539]" gridcol="0" gridrow="6" pageId="4" pageNumber="217">b</th>
<td id="BF3D5D9BFFD59B8BF015A15BFD51720D" box="[635,729,515,539]" gridcol="1" gridrow="6" pageId="4" pageNumber="217">4.2</td>
<td id="BF3D5D9BFFD59B8BF15DA15BFB98720D" box="[819,1040,515,539]" gridcol="2" gridrow="6" pageId="4" pageNumber="217">3.9 ± 0.3</td>
<td id="BF3D5D9BFFD59B8BF6C4A15BFADD720D" box="[1194,1365,515,539]" gridcol="3" gridrow="6" pageId="4" pageNumber="217">6.0 ± 0.2</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A17BFADD722D" box="[159,1365,547,571]" gridrow="7" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA17BFB98722D" box="[819,1040,547,571]" gridcol="2" gridrow="7" pageId="4" pageNumber="217">(3.64.4)</td>
<td id="BF3D5D9BFFD59B8BF6C4A17BFADD722D" box="[1194,1365,547,571]" gridcol="3" gridrow="7" pageId="4" pageNumber="217">(5.86.4)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A11BFADD724D" box="[159,1365,579,603]" gridrow="8" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A11BFADD724D" box="[1194,1365,579,603]" gridcol="3" gridrow="8" pageId="4" pageNumber="217">n = 13</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A138FADD726F" box="[159,1365,608,633]" gridrow="9" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A138FE4D726F" box="[159,453,608,633]" gridcol="0" gridrow="9" pageId="4" pageNumber="217">c</th>
<td id="BF3D5D9BFFD59B8BF015A138FD51726F" box="[635,729,608,633]" gridcol="1" gridrow="9" pageId="4" pageNumber="217">9.4</td>
<td id="BF3D5D9BFFD59B8BF15DA138FB98726F" box="[819,1040,608,633]" gridcol="2" gridrow="9" pageId="4" pageNumber="217">9.0 ± 0.9</td>
<td id="BF3D5D9BFFD59B8BF6C4A138FADD726F" box="[1194,1365,608,633]" gridcol="3" gridrow="9" pageId="4" pageNumber="217">2.5 ± 0.1</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A1D9FADD728F" box="[159,1365,641,665]" gridrow="10" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA1D9FB98728F" box="[819,1040,641,665]" gridcol="2" gridrow="10" pageId="4" pageNumber="217">(8.110.0)</td>
<td id="BF3D5D9BFFD59B8BF6C4A1D9FADD728F" box="[1194,1365,641,665]" gridcol="3" gridrow="10" pageId="4" pageNumber="217">(2.42.7)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A1F9FADD72AF" box="[159,1365,673,697]" gridrow="11" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A1F9FADD72AF" box="[1194,1365,673,697]" gridcol="3" gridrow="11" pageId="4" pageNumber="217">n = 13</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A1E6FADD72C0" box="[159,1365,702,726]" gridrow="12" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A1E6FE4D72C0" box="[159,453,702,726]" gridcol="0" gridrow="12" pageId="4" pageNumber="217">V%</th>
<td id="BF3D5D9BFFD59B8BF015A1E6FD5172C0" box="[635,729,702,726]" gridcol="1" gridrow="12" pageId="4" pageNumber="217"></td>
<td id="BF3D5D9BFFD59B8BF15DA1E6FB9872C0" box="[819,1040,702,726]" gridcol="2" gridrow="12" pageId="4" pageNumber="217"></td>
<td id="BF3D5D9BFFD59B8BF6C4A1E6FADD72C0" box="[1194,1365,702,726]" gridcol="3" gridrow="12" pageId="4" pageNumber="217">50.8 ± 1.6</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A186FADD72E0" box="[159,1365,734,758]" gridrow="13" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A186FADD72E0" box="[1194,1365,734,758]" gridcol="3" gridrow="13" pageId="4" pageNumber="217">(48.453.3)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A1A6FADD7300" box="[159,1365,766,790]" gridrow="14" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A1A6FADD7300" box="[1194,1365,766,790]" gridcol="3" gridrow="14" pageId="4" pageNumber="217">n = 13</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A044FADD7322" box="[159,1365,796,820]" gridrow="15" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A044FE4D7322" box="[159,453,796,820]" gridcol="0" gridrow="15" pageId="4" pageNumber="217">V´,%</th>
<td id="BF3D5D9BFFD59B8BF015A044FD517322" box="[635,729,796,820]" gridcol="1" gridrow="15" pageId="4" pageNumber="217"></td>
<td id="BF3D5D9BFFD59B8BF15DA044FB987322" box="[819,1040,796,820]" gridcol="2" gridrow="15" pageId="4" pageNumber="217"></td>
<td id="BF3D5D9BFFD59B8BF6C4A044FADD7322" box="[1194,1365,796,820]" gridcol="3" gridrow="15" pageId="4" pageNumber="217">84.7 ± 1.1</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A064FADD7342" box="[159,1365,828,852]" gridrow="16" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A064FADD7342" box="[1194,1365,828,852]" gridcol="3" gridrow="16" pageId="4" pageNumber="217">(82.286.9)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A004FADD7362" box="[159,1365,860,884]" gridrow="17" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A004FADD7362" box="[1194,1365,860,884]" gridcol="3" gridrow="17" pageId="4" pageNumber="217">n = 13</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A022FADD7384" box="[159,1365,890,914]" gridrow="18" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A022FE4D7384" box="[159,453,890,914]" gridcol="0" gridrow="18" pageId="4" pageNumber="217">Total length (mm)</th>
<td id="BF3D5D9BFFD59B8BF015A022FD517384" box="[635,729,890,914]" gridcol="1" gridrow="18" pageId="4" pageNumber="217">1.10</td>
<td id="BF3D5D9BFFD59B8BF15DA022FB987384" box="[819,1040,890,914]" gridcol="2" gridrow="18" pageId="4" pageNumber="217">0.97 ± 0.09</td>
<td id="BF3D5D9BFFD59B8BF6C4A022FADD7384" box="[1194,1365,890,914]" gridcol="3" gridrow="18" pageId="4" pageNumber="217">1.87 ± 0.07</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A0C2FADD73A4" box="[159,1365,922,946]" gridrow="19" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA0C2FB9873A4" box="[819,1040,922,946]" gridcol="2" gridrow="19" pageId="4" pageNumber="217">(0.861.10)</td>
<td id="BF3D5D9BFFD59B8BF6C4A0C2FADD73A4" box="[1194,1365,922,946]" gridcol="3" gridrow="19" pageId="4" pageNumber="217">(1.731.96)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A0E2FADD73C4" box="[159,1365,954,978]" gridrow="20" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A0E2FADD73C4" box="[1194,1365,954,978]" gridcol="3" gridrow="20" pageId="4" pageNumber="217">n = 13</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A08FFADD73F9" box="[159,1365,983,1007]" gridrow="21" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A08FFE4D73F9" box="[159,453,983,1007]" gridcol="0" gridrow="21" pageId="4" pageNumber="217">Maximum width</th>
<td id="BF3D5D9BFFD59B8BF015A08FFD5173F9" box="[635,729,983,1007]" gridcol="1" gridrow="21" pageId="4" pageNumber="217">85</td>
<td id="BF3D5D9BFFD59B8BF15DA08FFB9873F9" box="[819,1040,983,1007]" gridcol="2" gridrow="21" pageId="4" pageNumber="217">85 ± 14</td>
<td id="BF3D5D9BFFD59B8BF6C4A08FFADD73F9" box="[1194,1365,983,1007]" gridcol="3" gridrow="21" pageId="4" pageNumber="217">189 ± 15</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A0AFFADD7419" box="[159,1365,1015,1039]" gridrow="22" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA0AFFB987419" box="[819,1040,1015,1039]" gridcol="2" gridrow="22" pageId="4" pageNumber="217">(71107)</td>
<td id="BF3D5D9BFFD59B8BF6C4A0AFFADD7419" box="[1194,1365,1015,1039]" gridcol="3" gridrow="22" pageId="4" pageNumber="217">(163210)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A74DFADD743B" box="[159,1365,1045,1069]" gridrow="23" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A74DFE4D743B" box="[159,453,1045,1069]" gridcol="0" gridrow="23" pageId="4" pageNumber="217">Head capsule (length×width)</th>
<td id="BF3D5D9BFFD59B8BF015A74DFD51743B" box="[635,729,1045,1069]" gridcol="1" gridrow="23" pageId="4" pageNumber="217">60 × 56</td>
<td id="BF3D5D9BFFD59B8BF15DA74DFB98743B" box="[819,1040,1045,1069]" gridcol="2" gridrow="23" pageId="4" pageNumber="217">64 ± 2 × 56 ± 3</td>
<td id="BF3D5D9BFFD59B8BF6C4A74DFADD743B" box="[1194,1365,1045,1069]" gridcol="3" gridrow="23" pageId="4" pageNumber="217">20 ± 3 × 54 ± 2</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A76DFADD745B" box="[159,1365,1077,1101]" gridrow="24" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA76DFB98745B" box="[819,1040,1077,1101]" gridcol="2" gridrow="24" pageId="4" pageNumber="217">(6267 × 5259)</td>
<td id="BF3D5D9BFFD59B8BF6C4A76DFADD745B" box="[1194,1365,1077,1101]" gridcol="3" gridrow="24" pageId="4" pageNumber="217">(1826 × 5257)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A70BFADD747D" box="[159,1365,1107,1131]" gridrow="25" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A70BFE4D747D" box="[159,453,1107,1131]" gridcol="0" gridrow="25" pageId="4" pageNumber="217">Buccal cavity length</th>
<td id="BF3D5D9BFFD59B8BF015A70BFD51747D" box="[635,729,1107,1131]" gridcol="1" gridrow="25" pageId="4" pageNumber="217">9</td>
<td id="BF3D5D9BFFD59B8BF15DA70BFB98747D" box="[819,1040,1107,1131]" gridcol="2" gridrow="25" pageId="4" pageNumber="217">9 ± 2</td>
<td id="BF3D5D9BFFD59B8BF6C4A70BFADD747D" box="[1194,1365,1107,1131]" gridcol="3" gridrow="25" pageId="4" pageNumber="217">26 ± 2</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A72BFADD749D" box="[159,1365,1139,1163]" gridrow="26" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA72BFB98749D" box="[819,1040,1139,1163]" gridcol="2" gridrow="26" pageId="4" pageNumber="217">(712)</td>
<td id="BF3D5D9BFFD59B8BF6C4A72BFADD749D" box="[1194,1365,1139,1163]" gridcol="3" gridrow="26" pageId="4" pageNumber="217">(2329)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A7C8FADD74BE" box="[159,1365,1168,1192]" gridrow="27" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A7C8FE4D74BE" box="[159,453,1168,1192]" gridcol="0" gridrow="27" pageId="4" pageNumber="217">Procorpus length</th>
<td id="BF3D5D9BFFD59B8BF015A7C8FD5174BE" box="[635,729,1168,1192]" gridcol="1" gridrow="27" pageId="4" pageNumber="217">176</td>
<td id="BF3D5D9BFFD59B8BF15DA7C8FB9874BE" box="[819,1040,1168,1192]" gridcol="2" gridrow="27" pageId="4" pageNumber="217">178 ± 11</td>
<td id="BF3D5D9BFFD59B8BF6C4A7C8FADD74BE" box="[1194,1365,1168,1192]" gridcol="3" gridrow="27" pageId="4" pageNumber="217">213 ± 9</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A7E8FADD74DE" box="[159,1365,1200,1224]" gridrow="28" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA7E8FB9874DE" box="[819,1040,1200,1224]" gridcol="2" gridrow="28" pageId="4" pageNumber="217">(167193)</td>
<td id="BF3D5D9BFFD59B8BF6C4A7E8FADD74DE" box="[1194,1365,1200,1224]" gridcol="3" gridrow="28" pageId="4" pageNumber="217">(192225)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A796FADD74F0" box="[159,1365,1230,1254]" gridrow="29" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A796FE4D74F0" box="[159,453,1230,1254]" gridcol="0" gridrow="29" pageId="4" pageNumber="217">Isthmus length</th>
<td id="BF3D5D9BFFD59B8BF015A796FD5174F0" box="[635,729,1230,1254]" gridcol="1" gridrow="29" pageId="4" pageNumber="217">24</td>
<td id="BF3D5D9BFFD59B8BF15DA796FB9874F0" box="[819,1040,1230,1254]" gridcol="2" gridrow="29" pageId="4" pageNumber="217">26 ± 2</td>
<td id="BF3D5D9BFFD59B8BF6C4A796FADD74F0" box="[1194,1365,1230,1254]" gridcol="3" gridrow="29" pageId="4" pageNumber="217">30 ± 2</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A7B6FADD7510" box="[159,1365,1262,1286]" gridrow="30" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA7B6FB987510" box="[819,1040,1262,1286]" gridcol="2" gridrow="30" pageId="4" pageNumber="217">(2528)</td>
<td id="BF3D5D9BFFD59B8BF6C4A7B6FADD7510" box="[1194,1365,1262,1286]" gridcol="3" gridrow="30" pageId="4" pageNumber="217">(2734)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A653FADD7532" box="[159,1365,1291,1316]" gridrow="31" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A653FE4D7532" box="[159,453,1291,1316]" gridcol="0" gridrow="31" pageId="4" pageNumber="217">Basal bulb diameter</th>
