317 lines
36 KiB
XML
317 lines
36 KiB
XML
<document id="B33BE58F0D188D7AAF76E5BEECCB6BAE" ID="10.11646/zootaxa.4175.4.1" ID-DOI="10.11646/zootaxa.4175.4.1" ID-GBIF-Dataset="55c6a2a3-cf2b-41df-bb10-ee4ba1d423c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="256776" ID-ZooBank="5C11185B-E219-4214-8A01-517FBB8C99BA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476684690294" checkinUser="plazi" docAuthor="Ruta, Rafał" docDate="2016" docId="A8548C3CFFD9FFF2C7F3FAC8FA97FD75" docLanguage="en" docName="zootaxa.4175.4.1.pdf" docOrigin="Zootaxa 4175 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anticyphon oyonensis Ruta, 2016, sp. nov." docType="treatment" docUuid="3280F62D-4A81-40F6-91BF-2CE557C6891B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="310" masterDocId="546DF444FFD1FFFBC764FFB0FFBDF965" masterDocTitle="Anticyphon gen. nov., a new genus of Scirtidae (Coleoptera: Scirtoidea) inhabiting high altitude Andean cloud forests and páramo formation" masterLastPageNumber="318" masterPageNumber="301" pageNumber="309" updateTime="1698389907452" updateUser="plazi">
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<mods:title id="6E55EBF90089ACD3F3BB0EBC4019E309">Anticyphon gen. nov., a new genus of Scirtidae (Coleoptera: Scirtoidea) inhabiting high altitude Andean cloud forests and páramo formation</mods:title>
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<mods:name id="266869948471580A7143673696D97D16" type="personal">
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<mods:roleTerm id="3A70D0BED156888ABB4ECF80DEF0376C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3C1F7FF044477D50DABA2DF0DF2A64D8">Ruta, Rafał</mods:namePart>
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<mods:typeOfResource id="114978B1F33BED52E04F97F210C3A457">text</mods:typeOfResource>
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<mods:title id="BB360D245B37F2275CE0B8B4125EA5F8">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="48167C93D92CAA42918C2CB5A6307708">
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<mods:date id="EF54E8E58A7060CA4F9821D8454065EE">2016</mods:date>
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<mods:detail id="333A8FE986ECAC44A5FC131A444CFB3D" type="volume">
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<mods:number id="5DA3B6B53D602CB23BBD308A3B8403C6">4175</mods:number>
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<mods:detail id="AEDCCFC8FF58F53B623292D67F17B141" type="issue">
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<mods:number id="4239F1ED0A69122723C74646745CFFF2">4</mods:number>
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<mods:classification id="A2A5FE2E44989E0460DEA1EF826D82CA">journal article</mods:classification>
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<mods:identifier id="A0A9A3AC48A4C412CC7F5AF4B6A5F99C" type="DOI">10.11646/zootaxa.4175.4.1</mods:identifier>
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<mods:identifier id="6D023EED4642E3FBFB16F67ABF43C805" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="76D85E724F512454FE98E7437B894FD0" type="Zenodo-Dep">256776</mods:identifier>
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<mods:identifier id="3E61DF2CE0D98E19E957CED4E343D3AA" type="ZooBank">5C11185B-E219-4214-8A01-517FBB8C99BA</mods:identifier>
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<treatment id="A8548C3CFFD9FFF2C7F3FAC8FA97FD75" ID-DOI="http://doi.org/10.5281/zenodo.6060613" ID-GBIF-Taxon="124996813" ID-Zenodo-Dep="6060613" LSID="urn:lsid:zoobank.org:act:3280F62D-4A81-40F6-91BF-2CE557C6891B" httpUri="http://treatment.plazi.org/id/A8548C3CFFD9FFF2C7F3FAC8FA97FD75" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
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<subSubSection id="68E76EA1FFD9FFF3C7F3FAC8FE4FFCD6" pageId="8" pageNumber="309" type="nomenclature">
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<paragraph id="20423D2AFFD9FFF3C7F3FAC8FDBBFCF7" blockId="8.[151,518,1400,1460]" box="[151,518,1400,1426]" pageId="8" pageNumber="309">
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<heading id="7B0A8A46FFD9FFF3C7F3FAC8FDBBFCF7" bold="true" box="[151,518,1400,1426]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
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<emphasis id="1289E138FFD9FFF3C7F3FAC8FDBBFCF7" bold="true" box="[151,518,1400,1426]" pageId="8" pageNumber="309">
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<taxonomicName id="E7FD46A9FFD9FFF3C7F3FAC8FE1DFCF7" box="[151,416,1400,1426]" class="Insecta" family="Scirtidae" genus="Anticyphon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="oyonensis" status="sp. nov.">
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<emphasis id="1289E138FFD9FFF3C7F3FAC8FE1DFCF7" bold="true" box="[151,416,1400,1426]" italics="true" pageId="8" pageNumber="309">Anticyphon oyonensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="09BA5C43FFD9FFF3C6CCFAC8FDBBFCF7" box="[424,518,1400,1426]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="20423D2AFFD9FFF3C7F3FA2BFE4FFCD6" blockId="8.[151,518,1400,1460]" box="[151,498,1435,1460]" pageId="8" pageNumber="309">
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(
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<figureCitation id="B8C621AFFFD9FFF3C7FBFA2BFF5CFCD6" box="[159,225,1435,1460]" captionStart="FIGURE 1" captionStartId="2.[151,250,1122,1144]" captionTargetBox="[151,1436,193,1099]" captionTargetId="figure@2.[151,1436,193,1101]" captionTargetPageId="2" captionText="FIGURE 1. Anticyphon gen. nov., habitus. A) A. davidsoni sp. nov., B) A. ecuadorensis sp. nov., C) A. oyonensis sp. nov., female, D) A. oyonensis sp. nov., male E) A. paramoensis sp. nov., light form, F) A. paramoensis sp. nov., dark form, G) A. peruvianus sp. nov., H) A. santanderensis sp. nov. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/256777/files/figure.png" pageId="8" pageNumber="309">Figs 1</figureCitation>
