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<document id="F21A45CACCBD7663FE04555641AD1FF2" ID-DOI="10.24199/j.mmv.2012.69.05" ID-ISSN="1447-2554" ID-Zenodo-Dep="12212378" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718994422343" checkinUser="felipe" docAuthor="OLoughlin, P. Mark, Barmos, Shari &amp; VandenSpiegel, Didier" docDate="2012" docId="7E4A044D190EFFB79A9DFBDC941E88A1" docLanguage="en" docName="MemMusVictoria.69.269-308.pdf" docOrigin="Memoirs of Museum Victoria 69" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-69-2012/pages-269-308/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Thyone Oken 1815" docType="treatment" docVersion="2" lastPageNumber="295" masterDocId="82737C351914FFAD9A11FFCF956F8E08" masterDocTitle="The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae)" masterLastPageNumber="308" masterPageNumber="269" pageNumber="295" updateTime="1719008797983" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="55E1CA3E2344D0968126E37E72CE603B">The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae)</mods:title>
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<subSubSection id="BEF9E6D0190EFFB79A9DFBDC943E8A21" box="[140,337,1043,1065]" pageId="26" pageNumber="295" type="nomenclature">
<paragraph id="F65CB55B190EFFB79A9DFBDC943E8A21" blockId="26.[140,337,1043,1065]" box="[140,337,1043,1065]" pageId="26" pageNumber="295">
<heading id="AD140237190EFFB79A9DFBDC943E8A21" box="[140,337,1043,1065]" fontSize="9" level="2" pageId="26" pageNumber="295" reason="5">
<taxonomicName id="31E3CED8190EFFB79A9DFBDC943E8A21" ID-CoL="7Q4FF" authority="Oken, 1815" authorityName="Oken" authorityYear="1815" box="[140,337,1043,1065]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="genus">
<emphasis id="C4976949190EFFB79A9DFBDC95B58A21" bold="true" box="[140,218,1043,1065]" italics="true" pageId="26" pageNumber="295">Thyone</emphasis>
<bibRefCitation id="9272C8AA190EFFB79AF1FBDC943E8A21" author="Oken, L." box="[224,337,1043,1065]" pageId="26" pageNumber="295" pagination="417" refId="ref22942" refString="Oken, L. 1815. Lehrbuch der Naturgeschichte. Part 3: Zoologie 3. xxviii, 850, xviii. (Rejected by ICZN, 1956, opinion 417)." type="journal article" year="1815">Oken, 1815</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BEF9E6D0190EFFB79A9DFB8F941E88A1" pageId="26" pageNumber="295" type="diagnosis">
<paragraph id="F65CB55B190EFFB79A9DFB8F94B48AC1" blockId="26.[140,771,1087,1225]" pageId="26" pageNumber="295">
<emphasis id="C4976949190EFFB79A9DFB8F97F78A5C" box="[140,664,1087,1109]" italics="true" pageId="26" pageNumber="295">
Diagnosis (emended from
<bibRefCitation id="9272C8AA190EFFB79B89FB8F97FF8A5C" author="Pawson, D. L. &amp; Miller, J. E." box="[408,656,1087,1109]" pageId="26" pageNumber="295" pagination="391 - 403" refId="ref23051" refString="Pawson, D. L. and Miller, J. E. 1981. Western Atlantic sea cucumbers of the genus Thyone, with description of two new species (Echinodermata: Holothuroidea). Proceedings of the Biological Society of Washington 94 (2): 391 - 403." type="journal article" year="1981">Pawson and Miller 1981</bibRefCitation>
)
</emphasis>
. Tentacles 10; tube feet scattered on body wall, never restricted to radii; calcareous ring tubular with long posterior prolongations comprising a mosaic of small pieces; body wall ossicles tables with a spire of one or two pillars.
</paragraph>
<paragraph id="F65CB55B190EFFB79A9DFB2F97A68AFC" blockId="26.[140,713,1247,1269]" box="[140,713,1247,1269]" pageId="26" pageNumber="295">
<emphasis id="C4976949190EFFB79A9DFB2F94658AFD" box="[140,266,1248,1269]" italics="true" pageId="26" pageNumber="295">
<typeStatus id="29580BF9190EFFB79A9DFB2F95D48AFD" box="[140,187,1248,1269]" pageId="26" pageNumber="295">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="31E3CED8190EFFB79B07FB1097268AFD" authority="Muller, 1776" authorityName="Muller" authorityYear="1776" box="[278,585,1247,1269]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="species" species="fusus">
<emphasis id="C4976949190EFFB79B07FB1094AC8AFC" box="[278,451,1247,1268]" italics="true" pageId="26" pageNumber="295">Holothuria fusus</emphasis>
Müller, 1776
</taxonomicName>
(monotypy).
</paragraph>
<paragraph id="F65CB55B190EFFB79A9DFAC497658B9D" blockId="26.[140,771,1291,1429]" pageId="26" pageNumber="295">
<emphasis id="C4976949190EFFB79A9DFAC494C38B29" box="[140,428,1291,1313]" italics="true" pageId="26" pageNumber="295">Southern Australian species</emphasis>
.
<taxonomicName id="31E3CED8190EFFB79BABFAC497B08B29" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2012" box="[442,735,1291,1313]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="species" species="flindersi" status="sp. nov.">
<emphasis id="C4976949190EFFB79BABFAC4970E8B28" box="[442,609,1291,1312]" italics="true" pageId="26" pageNumber="295">Thyone flindersi</emphasis>
OLoughlin
</taxonomicName>
<taxonomicNameLabel id="DFA4D432190EFFB798F6FAC395D58B36" pageId="26" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="31E3CED8190EFFB79AD7FAE794828B36" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2012" box="[198,493,1320,1342]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="species" species="joshuai" status="sp. nov.">
<emphasis id="C4976949190EFFB79AD7FAE794068B35" box="[198,361,1320,1341]" italics="true" pageId="26" pageNumber="295">Thyone joshuai</emphasis>
OLoughlin
</taxonomicName>
,
<taxonomicNameLabel id="DFA4D432190EFFB79BEDFAE697208B36" box="[508,591,1321,1342]" pageId="26" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="31E3CED8190EFFB7984CFAE7946C8B53" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2012" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="species" species="kerkosa" status="sp. nov.">
<emphasis id="C4976949190EFFB7984CFAE7966C8B35" box="[605,771,1320,1341]" italics="true" pageId="26" pageNumber="295">Thyone kerkosa</emphasis>
OLoughlin
</taxonomicName>
<taxonomicNameLabel id="DFA4D432190EFFB79B1CFA89940F8B53" box="[269,352,1350,1371]" pageId="26" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="31E3CED8190EFFB79B7CFA8A97918B53" authority="Joshua and Creed, 1915" authorityName="Joshua and Creed" authorityYear="1915" box="[365,766,1349,1371]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="species" species="nigra">
<emphasis id="C4976949190EFFB79B7CFA8A94968B53" box="[365,505,1349,1371]" italics="true" pageId="26" pageNumber="295">Thyone nigra</emphasis>
<bibRefCitation id="9272C8AA190EFFB79812FA8A97918B53" author="Joshua E. C. &amp; Creed, E." box="[515,766,1349,1371]" pageId="26" pageNumber="295" pagination="16 - 24" refId="ref22814" refString="Joshua E. C. and Creed, E. 1915. South Australian Holothuroidea, with descriptions of new species. Transactions and Proceedings of the Royal Society of South Australia 39: 16 - 24, pls 2 - 4." type="journal article" year="1915">Joshua and Creed, 1915</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="31E3CED8190EFFB79A9DFAAD94EE8B7F" authority="Bell, 1884" authorityName="Bell. We" authorityYear="1884" box="[140,385,1378,1400]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="species" species="okeni">
<emphasis id="C4976949190EFFB79A9DFAAD947C8B7F" box="[140,275,1378,1399]" italics="true" pageId="26" pageNumber="295">Thyone okeni</emphasis>
<bibRefCitation id="9272C8AA190EFFB79B08FAAD94EE8B7F" author="Bell, F. J." box="[281,385,1378,1400]" pageId="26" pageNumber="295" refId="ref22242" refString="Bell, F. J. 1884. Echinodermata. Pp. 117 - 177 in Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. Alert 1881 - 2. Taylor and Francis: London." type="book" year="1884">Bell, 1884</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="31E3CED8190EFFB79B9BFAAD97C28B70" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2012" box="[394,685,1378,1400]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="species" species="spenceri" status="sp. nov.">
<emphasis id="C4976949190EFFB79B9BFAAD975F8B70" box="[394,560,1378,1400]" italics="true" pageId="26" pageNumber="295">Thyone spenceri</emphasis>
OLoughlin
</taxonomicName>
<taxonomicNameLabel id="DFA4D432190EFFB798A3FAAC97908B70" box="[690,767,1379,1400]" pageId="26" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="31E3CED8190EFFB79A9DFAB094D78B9D" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2012" box="[140,440,1407,1429]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="species" species="tourvillei" status="sp. nov.">
<emphasis id="C4976949190EFFB79A9DFAB094558B9C" box="[140,314,1407,1428]" italics="true" pageId="26" pageNumber="295">Thyone tourvillei</emphasis>
OLoughlin
</taxonomicName>
<taxonomicNameLabel id="DFA4D432190EFFB79BAFFA4F97658B9D" box="[446,522,1408,1429]" pageId="26" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="F65CB55B190EFFB79A9DFA64941E88A1" blockId="26.[140,771,1451,1705]" pageId="26" pageNumber="295">
<emphasis id="C4976949190EFFB79A9DFA64958B8BC8" box="[140,228,1451,1472]" italics="true" pageId="26" pageNumber="295">Remarks</emphasis>
. We acknowledge the current invalid status of
<taxonomicName id="31E3CED8190EFFB798A6FA64946F8BD6" authority="Oken, 1815" authorityName="Oken" authorityYear="1815" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="genus">
<emphasis id="C4976949190EFFB798A6FA64966D8BC8" box="[695,770,1451,1472]" italics="true" pageId="26" pageNumber="295">Thyone</emphasis>
<bibRefCitation id="9272C8AA190EFFB79A9DFA07946F8BD6" author="Oken, L." box="[140,256,1480,1502]" pageId="26" pageNumber="295" pagination="417" refId="ref22942" refString="Oken, L. 1815. Lehrbuch der Naturgeschichte. Part 3: Zoologie 3. xxviii, 850, xviii. (Rejected by ICZN, 1956, opinion 417)." type="journal article" year="1815">Oken, 1815</bibRefCitation>
</taxonomicName>
and our provisional retention is discussed in the Introduction. We emend the diagnosis of
<taxonomicName id="31E3CED8190EFFB79845FA2A978D8BF3" authority="Oken" authorityName="Oken" box="[596,738,1509,1531]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="genus">
<emphasis id="C4976949190EFFB79845FA2A97F08BF2" box="[596,671,1509,1530]" italics="true" pageId="26" pageNumber="295">Thyone</emphasis>
Oken
</taxonomicName>
to include tables with one pillar in the spire to accommodate a new species described in this work.
<bibRefCitation id="9272C8AA190EFFB79816F9EF966D883C" author="Pawson, D. L. &amp; Miller, J. E." box="[519,770,1567,1589]" pageId="26" pageNumber="295" pagination="391 - 403" refId="ref23051" refString="Pawson, D. L. and Miller, J. E. 1981. Western Atlantic sea cucumbers of the genus Thyone, with description of two new species (Echinodermata: Holothuroidea). Proceedings of the Biological Society of Washington 94 (2): 391 - 403." type="journal article" year="1981">Pawson and Miller 1981</bibRefCitation>
remarked on the need for a revision of the “supergenus”
<emphasis id="C4976949190EFFB798A0F9F3966C8859" box="[689,771,1596,1617]" italics="true" pageId="26" pageNumber="295">
<taxonomicName id="31E3CED8190EFFB798A0F9F397908859" box="[689,767,1596,1617]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="295" phylum="Echinodermata" rank="genus">Thyone</taxonomicName>
.
</emphasis>
This need continues for what will be a major undertaking. The revision should await further supportive evidence from emerging genetic data.
</paragraph>
</subSubSection>
</treatment>
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