<td id="BF3D5D9BFFD59B8BF015A653FD517532" box="[635,729,1291,1316]" gridcol="1" gridrow="31" pageId="4" pageNumber="217">42</td>
<td id="BF3D5D9BFFD59B8BF15DA653FB987532" box="[819,1040,1291,1316]" gridcol="2" gridrow="31" pageId="4" pageNumber="217">45 ± 3</td>
<td id="BF3D5D9BFFD59B8BF6C4A653FADD7532" box="[1194,1365,1291,1316]" gridcol="3" gridrow="31" pageId="4" pageNumber="217">75 ± 2</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A674FADD7552" box="[159,1365,1324,1348]" gridrow="32" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA674FB987552" box="[819,1040,1324,1348]" gridcol="2" gridrow="32" pageId="4" pageNumber="217">(4249)</td>
<td id="BF3D5D9BFFD59B8BF6C4A674FADD7552" box="[1194,1365,1324,1348]" gridcol="3" gridrow="32" pageId="4" pageNumber="217">(7179)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A611FADD7577" box="[159,1365,1353,1377]" gridrow="33" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A611FE4D7577" box="[159,453,1353,1377]" gridcol="0" gridrow="33" pageId="4" pageNumber="217">Oesophagus length</th>
<td id="BF3D5D9BFFD59B8BF015A611FD517577" box="[635,729,1353,1377]" gridcol="1" gridrow="33" pageId="4" pageNumber="217">244</td>
<td id="BF3D5D9BFFD59B8BF15DA611FB987577" box="[819,1040,1353,1377]" gridcol="2" gridrow="33" pageId="4" pageNumber="217">249 ± 1</td>
<td id="BF3D5D9BFFD59B8BF6C4A611FADD7577" box="[1194,1365,1353,1377]" gridcol="3" gridrow="33" pageId="4" pageNumber="217">312 ± 11</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A631FADD7597" box="[159,1365,1385,1409]" gridrow="34" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA631FB987597" box="[819,1040,1385,1409]" gridcol="2" gridrow="34" pageId="4" pageNumber="217">(238264)</td>
<td id="BF3D5D9BFFD59B8BF6C4A631FADD7597" box="[1194,1365,1385,1409]" gridcol="3" gridrow="34" pageId="4" pageNumber="217">(289330)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A6DFFADD7589" box="[159,1365,1415,1439]" gridrow="35" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A6DFFE4D7589" box="[159,453,1415,1439]" gridcol="0" gridrow="35" pageId="4" pageNumber="217">Nerve ring-anterior end</th>
<td id="BF3D5D9BFFD59B8BF015A6DFFD517589" box="[635,729,1415,1439]" gridcol="1" gridrow="35" pageId="4" pageNumber="217">165</td>
<td id="BF3D5D9BFFD59B8BF15DA6DFFB987589" box="[819,1040,1415,1439]" gridcol="2" gridrow="35" pageId="4" pageNumber="217">174 ± 10</td>
<td id="BF3D5D9BFFD59B8BF6C4A6DFFADD7589" box="[1194,1365,1415,1439]" gridcol="3" gridrow="35" pageId="4" pageNumber="217">147 ± 12</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A6FFFADD75A9" box="[159,1365,1447,1471]" gridrow="36" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA6FFFB9875A9" box="[819,1040,1447,1471]" gridcol="2" gridrow="36" pageId="4" pageNumber="217">(161184)</td>
<td id="BF3D5D9BFFD59B8BF6C4A6FFFADD75A9" box="[1194,1365,1447,1471]" gridcol="3" gridrow="36" pageId="4" pageNumber="217">(133173)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A69FFADD75C9" box="[159,1365,1479,1503]" gridrow="37" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A69FFADD75C9" box="[1194,1365,1479,1503]" gridcol="3" gridrow="37" pageId="4" pageNumber="217">n = 12</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A6BCFADD75EB" box="[159,1365,1508,1533]" gridrow="38" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A6BCFE4D75EB" box="[159,453,1508,1533]" gridcol="0" gridrow="38" pageId="4" pageNumber="217">Excretory pore-anterior end</th>
<td id="BF3D5D9BFFD59B8BF015A6BCFD5175EB" box="[635,729,1508,1533]" gridcol="1" gridrow="38" pageId="4" pageNumber="217">195</td>
<td id="BF3D5D9BFFD59B8BF15DA6BCFB9875EB" box="[819,1040,1508,1533]" gridcol="2" gridrow="38" pageId="4" pageNumber="217">198 ± 21</td>
<td id="BF3D5D9BFFD59B8BF6C4A6BCFADD75EB" box="[1194,1365,1508,1533]" gridcol="3" gridrow="38" pageId="4" pageNumber="217">257 ± 12</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A55CFADD760B" box="[159,1365,1540,1565]" gridrow="39" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA55CFB98760B" box="[819,1040,1540,1565]" gridcol="2" gridrow="39" pageId="4" pageNumber="217">(181224)</td>
<td id="BF3D5D9BFFD59B8BF6C4A55CFADD760B" box="[1194,1365,1540,1565]" gridcol="3" gridrow="39" pageId="4" pageNumber="217">(228275)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A57AFADD762C" box="[159,1365,1570,1594]" gridrow="40" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A57AFE4D762C" box="[159,453,1570,1594]" gridcol="0" gridrow="40" pageId="4" pageNumber="217">Vulva-anterior end (in mm)</th>
<td id="BF3D5D9BFFD59B8BF015A57AFD51762C" box="[635,729,1570,1594]" gridcol="1" gridrow="40" pageId="4" pageNumber="217"></td>
<td id="BF3D5D9BFFD59B8BF15DA57AFB98762C" box="[819,1040,1570,1594]" gridcol="2" gridrow="40" pageId="4" pageNumber="217"></td>
<td id="BF3D5D9BFFD59B8BF6C4A57AFADD762C" box="[1194,1365,1570,1594]" gridcol="3" gridrow="40" pageId="4" pageNumber="217">0.94 ± 0.04</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A51AFADD764C" box="[159,1365,1602,1626]" gridrow="41" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A51AFADD764C" box="[1194,1365,1602,1626]" gridcol="3" gridrow="41" pageId="4" pageNumber="217">(0.851.01)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A538FADD766E" box="[159,1365,1632,1656]" gridrow="42" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A538FE4D766E" box="[159,453,1632,1656]" gridcol="0" gridrow="42" pageId="4" pageNumber="217">Anus-anterior end (in mm)</th>
<td id="BF3D5D9BFFD59B8BF015A538FD51766E" box="[635,729,1632,1656]" gridcol="1" gridrow="42" pageId="4" pageNumber="217"></td>
<td id="BF3D5D9BFFD59B8BF15DA538FB98766E" box="[819,1040,1632,1656]" gridcol="2" gridrow="42" pageId="4" pageNumber="217"></td>
<td id="BF3D5D9BFFD59B8BF6C4A538FADD766E" box="[1194,1365,1632,1656]" gridcol="3" gridrow="42" pageId="4" pageNumber="217">1.12 ± 0.05</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A5D8FADD768E" box="[159,1365,1664,1688]" gridrow="43" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A5D8FADD768E" box="[1194,1365,1664,1688]" gridcol="3" gridrow="43" pageId="4" pageNumber="217">(1.031.20)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A5F8FADD76AE" box="[159,1365,1696,1720]" gridrow="44" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A5F8FADD76AE" box="[1194,1365,1696,1720]" gridcol="3" gridrow="44" pageId="4" pageNumber="217">n = 13</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A5E5FADD76C0" box="[159,1365,1725,1750]" gridrow="45" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A5E5FE4D76C0" box="[159,453,1725,1750]" gridcol="0" gridrow="45" pageId="4" pageNumber="217">Tail length</th>
<td id="BF3D5D9BFFD59B8BF015A5E5FD5176C0" box="[635,729,1725,1750]" gridcol="1" gridrow="45" pageId="4" pageNumber="217">109</td>
<td id="BF3D5D9BFFD59B8BF15DA5E5FB9876C0" box="[819,1040,1725,1750]" gridcol="2" gridrow="45" pageId="4" pageNumber="217">108 ± 7</td>
<td id="BF3D5D9BFFD59B8BF6C4A5E5FADD76C0" box="[1194,1365,1725,1750]" gridcol="3" gridrow="45" pageId="4" pageNumber="217">747 ± 45</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A585FADD76E0" box="[159,1365,1757,1782]" gridrow="46" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1">
<td id="BF3D5D9BFFD59B8BF15DA585FB9876E0" box="[819,1040,1757,1782]" gridcol="2" gridrow="46" pageId="4" pageNumber="217">(100118)</td>
<td id="BF3D5D9BFFD59B8BF6C4A585FADD76E0" box="[1194,1365,1757,1782]" gridcol="3" gridrow="46" pageId="4" pageNumber="217">(673828)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A5A6FADD7700" box="[159,1365,1790,1814]" gridrow="47" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A5A6FADD7700" box="[1194,1365,1790,1814]" gridcol="3" gridrow="47" pageId="4" pageNumber="217">n = 13</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A443FADD7725" box="[159,1365,1819,1843]" gridrow="48" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A443FE4D7725" box="[159,453,1819,1843]" gridcol="0" gridrow="48" pageId="4" pageNumber="217">Spicule length</th>
<td id="BF3D5D9BFFD59B8BF015A443FD517725" box="[635,729,1819,1843]" gridcol="1" gridrow="48" pageId="4" pageNumber="217">43</td>
<td id="BF3D5D9BFFD59B8BF15DA443FB987725" box="[819,1040,1819,1843]" gridcol="2" gridrow="48" pageId="4" pageNumber="217">42 ± 1</td>
<td id="BF3D5D9BFFD59B8BF6C4A443FADD7725" box="[1194,1365,1819,1843]" gridcol="3" gridrow="48" pageId="4" pageNumber="217"></td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A463FADD7745" box="[159,1365,1851,1875]" gridrow="49" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="BF3D5D9BFFD59B8BF15DA463FB987745" box="[819,1040,1851,1875]" gridcol="2" gridrow="49" pageId="4" pageNumber="217">(4144)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A401FADD7767" box="[159,1365,1881,1905]" gridrow="50" pageId="4" pageNumber="217">
<th id="BF3D5D9BFFD59B8BF2F1A401FE4D7767" box="[159,453,1881,1905]" gridcol="0" gridrow="50" pageId="4" pageNumber="217">Eggs</th>
<td id="BF3D5D9BFFD59B8BF015A401FD517767" box="[635,729,1881,1905]" gridcol="1" gridrow="50" pageId="4" pageNumber="217"></td>
<td id="BF3D5D9BFFD59B8BF15DA401FB987767" box="[819,1040,1881,1905]" gridcol="2" gridrow="50" pageId="4" pageNumber="217"></td>
<td id="BF3D5D9BFFD59B8BF6C4A401FADD7767" box="[1194,1365,1881,1905]" gridcol="3" gridrow="50" pageId="4" pageNumber="217">71 ± 5 × 50 ± 4</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A421FADD7787" box="[159,1365,1913,1937]" gridrow="51" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A421FADD7787" box="[1194,1365,1913,1937]" gridcol="3" gridrow="51" pageId="4" pageNumber="217">(5881 × 4055)</td>
</tr>
<tr id="FCEC34E7FFD59B8BF2F1A4C1FADD77A7" box="[159,1365,1945,1969]" gridrow="52" pageId="4" pageNumber="217" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="BF3D5D9BFFD59B8BF6C4A4C1FADD77A7" box="[1194,1365,1945,1969]" gridcol="3" gridrow="52" pageId="4" pageNumber="217">n = 15</td>
</tr>
</table>
</paragraph>
<caption id="16A3662DFFD46471F2F6A4CFFB4A77C5" ID-DOI="http://doi.org/10.5281/zenodo.10406829" ID-Zenodo-Dep="10406829" httpUri="https://zenodo.org/record/10406829/files/figure.png" pageId="5" pageNumber="218" startId="5.[152,255,1943,1967]" targetBox="[168,1419,181,1914]" targetPageId="5" targetType="figure">
<paragraph id="426336A5FFD46471F2F6A4CFFB4A77C5" blockId="5.[151,1437,1943,2004]" pageId="5" pageNumber="218">
<emphasis id="70A8EAB7FFD46471F2F6A4CFFE9477B9" bold="true" box="[152,284,1943,1967]" pageId="5" pageNumber="218">FIGURE 1.</emphasis>
<taxonomicName id="85DC4D26FFD46471F34BA4CFFD9B77B9" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[293,531,1943,1967]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="5" pageNumber="218" phylum="Nematoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="70A8EAB7FFD46471F34BA4CFFD9B77B9" box="[293,531,1943,1967]" italics="true" pageId="5" pageNumber="218">Desmicola ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD46471F075A4C0FDDC77B9" box="[539,596,1944,1967]" pageId="5" pageNumber="218" rank="species">n. sp.</taxonomicNameLabel>
Male. A. Oesophageal region, lateral view. B. Cephalic end, optical section. C. Cephalic end (reconstructed from SEM images). D. Tail, lateral view. E. Spicule. F. Habitus, lateral view.
</paragraph>
</caption>
<caption id="16A3662DFFD76472F2F6A40AFA1577C0" ID-DOI="http://doi.org/10.5281/zenodo.10406831" ID-Zenodo-Dep="10406831" httpUri="https://zenodo.org/record/10406831/files/figure.png" pageId="6" pageNumber="219" startId="6.[152,255,1874,1898]" targetBox="[151,1436,181,1850]" targetPageId="6" targetType="figure">
<paragraph id="426336A5FFD76472F2F6A40AFA1577C0" blockId="6.[151,1437,1874,2006]" pageId="6" pageNumber="219">
<emphasis id="70A8EAB7FFD76472F2F6A40AFE90777C" bold="true" box="[152,280,1874,1898]" pageId="6" pageNumber="219">FIGURE 2.</emphasis>
<taxonomicName id="85DC4D26FFD76472F373A40AFD8F777C" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[285,519,1874,1898]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="6" pageNumber="219" phylum="Nematoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="70A8EAB7FFD76472F373A40AFD8F777C" box="[285,519,1874,1898]" italics="true" pageId="6" pageNumber="219">Desmicola ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD76472F062A40AFDCC777F" box="[524,580,1874,1897]" pageId="6" pageNumber="219" rank="species">n. sp.</taxonomicNameLabel>
, SEM images. Male. A. Cephalic end and beginning of the lateral ala, lateral view. B. Cephalic end. C. Cephalic end,
<emphasis id="70A8EAB7FFD76472F3BCA42FFD927798" box="[466,538,1910,1934]" italics="true" pageId="6" pageNumber="219">en face</emphasis>
view. D. Tail, lateral view. E. Modified ventrally projected cuticular annuli, ventro-lateral view (anterior end to the top). F. Detail of the modified ventrally projected cuticular annuli, ventral view (anterior end to the top). G. Cloacal region, lateral view. H. Cloacal region, ventral view. Scale bars: A, D. 20 µm. B, E. 10 µm. C. 2 µm. F, G, H. 5 µm.
</paragraph>
</caption>
<caption id="16A3662DFFD66473F2F6A4CFFE9C77E1" ID-DOI="http://doi.org/10.5281/zenodo.10406833" ID-Zenodo-Dep="10406833" httpUri="https://zenodo.org/record/10406833/files/figure.png" pageId="7" pageNumber="220" startId="7.[152,255,1943,1967]" targetBox="[186,1401,181,1914]" targetPageId="7" targetType="figure">
<paragraph id="426336A5FFD66473F2F6A4CFFE9C77E1" blockId="7.[151,1437,1943,2039]" pageId="7" pageNumber="220">
<emphasis id="70A8EAB7FFD66473F2F6A4CFFE9277B9" bold="true" box="[152,282,1943,1967]" pageId="7" pageNumber="220">FIGURE 3.</emphasis>
<taxonomicName id="85DC4D26FFD66473F34CA4CFFD8777B9" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[290,527,1943,1967]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="7" pageNumber="220" phylum="Nematoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="70A8EAB7FFD66473F34CA4CFFD8777B9" box="[290,527,1943,1967]" italics="true" pageId="7" pageNumber="220">Desmicola ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD66473F078A4C0FDC677B9" box="[534,590,1944,1967]" pageId="7" pageNumber="220" rank="species">n. sp.</taxonomicNameLabel>
Female. A. Oesophageal region, lateral view. B. Cephalic end, optical section. C. Cephalic end (reconstructed from SEM images). D. Tail, lateral view. E.
<taxonomicName id="85DC4D26FFD66473F102A4E3FC3E77C5" authorityName="Megerle" authorityYear="1811" box="[876,950,1979,2003]" genus="Vagina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="221" phylum="Mollusca" rank="genus">Vagina</taxonomicName>
. F. Genital tract, lateral view. G. Egg. F. Habitus, lateral view.
</paragraph>
</caption>
<paragraph id="426336A5FFD9647CF2A9A3CFFA0B761F" blockId="8.[151,1437,151,1545]" pageId="8" pageNumber="221">
Female. Body comparatively robust, spindle-shaped, widening gradually from base of semispherical head capsule, body width uniform from base of oesophagus to near level of the vulva, then diminishing towards anus. Cervical cuticle unarmed (
<figureCitation id="DAE72A20FFD9647CF3D1A387FD9170EF" box="[447,537,223,249]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4A</figureCitation>
). Cuticle thick, markedly annulated by retrorse annuli from base of head capsule to just before level of anus (
<figureCitation id="DAE72A20FFD9647CF3BAA25BFDC6710B" box="[468,590,259,285]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4A, B</figureCitation>
). Annuli from base of head capsule to beginning of lateral alae
<emphasis id="70A8EAB7FFD9647CF772A25BFABE710A" box="[1308,1334,259,284]" italics="true" pageId="8" pageNumber="221">ca</emphasis>
. 25 µm wide and set-off each other by deep grooves (
<figureCitation id="DAE72A20FFD9647CF0CAA27FFC897157" box="[676,769,295,321]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4A</figureCitation>
). Annuli of the rest of body wider:
<emphasis id="70A8EAB7FFD9647CF6F7A27FFB3B7156" box="[1177,1203,295,320]" italics="true" pageId="8" pageNumber="221">ca</emphasis>
. 8 µm wide at level of first portion of lateral alae,
<emphasis id="70A8EAB7FFD9647CF383A213FD8F7172" box="[493,519,331,356]" italics="true" pageId="8" pageNumber="221">ca</emphasis>
. 12 µm wide at level of vulva and
<emphasis id="70A8EAB7FFD9647CF1FBA213FC3F7172" box="[917,951,331,356]" italics="true" pageId="8" pageNumber="221">ca.</emphasis>
8 µm wide near level of anus. Lateral alae well-developed from base of corpus (
<emphasis id="70A8EAB7FFD9647CF024A237FDE4719E" box="[586,620,367,392]" italics="true" pageId="8" pageNumber="221">ca.</emphasis>
23 annuli posterior to base of head capsule) terminating at anus level (
<figureCitation id="DAE72A20FFD9647CF2F1A2CBFE9571BB" box="[159,285,403,429]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4A, B</figureCitation>
). Oral opening triangular. Three large, isometric lips surrounding mouth, arranged as one dorsal and two sub-ventral lips, coinciding with sides of oral aperture (
<figureCitation id="DAE72A20FFD9647CF150A2EFFC1D71C7" box="[830,917,439,465]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4F</figureCitation>
). Lips triangular, partially fused at their bases (
<figureCitation id="DAE72A20FFD9647CF2F1A283FF7D71E3" box="[159,245,475,501]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4F</figureCitation>
). Distal tip of each lip bifurcated by a shallow cleavage (
<figureCitation id="DAE72A20FFD9647CF102A283FC4E71E3" box="[876,966,475,501]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4G</figureCitation>
). Complex ornamentations present in lips, consisting of cuticular ridges (
<emphasis id="70A8EAB7FFD9647CF398A2A7FD90720E" box="[502,536,511,536]" italics="true" pageId="8" pageNumber="221">ca.</emphasis>
1 µm wide) with rounded tips (
<figureCitation id="DAE72A20FFD9647CF1E3A2A7FC6E720F" box="[909,998,511,537]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4F</figureCitation>
). Ornamentations of dorsal lip consist of two symmetrical sets, each one with two asymmetrical horseshoe-like and one comma-shaped ridges (
<figureCitation id="DAE72A20FFD9647CF751A17BFA07722B" box="[1343,1423,547,573]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4I</figureCitation>
). Asymmetrical horseshoe-shaped ridges of each set arranged as one with convex side pointing to oral opening and other located dorsally to the first one, next to comma-like ridge with their concave sides facing each other (
<figureCitation id="DAE72A20FFD9647CF72FA133FA077293" box="[1345,1423,619,645]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4I</figureCitation>
). Sub-ventral lips presenting an hourglass-like ornamentation displaced to the ventral side of lips (
<figureCitation id="DAE72A20FFD9647CF6A7A1D7FAAB72BF" box="[1225,1315,655,681]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4H</figureCitation>
). One base of hourglass-like ornamentation (formed by one sigmoid ridge with a horseshoe-shaped ridge dorsal to it) pointing to center of oral opening, the other (formed by one straight ridge flanked by two colon-like ridges) points to basal side of lips (
<figureCitation id="DAE72A20FFD9647CF349A1A3FE0C7303" box="[295,388,763,789]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4H</figureCitation>
). A sigmoid ridge located in each interlabial space between sub-ventral lips and dorsal one. Surface of ornamentations covered in tiny, clustered, round protrusions (
<figureCitation id="DAE72A20FFD9647CF1BCA047FBAC732F" box="[978,1060,799,825]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4J</figureCitation>
). Amphids located at sub-ventral lips, lateral in position, horseshoe-like, their convex side directed to the external margin of the head capsule (
<figureCitation id="DAE72A20FFD9647CF701A01BFF367397" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4H</figureCitation>
). Small sensilla present in each interlabial space (
<figureCitation id="DAE72A20FFD9647CF088A03FFCBE7397" box="[742,822,871,897]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4J</figureCitation>
). Each sensillum presenting a dorsomedian ridge and a distal bifurcation forming two teeth (
<figureCitation id="DAE72A20FFD9647CF05BA0D3FD0E73B3" box="[565,646,907,933]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4J</figureCitation>
). Three large, triangular fang-like structures located in each interlabial space, beneath the sensilla (
<figureCitation id="DAE72A20FFD9647CF3BFA0F7FDA073DF" box="[465,552,943,969]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4F</figureCitation>
). Fang-like structures with two asymmetrical tips at their distal ends (
<figureCitation id="DAE72A20FFD9647CF750A0F7FA0773DF" box="[1342,1423,943,969]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4J</figureCitation>
). Three isomorphic, isometric triangular plates located beneath lips, slightly protrude towards mouth center. Margin of each plate bearing 14 teeth with rounded tips (
<figureCitation id="DAE72A20FFD9647CF0AFA0AFFC947407" box="[705,796,1015,1041]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4G</figureCitation>
). Tooth length
<emphasis id="70A8EAB7FFD9647CF1ABA0AFFC6F7406" box="[965,999,1015,1040]" italics="true" pageId="8" pageNumber="221">ca.</emphasis>
1 µm at both sides of plate, increasing in length to
<emphasis id="70A8EAB7FFD9647CF34DA743FECD7422" box="[291,325,1051,1076]" italics="true" pageId="8" pageNumber="221">ca.</emphasis>
2 µm towards center of plate. Buccal capsule pyriform, wide, notably cuticularized. Oesophagus consists of cylindrical, muscular corpus well set-off from isthmus. Isthmus slightly diminishing in diameter towards junction with rounded basal bulb. Valve-plate of basal bulb well-developed. Cardia well-developed, projecting into intestinal lumen. Intestine simple, sub-rectilinear, its anterior region dilated. Rectum short. Anus a crescent-like slit, its convex side anteriorly directed (
<figureCitation id="DAE72A20FFD9647CF031A7F3FD3274D3" box="[607,698,1195,1221]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4B</figureCitation>
). Posterior lip of anus depressed (
<figureCitation id="DAE72A20FFD9647CF62CA7F3FB1574D3" box="[1090,1181,1195,1221]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4B</figureCitation>
). Two V-like cuticular ridges located anterior to the anus, at level of endpoints of the anal slit, their vertices anteriorly directed (
<figureCitation id="DAE72A20FFD9647CF758A797FA0774FF" box="[1334,1423,1231,1257]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4B</figureCitation>
). Nerve ring encircling procorpus at its posterior half,
<emphasis id="70A8EAB7FFD9647CF093A7ABFC96751A" box="[765,798,1267,1292]" italics="true" pageId="8" pageNumber="221">ca.</emphasis>
65% of its length. Excretory pore ventral, slit-like,
<emphasis id="70A8EAB7FFD9647CF715A7ABFA14751A" box="[1403,1436,1267,1292]" italics="true" pageId="8" pageNumber="221">ca.</emphasis>
18 µm in length, located at level of isthmus, in a discontinuity of a cuticular annulus (
<figureCitation id="DAE72A20FFD9647CF616A64FFB5D7527" box="[1144,1237,1303,1329]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4K</figureCitation>
). Vulva a ventromedian transverse slit,
<emphasis id="70A8EAB7FFD9647CF3F5A663FE357542" box="[411,445,1339,1364]" italics="true" pageId="8" pageNumber="221">ca.</emphasis>
45 µm in length, located
<emphasis id="70A8EAB7FFD9647CF0B1A663FC897542" box="[735,769,1339,1364]" italics="true" pageId="8" pageNumber="221">ca.</emphasis>
midpoint of body (relative to total length of body). Lips of vulva prominent, posterior one hypertrophied, forming a cuticular flap, partially covering the contiguous annulus (
<figureCitation id="DAE72A20FFD9647CF2F1A6DBFF72758B" box="[159,250,1411,1437]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4C</figureCitation>
).
<taxonomicName id="85DC4D26FFD9647CF361A6DBFE1F758A" authorityName="Schallreuter" authorityYear="2001" box="[271,407,1411,1436]" genus="Vagina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="221" phylum="Mollusca" rank="species" species="vera">
<emphasis id="70A8EAB7FFD9647CF361A6DBFE1F758A" box="[271,407,1411,1436]" italics="true" pageId="8" pageNumber="221">Vagina vera</emphasis>
</taxonomicName>
muscular, anteriorly directed. Genital tract didelphic-amphidelphic. Both ovaries reflexed. Oocytes in single rows. Rounded seminal receptacle located between the basal end of anterior ovary and posterior uterus. Eggs broadly oval, with thin and smooth shell. Tail comparatively long,
<emphasis id="70A8EAB7FFD9647CF647A693FBC375F2" box="[1065,1099,1483,1508]" italics="true" pageId="8" pageNumber="221">ca.</emphasis>
half of body length, subulate with filiform terminal part. Phasmids pore-like, lateral in position, located
<emphasis id="70A8EAB7FFD9647CF1B5A6B7FC75761E" box="[987,1021,1519,1544]" italics="true" pageId="8" pageNumber="221">ca.</emphasis>
30 µm posterior to anus (
<figureCitation id="DAE72A20FFD9647CF773A6B7FAFE761F" box="[1309,1398,1519,1545]" captionStart="FIGURE 4" captionStartId="9.[152,255,1900,1924]" captionTargetBox="[178,1409,151,1889]" captionTargetId="figure-18@9.[178,1409,151,1889]" captionTargetPageId="9" captionText="FIGURE 4. Desmicola ryukyuensis n. sp., SEM images. Female.A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end, en face view. G. Oral opening, en face view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm." figureDoi="http://doi.org/10.5281/zenodo.10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="8" pageNumber="221">Fig. 4B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0AC6652EFFD9647EF2F9A56FFE7574D3" lastPageId="10" lastPageNumber="223" pageId="8" pageNumber="221" type="diagnosis">
<paragraph id="426336A5FFD9647CF2F9A56FFE037647" blockId="8.[151,1437,1591,2013]" box="[151,395,1591,1617]" pageId="8" pageNumber="221">
<emphasis id="70A8EAB7FFD9647CF2F9A56FFE037647" box="[151,395,1591,1617]" italics="true" pageId="8" pageNumber="221">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="426336A5FFD9647CF2F9A503FADB7713" blockId="8.[151,1437,1591,2013]" pageId="8" pageNumber="221">
The males of
<taxonomicName id="85DC4D26FFD9647CF346A503FE437663" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[296,459,1627,1653]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFD9647CF346A503FE437663" box="[296,459,1627,1653]" italics="true" pageId="8" pageNumber="221">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD9647CF3A1A503FD807663" box="[463,520,1627,1653]" pageId="8" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
resemble those of
<taxonomicName id="85DC4D26FFD9647CF0BEA503FC017663" authorityName="Sokolova" authorityYear="2019" box="[720,905,1627,1653]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="lamdongensis">
<emphasis id="70A8EAB7FFD9647CF0BEA503FC017663" box="[720,905,1627,1653]" italics="true" pageId="8" pageNumber="221">D. lamdongensis</emphasis>
</taxonomicName>
by having modified ventrally projected cuticular annuli in the posterior third of body, similar body length (
<quantity id="85249B40FFD9647CF178A527FC32768F" box="[790,954,1663,1689]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" metricValueMax="11.0" metricValueMin="8.6" pageId="8" pageNumber="221" unit="mm" value="0.98" valueMax="1.1" valueMin="0.86">0.861.10 mm</quantity>
<emphasis id="70A8EAB7FFD9647CF1AEA527FC56768E" box="[960,990,1663,1688]" italics="true" pageId="8" pageNumber="221">vs.</emphasis>
<quantity id="85249B40FFD9647CF18BA527FB00768F" box="[997,1160,1663,1689]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" metricValueMax="1.29" metricValueMin="0.99" pageId="8" pageNumber="221" unit="mm" value="1.14" valueMax="1.29" valueMin="0.99">0.991.29 mm</quantity>
) and comparative length of the oesophagus (b = 3.64.4
<emphasis id="70A8EAB7FFD9647CF381A5FBFD8676AA" box="[495,526,1699,1724]" italics="true" pageId="8" pageNumber="221">vs.</emphasis>
3.14.2) and tail (c = 8.110.0
<emphasis id="70A8EAB7FFD9647CF10BA5FBFC0B76AA" box="[869,899,1699,1724]" italics="true" pageId="8" pageNumber="221">vs.</emphasis>
6.49.9). They differ by the length of the spicule which is shorter in
<taxonomicName id="85DC4D26FFD9647CF300A59FFD9C76F7" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[366,532,1735,1761]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFD9647CF300A59FFD9C76F7" box="[366,532,1735,1761]" italics="true" pageId="8" pageNumber="221">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD9647CF075A59FFDDF76F7" box="[539,599,1735,1761]" pageId="8" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
(4144 µm
<emphasis id="70A8EAB7FFD9647CF08CA59FFC8876F6" box="[738,768,1735,1760]" italics="true" pageId="8" pageNumber="221">vs.</emphasis>
4855 µm). The lateral alae of
<taxonomicName id="85DC4D26FFD9647CF60DA59FFA8176F7" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[1123,1289,1735,1761]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFD9647CF60DA59FFA8176F7" box="[1123,1289,1735,1761]" italics="true" pageId="8" pageNumber="221">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD9647CF77EA59FFAC476F7" box="[1296,1356,1735,1761]" pageId="8" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
begins at the level of the 7
<superScript id="B5A99BEDFFD9647CF301A5B2FEF476EE" attach="none" box="[367,380,1770,1784]" fontSize="6" pageId="8" pageNumber="221">th</superScript>
8
<superScript id="B5A99BEDFFD9647CF3F9A5B2FE2C76EE" attach="left" box="[407,420,1770,1784]" fontSize="6" pageId="8" pageNumber="221">th</superScript>
annuli posterior to the cephalic capsule, rather than at the 4
<superScript id="B5A99BEDFFD9647CF62FA5B2FBC676EE" attach="left" box="[1089,1102,1770,1784]" fontSize="6" pageId="8" pageNumber="221">th</superScript>
as in
<emphasis id="70A8EAB7FFD9647CF6E1A5B3FADB7713" box="[1167,1363,1771,1797]" italics="true" pageId="8" pageNumber="221">
<taxonomicName id="85DC4D26FFD9647CF6E1A5B3FAD87713" authorityName="Sokolova" authorityYear="2019" box="[1167,1360,1771,1797]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="lamdongensis">D. lamdongensis</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="426336A5FFD9647EF2A9A457FE6170EF" blockId="8.[151,1437,1591,2013]" lastBlockId="10.[151,1437,150,1221]" lastPageId="10" lastPageNumber="223" pageId="8" pageNumber="221">
The spicule of
<taxonomicName id="85DC4D26FFD9647CF303A457FD9B773F" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[365,531,1807,1833]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFD9647CF303A457FD9B773F" box="[365,531,1807,1833]" italics="true" pageId="8" pageNumber="221">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD9647CF074A457FDDE773F" box="[538,598,1807,1833]" pageId="8" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
is shorter than
<taxonomicName id="85DC4D26FFD9647CF16FA457FC3F773F" authorityName="Hugot, Morand &amp; Van-Waerebeke" authorityYear="1993" box="[769,951,1807,1833]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="nhatrangenis">
<emphasis id="70A8EAB7FFD9647CF16FA457FC3F773F" box="[769,951,1807,1833]" italics="true" pageId="8" pageNumber="221">D. nhatrangenis</emphasis>
</taxonomicName>
and
<taxonomicName id="85DC4D26FFD9647CF182A457FB20773F" authorityName="Hugot, Morand &amp; Van Waerebeke" authorityYear="1991" box="[1004,1192,1807,1833]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="kinabaluensis">
<emphasis id="70A8EAB7FFD9647CF182A457FB20773F" box="[1004,1192,1807,1833]" italics="true" pageId="8" pageNumber="221">D. kinabaluensis</emphasis>
</taxonomicName>
(4144 µm
<emphasis id="70A8EAB7FFD9647CF75FA457FAD8773E" box="[1329,1360,1807,1832]" italics="true" pageId="8" pageNumber="221">vs.</emphasis>
4551 µm
<emphasis id="70A8EAB7FFD9647CF2ABA46BFF6C775A" box="[197,228,1843,1868]" italics="true" pageId="8" pageNumber="221">vs.</emphasis>
6265 µm) and its distal end is pointed
<emphasis id="70A8EAB7FFD9647CF0DEA46BFD46775A" box="[688,718,1843,1868]" italics="true" pageId="8" pageNumber="221">vs.</emphasis>
bifurcated in the other two species. Both
<taxonomicName id="85DC4D26FFD9647CF6C4A46BFAE4775B" authorityName="Hugot, Morand &amp; Van Waerebeke" authorityYear="1991" box="[1194,1388,1843,1869]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="nhatrangensis">
<emphasis id="70A8EAB7FFD9647CF6C4A46BFAE4775B" box="[1194,1388,1843,1869]" italics="true" pageId="8" pageNumber="221">D. nhatrangensis</emphasis>
</taxonomicName>
and
<taxonomicName id="85DC4D26FFD9647CF2F9A40FFEDC7767" authorityName="Hugot, Morand &amp; Van Waerebeke" authorityYear="1991" box="[151,340,1879,1905]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="kinabaluensis">
<emphasis id="70A8EAB7FFD9647CF2F9A40FFEDC7767" box="[151,340,1879,1905]" italics="true" pageId="8" pageNumber="221">D. kinabaluensis</emphasis>
</taxonomicName>
males possess a mamelon in the last third of the body (
<bibRefCitation id="264D4B54FFD9647CF1A9A40FFB187767" author="Hugot, J. P. &amp; Morand, S. &amp; Van Waerebeke, D." box="[967,1168,1879,1905]" pageId="8" pageNumber="221" pagination="65 - 74" refId="ref7869" refString="Hugot, J. P., Morand, S. &amp; Van Waerebeke, D. (1991) On genus Desmicola (Nematoda, Thelastomatidae) with description of two new species. Research and Reviews in Parasitology, 51 (1 - 4), 65 - 74." type="journal article" year="1991">
Hugot
<emphasis id="70A8EAB7FFD9647CF679A40FFBD87767" box="[1047,1104,1879,1905]" italics="true" pageId="8" pageNumber="221">et al.</emphasis>
1991
</bibRefCitation>
) consisting of ventrally dilated annuli with a transversal groove in the middle. The latter feature differentiates them from the males of
<taxonomicName id="85DC4D26FFD9647CF2F9A4C7FEC877AF" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[151,320,1951,1977]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFD9647CF2F9A4C7FEC877AF" box="[151,320,1951,1977]" italics="true" pageId="8" pageNumber="221">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD9647CF325A4C7FE0377AF" box="[331,395,1951,1977]" pageId="8" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
, with their modified ventrally projected cuticular annuli triangular in form. The tail of
<taxonomicName id="85DC4D26FFD9647CF7EFA4C7FE9277CB" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFD9647CF7EFA4C7FE9277CB" italics="true" pageId="8" pageNumber="221">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD9647CF34FA49BFED477CB" box="[289,348,1987,2013]" pageId="8" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
is comparatively shorter than in
<taxonomicName id="85DC4D26FFD9647CF0A7A49BFC0077CB" authorityName="Hugot, Morand &amp; Van Waerebeke" authorityYear="1991" box="[713,904,1987,2013]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="221" phylum="Nematoda" rank="species" species="nhatrangensis">
<emphasis id="70A8EAB7FFD9647CF0A7A49BFC0077CB" box="[713,904,1987,2013]" italics="true" pageId="8" pageNumber="221">D. nhatrangensis</emphasis>
</taxonomicName>
(c = 8.110.0
<emphasis id="70A8EAB7FFD9647CF644A49BFBC077CA" box="[1066,1096,1987,2012]" italics="true" pageId="8" pageNumber="221">vs.</emphasis>
2.74.4). Moreover, the males of
<taxonomicName id="85DC4D26FFDB647EF2D9A3CFFEFD70A7" authorityName="Hugot, Morand &amp; Van Waerebeke" authorityYear="1991" box="[183,373,151,177]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="kinabaluensis">
<emphasis id="70A8EAB7FFDB647EF2D9A3CFFEFD70A7" box="[183,373,151,177]" italics="true" pageId="10" pageNumber="223">D. kinabaluensis</emphasis>
</taxonomicName>
differs from the new species proposed herein by its longer (
<quantity id="85249B40FFDB647EF649A3CEFB4670A7" box="[1063,1230,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.475" metricValueMax="1.57" metricValueMin="1.38" pageId="10" pageNumber="223" unit="mm" value="1.475" valueMax="1.57" valueMin="1.38">1.381.57 mm</quantity>
<emphasis id="70A8EAB7FFDB647EF6B8A3CFFB7C70A6" box="[1238,1268,151,176]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
<quantity id="85249B40FFDB647EF693A3CFFA1C70A7" box="[1277,1428,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" metricValueMax="11.0" metricValueMin="8.6" pageId="10" pageNumber="223" unit="mm" value="0.98" valueMax="1.1" valueMin="0.86">0.861.1 mm</quantity>
) and less robust (a = 15.716.8
<emphasis id="70A8EAB7FFDB647EF39AA3E3FD9B70C2" box="[500,531,187,212]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
8.912.3) body, the shorter oesophagus (b = 4.55.2
<emphasis id="70A8EAB7FFDB647EF601A3E3FB0670C2" box="[1135,1166,187,212]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
3.64.4) and the longer tail (c = 4.04.6
<emphasis id="70A8EAB7FFDB647EF33EA387FEE770EE" box="[336,367,223,248]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
8.110.0).
</paragraph>
<caption id="16A3662DFFD8647DF2F6A434FBAF77E7" ID-DOI="http://doi.org/10.5281/zenodo.10406835" ID-Zenodo-Dep="10406835" httpUri="https://zenodo.org/record/10406835/files/figure.png" pageId="9" pageNumber="222" startId="9.[152,255,1900,1924]" targetBox="[178,1409,151,1889]" targetPageId="9" targetType="figure">
<paragraph id="426336A5FFD8647DF2F6A434FBAF77E7" blockId="9.[151,1437,1900,2033]" pageId="9" pageNumber="222">
<emphasis id="70A8EAB7FFD8647DF2F6A434FE907792" bold="true" box="[152,280,1900,1924]" pageId="9" pageNumber="222">FIGURE 4.</emphasis>
<taxonomicName id="85DC4D26FFD8647DF373A434FD807792" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[285,520,1900,1924]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="9" pageNumber="222" phylum="Nematoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="70A8EAB7FFD8647DF373A434FD807792" box="[285,520,1900,1924]" italics="true" pageId="9" pageNumber="222">Desmicola ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFD8647DF063A435FDCC7792" box="[525,580,1901,1924]" pageId="9" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
, SEM images. Female. A. Cervical region, lateral view. B. Anus region and phasmid, ventro-lateral view. C. Vulva, lateral view. D, E. Cephalic end, ventro-lateral view. F. Cephalic end,
<emphasis id="70A8EAB7FFD8647DF6D7A4C9FA8C77BE" box="[1209,1284,1936,1960]" italics="true" pageId="9" pageNumber="222">en face</emphasis>
view. G. Oral opening,
<emphasis id="70A8EAB7FFD8647DF29BA4EDFEB477DA" box="[245,316,1972,1996]" italics="true" pageId="9" pageNumber="222">en face</emphasis>
view. H. Sub-ventral lip. I. Dorsal lip. J. Sensilla and fang-like structure at interlabial space. K. Excretory pore, ventral view. Scale bars: A. 50 µm. B, C, D, E. 10 µm. F, I, K. 5 µm. G, H. 2 µm. J. 1 µm.
</paragraph>
</caption>
<paragraph id="426336A5FFDB647EF2A9A25BFEC87173" blockId="10.[151,1437,150,1221]" pageId="10" pageNumber="223">
The tail of the males of
<taxonomicName id="85DC4D26FFDB647EF3A1A25BFDFC710B" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[463,628,259,285]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF3A1A25BFDFC710B" box="[463,628,259,285]" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF015A25BFD3E710B" box="[635,694,259,285]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
is comparatively shorter than that of
<taxonomicName id="85DC4D26FFDB647EF63BA25BFB4D710B" authorityName="Spiridonov &amp; Ivanova" authorityYear="1998" box="[1109,1221,259,285]" class="Chromadorea" family="Thelastomatidae" genus="Posterovulva" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="danieli">
<emphasis id="70A8EAB7FFDB647EF63BA25BFB4D710B" box="[1109,1221,259,285]" italics="true" pageId="10" pageNumber="223">D. danieli</emphasis>
</taxonomicName>
and
<taxonomicName id="85DC4D26FFDB647EF694A25BFA14710A" authorityName="Van Waerebeke" authorityYear="1969" box="[1274,1436,259,284]" class="Chromadorea" family="Thelastomatidae" genus="Posterovulva" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="moramangi">
<emphasis id="70A8EAB7FFDB647EF694A25BFA14710A" box="[1274,1436,259,284]" italics="true" pageId="10" pageNumber="223">D. moramangi</emphasis>
</taxonomicName>
(c = 8.110.0
<emphasis id="70A8EAB7FFDB647EF35BA27FFEDB7156" box="[309,339,295,320]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
6.86.9
<emphasis id="70A8EAB7FFDB647EF3DBA27FFE5C7156" box="[437,468,295,320]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
4.05.4). Also, the spicule of
<taxonomicName id="85DC4D26FFDB647EF149A27FFC457157" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[807,973,295,321]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF149A27FFC457157" box="[807,973,295,321]" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF1BBA27FFB997157" box="[981,1041,295,321]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
is longer than in
<taxonomicName id="85DC4D26FFDB647EF6BBA27FFACE7157" authorityName="Spiridonov &amp; Ivanova" authorityYear="1998" box="[1237,1350,295,321]" class="Chromadorea" family="Thelastomatidae" genus="Posterovulva" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="danieli">
<emphasis id="70A8EAB7FFDB647EF6BBA27FFACE7157" box="[1237,1350,295,321]" italics="true" pageId="10" pageNumber="223">D. danieli</emphasis>
</taxonomicName>
(4144 µm
<emphasis id="70A8EAB7FFDB647EF2AAA213FF6B7172" box="[196,227,331,356]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
3841).
</paragraph>
<paragraph id="426336A5FFDB647EF2A9A237FD9072BF" blockId="10.[151,1437,150,1221]" pageId="10" pageNumber="223">
The females of
<taxonomicName id="85DC4D26FFDB647EF31DA237FD90719F" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[371,536,367,393]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF31DA237FD90719F" box="[371,536,367,393]" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF071A237FDD2719F" box="[543,602,367,393]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
are similar to
<taxonomicName id="85DC4D26FFDB647EF099A237FCEB719E" authorityName="Jex, Schneider, Rose &amp; Cribb" authorityYear="2005" box="[759,867,367,392]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ornata">
<emphasis id="70A8EAB7FFDB647EF099A237FCEB719E" box="[759,867,367,392]" italics="true" pageId="10" pageNumber="223">D. ornata</emphasis>
</taxonomicName>
by body length (
<quantity id="85249B40FFDB647EF673A237FB48719F" box="[1053,1216,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.97" metricValueMin="1.73" pageId="10" pageNumber="223" unit="mm" value="1.85" valueMax="1.97" valueMin="1.73">1.731.97 mm</quantity>
<emphasis id="70A8EAB7FFDB647EF6A8A237FB6C719E" box="[1222,1252,367,392]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
<quantity id="85249B40FFDB647EF684A237FA18719F" box="[1258,1424,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" metricValueMax="2.09" metricValueMin="1.77" pageId="10" pageNumber="223" unit="mm" value="1.93" valueMax="2.09" valueMin="1.77">1.772.09 mm</quantity>
), the robustness of the body (a = 8.612.0
<emphasis id="70A8EAB7FFDB647EF00BA2CBFD0C71BA" box="[613,644,403,428]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
9.817.1), the comparative lengths of the oesophagus (b = 5.86.4
<emphasis id="70A8EAB7FFDB647EF710A2CBFA1471BA" box="[1406,1436,403,428]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
5.77.4) and tail (c = 2.42.7
<emphasis id="70A8EAB7FFDB647EF38CA2EFFD8871C6" box="[482,512,439,464]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
2.33.9) as well as the position of the vulva (V% = 48.453.3
<emphasis id="70A8EAB7FFDB647EF6D4A2EFFB5071C6" box="[1210,1240,439,464]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
48.068.0). Both species feature triangular lips partially fused at their bases with their distal tips bifurcated. The ornamentations of the lips are similar in
<taxonomicName id="85DC4D26FFDB647EF3E4A2A7FDB8720F" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[394,560,511,537]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF3E4A2A7FDB8720F" box="[394,560,511,537]" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF056A2A7FDFC720F" box="[568,628,511,537]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
and
<emphasis id="70A8EAB7FFDB647EF0C4A2A7FC96720E" box="[682,798,511,536]" italics="true" pageId="10" pageNumber="223">
<taxonomicName id="85DC4D26FFDB647EF0C4A2A7FC92720E" authorityName="Jex, Schneider, Rose &amp; Cribb" authorityYear="2005" box="[682,794,511,536]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ornata">D. ornata</taxonomicName>
.
</emphasis>
They present the fang-like double-tipped structure in the interlabial spaces and three triangular plates beneath the lips with teeth at their margins.
<taxonomicName id="85DC4D26FFDB647EF612A17BFAF7722B" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[1148,1407,547,573]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="70A8EAB7FFDB647EF612A17BFAF7722B" box="[1148,1407,547,573]" italics="true" pageId="10" pageNumber="223">Desmicola ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF7E9A17BFF3F7277" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="85DC4D26FFDB647EF322A11FFE327276" authorityName="Jex, Schneider, Rose &amp; Cribb" authorityYear="2005" box="[332,442,583,608]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ornata">
<emphasis id="70A8EAB7FFDB647EF322A11FFE327276" box="[332,442,583,608]" italics="true" pageId="10" pageNumber="223">D. ornata</emphasis>
</taxonomicName>
by the shape of the small sensilla in the interlabial space, with the distal tip bifurcate
<emphasis id="70A8EAB7FFDB647EF710A11FFA147276" box="[1406,1436,583,608]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
not bifurcate. Lateral alae are absent in
<taxonomicName id="85DC4D26FFDB647EF03AA133FD4A7292" authorityName="Jex, Schneider, Rose &amp; Cribb" authorityYear="2005" box="[596,706,619,644]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ornata">
<emphasis id="70A8EAB7FFDB647EF03AA133FD4A7292" box="[596,706,619,644]" italics="true" pageId="10" pageNumber="223">D. ornata</emphasis>
</taxonomicName>
and in
<taxonomicName id="85DC4D26FFDB647EF178A133FC347293" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[790,956,619,645]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF178A133FC347293" box="[790,956,619,645]" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF1AAA133FB887293" box="[964,1024,619,645]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
they are present, from to the base of the corpus to the level of the anus.
</paragraph>
<paragraph id="426336A5FFDB647EF2A9A1EBFB297397" blockId="10.[151,1437,150,1221]" pageId="10" pageNumber="223">
The females of
<taxonomicName id="85DC4D26FFDB647EF313A1EBFDAE72DB" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[381,550,691,717]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF313A1EBFDAE72DB" box="[381,550,691,717]" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF041A1EBFDE572DB" box="[559,621,691,717]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
are also similar to
<taxonomicName id="85DC4D26FFDB647EF121A1EBFB8572DB" authorityName="Sokolova" authorityYear="2019" box="[847,1037,691,717]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="lamdongensis">
<emphasis id="70A8EAB7FFDB647EF121A1EBFB8572DB" box="[847,1037,691,717]" italics="true" pageId="10" pageNumber="223">D. lamdongensis</emphasis>
</taxonomicName>
in body length (
<quantity id="85249B40FFDB647EF6A0A1EBFAFD72DB" box="[1230,1397,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.97" metricValueMin="1.73" pageId="10" pageNumber="223" unit="mm" value="1.85" valueMax="1.97" valueMin="1.73">1.731.97 mm</quantity>
<emphasis id="70A8EAB7FFDB647EF710A1EBFA1472DA" box="[1406,1436,691,716]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
<quantity id="85249B40FFDB647EF2F9A18FFEB772E7" box="[151,319,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" metricValueMax="2.36" metricValueMin="1.78" pageId="10" pageNumber="223" unit="mm" value="2.07" valueMax="2.36" valueMin="1.78">1.782.36 mm</quantity>
) and comparative lengths of the oesophagus (b = 5.86.4
<emphasis id="70A8EAB7FFDB647EF19EA18FFB8672E6" box="[1008,1038,727,752]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
5.16.6) and tail (c = 2.42.7
<emphasis id="70A8EAB7FFDB647EF710A18FFA1472E6" box="[1406,1436,727,752]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
2.12.6). They feature horseshoe-like amphids and bifurcate sensilla at the interlabial space. Both taxa can be differentiated by the presence of fused lips in
<emphasis id="70A8EAB7FFDB647EF0FCA047FCFA732E" box="[658,882,799,825]" italics="true" pageId="10" pageNumber="223">
<taxonomicName id="85DC4D26FFDB647EF0FCA047FCC5732F" authorityName="Sokolova" authorityYear="2019" box="[658,845,799,825]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="lamdongensis">D. lamdongensis</taxonomicName>
vs.
</emphasis>
partially fused lips in the new species.
<taxonomicName id="85DC4D26FFDB647EF74AA047FEB9734B" authorityName="Sokolova" authorityYear="2019" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="lamdongensis">
<emphasis id="70A8EAB7FFDB647EF74AA047FEB9734B" italics="true" pageId="10" pageNumber="223">Desmicola lamdongensis</emphasis>
</taxonomicName>
lacks the double-tipped fang-like structures at the interlabial spaces and the toothed plates. The eggs of
<taxonomicName id="85DC4D26FFDB647EF2D8A03FFED47397" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[182,348,871,897]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF2D8A03FFED47397" box="[182,348,871,897]" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF30DA03FFE177397" box="[355,415,871,897]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
are smaller than
<taxonomicName id="85DC4D26FFDB647EF00EA03FFC947397" authorityName="Sokolova" authorityYear="2019" box="[608,796,871,897]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="lamdongensis">
<emphasis id="70A8EAB7FFDB647EF00EA03FFC947397" box="[608,796,871,897]" italics="true" pageId="10" pageNumber="223">D. lamdongensis</emphasis>
</taxonomicName>
(5881×4055
<emphasis id="70A8EAB7FFDB647EF1A1A03FFC657396" box="[975,1005,871,896]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
8898×5365).
</paragraph>
<paragraph id="426336A5FFDB647EF2A9A0D3FE7574D3" blockId="10.[151,1437,150,1221]" pageId="10" pageNumber="223">
The body of
<taxonomicName id="85DC4D26FFDB647EF330A0D3FD8E73B3" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[350,518,907,933]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF330A0D3FD8E73B3" box="[350,518,907,933]" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF061A0D3FDC673B3" box="[527,590,907,933]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
is shorter than the rest of the species of the genus, namely
<taxonomicName id="85DC4D26FFDB647EF691A0D3FAF973B3" authorityName="Spiridonov &amp; Ivanova" authorityYear="1998" box="[1279,1393,907,933]" class="Chromadorea" family="Thelastomatidae" genus="Posterovulva" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="danieli">
<emphasis id="70A8EAB7FFDB647EF691A0D3FAF973B3" box="[1279,1393,907,933]" italics="true" pageId="10" pageNumber="223">D. danieli</emphasis>
</taxonomicName>
,
<taxonomicName id="85DC4D26FFDB647EF7EFA0D3FEB973DF" authorityName="Hugot, Morand &amp; Van Waerebeke" authorityYear="1991" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="kinabaluensis">
<emphasis id="70A8EAB7FFDB647EF7EFA0D3FEB973DF" italics="true" pageId="10" pageNumber="223">D. kinabaluensis</emphasis>
</taxonomicName>
,
<taxonomicName id="85DC4D26FFDB647EF32FA0F7FE2B73DF" authorityName="Skrjabin" authorityYear="1916" box="[321,419,943,969]" class="Chromadorea" family="Oxyuridae" genus="Oxyuris" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="leidyi">
<emphasis id="70A8EAB7FFDB647EF32FA0F7FE2B73DF" box="[321,419,943,969]" italics="true" pageId="10" pageNumber="223">D. leidyi</emphasis>
</taxonomicName>
,
<taxonomicName id="85DC4D26FFDB647EF3DDA0F7FDDF73DE" authorityName="Van Waerebeke" authorityYear="1969" box="[435,599,943,968]" class="Chromadorea" family="Thelastomatidae" genus="Posterovulva" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="moramangi">
<emphasis id="70A8EAB7FFDB647EF3DDA0F7FDDF73DE" box="[435,599,943,968]" italics="true" pageId="10" pageNumber="223">D. moramangi</emphasis>
</taxonomicName>
,
<taxonomicName id="85DC4D26FFDB647EF009A0F7FCA273DF" authorityName="Hugot, Morand &amp; Van Waerebeke" authorityYear="1991" box="[615,810,943,969]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="nhatrangensis">
<emphasis id="70A8EAB7FFDB647EF009A0F7FCA273DF" box="[615,810,943,969]" italics="true" pageId="10" pageNumber="223">D. nhatrangensis</emphasis>
</taxonomicName>
and
<taxonomicName id="85DC4D26FFDB647EF10AA0F7FC6473DF" authorityName="Adamson" authorityYear="1984" box="[868,1004,943,969]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="skrjabini">
<emphasis id="70A8EAB7FFDB647EF10AA0F7FC6473DF" box="[868,1004,943,969]" italics="true" pageId="10" pageNumber="223">D. skrjabini</emphasis>
</taxonomicName>
(
<quantity id="85249B40FFDB647EF192A0F7FB2A73DF" box="[1020,1186,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.97" metricValueMin="1.73" pageId="10" pageNumber="223" unit="mm" value="1.85" valueMax="1.97" valueMin="1.73">1.731.97 mm</quantity>
<emphasis id="70A8EAB7FFDB647EF6C4A0F7FB4073DE" box="[1194,1224,943,968]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
<quantity id="85249B40FFDB647EF6BFA0F7FAFE73DF" box="[1233,1398,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="2.46" metricValueMax="2.69" metricValueMin="2.23" pageId="10" pageNumber="223" unit="mm" value="2.46" valueMax="2.69" valueMin="2.23">2.232.69 mm</quantity>
<emphasis id="70A8EAB7FFDB647EF710A0F7FA1473DE" box="[1406,1436,943,968]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
<quantity id="85249B40FFDB647EF2F9A08BFEB373FB" box="[151,315,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.765" metricValueMax="4.23" metricValueMin="3.3" pageId="10" pageNumber="223" unit="mm" value="3.765" valueMax="4.23" valueMin="3.3">3.304.23 mm</quantity>
<emphasis id="70A8EAB7FFDB647EF32DA08BFEE973FA" box="[323,353,979,1004]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
<quantity id="85249B40FFDB647EF307A08BFE4673FB" box="[361,462,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.46" pageId="10" pageNumber="223" unit="mm" value="3.46">3.46 mm</quantity>
<emphasis id="70A8EAB7FFDB647EF3B8A08BFE7C73FA" box="[470,500,979,1004]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
<quantity id="85249B40FFDB647EF392A08AFD1773FB" box="[508,671,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.315" metricValueMax="2.52" metricValueMin="2.11" pageId="10" pageNumber="223" unit="mm" value="2.315" valueMax="2.52" valueMin="2.11">2.112.52 mm</quantity>
<emphasis id="70A8EAB7FFDB647EF0C9A08BFD4D73FA" box="[679,709,979,1004]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
<quantity id="85249B40FFDB647EF0A3A08BFCF973FB" box="[717,881,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.305" metricValueMax="2.43" metricValueMin="2.18" pageId="10" pageNumber="223" unit="mm" value="2.305" valueMax="2.43" valueMin="2.18">2.182.43 mm</quantity>
<emphasis id="70A8EAB7FFDB647EF117A08BFC1F73FA" box="[889,919,979,1004]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
<quantity id="85249B40FFDB647EF1F1A08BFBCF73FB" box="[927,1095,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.21" metricValueMax="3.44" metricValueMin="2.98" pageId="10" pageNumber="223" unit="mm" value="3.21" valueMax="3.44" valueMin="2.98">2.983.44 mm</quantity>
). However, despite its smaller body length the oesophagus of
<taxonomicName id="85DC4D26FFDB647EF068A0AFFD277407" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[518,687,1015,1041]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF068A0AFFD277407" box="[518,687,1015,1041]" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF0D4A0AFFD727407" box="[698,762,1015,1041]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
is comparatively longer than some of the aforementioned species:
<taxonomicName id="85DC4D26FFDB647EF294A743FE317423" authorityName="Hugot, Morand &amp; Van Waerebeke" authorityYear="1991" box="[250,441,1051,1077]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="kinabaluensis">
<emphasis id="70A8EAB7FFDB647EF294A743FE317423" box="[250,441,1051,1077]" italics="true" pageId="10" pageNumber="223">D. kinabaluensis</emphasis>
</taxonomicName>
,
<taxonomicName id="85DC4D26FFDB647EF3A4A743FDA57423" authorityName="Skrjabin" authorityYear="1916" box="[458,557,1051,1077]" class="Chromadorea" family="Oxyuridae" genus="Oxyuris" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="leidyi">
<emphasis id="70A8EAB7FFDB647EF3A4A743FDA57423" box="[458,557,1051,1077]" italics="true" pageId="10" pageNumber="223">D. leidyi</emphasis>
</taxonomicName>
,
<taxonomicName id="85DC4D26FFDB647EF050A743FD6C7422" authorityName="Van Waerebeke" authorityYear="1969" box="[574,740,1051,1076]" class="Chromadorea" family="Thelastomatidae" genus="Posterovulva" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="moramangi">
<emphasis id="70A8EAB7FFDB647EF050A743FD6C7422" box="[574,740,1051,1076]" italics="true" pageId="10" pageNumber="223">D. moramangi</emphasis>
</taxonomicName>
,
<taxonomicName id="85DC4D26FFDB647EF09BA743FC317423" authorityName="Hugot, Morand &amp; Van Waerebeke" authorityYear="1991" box="[757,953,1051,1077]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="nhatrangensis">
<emphasis id="70A8EAB7FFDB647EF09BA743FC317423" box="[757,953,1051,1077]" italics="true" pageId="10" pageNumber="223">D. nhatrangensis</emphasis>
</taxonomicName>
and
<taxonomicName id="85DC4D26FFDB647EF19BA743FBF67423" authorityName="Adamson" authorityYear="1984" box="[1013,1150,1051,1077]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="skrjabini">
<emphasis id="70A8EAB7FFDB647EF19BA743FBF67423" box="[1013,1150,1051,1077]" italics="true" pageId="10" pageNumber="223">D. skrjabini</emphasis>
</taxonomicName>
(b = 5.86.4
<emphasis id="70A8EAB7FFDB647EF74EA743FAB67422" box="[1312,1342,1051,1076]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
8.49.8
<emphasis id="70A8EAB7FFDB647EF2F9A767FF3E744E" box="[151,182,1087,1112]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
6.9
<emphasis id="70A8EAB7FFDB647EF286A767FE8F744E" box="[232,263,1087,1112]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
6.77.5
<emphasis id="70A8EAB7FFDB647EF304A767FE01744E" box="[362,393,1087,1112]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
7.58.4
<emphasis id="70A8EAB7FFDB647EF382A767FD83744E" box="[492,523,1087,1112]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
7.8). The body of
<taxonomicName id="85DC4D26FFDB647EF08EA767FC0F744F" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[736,903,1087,1113]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF08EA767FC0F744F" box="[736,903,1087,1113]" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF1E1A767FC44744F" box="[911,972,1087,1113]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
is more robust than
<taxonomicName id="85DC4D26FFDB647EF6DBA767FAFA744F" authorityName="Hugot, Morand &amp; Van Waerebeke" authorityYear="1991" box="[1205,1394,1087,1113]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="kinabaluensis">
<emphasis id="70A8EAB7FFDB647EF6DBA767FAFA744F" box="[1205,1394,1087,1113]" italics="true" pageId="10" pageNumber="223">D. kinabaluensis</emphasis>
</taxonomicName>
,
<taxonomicName id="85DC4D26FFDB647EF7EFA767FE90746A" authorityName="Van Waerebeke" authorityYear="1969" class="Chromadorea" family="Thelastomatidae" genus="Posterovulva" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="moramangi">
<emphasis id="70A8EAB7FFDB647EF7EFA767FE90746A" italics="true" pageId="10" pageNumber="223">D. moramangi</emphasis>
</taxonomicName>
,
<taxonomicName id="85DC4D26FFDB647EF34BA73BFE6D746B" authorityName="Hugot, Morand &amp; Van Waerebeke" authorityYear="1991" box="[293,485,1123,1149]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="nhatrangensis">
<emphasis id="70A8EAB7FFDB647EF34BA73BFE6D746B" box="[293,485,1123,1149]" italics="true" pageId="10" pageNumber="223">D. nhatrangensis</emphasis>
</taxonomicName>
and
<taxonomicName id="85DC4D26FFDB647EF074A73BFD17746B" authorityName="Adamson" authorityYear="1984" box="[538,671,1123,1149]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="skrjabini">
<emphasis id="70A8EAB7FFDB647EF074A73BFD17746B" box="[538,671,1123,1149]" italics="true" pageId="10" pageNumber="223">D. skrjabini</emphasis>
</taxonomicName>
(a = 8.612.0
<emphasis id="70A8EAB7FFDB647EF150A73BFCD4746A" box="[830,860,1123,1148]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
20.226.5
<emphasis id="70A8EAB7FFDB647EF1B6A73BFC7F746A" box="[984,1015,1123,1148]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
12.517.0
<emphasis id="70A8EAB7FFDB647EF61CA73BFB19746A" box="[1138,1169,1123,1148]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
14.516.2
<emphasis id="70A8EAB7FFDB647EF763A73BFAA3746A" box="[1293,1323,1123,1148]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
12.6), but comparatively slender than
<taxonomicName id="85DC4D26FFDB647EF3A3A7DFFDA574B7" authorityName="Skrjabin" authorityYear="1916" box="[461,557,1159,1185]" class="Chromadorea" family="Oxyuridae" genus="Oxyuris" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="leidyi">
<emphasis id="70A8EAB7FFDB647EF3A3A7DFFDA574B7" box="[461,557,1159,1185]" italics="true" pageId="10" pageNumber="223">D. leidyi</emphasis>
</taxonomicName>
(a = 8.612.0
<emphasis id="70A8EAB7FFDB647EF0BEA7DFFD6674B6" box="[720,750,1159,1184]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
8.0). The index V´ of
<taxonomicName id="85DC4D26FFDB647EF186A7DFFBDF74B7" authorityName="Spiridonov &amp; Ivanova" authorityYear="1998" box="[1000,1111,1159,1185]" class="Chromadorea" family="Thelastomatidae" genus="Posterovulva" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="danieli">
<emphasis id="70A8EAB7FFDB647EF186A7DFFBDF74B7" box="[1000,1111,1159,1185]" italics="true" pageId="10" pageNumber="223">D. danieli</emphasis>
</taxonomicName>
is larger than
<taxonomicName id="85DC4D26FFDB647EF698A7DFFA1474B7" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[1270,1436,1159,1185]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF698A7DFFA1474B7" box="[1270,1436,1159,1185]" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF2F9A7F3FF5B74D3" box="[151,211,1195,1221]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
(V´ = 90.0
<emphasis id="70A8EAB7FFDB647EF339A7F3FEFD74D2" box="[343,373,1195,1220]" italics="true" pageId="10" pageNumber="223">vs.</emphasis>
82.286.9).
</paragraph>
</subSubSection>
<subSubSection id="0AC6652EFFDB647EF2F9A7ABFEB1756F" pageId="10" pageNumber="223" type="distribution">
<paragraph id="426336A5FFDB647EF2F9A7ABFEFF7527" blockId="10.[151,1437,1267,1437]" pageId="10" pageNumber="223">
<materialsCitation id="F2B43CF8FFDB647EF2F9A7ABFEFB7527" country="Japan" county="Ryukyu Archipelago" location="Aira River" municipality="Iriomote Island" pageId="10" pageNumber="223" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="70A8EAB7FFDB647EF2F9A7ABFEBC751B" box="[151,308,1267,1293]" italics="true" pageId="10" pageNumber="223">
<typeStatus id="9D678807FFDB647EF2F9A7ABFF43751A" box="[151,203,1267,1292]" pageId="10" pageNumber="223">Type</typeStatus>
locality.
</emphasis>
<location id="4703607EFFDB647EF32CA7ABFE4B751B" LSID="urn:lsid:plazi:treatment:CA7587B3FFD2647FF2F9A237FD22719F:4703607EFFDB647EF32CA7ABFE4B751B" box="[322,451,1267,1293]" country="Japan" county="Ryukyu Archipelago" municipality="Iriomote Island" name="Aira River" pageId="10" pageNumber="223" stateProvince="Okinawa">Aira River</location>
,
<location id="4703607EFFDB647EF3BBA7ABFD06751B" LSID="urn:lsid:plazi:treatment:CA7587B3FFD2647FF2F9A237FD22719F:4703607EFFDB647EF3BBA7ABFD06751B" box="[469,654,1267,1293]" country="Japan" county="Ryukyu Archipelago" municipality="Iriomote Island" name="Taketomi Town" pageId="10" pageNumber="223" stateProvince="Okinawa">Taketomi Town</location>
,
<location id="4703607EFFDB647EF0CEA7ABFCE5751B" LSID="urn:lsid:plazi:treatment:CA7587B3FFD2647FF2F9A237FD22719F:4703607EFFDB647EF0CEA7ABFCE5751B" box="[672,877,1267,1293]" country="Japan" county="Ryukyu Archipelago" municipality="Iriomote Island" name="Yaeyama District" pageId="10" pageNumber="223" stateProvince="Okinawa">Yaeyama District</location>
,
<collectingMunicipality id="A207ACDFFFDB647EF111A7ABFBBD751B" box="[895,1077,1267,1293]" pageId="10" pageNumber="223">Iriomote Island</collectingMunicipality>
,
<collectingRegion id="8018F847FFDB647EF626A7ABFABA751B" box="[1096,1330,1267,1293]" country="Japan" name="Okinawa" pageId="10" pageNumber="223">Okinawa Prefecture</collectingRegion>
,
<collectingCounty id="AB024E29FFDB647EF72BA7ABFEAC7527" pageId="10" pageNumber="223">Ryukyu Archipelago</collectingCounty>
,
<collectingCountry id="3ACB7635FFDB647EF35EA64FFEFB7527" box="[304,371,1303,1329]" name="Japan" pageId="10" pageNumber="223">Japan</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="426336A5FFDB647EF2F9A663FB5B7543" blockId="10.[151,1437,1267,1437]" box="[151,1235,1339,1365]" pageId="10" pageNumber="223">
<emphasis id="70A8EAB7FFDB647EF2F9A663FE817543" box="[151,265,1339,1365]" italics="true" pageId="10" pageNumber="223">
<typeStatus id="9D678807FFDB647EF2F9A663FF437542" box="[151,203,1339,1364]" pageId="10" pageNumber="223">Type</typeStatus>
host.
</emphasis>
<taxonomicName id="85DC4D26FFDB647EF361A663FCE37543" authority="Asahina, 1988" authorityName="Asahina" authorityYear="1988" box="[271,875,1339,1365]" class="Insecta" family="Blaberidae" genus="Panesthia" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="223" phylum="Arthropoda" rank="subSpecies" species="angustipennis" subSpecies="yayeyamensis">
<emphasis id="70A8EAB7FFDB647EF361A663FD4B7542" box="[271,707,1339,1365]" italics="true" pageId="10" pageNumber="223">Panesthia angustipennis yayeyamensis</emphasis>
Asahina, 1988
</taxonomicName>
(
<taxonomicName id="85DC4D26FFDB647EF114A663FC477543" authorityName="Linnaeus" authorityYear="1758" box="[890,975,1339,1365]" class="Insecta" kingdom="Animalia" pageId="10" pageNumber="223" phylum="Arthropoda" rank="class">Insecta</taxonomicName>
: Blattaria:
<taxonomicName id="85DC4D26FFDB647EF627A663FB407543" box="[1097,1224,1339,1365]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="223" phylum="Arthropoda" rank="family">Blaberidae</taxonomicName>
).
</paragraph>
<paragraph id="426336A5FFDB647EF2F9A607FEB1756F" blockId="10.[151,1437,1267,1437]" box="[151,313,1375,1401]" pageId="10" pageNumber="223">
<taxonomicName id="85DC4D26FFDB647EF2F9A607FF49756F" box="[151,193,1375,1401]" class="Bacteroidia" family="Flavobacteriaceae" genus="Site" higherTaxonomySource="GBIF" kingdom="Bacteria" order="Flavobacteriales" pageId="10" pageNumber="223" phylum="Bacteroidota" rank="genus">
<emphasis id="70A8EAB7FFDB647EF2F9A607FF49756F" box="[151,193,1375,1401]" italics="true" pageId="10" pageNumber="223">Site</emphasis>
</taxonomicName>
. Hind gut.
</paragraph>
</subSubSection>
<subSubSection id="0AC6652EFFDB647EF2F9A6DBFB6E758B" box="[151,1254,1411,1437]" pageId="10" pageNumber="223" type="etymology">
<paragraph id="426336A5FFDB647EF2F9A6DBFB6E758B" blockId="10.[151,1437,1267,1437]" box="[151,1254,1411,1437]" pageId="10" pageNumber="223">
<emphasis id="70A8EAB7FFDB647EF2F9A6DBFE9D758B" box="[151,277,1411,1437]" italics="true" pageId="10" pageNumber="223">Etymology.</emphasis>
Specific epithet referred to the Ryukyu Archipelago, to which Iriomote Island belongs.
</paragraph>
</subSubSection>
<subSubSection id="0AC6652EFFDB647FF2F9A693FD22719F" lastPageId="11" lastPageNumber="224" pageId="10" pageNumber="223" type="description">
<paragraph id="426336A5FFDB647EF2F9A693FDAC75F3" blockId="10.[151,1436,1483,1581]" box="[151,548,1483,1509]" pageId="10" pageNumber="223">
<emphasis id="70A8EAB7FFDB647EF2F9A693FDAC75F3" box="[151,548,1483,1509]" italics="true" pageId="10" pageNumber="223">Molecular identification of the host</emphasis>
</paragraph>
<paragraph id="426336A5FFDB647EF2F9A6B7FC29763B" blockId="10.[151,1436,1483,1581]" pageId="10" pageNumber="223">
The BLAST alignment resulted in a high percent identity (99.71%) of the COII sequence of the host with the sequence AB007541 of
<emphasis id="70A8EAB7FFDB647EF3CDA54BFDFD763A" box="[419,629,1555,1580]" italics="true" pageId="10" pageNumber="223">P. a. yayeyamensis</emphasis>
also from Iriomote Island.
</paragraph>
<paragraph id="426336A5FFDB647EF2F9A503FEAC7663" blockId="10.[151,1437,1627,1977]" box="[151,292,1627,1653]" pageId="10" pageNumber="223">
<emphasis id="70A8EAB7FFDB647EF2F9A503FEAC7663" box="[151,292,1627,1653]" italics="true" pageId="10" pageNumber="223">DNA studies</emphasis>
</paragraph>
<paragraph id="426336A5FFDB647EF2F9A527FBA6773F" blockId="10.[151,1437,1627,1977]" pageId="10" pageNumber="223">
Two partial sequences of the D2-D3 region of the 28S rDNA were obtained from a female and a male of
<taxonomicName id="85DC4D26FFDB647EF7EFA527FE9376AB" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDB647EF7EFA527FE9376AB" italics="true" pageId="10" pageNumber="223">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDB647EF34BA5FBFEEB76AB" box="[293,355,1699,1725]" pageId="10" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
(743 bp and 742 bp, respectively). Both sequences were identical, in an alignment of 739 bp. The new species differs in 28 homologous positions (in an alignment of 634 bp) from a sequence of an unidentified
<taxonomicName id="85DC4D26FFDB647EF2F9A5B3FE877713" authorityName="Basir" authorityYear="1956" box="[151,271,1771,1797]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="genus">
<emphasis id="70A8EAB7FFDB647EF2F9A5B3FE877713" box="[151,271,1771,1797]" italics="true" pageId="10" pageNumber="223">Desmicola</emphasis>
</taxonomicName>
(GQ368463) from a Vietnamese
<taxonomicName id="85DC4D26FFDB647EF0F8A5B3FBF97713" authority="(Spiridonov &amp; Guzeeva 2009)" baseAuthorityName="Spiridonov &amp; Guzeeva" baseAuthorityYear="2009" box="[662,1137,1771,1797]" class="Insecta" family="Blaberidae" genus="Panesthia" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="70A8EAB7FFDB647EF0F8A5B3FC8E7713" box="[662,774,1771,1797]" italics="true" pageId="10" pageNumber="223">Panesthia</emphasis>
(
<bibRefCitation id="264D4B54FFDB647EF176A5B3FBE17713" author="Spiridonov, S. E. &amp; Guzeeva, E. A." box="[792,1129,1771,1797]" pageId="10" pageNumber="223" pagination="127 - 134" refId="ref9097" refString="Spiridonov, S. E. &amp; Guzeeva, E. A. (2009) Phylogeny of nematodes of the superfamily Thelastomatoidea (Oxyurida) inferred from LSU rDNA sequence. Russian Journal of Nematology, 17, 127 - 134." type="journal article" year="2009">Spiridonov &amp; Guzeeva 2009</bibRefCitation>
)
</taxonomicName>
, the only sequence of the genus available in GenBank that comprises the D2-D3 domains of the 28S rDNA.
</paragraph>
<paragraph id="426336A5FFDB647EF2A9A46BFDAF77AF" blockId="10.[151,1437,1627,1977]" pageId="10" pageNumber="223">
The new species differs in four homologous positions from a sequence (AM232760) of the D3 domain of the 28S rDNA from
<taxonomicName id="85DC4D26FFDB647EF3EDA40FFE7A7766" authorityName="Jex, Schneider, Rose &amp; Cribb" authorityYear="2005" box="[387,498,1879,1904]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ornata">
<emphasis id="70A8EAB7FFDB647EF3EDA40FFE7A7766" box="[387,498,1879,1904]" italics="true" pageId="10" pageNumber="223">D. ornata</emphasis>
</taxonomicName>
, parasite of the Australian
<taxonomicName id="85DC4D26FFDB647EF147A40FFB327767" authority="(Jex et al. 2006)" baseAuthorityName="Jex" baseAuthorityYear="2006" box="[809,1210,1879,1905]" class="Insecta" family="Blaberidae" genus="Panesthia" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="cribata">
<emphasis id="70A8EAB7FFDB647EF147A40FFC7A7767" box="[809,1010,1879,1905]" italics="true" pageId="10" pageNumber="223">Panesthia cribata</emphasis>
(
<bibRefCitation id="264D4B54FFDB647EF66DA40FFB3A7767" author="Jex, A. R. &amp; Hu, M. &amp; Rose, H. A. &amp; Schneider, M. &amp; Cribb, T. H. &amp; Gasser, R. B." box="[1027,1202,1879,1905]" pageId="10" pageNumber="223" pagination="123 - 129" refId="ref7921" refString="Jex, A. R., Hu, M., Rose, H. A., Schneider, M., Cribb, T. H. &amp; Gasser, R. B. (2006) Molecular characterization of Thelastomatoidea (Nematoda: Oxyurida) from cockroaches in Australia. Parasitology, 133, 123 - 129." type="journal article" year="2006">
Jex
<emphasis id="70A8EAB7FFDB647EF65DA40FFBE57767" box="[1075,1133,1879,1905]" italics="true" pageId="10" pageNumber="223">et al.</emphasis>
2006
</bibRefCitation>
)
</taxonomicName>
. On the other hand,
<taxonomicName id="85DC4D26FFDB647EF2F9A423FEBE7783" box="[151,310,1915,1941]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="undetermined">
<emphasis id="70A8EAB7FFDB647EF2F9A423FE877783" box="[151,271,1915,1941]" italics="true" pageId="10" pageNumber="223">Desmicola</emphasis>
sp.
</taxonomicName>
from
<collectingCountry id="3ACB7635FFDB647EF314A423FE517783" box="[378,473,1915,1941]" name="Vietnam" pageId="10" pageNumber="223">Vietnam</collectingCountry>
(GQ368463) differs in two homologous positions from
<taxonomicName id="85DC4D26FFDB647EF63FA423FB367782" authorityName="Jex, Schneider, Rose &amp; Cribb" authorityYear="2005" box="[1105,1214,1915,1940]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="223" phylum="Nematoda" rank="species" species="ornata">
<emphasis id="70A8EAB7FFDB647EF63FA423FB367782" box="[1105,1214,1915,1940]" italics="true" pageId="10" pageNumber="223">D. ornata</emphasis>
</taxonomicName>
. These results were obtained in an alignment of 242 bp.
</paragraph>
<paragraph id="426336A5FFDA647FF2A9A3CFFAA170C3" blockId="11.[151,1436,151,393]" pageId="11" pageNumber="224">In addition, two partial sequences of the 18S rDNA were obtained from the female and male of the present species (1683 bp and 1680 bp, respectively). Both sequences were identical, in an alignment of 1680 bp.</paragraph>
<paragraph id="426336A5FFDA647FF2A9A387FD22719F" blockId="11.[151,1436,151,393]" pageId="11" pageNumber="224">
In both ML and BI phylograms both sequences of
<taxonomicName id="85DC4D26FFDA647FF094A387FC2870EF" authority="Morffe &amp; García &amp; Hasegawa, 2023" authorityName="Morffe &amp; García &amp; Hasegawa" authorityYear="2023" box="[762,928,223,249]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="224" phylum="Nematoda" rank="species" species="ryukyuensis" status="n. sp.">
<emphasis id="70A8EAB7FFDA647FF094A387FC2870EF" box="[762,928,223,249]" italics="true" pageId="11" pageNumber="224">D. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B9B57CCFFDA647FF1C6A387FC6B70EF" box="[936,995,223,249]" pageId="11" pageNumber="224" rank="species">n. sp.</taxonomicNameLabel>
form a monophyletic clade with strong nodal support, with
<taxonomicName id="85DC4D26FFDA647FF3ECA25BFDAC710A" box="[386,548,259,285]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="224" phylum="Nematoda" rank="species" species="undetermined">
<emphasis id="70A8EAB7FFDA647FF3ECA25BFE72710B" box="[386,506,259,285]" italics="true" pageId="11" pageNumber="224">Desmicola</emphasis>
sp.
</taxonomicName>
The genus
<taxonomicName id="85DC4D26FFDA647FF0DEA25BFCA0710B" authorityName="Basir" authorityYear="1956" box="[688,808,259,285]" class="Chromadorea" family="Thelastomatidae" genus="Desmicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="224" phylum="Nematoda" rank="genus">
<emphasis id="70A8EAB7FFDA647FF0DEA25BFCA0710B" box="[688,808,259,285]" italics="true" pageId="11" pageNumber="224">Desmicola</emphasis>
</taxonomicName>
form a clade containing
<taxonomicName id="85DC4D26FFDA647FF63FA25BFA10710B" authority="(Leidy, 1849)" baseAuthorityName="Leidy" baseAuthorityYear="1849" box="[1105,1432,259,285]" class="Chromadorea" family="Thelastomatidae" genus="Aorurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="224" phylum="Nematoda" rank="species" species="agile">
<emphasis id="70A8EAB7FFDA647FF63FA25BFB67710B" box="[1105,1263,259,285]" italics="true" pageId="11" pageNumber="224">Aorurus agile</emphasis>
(Leidy, 1849)
</taxonomicName>
,
<taxonomicName id="85DC4D26FFDA647FF2F9A27FFEC57157" box="[151,333,295,321]" class="Chromadorea" family="Thelastomatidae" genus="Cordonicola" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="224" phylum="Nematoda" rank="species" species="undetermined">
<emphasis id="70A8EAB7FFDA647FF2F9A27FFEAC7157" box="[151,292,295,321]" italics="true" pageId="11" pageNumber="224">Cordonicola</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="85DC4D26FFDA647FF334A27FFC097157" authority="Phillips &amp; Bernard, 2018" authorityName="Phillips &amp; Bernard" authorityYear="2018" box="[346,897,295,321]" class="Chromadorea" family="Thelastomatidae" genus="Stauratostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="224" phylum="Nematoda" rank="species" species="shelleyi">
<emphasis id="70A8EAB7FFDA647FF334A27FFDD37157" box="[346,603,295,321]" italics="true" pageId="11" pageNumber="224">Stauratostoma shelleyi</emphasis>
<bibRefCitation id="264D4B54FFDA647FF00CA27FFC097157" author="Phillips, G. &amp; Pivar, R. J. &amp; Sun, X. &amp; Moulton, J. K. &amp; Bernard, E. C." box="[610,897,295,321]" pageId="11" pageNumber="224" pagination="133 - 146" refId="ref8704" refString="Phillips, G., Pivar, R. J., Sun, X., Moulton, J. K. &amp; Bernard, E. C. (2018) Stauratostoma shelleyi n. gen., n. sp. (Nematoda: Rhabditida: Thelastomatidae) from Appalachian Polydesmid Millipedes (Polydesmida: Xystodesmidae). Journal of Nematology, 50 (2), 133 - 146. https: // doi. org / 10.21307 / jofnem- 2018 - 023" type="journal article" year="2018">Phillips &amp; Bernard, 2018</bibRefCitation>
</taxonomicName>
, several species of
<taxonomicName id="85DC4D26FFDA647FF635A27FFAE47157" authority="Leidy, 1849" authorityName="Leidy" authorityYear="1849" box="[1115,1388,295,321]" class="Chromadorea" family="Thelastomatidae" genus="Thelastoma" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="224" phylum="Nematoda" rank="genus">
<emphasis id="70A8EAB7FFDA647FF635A27FFB567157" box="[1115,1246,295,321]" italics="true" pageId="11" pageNumber="224">Thelastoma</emphasis>
Leidy, 1849
</taxonomicName>
and unidentified thelastomatids. This clade is supported by maximum value of posterior probability for the BI analysis, but has low bootstrap value in the ML analysis.
</paragraph>
</subSubSection>
</treatment>
</document>