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C–D, 2D–F, 3B, 5, 8, 9)
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</paragraph>
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</subSubSection>
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<subSubSection id="68E76EA1FFD9FFF3C7F3FA53FDBAFF21" pageId="8" pageNumber="309" type="materials_examined">
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<paragraph id="20423D2AFFD9FFF3C7F3FA53FDBAFF21" blockId="8.[151,1437,1507,2036]" pageId="8" pageNumber="309">
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<materialsCitation id="90953777FFD9FFF3C7F3FA53FB80FF7A" ID-GBIF-Occurrence="1323865283" collectingDate="1987-02-24" collectingDateMax="1987-02-26" collectingDateMin="1987-02-24" collectionCode="ZMUC" collectorName="O. Karsholt & Zool. Mus. Copenhagen" country="Peru" elevation="4000" location="Pueblo Quichas" municipality="Quabrada Quichas" pageId="8" pageNumber="309" specimenCount="2" stateProvince="Lima" typeStatus="holotype">
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<emphasis id="1289E138FFD9FFF3C7F3FA53FEFBFC99" bold="true" box="[151,326,1507,1532]" pageId="8" pageNumber="309">
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<typeStatus id="FF468388FFD9FFF3C7F3FA53FF6FFC99" box="[151,210,1507,1532]" pageId="8" pageNumber="309">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="FF468388FFD9FFF3C62AFA53FE04FC99" box="[334,441,1507,1532]" pageId="8" pageNumber="309" type="holotype">Holotype</typeStatus>
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, male (
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<collectionCode id="46ECA5EFFFD9FFF3C568FA53FDD9FC9E" box="[524,612,1507,1531]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="8" pageNumber="309">ZMUC</collectionCode>
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): “
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<collectingCountry id="58EA7DBAFFD9FFF3C5E6FA53FD72FC9E" box="[642,719,1507,1531]" name="Peru" pageId="8" pageNumber="309">PERU</collectingCountry>
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, Dept.
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<collectingRegion id="E239F3C8FFD9FFF3C445FA53FCE1FC99" box="[801,860,1507,1532]" country="Peru" name="Lima" pageId="8" pageNumber="309">Lima</collectingRegion>
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\ 30:
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<quantity id="E70590CFFFD9FFF3C4C5FA53FC57FC99" box="[929,1002,1507,1532]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="309" unit="km" value="10.0">10 km</quantity>
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N Oyón
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</locationDeviation>
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\
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<collectingMunicipality id="C026A750FFD9FFF3C301FA53FA8BFC99" box="[1125,1334,1507,1532]" pageId="8" pageNumber="309">Quabrada Quichas</collectingMunicipality>
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\
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<location id="25226BF1FFD9FFF3C22AFA53FF4BFF7A" LSID="urn:lsid:plazi:treatment:A8548C3CFFD9FFF2C7F3FAC8FA97FD75:25226BF1FFD9FFF3C22AFA53FF4BFF7A" country="Peru" municipality="Quabrada Quichas" name="Pueblo Quichas" pageId="8" pageNumber="309" stateProvince="Lima">Pueblo Quichas</location>
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,
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<quantity id="E70590CFFFD9FFF3C661F9B7FEE3FF7A" box="[261,350,1543,1567]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="309" unit="m" value="4000.0">
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<elevation id="ABD0DA19FFD9FFF3C661F9B7FEE3FF7A" box="[261,350,1543,1567]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="309" unit="m" value="4000.0">4000 m</elevation>
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</quantity>
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\
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<date id="54431BEAFFD9FFF3C618F9B6FDA3FF7A" box="[380,542,1542,1567]" pageId="8" pageNumber="309" value="1987-02-24" valueMax="1987-02-26" valueMin="1987-02-24">
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<collectingDate id="4407E202FFD9FFF3C618F9B6FDA3FF7A" box="[380,542,1542,1567]" pageId="8" pageNumber="309" value="1987-02-24" valueMax="1987-02-26" valueMin="1987-02-24">24.–26.ii.1987</collectingDate>
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</date>
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\
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<collectorName id="8D0858FCFFD9FFF3C558F9B7FD79FF7A" box="[572,708,1542,1567]" pageId="8" pageNumber="309">O. Karsholt</collectorName>
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leg. \
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||
<collectorName id="8D0858FCFFD9FFF3C472F9B6FB8EFF7A" box="[790,1075,1542,1567]" pageId="8" pageNumber="309">Zool. Mus. Copenhagen</collectorName>
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”
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</materialsCitation>
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.
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||
<materialsCitation id="90953777FFD9FFF3C32FF9B8FDB9FF21" ID-GBIF-Occurrence="1323865284" collectingDate="1987-02-24" collectingDateMax="1987-02-26" collectingDateMin="1987-02-24" collectionCode="ZMUC" collectorName="O. Karsholt & Zool. Mus. Copenhagen" country="Peru" elevation="4000" location="Pueblo Quichas" municipality="Quabrada Quichas" pageId="8" pageNumber="309" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Lima" typeStatus="paratype">
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||
<typeStatus id="FF468388FFD9FFF3C32FF9B8FB06FF45" box="[1099,1211,1544,1568]" pageId="8" pageNumber="309" type="paratype">Paratypes</typeStatus>
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||
,
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||
<specimenCount id="36FBF6A3FFD9FFF3C3AEF9B7FA95FF7A" box="[1226,1320,1542,1567]" pageId="8" pageNumber="309" type="male">3 males</specimenCount>
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,
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<specimenCount id="36FBF6A3FFD9FFF3C253F9B7FA21FF7A" box="[1335,1436,1542,1567]" pageId="8" pageNumber="309" type="female">1 female</specimenCount>
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||
(
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||
<collectionCode id="46ECA5EFFFD9FFF3C7FBF99BFF4AFF26" box="[159,247,1579,1603]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="8" pageNumber="309">ZMUC</collectionCode>
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): same data as holotype
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="68E76EA1FFD9FFF3C7A3F9FEFBD5FF92" pageId="8" pageNumber="309" type="diagnosis">
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||
<paragraph id="20423D2AFFD9FFF3C7A3F9FEFBD5FF92" blockId="8.[151,1437,1507,2036]" pageId="8" pageNumber="309">
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<emphasis id="1289E138FFD9FFF3C7A3F9FEFEFCFF02" bold="true" box="[199,321,1614,1639]" pageId="8" pageNumber="309">Diagnosis.</emphasis>
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Body shape, size and granulate punctation of pronotal disc make this species similar to
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<taxonomicName id="E7FD46A9FFD9FFF3C2E0F9FFFE97FFEE" class="Insecta" family="Scirtidae" genus="Anticyphon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="ecuadorensis" status="sp. nov.">
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<emphasis id="1289E138FFD9FFF3C2E0F9FFFE97FFEE" italics="true" pageId="8" pageNumber="309">A. ecuadorensis</emphasis>
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</taxonomicName>
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<emphasis id="1289E138FFD9FFF3C654F9C3FE34FFE9" bold="true" box="[304,393,1651,1676]" pageId="8" pageNumber="309">
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<taxonomicNameLabel id="09BA5C43FFD9FFF3C654F9C3FE34FFE9" box="[304,393,1651,1676]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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and
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<taxonomicName id="E7FD46A9FFD9FFF3C6DAF9C3FDE7FFEE" box="[446,602,1651,1675]" class="Insecta" family="Scirtidae" genus="Anticyphon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="peruvianus" status="sp. nov.">
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<emphasis id="1289E138FFD9FFF3C6DAF9C3FDE7FFEE" box="[446,602,1651,1675]" italics="true" pageId="8" pageNumber="309">A. peruvianus</emphasis>
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</taxonomicName>
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<emphasis id="1289E138FFD9FFF3C53BF9C3FD05FFE9" bold="true" box="[607,696,1651,1676]" pageId="8" pageNumber="309">
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||
<taxonomicNameLabel id="09BA5C43FFD9FFF3C53BF9C3FD05FFE9" box="[607,696,1651,1676]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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Identifiable on the basis of male genitalia morphology: penis with curved parameroids, distinctly shorter than median process of trigonium. Tegmen with subtriangular parameres. Female of
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<taxonomicName id="E7FD46A9FFD9FFF3C674F90BFE23FFB6" box="[272,414,1723,1747]" class="Insecta" family="Scirtidae" genus="Anticyphon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="oyonensis" status="sp. nov.">
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<emphasis id="1289E138FFD9FFF3C674F90BFE23FFB6" box="[272,414,1723,1747]" italics="true" pageId="8" pageNumber="309">A. oyonensis</emphasis>
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</taxonomicName>
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||
<emphasis id="1289E138FFD9FFF3C6C1F90BFE42FFB1" bold="true" box="[421,511,1723,1748]" pageId="8" pageNumber="309">
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<taxonomicNameLabel id="09BA5C43FFD9FFF3C6C1F90BFE42FFB1" box="[421,511,1723,1748]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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can probably be distinguished from other—currently unknown—females of other species of
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<taxonomicName id="E7FD46A9FFD9FFF3C669F96FFE30FF92" box="[269,397,1759,1783]" class="Insecta" family="Scirtidae" genus="Anticyphon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="1289E138FFD9FFF3C669F96FFE30FF92" box="[269,397,1759,1783]" italics="true" pageId="8" pageNumber="309">Anticyphon</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1289E138FFD9FFF3C6F0F96EFE40FF92" bold="true" box="[404,509,1758,1783]" pageId="8" pageNumber="309">
|
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<taxonomicNameLabel id="09BA5C43FFD9FFF3C6F0F96EFE40FF92" box="[404,509,1758,1783]" pageId="8" pageNumber="309" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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on the basis of vulvar and bursal sclerites morphology.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68E76EA1FFD9FFF2C7A3F8B3FAE7FAAD" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309" type="description">
|
||
<paragraph id="20423D2AFFD9FFF2C7A3F8B3FC22F890" blockId="8.[151,1437,1507,2036]" lastBlockId="9.[151,1437,151,1040]" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
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||
<emphasis id="1289E138FFD9FFF3C7A3F8B3FEE4FE79" bold="true" box="[199,345,1795,1820]" pageId="8" pageNumber="309">Description.</emphasis>
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Male. Body oval, small, slightly flattened, clothed with brownish procumbent setae. Colouration of dorsum brown, antennae and legs yellowish brown. Head small, 1.6× wider than interocular space, covered with granulate punctures; eyes relatively big, protuberant; frons without depressions. Each mandible with small subtriangular denticle on inner margin. Antennae as in genus description. Pronotum small, transversely rectangular, sides slightly curved, widest at posterior angles, anterolateral angles broadly rounded, not produced; disc moderately convex. Punctation of pronotum granulate, both on lateral and central portions. Pronotum without distinct pits along basal margin. Each elytron with 2 subtly marked longitudinal carinae. Elytral punctation irregular, relatively dense; punctures separated by ca. 0.7 diameter. Penis (
|
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<figureCitation id="B8C621AFFFD8FFF2C484FF27FB9FF9D5" box="[992,1058,151,176]" captionStart="FIGURE 8" captionStartId="9.[151,250,1965,1987]" captionTargetBox="[151,1436,1093,1943]" captionTargetId="figure@9.[151,1436,1093,1944]" captionTargetPageId="9" captionText="FIGURE 8. Anticyphon oyonensis sp. nov., male genitalia. A) penis, B) tegmen, C) sternite VIII, D) sternite IX, E) tergite VIII, F) tergite IX. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/256783/files/figure.png" pageId="9" pageNumber="310">Fig. 8</figureCitation>
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A) moderately large (L
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||
<quantity id="E70590CFFFD8FFF2C255FF28FA25F9CA" box="[1329,1432,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="9" pageNumber="310" unit="mm" value="1.22">1.22 mm</quantity>
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||
,
|
||
<geoCoordinate id="45C95BEDFFD8FFF2C7F3FF0DFF54F9B1" box="[151,233,188,212]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="555" value="-0.42">
|
||
W
|
||
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|
||
</geoCoordinate>
|
||
mm), parameroids relatively short, curved, pointed at apices; trigonium with long and very narrow median process and two short lateral lobes, median process longer than parameroids, pala much longer than parameroids, narrow; tegmen (
|
||
<figureCitation id="B8C621AFFFD8FFF2C63DFEB4FE26F879" box="[345,411,260,285]" captionStart="FIGURE 8" captionStartId="9.[151,250,1965,1987]" captionTargetBox="[151,1436,1093,1943]" captionTargetId="figure@9.[151,1436,1093,1944]" captionTargetPageId="9" captionText="FIGURE 8. Anticyphon oyonensis sp. nov., male genitalia. A) penis, B) tegmen, C) sternite VIII, D) sternite IX, E) tergite VIII, F) tergite IX. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/256783/files/figure.png" pageId="9" pageNumber="310">Fig. 8</figureCitation>
|
||
B) relatively small (L
|
||
<quantity id="E70590CFFFD8FFF2C5FCFEB4FD42F879" box="[664,767,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="310" unit="mm" value="0.85">0.85 mm</quantity>
|
||
,
|
||
<geoCoordinate id="45C95BEDFFD8FFF2C468FEB5FCDDF879" box="[780,864,260,284]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="555" value="-0.44">
|
||
W
|
||
<quantity id="E70590CFFFD8FFF2C44BFEB4FCDDF879" box="[815,864,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="310" unit="mm" value="0.44">0.44</quantity>
|
||
</geoCoordinate>
|
||
mm), with wide, subtriangular parameres, pointed at apices; sternite VIII (
|
||
<figureCitation id="B8C621AFFFD8FFF2C6CFFE97FE52F825" box="[427,495,295,320]" captionStart="FIGURE 8" captionStartId="9.[151,250,1965,1987]" captionTargetBox="[151,1436,1093,1943]" captionTargetId="figure@9.[151,1436,1093,1944]" captionTargetPageId="9" captionText="FIGURE 8. Anticyphon oyonensis sp. nov., male genitalia. A) penis, B) tegmen, C) sternite VIII, D) sternite IX, E) tergite VIII, F) tergite IX. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/256783/files/figure.png" pageId="9" pageNumber="310">Fig. 8</figureCitation>
|
||
C) small (L
|
||
<quantity id="E70590CFFFD8FFF2C5E5FE98FD57F85A" box="[641,746,296,320]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="310" unit="mm" value="0.24">0.24 mm</quantity>
|
||
,
|
||
<geoCoordinate id="45C95BEDFFD8FFF2C59CFE98FCF0F825" box="[760,845,296,320]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="555" value="-0.38">
|
||
W
|
||
<quantity id="E70590CFFFD8FFF2C478FE98FCF0F825" box="[796,845,296,320]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="9" pageNumber="310" unit="mm" value="0.38">0.38</quantity>
|
||
</geoCoordinate>
|
||
mm), widely V-shaped, with sparse setae in apical portions; sternite IX (
|
||
<figureCitation id="B8C621AFFFD8FFF2C6F8FEFCFE59F801" box="[412,484,332,357]" captionStart="FIGURE 8" captionStartId="9.[151,250,1965,1987]" captionTargetBox="[151,1436,1093,1943]" captionTargetId="figure@9.[151,1436,1093,1944]" captionTargetPageId="9" captionText="FIGURE 8. Anticyphon oyonensis sp. nov., male genitalia. A) penis, B) tegmen, C) sternite VIII, D) sternite IX, E) tergite VIII, F) tergite IX. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/256783/files/figure.png" pageId="9" pageNumber="310">Fig. 8</figureCitation>
|
||
D) relatively small (L
|
||
<quantity id="E70590CFFFD8FFF2C598FEFCFCD5F801" box="[764,872,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="310" unit="mm" value="0.37">0.37 mm</quantity>
|
||
,
|
||
<geoCoordinate id="45C95BEDFFD8FFF2C41FFEFDFC69F801" box="[891,980,332,356]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="555" value="-0.37">
|
||
W
|
||
<quantity id="E70590CFFFD8FFF2C4C7FEFCFC69F801" box="[931,980,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="310" unit="mm" value="0.37">0.37</quantity>
|
||
</geoCoordinate>
|
||
mm), consisting of two subtriangular hemisternites, lightly sclerotized, with setose apical portion; tergite VIII (
|
||
<figureCitation id="B8C621AFFFD8FFF2C4B2FEDFFBA5F8ED" box="[982,1048,367,392]" captionStart="FIGURE 8" captionStartId="9.[151,250,1965,1987]" captionTargetBox="[151,1436,1093,1943]" captionTargetId="figure@9.[151,1436,1093,1944]" captionTargetPageId="9" captionText="FIGURE 8. Anticyphon oyonensis sp. nov., male genitalia. A) penis, B) tegmen, C) sternite VIII, D) sternite IX, E) tergite VIII, F) tergite IX. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/256783/files/figure.png" pageId="9" pageNumber="310">Fig. 8</figureCitation>
|
||
E) (L
|
||
<quantity id="E70590CFFFD8FFF2C33AFEDFFB78F8E2" box="[1118,1221,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="9" pageNumber="310" unit="mm" value="0.57">0.57 mm</quantity>
|
||
,
|
||
<geoCoordinate id="45C95BEDFFD8FFF2C3B6FEC0FA98F8ED" box="[1234,1317,367,392]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="555" value="-0.54">
|
||
W
|
||
<quantity id="E70590CFFFD8FFF2C390FEDFFA98F8ED" box="[1268,1317,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="310" unit="mm" value="0.54">0.54</quantity>
|
||
</geoCoordinate>
|
||
mm) with square apical plate, apical portion covered with dense microsetae, apical margin with row of dense, short setae intermixed with sparse, longer ones, apodemes much shorter than apical portion; tergite IX (
|
||
<figureCitation id="B8C621AFFFD8FFF2C3CFFE07FB50F8B5" box="[1195,1261,439,464]" captionStart="FIGURE 8" captionStartId="9.[151,250,1965,1987]" captionTargetBox="[151,1436,1093,1943]" captionTargetId="figure@9.[151,1436,1093,1944]" captionTargetPageId="9" captionText="FIGURE 8. Anticyphon oyonensis sp. nov., male genitalia. A) penis, B) tegmen, C) sternite VIII, D) sternite IX, E) tergite VIII, F) tergite IX. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/256783/files/figure.png" pageId="9" pageNumber="310">Fig. 8</figureCitation>
|
||
F) (L
|
||
<quantity id="E70590CFFFD8FFF2C255FE07FA25F8AA" box="[1329,1432,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="310" unit="mm" value="0.54">0.54 mm</quantity>
|
||
,
|
||
<geoCoordinate id="45C95BEDFFD8FFF2C7F3FE6DFF54F891" box="[151,233,476,500]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="555" value="-0.41">
|
||
W
|
||
<quantity id="E70590CFFFD8FFF2C7DCFE6CFF54F891" box="[184,233,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="9" pageNumber="310" unit="mm" value="0.41">0.41</quantity>
|
||
</geoCoordinate>
|
||
mm) narrower than tergite VIII, central portion membranous.
|
||
</paragraph>
|
||
<paragraph id="20423D2AFFD8FFF2C7A3FE4FFC26FA71" blockId="9.[151,1437,151,1040]" pageId="9" pageNumber="310">
|
||
Female. Externally not differing from males. Sternite VIII elongate (L
|
||
<quantity id="E70590CFFFD8FFF2C493FDB0FBDDFB72" box="[1015,1120,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="310" unit="mm" value="1.3">1.30 mm</quantity>
|
||
,
|
||
<geoCoordinate id="45C95BEDFFD8FFF2C30AFDB0FB7EFB7D" box="[1134,1219,512,536]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="555" value="-0.3">
|
||
W
|
||
<quantity id="E70590CFFFD8FFF2C3F6FDB0FB7EFB7D" box="[1170,1219,512,536]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="310" unit="mm" value="0.3">0.30</quantity>
|
||
</geoCoordinate>
|
||
mm), with sinuate apodemes which are not fused basally, apex with sparse setae. Tergite VIII very long (L
|
||
<quantity id="E70590CFFFD8FFF2C3EAFD94FB48FB59" box="[1166,1269,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.86" pageId="9" pageNumber="310" unit="mm" value="1.86">1.86 mm</quantity>
|
||
,
|
||
<geoCoordinate id="45C95BEDFFD8FFF2C266FD95FAEAFB59" box="[1282,1367,548,572]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="555" value="-0.72">
|
||
W
|
||
<quantity id="E70590CFFFD8FFF2C242FD94FAEAFB59" box="[1318,1367,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="9" pageNumber="310" unit="mm" value="0.72">0.72</quantity>
|
||
</geoCoordinate>
|
||
mm); apical portion subtrapezoidal, with sparse setation on apical margin; apodemes long, 4× longer than apical portion. Ovipositor (
|
||
<figureCitation id="B8C621AFFFD8FFF2C646FDDCFED4FBE1" box="[290,361,620,645]" captionStart="FIGURE 9" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[151,1436,195,1779]" captionTargetId="figure@10.[151,1436,195,1781]" captionTargetPageId="10" captionText="FIGURE 9. Anticyphon oyonensis sp. nov., female genitalia. A) genitalia, B) vaginal sclerites, C) bursal sclerites, D) bursella. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/256784/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
|
||
A) long (L ca.
|
||
<quantity id="E70590CFFFD8FFF2C544FDDCFDC3FBE1" box="[544,638,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="310" unit="mm" value="2.8">2.8 mm</quantity>
|
||
); branchlets present; coxites (L
|
||
<quantity id="E70590CFFFD8FFF2C49EFDDCFBDEFBE1" box="[1018,1123,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="9" pageNumber="310" unit="mm" value="0.48">0.48 mm</quantity>
|
||
) membranous, triangularly elongate, narrow, not covered with setae; styli with two tufts of setae at apex. Vagina (
|
||
<figureCitation id="B8C621AFFFD8FFF2C337FD3FFB28FBCD" box="[1107,1173,655,680]" captionStart="FIGURE 9" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[151,1436,195,1779]" captionTargetId="figure@10.[151,1436,195,1781]" captionTargetPageId="10" captionText="FIGURE 9. Anticyphon oyonensis sp. nov., female genitalia. A) genitalia, B) vaginal sclerites, C) bursal sclerites, D) bursella. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/256784/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
|
||
B) membranous, with 5 small, irregular sclerites; two bursal sclerites (
|
||
<figureCitation id="B8C621AFFFD8FFF2C5CBFD04FD49FBA9" box="[687,756,692,717]" captionStart="FIGURE 9" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[151,1436,195,1779]" captionTargetId="figure@10.[151,1436,195,1781]" captionTargetPageId="10" captionText="FIGURE 9. Anticyphon oyonensis sp. nov., female genitalia. A) genitalia, B) vaginal sclerites, C) bursal sclerites, D) bursella. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/256784/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
|
||
C) present: larger, rod-like (L
|
||
<quantity id="E70590CFFFD8FFF2C332FD04FB02FBA9" box="[1110,1215,692,717]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="9" pageNumber="310" unit="mm" value="0.61">0.61 mm</quantity>
|
||
,
|
||
<geoCoordinate id="45C95BEDFFD8FFF2C3ABFD05FA9EFBA9" box="[1231,1315,692,716]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="555" value="-0.11">
|
||
W
|
||
<quantity id="E70590CFFFD8FFF2C397FD04FA9EFBA9" box="[1267,1315,692,716]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="310" unit="mm" value="0.11">0.11</quantity>
|
||
</geoCoordinate>
|
||
mm), and smaller, H-shaped (L
|
||
<quantity id="E70590CFFFD8FFF2C6EEFD68FE4CFB8A" box="[394,497,728,752]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="310" unit="mm" value="0.2">0.20 mm</quantity>
|
||
,
|
||
<geoCoordinate id="45C95BEDFFD8FFF2C698FD68FDF2FB95" box="[508,591,728,752]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="555" value="-0.1">
|
||
W
|
||
<quantity id="E70590CFFFD8FFF2C57AFD68FDF2FB95" box="[542,591,728,752]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="310" unit="mm" value="0.1">0.10</quantity>
|
||
</geoCoordinate>
|
||
mm), both in membranous socket (L
|
||
<quantity id="E70590CFFFD8FFF2C493FD68FBE0FB8A" box="[1015,1117,728,752]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="310" unit="mm" value="0.8">0.80 mm</quantity>
|
||
,
|
||
<geoCoordinate id="45C95BEDFFD8FFF2C30DFD68FB01FB95" box="[1129,1212,727,752]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="555" value="-0.35">
|
||
W
|
||
<quantity id="E70590CFFFD8FFF2C3EFFD67FB01FB95" box="[1163,1212,727,752]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="310" unit="mm" value="0.35">0.35</quantity>
|
||
</geoCoordinate>
|
||
mm); bursella (
|
||
<figureCitation id="B8C621AFFFD8FFF2C20BFD67FF18FA71" captionStart="FIGURE 9" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[151,1436,195,1779]" captionTargetId="figure@10.[151,1436,195,1781]" captionTargetPageId="10" captionText="FIGURE 9. Anticyphon oyonensis sp. nov., female genitalia. A) genitalia, B) vaginal sclerites, C) bursal sclerites, D) bursella. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/256784/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
|
||
D) covered with oval, petagonal and hexagonal cuticular structures.
|
||
</paragraph>
|
||
<paragraph id="20423D2AFFD8FFF2C7A3FCAFFAE7FAAD" blockId="9.[151,1437,151,1040]" pageId="9" pageNumber="310">
|
||
<emphasis id="1289E138FFD8FFF2C7A3FCAFFDBDFA5D" bold="true" box="[199,512,799,824]" pageId="9" pageNumber="310">Measurements and ratios.</emphasis>
|
||
Males (n = 3): TL
|
||
<quantity id="E70590CFFFD8FFF2C584FCAFFC38FA52" box="[736,901,799,824]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.65" metricValueMin="4.25" pageId="9" pageNumber="310" unit="mm" value="4.45" valueMax="4.65" valueMin="4.25">4.25–4.65 mm</quantity>
|
||
(
|
||
<quantity id="E70590CFFFD8FFF2C4F1FCAFFBBDFA5D" box="[917,1024,799,824]" metricMagnitude="-3" metricUnit="m" metricValue="4.52" pageId="9" pageNumber="310" unit="mm" value="4.52">4.52 mm</quantity>
|
||
), PL
|
||
<quantity id="E70590CFFFD8FFF2C35AFCAFFB5EFA52" box="[1086,1251,799,824]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="9" pageNumber="310" unit="mm" value="0.725" valueMax="0.75" valueMin="0.7">0.70–0.75 mm</quantity>
|
||
(
|
||
<quantity id="E70590CFFFD8FFF2C396FC90FAE3FA5D" box="[1266,1374,800,824]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="9" pageNumber="310" unit="mm" value="0.72">0.72 mm</quantity>
|
||
), PW
|
||
<quantity id="E70590CFFFD8FFF2C7F3FCF4FE87FA39" box="[151,314,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.65" metricValueMin="1.55" pageId="9" pageNumber="310" unit="mm" value="1.6" valueMax="1.65" valueMin="1.55">1.55–1.65 mm</quantity>
|
||
(
|
||
<quantity id="E70590CFFFD8FFF2C62DFCF4FE0FFA39" box="[329,434,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.61" pageId="9" pageNumber="310" unit="mm" value="1.61">1.61 mm</quantity>
|
||
), EL 3.65–4.00 mm (
|
||
<quantity id="E70590CFFFD8FFF2C5C4FCF4FCB7FA39" box="[672,778,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="3.87" pageId="9" pageNumber="310" unit="mm" value="3.87">3.87 mm</quantity>
|
||
), EW
|
||
<quantity id="E70590CFFFD8FFF2C434FCF4FC4EFA39" box="[848,1011,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="2.525" metricValueMax="2.6" metricValueMin="2.45" pageId="9" pageNumber="310" unit="mm" value="2.525" valueMax="2.6" valueMin="2.45">2.45–2.60 mm</quantity>
|
||
(
|
||
<quantity id="E70590CFFFD8FFF2C365FCF4FBD6FA39" box="[1025,1131,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" pageId="9" pageNumber="310" unit="mm" value="2.55">2.55 mm</quantity>
|
||
), TL/EW 1.73–1.79 (1.77), PW/PL 2.20–2.33 (2.25), EL/EW 1.49–1.54 (1.52), EL/PL 5.21–5.71 (5.40). Females (n = 2) TL
|
||
<quantity id="E70590CFFFD8FFF2C393FCD7FA21FA1A" box="[1271,1436,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="4.375" metricValueMax="4.5" metricValueMin="4.25" pageId="9" pageNumber="310" unit="mm" value="4.375" valueMax="4.5" valueMin="4.25">4.25–4.50 mm</quantity>
|
||
(
|
||
<quantity id="E70590CFFFD8FFF2C7FAFC3CFEB7FAC1" box="[158,266,908,932]" metricMagnitude="-3" metricUnit="m" metricValue="4.38" pageId="9" pageNumber="310" unit="mm" value="4.38">4.38 mm</quantity>
|
||
), PL
|
||
<quantity id="E70590CFFFD8FFF2C623FC3CFE12FAC1" box="[327,431,908,933]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="310" unit="mm" value="0.75">0.75 mm</quantity>
|
||
, PW
|
||
<quantity id="E70590CFFFD8FFF2C68AFC3CFD2EFAC1" box="[494,659,908,933]" metricMagnitude="-3" metricUnit="m" metricValue="1.575" metricValueMax="1.6" metricValueMin="1.55" pageId="9" pageNumber="310" unit="mm" value="1.575" valueMax="1.6" valueMin="1.55">1.55–1.60 mm</quantity>
|
||
(
|
||
<quantity id="E70590CFFFD8FFF2C5C7FC3CFCB3FAC1" box="[675,782,908,933]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="9" pageNumber="310" unit="mm" value="1.58">1.58 mm</quantity>
|
||
), EL
|
||
<quantity id="E70590CFFFD8FFF2C429FC3CFC4FFAC1" box="[845,1010,908,933]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.85" metricValueMin="3.65" pageId="9" pageNumber="310" unit="mm" value="3.75" valueMax="3.85" valueMin="3.65">3.65–3.85 mm</quantity>
|
||
(
|
||
<quantity id="E70590CFFFD8FFF2C366FC3CFBD0FAC1" box="[1026,1133,908,933]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" pageId="9" pageNumber="310" unit="mm" value="3.75">3.75 mm</quantity>
|
||
), EW
|
||
<quantity id="E70590CFFFD8FFF2C3D1FC3CFAE7FAC1" box="[1205,1370,908,933]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="9" pageNumber="310" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.30–2.50 mm</quantity>
|
||
(
|
||
<quantity id="E70590CFFFD8FFF2C20EFC3CFF7AFAAD" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="310" unit="mm" value="2.4">2.40 mm</quantity>
|
||
), TL/EW 1.80–1.85 (1.82), PW/PL 2.07–2.13 (2.10), EL/EW 1.54–1.59 (1.56), EL/PL 4.87–5.13 (5.00).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68E76EA1FFD8FFF2C7A3FC64FCE6FA89" box="[199,859,980,1005]" pageId="9" pageNumber="310" type="distribution">
|
||
<paragraph id="20423D2AFFD8FFF2C7A3FC64FCE6FA89" blockId="9.[151,1437,151,1040]" box="[199,859,980,1005]" pageId="9" pageNumber="310">
|
||
<emphasis id="1289E138FFD8FFF2C7A3FC64FEDDFA88" bold="true" box="[199,352,980,1005]" pageId="9" pageNumber="310">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="FF468388FFD8FFF2C504FC65FD2DFA88" box="[608,656,981,1005]" pageId="9" pageNumber="310">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="B8C621AFFFD8FFF2C59EFC64FCF3FA89" box="[762,846,980,1005]" captionStart="FIGURE 14" captionStartId="16.[151,250,1430,1452]" captionTargetBox="[341,1251,195,1398]" captionTargetId="figure@16.[331,1251,195,1398]" captionTargetPageId="16" captionText="FIGURE 14. Distribution of Anticyphon gen. nov. Please note that precise locality of A. peruvianus sp. nov. is unknown (see note under the species description)." httpUri="https://zenodo.org/record/256789/files/figure.png" pageId="9" pageNumber="310">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68E76EA1FFD8FFF2C7A3FC47FA97FD75" box="[199,1322,1015,1040]" pageId="9" pageNumber="310" type="etymology">
|
||
<paragraph id="20423D2AFFD8FFF2C7A3FC47FA97FD75" blockId="9.[151,1437,151,1040]" box="[199,1322,1015,1040]" pageId="9" pageNumber="310">
|
||
<emphasis id="1289E138FFD8FFF2C7A3FC47FEF3FD75" bold="true" box="[199,334,1015,1040]" pageId="9" pageNumber="310">Etymology.</emphasis>
|
||
Named after Oyón Province, where the
|
||
<typeStatus id="FF468388FFD8FFF2C476FC48FCFFFD75" box="[786,834,1016,1040]" pageId="9" pageNumber="310">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="E7FD46A9FFD8FFF2C4A6FC48FBEDFD6A" box="[962,1104,1016,1039]" class="Insecta" family="Scirtidae" genus="Anticyphon" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="oyonensis" status="sp. nov.">
|
||
<emphasis id="1289E138FFD8FFF2C4A6FC48FBEDFD6A" box="[962,1104,1016,1039]" italics="true" pageId="9" pageNumber="310">A. oyonensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1289E138FFD8FFF2C333FC47FB0DFD75" bold="true" box="[1111,1200,1015,1040]" pageId="9" pageNumber="310">
|
||
<taxonomicNameLabel id="09BA5C43FFD8FFF2C333FC47FB0DFD75" box="[1111,1200,1015,1040]" pageId="9" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is located.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |