300 lines
38 KiB
XML
300 lines
38 KiB
XML
<document id="7CD2721FF9C23D455A5653D34AB91687" ID-CLB-Dataset="7896" ID-DOI="10.11646/zootaxa.4875.1.1" ID-GBIF-Dataset="a75d6e42-221a-4df9-9a7d-5f33dbff6a56" ID-ISSN="1175-5326" ID-Zenodo-Dep="4578779" ID-ZooBank="278200CE-E16F-45B4-9A89-60C2052415C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610024385246" checkinUser="plazi" docAuthor="Görn, Sebastian" docDate="2020" docId="807787F6B81CFFFB49A1D55DFF413D86" docLanguage="en" docName="zootaxa.4875.1.1.pdf" docOrigin="Zootaxa 4875 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heptodonta horii, sp. nov." docType="treatment" docVersion="8" lastPageNumber="36" masterDocId="7C4EFF8EB83EFFD84936D36BFFA83A3C" masterDocTitle="Revision of the Oriental tiger beetle genus Heptodonta Hope, 1838 (Coleoptera: Cicindelidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="35" updateTime="1698923305811" updateUser="ExternalLinkService">
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<mods:titleInfo id="8428B93E1508596E027C9C7EBE39413F">
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<mods:title id="DCC6D833393C16B8736D387A48485488">Revision of the Oriental tiger beetle genus Heptodonta Hope, 1838 (Coleoptera: Cicindelidae)</mods:title>
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<mods:name id="05ECCCA9A88361CC734BEC0D0120A702" type="personal">
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<mods:namePart id="96C72517B1F149DDCE4FF298D5EA4B9C">Görn, Sebastian</mods:namePart>
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<mods:title id="816B3D01872732CFEDB3A9D2E6C1A71B">Zootaxa</mods:title>
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<mods:part id="8FE9EE7FBBF6615DFF4BE4B9DBF65544">
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<mods:date id="13AC99E0E55D7A69D3104E3761B20EE1">2020</mods:date>
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<mods:number id="2234A79D4780F779F153831C62B1FDEB">2020-11-06</mods:number>
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<mods:number id="A548CDD0F1F20D83A0D3652D5ED29F64">4875</mods:number>
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<mods:classification id="998CD51CA633961E7EBBDF06DF5D830C">journal article</mods:classification>
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<mods:identifier id="24C954215494A302239592756E066F63" type="CLB-Dataset">7896</mods:identifier>
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<mods:identifier id="C9C40CA2C64922FC00D9F711B88507D0" type="DOI">10.11646/zootaxa.4875.1.1</mods:identifier>
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<mods:identifier id="B08B3F2B2E72385FDACAE69229C12544" type="GBIF-Dataset">a75d6e42-221a-4df9-9a7d-5f33dbff6a56</mods:identifier>
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<mods:identifier id="3341ED5694E6BD656D3961C3E7B2396D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="98C8791DC5AAD39D97726C33EBC476AB" type="Zenodo-Dep">4578779</mods:identifier>
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<treatment id="807787F6B81CFFFB49A1D55DFF413D86" ID-DOI="http://doi.org/10.5281/zenodo.4579803" ID-GBIF-Taxon="179988685" ID-Zenodo-Dep="4579803" LSID="urn:lsid:plazi:treatment:807787F6B81CFFFB49A1D55DFF413D86" httpUri="http://treatment.plazi.org/id/807787F6B81CFFFB49A1D55DFF413D86" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
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<subSubSection id="40C4656BB81CFFFA49A1D55DFE643C6D" box="[151,460,1590,1617]" pageId="34" pageNumber="35" type="nomenclature">
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<paragraph id="086136E0B81CFFFA49A1D55DFE643C6D" blockId="34.[151,460,1590,1653]" box="[151,460,1590,1617]" pageId="34" pageNumber="35">
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<heading id="5329818CB81CFFFA49A1D55DFE643C6D" bold="true" box="[151,460,1590,1617]" fontSize="11" level="2" pageId="34" pageNumber="35" reason="4">
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<emphasis id="3AAAEAF2B81CFFFA49A1D55DFE643C6D" bold="true" box="[151,460,1590,1617]" pageId="34" pageNumber="35">
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<taxonomicName id="CFDE4D63B81CFFFA49A1D55DFECF3C6D" ID-CoL="8MD4C" authority="Gorn. 2020" box="[151,359,1590,1617]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="horii" status="sp. nov.">
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<emphasis id="3AAAEAF2B81CFFFA49A1D55DFECF3C6D" bold="true" box="[151,359,1590,1617]" italics="true" pageId="34" pageNumber="35">Heptodonta horii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="21995789B81CFFFA4858D55CFE643C6D" box="[366,460,1591,1617]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="40C4656BB81CFFFA49A1D530FEED3C49" box="[151,325,1626,1653]" pageId="34" pageNumber="35" type="description">
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<paragraph id="086136E0B81CFFFA49A1D530FEED3C49" blockId="34.[151,460,1590,1653]" box="[151,325,1626,1653]" pageId="34" pageNumber="35">
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<figureCitation id="90E52A65B81CFFFA49A1D530FF4D3C49" box="[151,229,1627,1653]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1827,1851]" captionTargetBox="[177,1392,191,1791]" captionTargetId="figure-15@5.[177,1410,181,1803]" captionTargetPageId="5" captionText="FIGURES 1–7. 1–2—anterior elytra: 1—Heptodonta melanopyga (Schaum, 1862), ♀, Philippines, Luzon, Banaue (PSCH); 2—H. mindoroensis Cassola, 2000, ♂, Philippines, Mindoro, Mt. Halcon (JWCW). 3–4—pronotum: 3—H. arrowi W. Horn, 1900, ♂, Myanmar, Magway Region, Pyon Village (SGCH); 4—Heptodonta horii sp. nov., ♂, holotype, Myanmar, Bago Re- gion, Chanthakwin (JWCW); 5–7—head: 5—Heptodonta tempesta sp. nov., ♂, holotype, India, Meghalaya, Balpakram NP (JWCW); 6—Heptodonta pulchella (Hope, 1831), ♂, Nepal, Bagmati, Sollenta-Dhading (JWCW); 7—Heptodonta schuelei sp. nov., ♂, holotype, India, Meghalaya, Cherrapunjee (JWCW). Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4578783" httpUri="https://zenodo.org/record/4578783/files/figure.png" pageId="34" pageNumber="35">Figs. 4</figureCitation>
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,
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<figureCitation id="90E52A65B81CFFFA49C7D531FEED3C49" box="[241,325,1626,1653]" captionStart="FIGURES 99–105" captionStartId="38.[151,264,1921,1945]" captionTargetBox="[152,1435,184,1869]" captionTargetId="figure-20@38.[151,1435,181,1897]" captionTargetPageId="38" captionText="FIGURES 99–105. Heptodonta horii sp. nov., Myanmar, Bago Region, Chanthakwin (JWCW): 99, 101, 103–105—♂, holo- type: 99—habitus, 101—head, 103—lateral body, 104—aedeagus in left lateral aspect, 105—aedeagus in right lateral aspect; 100, 102—♀, paratype: 100—habitus, 102—head. Scales: 99–103 = 1 mm, 104–105 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578824" httpUri="https://zenodo.org/record/4578824/files/figure.png" pageId="34" pageNumber="35">99–105</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="40C4656BB81CFFFB49A1D5C8FAFC3A8D" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="materials_examined">
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<paragraph id="086136E0B81CFFFA49A1D5C8FD7C3C81" blockId="34.[151,1437,1698,2013]" box="[151,724,1698,1725]" pageId="34" pageNumber="35">
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<materialsCitation id="B8B63CBDB81CFFFA49A1D5C8FD793C81" ID-GBIF-Occurrence="3045076596" box="[151,721,1698,1725]" country="Myanmar" location="Mon State" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Bago" typeStatus="holotype">
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<emphasis id="3AAAEAF2B81CFFFA49A1D5C8FE9E3C80" bold="true" box="[151,310,1698,1725]" pageId="34" pageNumber="35">
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<typeStatus id="D7658842B81CFFFA49A1D5C8FF7A3C81" box="[151,210,1699,1725]" pageId="34" pageNumber="35">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="70C97670B81CFFFA480AD5C8FE053C81" box="[316,429,1699,1725]" name="Myanmar" pageId="34" pageNumber="35">Myanmar</collectingCountry>
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:
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<collectingRegion id="CA1AF802B81CFFFA488ED5C8FDE63C81" box="[440,590,1699,1725]" country="Myanmar" name="Bago" pageId="34" pageNumber="35">Bago Region</collectingRegion>
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,
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<location id="0D01603BB81CFFFA4B6FD5C8FD793C81" LSID="urn:lsid:plazi:treatment:807787F6B81CFFFB49A1D55DFF413D86:0D01603BB81CFFFA4B6FD5C8FD793C81" box="[601,721,1699,1725]" country="Myanmar" name="Mon State" pageId="34" pageNumber="35" stateProvince="Bago">Mon State</location>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="086136E0B81CFFFA49F1D5ACFA343DE1" blockId="34.[151,1437,1698,2013]" pageId="34" pageNumber="35">
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<emphasis id="3AAAEAF2B81CFFFA49F1D5ACFEDE3CDC" bold="true" box="[199,374,1734,1761]" pageId="34" pageNumber="35">Type material.</emphasis>
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<materialsCitation id="B8B63CBDB81CFFFA4848D5ACFAB43D15" ID-GBIF-Occurrence="3045076456" collectingDate="1997-05" collectorName="M. Lgt & Det. F. Cassola" country="Myanmar" determinerName="Det. F. Cassola" location="PEGU" pageId="34" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Bago" typeStatus="holotype">
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<emphasis id="3AAAEAF2B81CFFFA4848D5ACFE433CDD" bold="true" box="[382,491,1735,1761]" pageId="34" pageNumber="35">
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<typeStatus id="D7658842B81CFFFA4848D5ACFE433CDD" box="[382,491,1735,1761]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
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</emphasis>
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:
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<specimenCount id="1ED8FD69B81CFFFA48CDD5ACFDB83CDD" box="[507,528,1735,1761]" pageId="34" pageNumber="35" type="male">♂</specimenCount>
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in JWCW, labelled: “
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<date id="7C601020B81CFFFA4A31D5ACFCC93CDD" box="[775,865,1735,1761]" pageId="34" pageNumber="35" value="1997-05">
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<collectingDate id="6C24E9C8B81CFFFA4A31D5ACFCC93CDD" box="[775,865,1735,1761]" pageId="34" pageNumber="35" value="1997-05">V. 1997</collectingDate>
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</date>
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<collectingCountry id="70C97670B81CFFFA4A5FD5ACFC663CDD" box="[873,974,1735,1761]" name="Myanmar" pageId="34" pageNumber="35">BURMA</collectingCountry>
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/
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<collectingRegion id="CA1AF802B81CFFFA4AD0D5ACFB873CDD" box="[998,1071,1735,1761]" country="Myanmar" name="Bago" pageId="34" pageNumber="35">PEGU</collectingRegion>
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/ CHANTHAKWIN / KLÍCHA
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<collectorName id="A52B5336B81CFFFA49A1D580FF403D39" box="[151,232,1771,1797]" pageId="34" pageNumber="35">M. Lgt</collectorName>
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” [with yellow frame, printed] // “SANDY WALLS OF / RAVINERS - IN / SHADE OF BUSHES - / NOT IN OPEN SUN” [printed] // “
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<taxonomicName id="CFDE4D63B81CFFFA48EFD464FD3C3D15" authorityName="Arrowi W. Horn" authorityYear="1900" box="[473,660,1807,1833]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">HEPTODONTA</taxonomicName>
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/ arrowi / W.
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<bibRefCitation id="6C4F4B11B81CFFFA4A1ED464FC0E3D15" author="Horn, W." box="[808,934,1807,1833]" pageId="34" pageNumber="35" pagination="361 - 365" refId="ref39094" refString="Horn, W. (1900) 7 neue Cicindeliden. Deutsche Entomologische Zeitschrift, 2, 361 - 365." type="journal article" year="1900">Horn, 1900</bibRefCitation>
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/
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<determinerName id="FDBDDA24B81CFFFA4A8FD464FBCA3D15" box="[953,1122,1807,1833]" pageId="34" pageNumber="35">
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<collectorName id="A52B5336B81CFFFA4A8FD464FBCA3D15" box="[953,1122,1807,1833]" pageId="34" pageNumber="35">Det. F. Cassola</collectorName>
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</determinerName>
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, 2004” [printed]
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</materialsCitation>
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.
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<emphasis id="3AAAEAF2B81CFFFA4C12D464FA343D15" bold="true" box="[1316,1436,1807,1833]" pageId="34" pageNumber="35">
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<typeStatus id="D7658842B81CFFFA4C12D464FA343D15" box="[1316,1436,1807,1833]" pageId="34" pageNumber="35" type="paratype">Paratypes</typeStatus>
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</emphasis>
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(7):
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<specimenCount id="1ED8FD69B81CFFFA49FED458FF433D71" box="[200,235,1843,1869]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
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<specimenCount id="1ED8FD69B81CFFFA49C2D458FE9C3D71" box="[244,308,1843,1869]" pageId="34" pageNumber="35" type="female">2 ♀♀</specimenCount>
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in JWCW,
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<materialsCitation id="B8B63CBDB81CFFFA488DD458FD6D3D4D" ID-GBIF-Occurrence="3045076537" collectingDate="1997-05" collectorName="M. Lgt & Det. F. Cassola" country="Myanmar" determinerName="Det. F. Cassola" location="PEGU" pageId="34" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Bago" typeStatus="holotype">
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<specimenCount id="1ED8FD69B81CFFFA488DD458FE4E3D71" box="[443,486,1843,1869]" pageId="34" pageNumber="35" type="male">1 ♂</specimenCount>
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in SMNS with same locality labels as
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<typeStatus id="D7658842B81CFFFA4A93D458FBAF3D71" box="[933,1031,1843,1869]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
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plus: “
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<taxonomicName id="CFDE4D63B81CFFFA4D6ED458FABB3D71" authorityName="Arrowi W. Horn" authorityYear="1900" box="[1112,1299,1843,1869]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">HEPTODONTA</taxonomicName>
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/ ARROWI / (W.
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<bibRefCitation id="6C4F4B11B81CFFFA49EFD43CFEFD3D4D" author="Horn, W." box="[217,341,1879,1905]" pageId="34" pageNumber="35" pagination="361 - 365" refId="ref39094" refString="Horn, W. (1900) 7 neue Cicindeliden. Deutsche Entomologische Zeitschrift, 2, 361 - 365." type="journal article" year="1900">Horn 1900</bibRefCitation>
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) / K. WERNER Det.” [printed]
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</materialsCitation>
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.
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<materialsCitation id="B8B63CBDB81CFFFA4BE7D43CFDEA3D85" ID-GBIF-Occurrence="3045076367" collectingDate="1997-05" collectorName="M. Lgt & K. Werner" country="Myanmar" location="PEGU" pageId="34" pageNumber="35" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Bago" typeStatus="paratype">
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<specimenCount id="1ED8FD69B81CFFFA4BE7D43CFD563D4D" box="[721,766,1879,1905]" pageId="34" pageNumber="35" type="male">1 ♂</specimenCount>
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<specimenCount id="1ED8FD69B81CFFFA4A31D43CFC9B3D4D" box="[775,819,1879,1905]" pageId="34" pageNumber="35" type="female">1 ♀</specimenCount>
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in ZSM: “
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<date id="7C601020B81CFFFA4A86D43CFBA23D4D" box="[944,1034,1879,1905]" pageId="34" pageNumber="35" value="1997-05">
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<collectingDate id="6C24E9C8B81CFFFA4A86D43CFBA23D4D" box="[944,1034,1879,1905]" pageId="34" pageNumber="35" value="1997-05">V. 1997</collectingDate>
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</date>
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<collectingCountry id="70C97670B81CFFFA4D25D43CFBD03D4D" box="[1043,1144,1879,1905]" name="Myanmar" pageId="34" pageNumber="35">BURMA</collectingCountry>
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/
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<collectingRegion id="CA1AF802B81CFFFA4DA5D43CFB743D4D" box="[1171,1244,1879,1905]" country="Myanmar" name="Bago" pageId="34" pageNumber="35">PEGU</collectingRegion>
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/ CHANTHAK- WIN / KLÍCHA
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<collectorName id="A52B5336B81CFFFA4863D410FE0D3DA9" box="[341,421,1915,1941]" pageId="34" pageNumber="35">M. Lgt</collectorName>
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” [printed] // “
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<taxonomicName id="CFDE4D63B81CFFFA4B76D410FD6E3DA9" authorityName="Arrowi W. Horn" authorityYear="1900" box="[576,710,1915,1941]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
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/
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||
<taxonomicName id="CFDE4D63B81CFFFA4BEAD410FC713DA9" authority="W. HORN 1900" authorityName="W. Horn" authorityYear="1900" box="[732,985,1915,1941]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="arrowi">
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arrowi W.
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<bibRefCitation id="6C4F4B11B81CFFFA4A7CD410FC713DA9" author="Horn, W." box="[842,985,1915,1941]" pageId="34" pageNumber="35" pagination="361 - 365" refId="ref39094" refString="Horn, W. (1900) 7 neue Cicindeliden. Deutsche Entomologische Zeitschrift, 2, 361 - 365." type="journal article" year="1900">HORN 1900</bibRefCitation>
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</taxonomicName>
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/ WERNER det. ’97” [printed] // “ZSM – Samml. /
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<collectorName id="A52B5336B81CFFFA4821D4F4FE233D85" box="[279,395,1951,1977]" pageId="34" pageNumber="35">K. Werner</collectorName>
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2009” [printed]
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</materialsCitation>
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.
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<materialsCitation id="B8B63CBDB81CFFFA4B7AD4F4FA343DE1" ID-GBIF-Occurrence="3045076535" collectingDate="2018-06-02" collectorName="Thanda Moe & All" country="Myanmar" location="Mon State" municipality="Mokkamu" pageId="34" pageNumber="35" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<specimenCount id="1ED8FD69B81CFFFA4B7AD4F4FDDD3D85" box="[588,629,1951,1977]" pageId="34" pageNumber="35" type="female">1 ♀</specimenCount>
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in SGCH: “vertical clay bank of a small trench, /
|
||
<collectingMunicipality id="E805AC9AB81CFFFA4D97D4F4FABF3D85" box="[1185,1303,1951,1977]" pageId="34" pageNumber="35">Mokkamu</collectingMunicipality>
|
||
,
|
||
<location id="0D01603BB81CFFFA4C14D4F4FA303D85" LSID="urn:lsid:plazi:treatment:807787F6B81CFFFB49A1D55DFF413D86:0D01603BB81CFFFA4C14D4F4FA303D85" box="[1314,1432,1951,1977]" country="Myanmar" municipality="Mokkamu" name="Mon State" pageId="34" pageNumber="35">Mon State</location>
|
||
, /
|
||
<collectingCountry id="70C97670B81CFFFA4990D4A8FE903DE1" box="[166,312,1987,2013]" name="Myanmar" pageId="34" pageNumber="35">MYANMAR</collectingCountry>
|
||
/
|
||
<date id="7C601020B81CFFFA4878D4A8FE753DE1" box="[334,477,1987,2013]" pageId="34" pageNumber="35" value="2018-06-02">
|
||
<collectingDate id="6C24E9C8B81CFFFA4878D4A8FE753DE1" box="[334,477,1987,2013]" pageId="34" pageNumber="35" value="2018-06-02">June 2, 2018</collectingDate>
|
||
</date>
|
||
/ Michio HORI &
|
||
<collectorName id="A52B5336B81CFFFA4B85D4A8FCE93DE1" box="[691,833,1987,2013]" pageId="34" pageNumber="35">Thanda Moe</collectorName>
|
||
leg.” [printed].
|
||
<collectorName id="A52B5336B81CFFFA4AC5D4A8FBBF3DE1" box="[1011,1047,1987,2013]" pageId="34" pageNumber="35">All</collectorName>
|
||
type specimens labelled: “HOLO-
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="086136E0B81DFFFB49A1D3FCFAFC3A8D" blockId="35.[151,1437,150,1978]" box="[151,1364,150,178]" pageId="35" pageNumber="36">
|
||
TYPE (or
|
||
<typeStatus id="D7658842B81DFFFB483AD3FCFE3F3A8D" box="[268,407,151,177]" pageId="35" pageNumber="36" type="paratype">PARATYPE</typeStatus>
|
||
respectively) /
|
||
<taxonomicName id="CFDE4D63B81DFFFB4B72D3FCFD6F3A8D" authorityName="Arrowi W. Horn" authorityYear="1900" box="[580,711,151,177]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
|
||
/ horii
|
||
<taxonomicNameLabel id="21995789B81DFFFB4A2FD3FCFCC73A8D" box="[793,879,151,177]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
/ ded. Sebastian Görn 2020” [red, printed].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="40C4656BB81DFFFB49F1D3D1FE7C3BC9" pageId="35" pageNumber="36" type="diagnosis">
|
||
<paragraph id="086136E0B81DFFFB49F1D3D1FE7C3BC9" blockId="35.[151,1437,150,1978]" pageId="35" pageNumber="36">
|
||
<emphasis id="3AAAEAF2B81DFFFB49F1D3D1FEEA3AE8" bold="true" box="[199,322,186,212]" pageId="35" pageNumber="36">Diagnosis.</emphasis>
|
||
Small slender species with typical labral shape. Resembling
|
||
<taxonomicName id="CFDE4D63B81DFFFB4ADCD3D0FBFE3AE8" authorityName="W. Horn" authorityYear="1900" box="[1002,1110,187,212]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="arrowi">
|
||
<emphasis id="3AAAEAF2B81DFFFB4ADCD3D0FBFE3AE8" box="[1002,1110,187,212]" italics="true" pageId="35" pageNumber="36">H. arrowi</emphasis>
|
||
</taxonomicName>
|
||
, but distinguishable by acute aedeagal apex, wavy diagonal spines row within aedeagal inner sac (best seen in right lateral aspect), slender elytral and pronotal shape, finer pronotal striation (
|
||
<figureCitation id="90E52A65B81DFFFB4BA5D268FD723B21" box="[659,730,259,285]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1827,1851]" captionTargetBox="[177,1392,191,1791]" captionTargetId="figure-15@5.[177,1410,181,1803]" captionTargetPageId="5" captionText="FIGURES 1–7. 1–2—anterior elytra: 1—Heptodonta melanopyga (Schaum, 1862), ♀, Philippines, Luzon, Banaue (PSCH); 2—H. mindoroensis Cassola, 2000, ♂, Philippines, Mindoro, Mt. Halcon (JWCW). 3–4—pronotum: 3—H. arrowi W. Horn, 1900, ♂, Myanmar, Magway Region, Pyon Village (SGCH); 4—Heptodonta horii sp. nov., ♂, holotype, Myanmar, Bago Re- gion, Chanthakwin (JWCW); 5–7—head: 5—Heptodonta tempesta sp. nov., ♂, holotype, India, Meghalaya, Balpakram NP (JWCW); 6—Heptodonta pulchella (Hope, 1831), ♂, Nepal, Bagmati, Sollenta-Dhading (JWCW); 7—Heptodonta schuelei sp. nov., ♂, holotype, India, Meghalaya, Cherrapunjee (JWCW). Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4578783" httpUri="https://zenodo.org/record/4578783/files/figure.png" pageId="35" pageNumber="36">Fig. 4</figureCitation>
|
||
), and considerably reduced third labral teeth. There might be some confusion with exceptionally small specimens of
|
||
<taxonomicName id="CFDE4D63B81DFFFB4A35D24CFC253B7D" baseAuthorityName="Hope" baseAuthorityYear="1831" box="[771,909,295,321]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="3AAAEAF2B81DFFFB4A35D24CFC253B7D" box="[771,909,295,321]" italics="true" pageId="35" pageNumber="36">H. pulchella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="CFDE4D63B81DFFFB4AADD24CFBBE3B7C" authorityName="Chaudoir" authorityYear="1865" box="[923,1046,295,320]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="eugenia">
|
||
<emphasis id="3AAAEAF2B81DFFFB4AADD24CFBBE3B7C" box="[923,1046,295,320]" italics="true" pageId="35" pageNumber="36">H. eugenia</emphasis>
|
||
</taxonomicName>
|
||
, or
|
||
<taxonomicName id="CFDE4D63B81DFFFB4D74D24CFB793B7D" authorityName="W. Horn" authorityYear="1908" box="[1090,1233,295,321]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="vermifera">
|
||
<emphasis id="3AAAEAF2B81DFFFB4D74D24CFB793B7D" box="[1090,1233,295,321]" italics="true" pageId="35" pageNumber="36">H. vermifera</emphasis>
|
||
</taxonomicName>
|
||
, but in contrast to those species the labrum of
|
||
<taxonomicName id="CFDE4D63B81DFFFB48FDD220FD8B3B59" box="[459,547,331,357]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="horii" status="sp. nov.">
|
||
<emphasis id="3AAAEAF2B81DFFFB48FDD220FD8B3B59" box="[459,547,331,357]" italics="true" pageId="35" pageNumber="36">H. horii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3AAAEAF2B81DFFFB4B1FD220FD2A3B59" bold="true" box="[553,642,331,357]" pageId="35" pageNumber="36">
|
||
<taxonomicNameLabel id="21995789B81DFFFB4B1FD220FD2A3B59" box="[553,642,331,357]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a specific semicircular shape in males and semicircular-trapezoidal shape in females, and the elytra have slightly reduced microsculpture and dense, shallowly fused punctures. Unlike
|
||
<taxonomicName id="CFDE4D63B81DFFFB49FFD2F8FEFC3B91" baseAuthorityName="Hope" baseAuthorityYear="1831" box="[201,340,403,429]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="3AAAEAF2B81DFFFB49FFD2F8FEFC3B91" box="[201,340,403,429]" italics="true" pageId="35" pageNumber="36">H. pulchella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="CFDE4D63B81DFFFB48BAD2F8FDB43B91" authorityName="W. Horn" authorityYear="1908" box="[396,540,403,429]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="vermifera">
|
||
<emphasis id="3AAAEAF2B81DFFFB48BAD2F8FDB43B91" box="[396,540,403,429]" italics="true" pageId="35" pageNumber="36">H. vermifera</emphasis>
|
||
</taxonomicName>
|
||
, the mesepisternum is violet. It is distinct from
|
||
<taxonomicName id="CFDE4D63B81DFFFB4D0AD2F8FB103B90" authorityName="Chaudoir" authorityYear="1865" box="[1084,1208,403,428]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="eugenia">
|
||
<emphasis id="3AAAEAF2B81DFFFB4D0AD2F8FB103B90" box="[1084,1208,403,428]" italics="true" pageId="35" pageNumber="36">H. eugenia</emphasis>
|
||
</taxonomicName>
|
||
by dense anteapical punctures on elytra, and from
|
||
<taxonomicName id="CFDE4D63B81DFFFB48D8D2DCFDD43BED" baseAuthorityName="White" baseAuthorityYear="1844" box="[494,636,439,465]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="posticalis">
|
||
<emphasis id="3AAAEAF2B81DFFFB48D8D2DCFDD43BED" box="[494,636,439,465]" italics="true" pageId="35" pageNumber="36">H. posticalis</emphasis>
|
||
</taxonomicName>
|
||
, which has a similar elytral surface, by different labral shape and total body length under
|
||
<quantity id="CF269B05B81DFFFB485CD2B1FE783BC9" box="[362,464,474,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="35" pageNumber="36" unit="mm" value="11.5">11.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="40C4656BB81DFFFB49F1D294FEDE3801" pageId="35" pageNumber="36" type="etymology">
|
||
<paragraph id="086136E0B81DFFFB49F1D294FEDE3801" blockId="35.[151,1437,150,1978]" pageId="35" pageNumber="36">
|
||
<emphasis id="3AAAEAF2B81DFFFB49F1D294FEE63825" bold="true" box="[199,334,511,537]" pageId="35" pageNumber="36">Etymology.</emphasis>
|
||
This new species is dedicated to Michio Hori for his essential support with material, images, and habitat information.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="40C4656BB81DFFFB49F1D12DFCB93D71" pageId="35" pageNumber="36" type="description">
|
||
<paragraph id="086136E0B81DFFFB49F1D12DFAB5385D" blockId="35.[151,1437,150,1978]" box="[199,1309,582,610]" pageId="35" pageNumber="36">
|
||
<emphasis id="3AAAEAF2B81DFFFB49F1D12DFEF1385C" bold="true" box="[199,345,582,608]" pageId="35" pageNumber="36">Description.</emphasis>
|
||
Body size: Length
|
||
<quantity id="CF269B05B81DFFFB4B02D12CFD61385D" box="[564,713,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="10.6" metricValueMin="8.9" pageId="35" pageNumber="36" unit="mm" value="9.75" valueMax="10.6" valueMin="8.9">8.9–10.6 mm</quantity>
|
||
(without labrum), width
|
||
<quantity id="CF269B05B81DFFFB4AD5D12DFBC2385D" box="[995,1130,582,609]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.1" metricValueMin="2.5" pageId="35" pageNumber="36" unit="mm" value="2.8" valueMax="3.1" valueMin="2.5">2.5–3.1 mm</quantity>
|
||
(
|
||
<figureCitation id="90E52A65B81DFFFB4D4FD12CFAB8385D" box="[1145,1296,583,609]" captionStart="FIGURES 99–105" captionStartId="38.[151,264,1921,1945]" captionTargetBox="[152,1435,184,1869]" captionTargetId="figure-20@38.[151,1435,181,1897]" captionTargetPageId="38" captionText="FIGURES 99–105. Heptodonta horii sp. nov., Myanmar, Bago Region, Chanthakwin (JWCW): 99, 101, 103–105—♂, holo- type: 99—habitus, 101—head, 103—lateral body, 104—aedeagus in left lateral aspect, 105—aedeagus in right lateral aspect; 100, 102—♀, paratype: 100—habitus, 102—head. Scales: 99–103 = 1 mm, 104–105 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578824" httpUri="https://zenodo.org/record/4578824/files/figure.png" pageId="35" pageNumber="36">Figs. 99–100</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="086136E0B81DFFFB49F1D100FD2F3E65" blockId="35.[151,1437,150,1978]" pageId="35" pageNumber="36">
|
||
Dorsal surface of head metallic reddish and green. Frons transversely grooved. Vertex anterior and in transition to orbital plates with rather straight longitudinal grooves, posterior with crumpled wrinkles. Orbital plates with fine striae that are wrinkled anteriorly, and with two setae on each side. Occiput with wrinkled transverse grooves, anteromedian irregularly rugose. Genae glabrous and considerably grooved, from anterior to posterior yellow, green, blue, and violet. Clypeus glabrous, copper and green with two anterior blue spots. Labrum testaceous, with four setae, five apical teeth, and one lateral tooth on each side, male labrum semicircular (
|
||
<quantity id="CF269B05B81DFFFB4D72D074FB403905" box="[1092,1256,799,825]" metricMagnitude="-4" metricUnit="m" metricValue="7.1499999999999995" metricValueMax="7.3" metricValueMin="7.0" pageId="35" pageNumber="36" unit="mm" value="0.715" valueMax="0.73" valueMin="0.7">0.70–0.73 mm</quantity>
|
||
long,
|
||
<quantity id="CF269B05B81DFFFB4C1AD074FF6B3961" metricMagnitude="-3" metricUnit="m" metricValue="1.265" metricValueMax="1.3" metricValueMin="1.23" pageId="35" pageNumber="36" unit="mm" value="1.265" valueMax="1.3" valueMin="1.23">1.23–1.30 mm</quantity>
|
||
wide,
|
||
<figureCitation id="90E52A65B81DFFFB4823D028FED03961" box="[277,376,835,861]" captionStart="FIGURES 99–105" captionStartId="38.[151,264,1921,1945]" captionTargetBox="[152,1435,184,1869]" captionTargetId="figure-20@38.[151,1435,181,1897]" captionTargetPageId="38" captionText="FIGURES 99–105. Heptodonta horii sp. nov., Myanmar, Bago Region, Chanthakwin (JWCW): 99, 101, 103–105—♂, holo- type: 99—habitus, 101—head, 103—lateral body, 104—aedeagus in left lateral aspect, 105—aedeagus in right lateral aspect; 100, 102—♀, paratype: 100—habitus, 102—head. Scales: 99–103 = 1 mm, 104–105 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578824" httpUri="https://zenodo.org/record/4578824/files/figure.png" pageId="35" pageNumber="36">Fig. 101</figureCitation>
|
||
) with reduced apical teeth; female labrum trapezoidal (1.00–
|
||
<quantity id="CF269B05B81DFFFB4D77D029FB003961" box="[1089,1192,834,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="35" pageNumber="36" unit="mm" value="1.05">1.05 mm</quantity>
|
||
long,
|
||
<quantity id="CF269B05B81DFFFB4DC0D029FA353961" box="[1270,1437,834,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.415" metricValueMax="1.48" metricValueMin="1.35" pageId="35" pageNumber="36" unit="mm" value="1.415" valueMax="1.48" valueMin="1.35">1.35–1.48 mm</quantity>
|
||
wide,
|
||
<figureCitation id="90E52A65B81DFFFB49EAD00CFE9239BD" box="[220,314,871,897]" captionStart="FIGURES 99–105" captionStartId="38.[151,264,1921,1945]" captionTargetBox="[152,1435,184,1869]" captionTargetId="figure-20@38.[151,1435,181,1897]" captionTargetPageId="38" captionText="FIGURES 99–105. Heptodonta horii sp. nov., Myanmar, Bago Region, Chanthakwin (JWCW): 99, 101, 103–105—♂, holo- type: 99—habitus, 101—head, 103—lateral body, 104—aedeagus in left lateral aspect, 105—aedeagus in right lateral aspect; 100, 102—♀, paratype: 100—habitus, 102—head. Scales: 99–103 = 1 mm, 104–105 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578824" httpUri="https://zenodo.org/record/4578824/files/figure.png" pageId="35" pageNumber="36">Fig. 102</figureCitation>
|
||
) with three acute median teeth and reduced third apical teeth. Mandibles testaceous in females, pale testaceous in males, with rufous-testaceous teeth. Labial and maxillary palpi testaceous, terminal palpomeres in females entirely dark-brown, in males apically or entirely dark-brown; female pre-terminal maxillary palpomeres dark-testaceous. Antennae slender, extending back over the first half of the elytra in males, dark-brown, but first six antennomeres frequently dark-testaceous, scape with a single apical seta (one female with two setae on a scape), antennomeres 3–4 with few scattered setae, antennomeres 5–11 finely and evenly pubescent; in males terminal antennomere slightly to conspicuously dilated.
|
||
</paragraph>
|
||
<paragraph id="086136E0B81DFFFB49F1D708FC833F31" blockId="35.[151,1437,150,1978]" pageId="35" pageNumber="36">
|
||
Thorax entirely glabrous. Pronotum metallic green with expanded reddish-bronze areas on anterior lobe, posterior lobe, and disc, slightly elongated (1.75–2.00 mm long,
|
||
<quantity id="CF269B05B81DFFFB4A10D7EDFC6F3E9D" box="[806,967,1158,1185]" metricMagnitude="-3" metricUnit="m" metricValue="1.7149999999999999" metricValueMax="1.88" metricValueMin="1.55" pageId="35" pageNumber="36" unit="mm" value="1.7149999999999999" valueMax="1.88" valueMin="1.55">1.55–1.88 mm</quantity>
|
||
wide), moderate median line blue, moderate transverse grooves wrinkled on disc, pronounced anterior and posterior sulci, posterior lobe almost as wide as anterior, lateral margins of median lobe rather convex. Proepisternum and mesepisternum violet, metepisternum, sterna and epimera bluish-green with some greenish lustre.
|
||
</paragraph>
|
||
<paragraph id="086136E0B81DFFFB49F1D67CFCBE3FFD" blockId="35.[151,1437,150,1978]" pageId="35" pageNumber="36">
|
||
Elytra elongate, length
|
||
<quantity id="CF269B05B81DFFFB48FBD67DFDFF3F0D" box="[461,599,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="6.5" metricValueMin="5.3" pageId="35" pageNumber="36" unit="mm" value="5.9" valueMax="6.5" valueMin="5.3">5.3–6.5 mm</quantity>
|
||
, slightly dilated laterally in females and almost parallel in males, maximal width at or a little anterior to the middle, metallic reddish-bronze with some greenish lateral colouration, lateral margin violet, reduced (but still visible) microsculpture, juxtahumeral and apical impression slightly pronounced, discal impression and basodiscal convexity moderate, posterior gibbosity lacking, dense punctures slightly deeper anterior and forming shallowly fused transverse grooves.
|
||
</paragraph>
|
||
<paragraph id="086136E0B81DFFFB49F1D6A0FADF3C49" blockId="35.[151,1437,150,1978]" pageId="35" pageNumber="36">Legs elongated, procoxae and mesocoxae testaceous with gently dark-testaceous and iridescent green colouration, metacoxae metallic blue to green and with testaceous posterior apex, pro- and mesocoxae anteriorly, metacoxae laterally densely covered with long white setae, mesocoxae posteriorly with few white setae. Trochanters pale-testaceous. Femora testaceous with dark-brown distal apices, dark colouration dorsally slightly extended. Tarsi darktestaceous with dark-brown distal apex or dark-brown with lightened proximal apex, slender. Claws testaceous.</paragraph>
|
||
<paragraph id="086136E0B81DFFFB49F1D514FD923C81" blockId="35.[151,1437,150,1978]" pageId="35" pageNumber="36">
|
||
Abdominal sternites glabrous (apart from few long setae at posterior margins), dark-brown with bluish reflections on first five sternites (
|
||
<figureCitation id="90E52A65B81DFFFB48FDD5C8FD853C81" box="[459,557,1699,1725]" captionStart="FIGURES 99–105" captionStartId="38.[151,264,1921,1945]" captionTargetBox="[152,1435,184,1869]" captionTargetId="figure-20@38.[151,1435,181,1897]" captionTargetPageId="38" captionText="FIGURES 99–105. Heptodonta horii sp. nov., Myanmar, Bago Region, Chanthakwin (JWCW): 99, 101, 103–105—♂, holo- type: 99—habitus, 101—head, 103—lateral body, 104—aedeagus in left lateral aspect, 105—aedeagus in right lateral aspect; 100, 102—♀, paratype: 100—habitus, 102—head. Scales: 99–103 = 1 mm, 104–105 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578824" httpUri="https://zenodo.org/record/4578824/files/figure.png" pageId="35" pageNumber="36">Fig. 103</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="086136E0B81DFFFB49F1D5ACFCB93D71" blockId="35.[151,1437,150,1978]" pageId="35" pageNumber="36">
|
||
Aedeagus arched, length
|
||
<quantity id="CF269B05B81DFFFB48E8D5ACFD2A3CDD" box="[478,642,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="2.265" metricValueMax="2.33" metricValueMin="2.2" pageId="35" pageNumber="36" unit="mm" value="2.265" valueMax="2.33" valueMin="2.2">2.20–2.33 mm</quantity>
|
||
, slight medioventral convexity, pointed apex elongated ventrally, dorsal continuously curved, aedeagal base rather steeply sloping. Inner sac in right lateral aspect with broad wavy darktestaceous apical-anteromedian spine row that is diagonally directed, in left lateral aspect inner sac median with diagonal anterodorsal flabellate sclerite (
|
||
<figureCitation id="90E52A65B81DFFFB4B69D458FCAD3D71" box="[607,773,1842,1869]" captionStart="FIGURES 99–105" captionStartId="38.[151,264,1921,1945]" captionTargetBox="[152,1435,184,1869]" captionTargetId="figure-20@38.[151,1435,181,1897]" captionTargetPageId="38" captionText="FIGURES 99–105. Heptodonta horii sp. nov., Myanmar, Bago Region, Chanthakwin (JWCW): 99, 101, 103–105—♂, holo- type: 99—habitus, 101—head, 103—lateral body, 104—aedeagus in left lateral aspect, 105—aedeagus in right lateral aspect; 100, 102—♀, paratype: 100—habitus, 102—head. Scales: 99–103 = 1 mm, 104–105 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578824" httpUri="https://zenodo.org/record/4578824/files/figure.png" pageId="35" pageNumber="36">Figs. 104–105</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="40C4656BB81DFFFB49F1D43DFC263D4D" box="[199,910,1878,1905]" pageId="35" pageNumber="36" type="distribution">
|
||
<paragraph id="086136E0B81DFFFB49F1D43DFC263D4D" blockId="35.[151,1437,150,1978]" box="[199,910,1878,1905]" pageId="35" pageNumber="36">
|
||
<emphasis id="3AAAEAF2B81DFFFB49F1D43DFEC93D4C" bold="true" box="[199,353,1878,1904]" pageId="35" pageNumber="36">Distribution.</emphasis>
|
||
Central
|
||
<emphasis id="3AAAEAF2B81DFFFB48F7D43DFDF03D4C" bold="true" box="[449,600,1878,1904]" pageId="35" pageNumber="36">
|
||
<collectingCountry id="70C97670B81DFFFB48F7D43DFDF03D4C" box="[449,600,1878,1904]" name="Myanmar" pageId="35" pageNumber="36">MYANMAR</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="CA1AF802B81DFFFB4B51D43CFD573D4D" box="[615,767,1879,1905]" country="Myanmar" name="Bago" pageId="35" pageNumber="36">Bago Region</collectingRegion>
|
||
, Mon State).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="40C4656BB81DFFFB49F1D411FF413D86" pageId="35" pageNumber="36" type="biology_ecology">
|
||
<paragraph id="086136E0B81DFFFB49F1D411FF413D86" blockId="35.[151,1437,150,1978]" pageId="35" pageNumber="36">
|
||
<emphasis id="3AAAEAF2B81DFFFB49F1D411FE8F3DA8" bold="true" box="[199,295,1914,1940]" pageId="35" pageNumber="36">Biology.</emphasis>
|
||
In shaded habitats on vertical surfaces such as sandy walls and clay banks, along shaded paths (see labels).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |