treatments-xml/data/7E/11/33/7E11335ADD9554919F42A0A0D34D00F1.xml

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<mods:title id="8B4A2B2BB0D91E9B07FC54CA88D11D10">Impatiens wutaishanensis (Balsaminaceae), a new species from Southeast Yunnan, China</mods:title>
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<mods:namePart id="9DD3664CBFA0DBFAB21862899BFB93BF">Liao, Rong-Li</mods:namePart>
<mods:affiliation id="96B5391CE71F2AC9152C44E9145D9F08">School of Life Sciences, Yunnan University, Kunming 650091, Yunnan, China &amp; Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations / Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, Yunnan, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
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<mods:namePart id="F76F3BD71EA8D30C5F57E6152D1A89FB">Cai, Lei</mods:namePart>
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<mods:affiliation id="FAA60FA228150410A8FB915BB98F51D5">School of Life Sciences, Yunnan University, Kunming 650091, Yunnan, China</mods:affiliation>
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<mods:namePart id="C366F036396FDD224DE73296EDD42190">Yu, Zhi-Yong</mods:namePart>
<mods:affiliation id="CAA25F009A5F9C08616E0195C7634A7B">Management Bureau of Jinping Fenshuiling National Nature Reserve, Jingping 661500, Yunnan, China</mods:affiliation>
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<mods:namePart id="136E33A1C64DE0231FDE76F3BADADF1A">Wang, Yue-Hua</mods:namePart>
<mods:affiliation id="82BA74BA0149D6A4DD8D4992EB3C6376">School of Life Sciences, Yunnan University, Kunming 650091, Yunnan, China</mods:affiliation>
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<mods:namePart id="1954A6D0032AAE909B930883EA75848E">Sun, Wei-Bang</mods:namePart>
<mods:affiliation id="C2A200C784CA9FBEB98B7BF65DEF0D1D">Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations / Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, Yunnan, China</mods:affiliation>
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<treatment id="7E11335ADD9554919F42A0A0D34D00F1" ID-GBIF-Taxon="181024617" LSID="urn:lsid:plazi:treatment:7E11335ADD9554919F42A0A0D34D00F1" httpUri="http://treatment.plazi.org/id/7E11335ADD9554919F42A0A0D34D00F1" lastPageNumber="43" pageId="0" pageNumber="43" scope_family="Balsaminaceae">
<subSubSection id="46FC27BAE06E8F0CD0CC23C421DCDFF2" pageId="0" pageNumber="43" type="nomenclature">
<paragraph id="79D293C547E64E1413079C50E6DC46B1" pageId="0" pageNumber="43">
<taxonomicName id="0DF747938201CF5AC9B09DA37FF13F05" ID-CoL="8H4V5" LSID="7E11335A-DD95-5491-9F42-A0A0D34D00F1" authority="R. L. Liao &amp; Lei Cai" authorityName="R. L. Liao &amp; Lei Cai" authorityYear="2021" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens wutaishanensis" order="Ericales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="wutaishanensis" status="sp. nov.">Impatiens wutaishanensis R.L. Liao &amp; Lei Cai</taxonomicName>
<taxonomicNameLabel id="80636AA68069E392CD751493B2527C2D" pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
<figureCitation id="B76AA06B925ABDA4BCA6F3C83894B40D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Impatiens wutaishanensis R. L. Liao &amp; Lei Cai A habit B whole plant C inflorescence D capsule E flower in front view F flower in lateral view G different parts of the flower." figureDoi="10.3897/phytokeys.176.58825.figure1" httpUri="https://binary.pensoft.net/fig/531547" pageId="0" pageNumber="43">Figs 1</figureCitation>
<figureCitation id="B2582AEDD9FBBD656B4F50F17363B2F7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Impatiens wutaishanensis R. L. Liao &amp; Lei Cai (drawn by R. M. Zhang) A habit B root C inner lateral sepal D outer lateral sepal E dorsal petal F lateral united petals (F 1: upper petal F 2: lower petal) G lower sepal H ovary and stamens I flower in front view J flower in lateral view K partial view of leaf margin." figureDoi="10.3897/phytokeys.176.58825.figure2" httpUri="https://binary.pensoft.net/fig/531548" pageId="0" pageNumber="43">, 2</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="E0AC28A7C4F6C40168CF2DA01485B165" pageId="0" pageNumber="43">
<typeStatus id="D31A3BF921D7E099CA9B8DB9EAEEA868">Type</typeStatus>
.
</paragraph>
<paragraph id="23DB4F14EC92E94AC3C96A752DC00E63" pageId="0" pageNumber="43">
<materialsCitation id="D0A638523973636A042FCD110877678B" ID-GBIF-Occurrence="3079226301" collectingDate="2016-09-10" collectorName="Lei Cai, Z. Y. Yu" country="China" county="Jinping County" elevation="1650" latitude="22.772213" location="Maandi" longLatPrecision="1" longitude="103.47494" specimenCount="1" stateProvince="Yunnan Province" typeStatus="holotype">
<collectingCountry id="7C8F24F5BED80BA84EB395DF962876D5" name="China">China</collectingCountry>
, 
<collectingRegion id="F82EFA75BC031C7EB2892F993E8CEB38" country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty id="49F00322ED179A768E26DD89D12B57EF">Jinping County</collectingCounty>
, 
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, alt. 
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<quantity id="8F0F18A4F885F72B4B2FB1B2AF28309E" metricMagnitude="3" metricUnit="m" metricValue="1.65" unit="m" value="1650.0">1650 m</quantity>
a.s.l.
</elevation>
, 
<geoCoordinate id="AB1BA4977600EA3E643A427A3B202F4A" degrees="22" direction="north" minutes="46" orientation="latitude" precision="1" seconds="19.97" value="22.772213">22°46'19.97&quot;N</geoCoordinate>
, 
<geoCoordinate id="047713F8186C1ED67E2482BD40D0A964" degrees="103" direction="east" minutes="28" orientation="longitude" precision="1" seconds="29.78" value="103.47494">103°28'29.78&quot;E</geoCoordinate>
, 
<collectingDate id="6E38FEB526E6BC8C1EE549624B73555D" value="2016-09-10">10 September 2016</collectingDate>
, 
<collectorName id="AE869C47A17D22F8861317D668AD4144">Lei Cai</collectorName>
&amp; 
<collectorName id="ADA83208FE7DE38D1F0513629CB48BDF">Z.Y. Yu</collectorName>
CL16050 (
<typeStatus id="5F0D1340353ED50AF3F093F541DCCCEA">holotype</typeStatus>
:1498854, KUN!; isotypes:1498855, KUN!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B961B62862BDD4DA98F4817F957829B4" pageId="0" pageNumber="43" type="diagnosis">
<paragraph id="D76F6853A5222CBCA29612269D0AEAC7" pageId="0" pageNumber="43">Diagnosis.</paragraph>
<paragraph id="E5D0C2BB4F10C810312DCA8E01F85AED" pageId="0" pageNumber="43">
This species is similar to 
<taxonomicName id="F9991BA3C735FB3655B40068AFC976F3" authorityName="S.X. Yu &amp; Y.T. Hou" authorityYear="2011" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens parvisepala" order="Ericales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="parvisepala">
<emphasis id="C69C65C41917EE8169696E2CFBE51FAC" italics="true" pageId="0" pageNumber="43">Impatiens parvisepala</emphasis>
</taxonomicName>
S.X. Yu &amp; Y.T. Hou (2011: 57) (
<bibRefCitation id="9770C042DE7267B0C8A6CAEAA7E3D871" DOI="https://doi.org/10.5735/085.048.0108" author="Hou, YT" journalOrPublisher="Annales Botanici Fennici" pageId="0" pageNumber="43" pagination="57 - 62" refId="B11" refString="Hou, YT, Zhou, XR, Yu, SX, 2011. Impatiens parvisepala (Balsaminaceae), a new species from Guangxi, China. Annales Botanici Fennici 48 (1): 57 - 62, DOI: https://doi.org/10.5735/085.048.0108" title="Impatiens parvisepala (Balsaminaceae), a new species from Guangxi, China." url="https://doi.org/10.5735/085.048.0108" volume="48" year="2011">Hou et al. 2011</bibRefCitation>
) in its racemose inflorescences, its alternate and aggregated or subverticillate at stem apex arrangement leaf, and its yellow flowers, but it can be distinguished by its conspicuous 0.5-2.4 cm long petiole (vs. sub-sessile or sessile), its shorter 10-35 cm high plants (vs. 35-60 cm high), its 3.5-12 
<normalizedToken id="BAD3BAE2C7E035534B94A9521C20B094" originalValue="×">x</normalizedToken>
1.5-4 cm elliptic to lanceolate-oblong leaf blades with cuneate bases (vs. 12-20 
<normalizedToken id="C589992207FDACB4B223F7C237E01973" originalValue="×">x</normalizedToken>
3.5-6 cm obovate or obovate-lanceolate blade with attenuate bases), its slightly incurved or narrowing to incurved spur (vs. nearly straight spur), and its racemose inflorescences with up to 22 flowers (vs. 6-8 flowers per inflorescence).
</paragraph>
<caption id="2D91B594307EA0117FE4570D28FB2340" doi="10.3897/phytokeys.176.58825.figure1" httpUri="https://binary.pensoft.net/fig/531547" pageId="0" pageNumber="43" start="Figure 1" startId="F1">
<paragraph id="DB3068320D8591FEB0DA7E5F8B1F7BC4" pageId="0" pageNumber="43">
<emphasis id="DB72BB073341197F795EDAC11F325F15" bold="true" pageId="0" pageNumber="43">Figure 1.</emphasis>
<taxonomicName id="6C8DE05C7DF04DE7F7AFDBC536A5DC82" authorityName="R. L. Liao &amp; Lei Cai" authorityYear="2021" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens wutaishanensis" order="Ericales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="wutaishanensis">
<emphasis id="0AD4B33A207E603F19290517774D2DC8" italics="true" pageId="0" pageNumber="43">Impatiens wutaishanensis</emphasis>
</taxonomicName>
R.L. Liao &amp; Lei Cai 
<emphasis id="306D914E3AEA123C1108C8388451169B" bold="true" pageId="0" pageNumber="43">A</emphasis>
habit 
<emphasis id="135238B7DFF6056F8EE0308AB898162E" bold="true" pageId="0" pageNumber="43">B</emphasis>
whole plant 
<emphasis id="307DA47902784E0A5FCFD6A39703B91F" bold="true" pageId="0" pageNumber="43">C</emphasis>
inflorescence 
<emphasis id="AF564CD16939BA9559A6AD9F7F33B832" bold="true" pageId="0" pageNumber="43">D</emphasis>
capsule 
<emphasis id="CEF455CED7DC244791AB6E357D3D623F" bold="true" pageId="0" pageNumber="43">E</emphasis>
flower in front view 
<emphasis id="AE9978061B697188B6DD75E86E795FBE" bold="true" pageId="0" pageNumber="43">F</emphasis>
flower in lateral view 
<emphasis id="66066D4A9F97A073CFA12AB765BCE6DC" bold="true" pageId="0" pageNumber="43">G</emphasis>
different parts of the flower.
</paragraph>
</caption>
<caption id="F1367A92D9BDE3F9C2F018CF196EEF77" doi="10.3897/phytokeys.176.58825.figure2" httpUri="https://binary.pensoft.net/fig/531548" pageId="0" pageNumber="43" start="Figure 2" startId="F2">
<paragraph id="B33445B023228647365CF349B84EB1B7" pageId="0" pageNumber="43">
<emphasis id="023CEE53CA788B41293F95196E40F861" bold="true" pageId="0" pageNumber="43">Figure 2.</emphasis>
<taxonomicName id="341B4B7F3DB103DEC5B6711C099B00C5" authorityName="R. L. Liao &amp; Lei Cai" authorityYear="2021" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens wutaishanensis" order="Ericales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="wutaishanensis">
<emphasis id="54FE490D81AB92DCB704E4909063C41C" italics="true" pageId="0" pageNumber="43">Impatiens wutaishanensis</emphasis>
</taxonomicName>
R.L. Liao &amp; Lei Cai (drawn by R.M. Zhang) 
<emphasis id="5B669F4CE21F2B4E3A175F721665F572" bold="true" pageId="0" pageNumber="43">A</emphasis>
habit 
<emphasis id="60CA186F72BC8AA463CE3FE589304921" bold="true" pageId="0" pageNumber="43">B</emphasis>
root 
<emphasis id="AD917AF5E1372F19E900E6112B207C31" bold="true" pageId="0" pageNumber="43">C</emphasis>
inner lateral sepal 
<emphasis id="DC2F8BF56B88695CD0A4D1498BD10D92" bold="true" pageId="0" pageNumber="43">D</emphasis>
outer lateral sepal 
<emphasis id="77353495D59163B5D8A96409C500C15C" bold="true" pageId="0" pageNumber="43">E</emphasis>
dorsal petal 
<emphasis id="60437D96620753078CBECF22C4F53F69" bold="true" pageId="0" pageNumber="43">F</emphasis>
lateral united petals (
<emphasis id="E4BCAD8C6B5F7C59A00CDED5A737ABD0" bold="true" pageId="0" pageNumber="43">F1</emphasis>
: upper petal 
<emphasis id="2534979F5E7741663D0C7B55E898E84A" bold="true" pageId="0" pageNumber="43">F2</emphasis>
: lower petal) 
<emphasis id="52E049DC812089CB80D2A36C2FB144CC" bold="true" pageId="0" pageNumber="43">G</emphasis>
lower sepal 
<emphasis id="AC8A538A5ED729C8E178053F975624D7" bold="true" pageId="0" pageNumber="43">H</emphasis>
ovary and stamens 
<emphasis id="12C9CD09EE365AB9D1831E61E4A38653" bold="true" pageId="0" pageNumber="43">I</emphasis>
flower in front view 
<emphasis id="817D24208200F4B78C6AF9BFD6A16600" bold="true" pageId="0" pageNumber="43">J</emphasis>
flower in lateral view 
<emphasis id="BAD770CE60C4DC019A9D3711C9A6B4B4" bold="true" pageId="0" pageNumber="43">K</emphasis>
partial view of leaf margin.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F66BBFFC6E82002045B304CF3CF6F010" pageId="0" pageNumber="43" type="description">
<paragraph id="750A52A7ABCD5DEEE7931AEA65487850" pageId="0" pageNumber="43">Description.</paragraph>
<paragraph id="7073495F6E4BB0E6B9D891B5DB1D6C67" pageId="0" pageNumber="43">
Perennial herb, 10-35 cm tall, glabrous. Root-system shallow, with runners and adventitious roots. Stem fleshy, erect, simple, lower nodes swollen. Leaves simple, alternate, aggregated or subverticillate at stem apex; petiole 0.5-2.4 cm long, leaf blade 3.5-12 
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1.5-4 cm, elliptic, lanceolate, or lanceolate-oblong, base cuneate, margin roughly crenate, mucronulate, apex acuminate, adaxial surface dark green, abaxial surface pale green, lateral veins 4-8 pairs. Racemes in the upper leaf axils, 1-4, 4.2-28 cm long, erect, each with up to 22 flowers. Pedicels 0.6-2.5 cm long, bracteate at base. Bracts ca. 8 mm long, lanceolate to subulate, apex acute, base obtuse, margin entire. Flowers yellow. Lateral sepals 4, light green, the outer pair ca. 5-8 
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3-4 mm, ovate or obovate; the inner pair ca. 8-13 
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1-3 mm, sickle-shaped, obliquely lanceolate, apex acuminate or caudate. Lower sepal 1.5-2.2 
<normalizedToken id="2285B912DC8ABAD6927BF32A411BDF32" originalValue="×">x</normalizedToken>
1.2-2 cm excluding the spur, yellow to yellowish green with reddish patches, obliquely infundibuliform, base gradually constricted into a spur, spur 1.2-3 cm long, slightly incurved or narrowing to incurved, apex rostellate. Dorsal petal 1.3-2.3 
<normalizedToken id="5FF0BA3C11109BCF0DD275595FDB0A01" originalValue="×">x</normalizedToken>
0.7-1.6 cm, yellow with nearly transparent base, obovate to ovate, apex acuminate, with an inconspicuous dorsal crest, base truncate or cuneate. Lateral united petals ca. 1.6-3 cm long, yellow with nearly transparent base and reddish patches; the upper petals ca. 1.5-2.3 
<normalizedToken id="D5C98A3AE09E2D0F5B3267BABD9B7958" originalValue="×">x</normalizedToken>
0.7-1 cm, oblong; the lower petal ca.1.2-1.8 
<normalizedToken id="3886E58C59A965CA0EAD1B4164BED057" originalValue="×">x</normalizedToken>
0.6-0.8 cm, reniform. Stamens 5, filaments linear, ca. 3 mm long, anthers obtuse. Ovary fusiform, slightly curved. Placentation axile with four locules. Capsule (immature) clavate, 2-2.5 cm long.
</paragraph>
</subSubSection>
<subSubSection id="FC0D952E1E9CE36B8CA7A5C8A0322C8F" pageId="0" pageNumber="43" type="pollen morphology">
<paragraph id="12385051F806D8EE96C516C72B63A2EC" pageId="0" pageNumber="43">Pollen morphology.</paragraph>
<paragraph id="37EFD248BE67E02462AFD45CE6C61909" pageId="0" pageNumber="43">
Pollen grains triangular-round with three equal sides in polar view, and the equatorial view is elliptic, long-elliptic, 
<normalizedToken id="CE6C784D5280B249B97DD05567169D93" originalValue="P×E=16.56±">PxE=16.56+/-</normalizedToken>
1.78 (14.51-21.73) 
<normalizedToken id="39010395633DCB766C6B0827510AA6BD" originalValue="×">x</normalizedToken>
30.00 
<normalizedToken id="ECB80F750773E33BFE25D866E75D88D9" originalValue="±">+/-</normalizedToken>
0.98 (28.35-32.11) 
<normalizedToken id="84E78C5DDA6734EEE1C3B7BA3D8908FF" originalValue="μm">μm</normalizedToken>
. 3-colpate, linear, the entire surface is covered with reticulate ornamentation, granules in lumina (Fig. 
<figureCitation id="D6C5EC249F4DF6A2B6E219C38FE1B02F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micromorphology of Impatiens wutaishanensis R. L. Liao &amp; Lei Cai under cryo-SEM A-D pollen A polar view B equatorial view C partial view D total view E-F leaf surface E adaxial epidermis F abaxial epidermis. Scale bar: 2 μm (A, B); 1 μm (C); 20 μm (D-F)." figureDoi="10.3897/phytokeys.176.58825.figure3" httpUri="https://binary.pensoft.net/fig/531549" pageId="0" pageNumber="43">3</figureCitation>
).
</paragraph>
<caption id="6D90693BA49392DE7B8B39B828C44D42" doi="10.3897/phytokeys.176.58825.figure3" httpUri="https://binary.pensoft.net/fig/531549" pageId="0" pageNumber="43" start="Figure 3" startId="F3">
<paragraph id="AB74BB2E7B400AA639BDEDB7364189E9" pageId="0" pageNumber="43">
<emphasis id="230D0346B946276603AB50D5FE6CFC79" bold="true" pageId="0" pageNumber="43">Figure 3.</emphasis>
Micromorphology of 
<taxonomicName id="567FF2CBC716F1C8F786135200C3D4ED" authorityName="R. L. Liao &amp; Lei Cai" authorityYear="2021" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens wutaishanensis" order="Ericales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="wutaishanensis">
<emphasis id="68AC48339DDA425EF7ECCB8A0B94FE46" italics="true" pageId="0" pageNumber="43">Impatiens wutaishanensis</emphasis>
</taxonomicName>
R.L. Liao &amp; Lei Cai under cryo-SEM 
<emphasis id="5D328F6419B851FBD98F7A824AE2F22B" bold="true" pageId="0" pageNumber="43">A-D</emphasis>
pollen 
<emphasis id="9CAC00256CA1FF447E1CB5580425B5ED" bold="true" pageId="0" pageNumber="43">A</emphasis>
polar view 
<emphasis id="194CECBBF11137D8197F1724512CD95F" bold="true" pageId="0" pageNumber="43">B</emphasis>
equatorial view 
<emphasis id="3EFAF0580DE7042388609829CA88D23C" bold="true" pageId="0" pageNumber="43">C</emphasis>
partial view 
<emphasis id="EA6C1DBF74B087715EFDAFAB1AB09930" bold="true" pageId="0" pageNumber="43">D</emphasis>
total view 
<emphasis id="31FC38D90BD3ADD950B825C812BF77EB" bold="true" pageId="0" pageNumber="43">E-F</emphasis>
leaf surface 
<emphasis id="E52EE7A7C1985CBBBCEC69D3FECAAFDF" bold="true" pageId="0" pageNumber="43">E</emphasis>
adaxial epidermis 
<emphasis id="50AD823D00A2E7D4C8EEB482A98B7476" bold="true" pageId="0" pageNumber="43">F</emphasis>
abaxial epidermis. Scale bar: 2 
<normalizedToken id="20488801A8F9CC9BFBC3CED13EDC28FB" originalValue="μm">μm</normalizedToken>
(
<emphasis id="0AC2C2953EA0E877C9642A4F034C82C9" bold="true" pageId="0" pageNumber="43">A, B</emphasis>
); 1 
<normalizedToken id="787FC328A47442FA8E2FD6EE18CD724F" originalValue="μm">μm</normalizedToken>
(
<emphasis id="4A7B165DA66117791F4B87DF2C481825" bold="true" pageId="0" pageNumber="43">C</emphasis>
); 20 
<normalizedToken id="757486EA59F03F5306C2F4359135F167" originalValue="μm">μm</normalizedToken>
(
<emphasis id="9218BE082E7C75C7174B9F54B6EB27DE" bold="true" pageId="0" pageNumber="43">D-F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="436735D6863CE2D2CC4A12D452B57849" pageId="0" pageNumber="43" type="leaf surface micromorphology">
<paragraph id="0B2D9E9A22903B33884F45B6B4A2BDD3" pageId="0" pageNumber="43">Leaf surface micromorphology.</paragraph>
<paragraph id="EEC0FD565D153D32A12373117E47E29B" pageId="0" pageNumber="43">
The anticlinal walls of the epidermal cells on the adaxial surfaces are straight, while those of the abaxial surface are sinuate. The stomata only appeared on the abaxial surface of the leaf, the shape was anomocytic and the outline of the guard cells was suborbiculate (Fig. 
<figureCitation id="2D77321C078CBC25584B35A4192A67FB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micromorphology of Impatiens wutaishanensis R. L. Liao &amp; Lei Cai under cryo-SEM A-D pollen A polar view B equatorial view C partial view D total view E-F leaf surface E adaxial epidermis F abaxial epidermis. Scale bar: 2 μm (A, B); 1 μm (C); 20 μm (D-F)." figureDoi="10.3897/phytokeys.176.58825.figure3" httpUri="https://binary.pensoft.net/fig/531549" pageId="0" pageNumber="43">3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="158777B60E581EC4D840925BF7F136DD" pageId="0" pageNumber="43" type="phenology">
<paragraph id="627C16FDC4BFBFDC0308C3E4736842E2" pageId="0" pageNumber="43">Phenology.</paragraph>
<paragraph id="B970B9F73E1D8C1B68919B0385851BA1" pageId="0" pageNumber="43">This new species was observed flowering from August to November, and fruiting from September to December.</paragraph>
</subSubSection>
<subSubSection id="5D8C740CA73C22CA656A44B9F8E8E68F" pageId="0" pageNumber="43" type="distribution">
<paragraph id="45962605EB85D2AC1CCBAC643F844350" pageId="0" pageNumber="43">Distribution.</paragraph>
<paragraph id="51965D8811B9076DBAFB7D4F07574B86" pageId="0" pageNumber="43">
This species is currently known from only two small subpopulations less than 10 km away (Shidong and Biaoshuiyan) in Jinping County of Southeast Yunnan, China (Fig. 
<figureCitation id="7DBF3157A4601F3E2F85FC84D080365F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of Impatiens wutaishanensis R. L. Liao &amp; Lei Cai and I. parvisepala S. X. Yu &amp; Y. T. Hou." figureDoi="10.3897/phytokeys.176.58825.figure4" httpUri="https://binary.pensoft.net/fig/531550" pageId="0" pageNumber="43">4</figureCitation>
). The distribution area is very close to the border between China and Vietnam. We assume that this species should be distributed in Vietnam due to its similar habitat and proximity to the type locality, which will be verified by future investigation.
</paragraph>
<caption id="1D534C5F2B16D3536D3AD3BF86FE57BB" doi="10.3897/phytokeys.176.58825.figure4" httpUri="https://binary.pensoft.net/fig/531550" pageId="0" pageNumber="43" start="Figure 4" startId="F4">
<paragraph id="9632E4872A352B9C58DA90BC3C012774" pageId="0" pageNumber="43">
<emphasis id="CCBA0DB37E6567454715759DAA581698" bold="true" pageId="0" pageNumber="43">Figure 4.</emphasis>
Distribution of 
<taxonomicName id="D69C2C56D7E399A1195E413EFA7926D6" authorityName="R. L. Liao &amp; Lei Cai" authorityYear="2021" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens wutaishanensis" order="Ericales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="wutaishanensis">
<emphasis id="56F2C7CEA7D03E7F1D330F88E2DB1863" italics="true" pageId="0" pageNumber="43">Impatiens wutaishanensis</emphasis>
</taxonomicName>
R.L. Liao &amp; Lei Cai and 
<taxonomicName id="701B28FE3C0F80288B31BD785F32F0F3" lsidName="I. parvisepala" pageId="0" pageNumber="43" rank="species" species="parvisepala">
<emphasis id="A0E1FC4D1EBA9A8784B67E24B9D4EA27" italics="true" pageId="0" pageNumber="43">I. parvisepala</emphasis>
</taxonomicName>
S.X. Yu &amp; Y.T. Hou.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="597CB8B9B018C5B19CAFE08271AFF49C" pageId="0" pageNumber="43" type="ecology">
<paragraph id="55DD2E5471692EF1D9E62D81C99EE6F0" pageId="0" pageNumber="43">Ecology.</paragraph>
<paragraph id="FEF4E2683ED9B4FAE48E15F6DBFD3088" pageId="0" pageNumber="43">
This new species has a runner with adventitious root produced from the nodes and was collected growing in the understory of forest at an elevation of 1300-1650 m. In the subpopulation in Shidong, the plants found coexist with the cultivated 
<taxonomicName id="6B47D74276E669F1AFEAFDE18C300BB4" authorityName="Crevost &amp; Lemarie" authorityYear="1917" class="Monocotyledoneae" family="Zingiberaceae" genus="Amomum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Amomum tsaoko" order="Scitamineae" pageId="0" pageNumber="43" phylum="Angiospermae" rank="species" species="tsaoko">
<emphasis id="B1700B2FAEAB996A6758434D9CEDD037" italics="true" pageId="0" pageNumber="43">Amomum tsaoko</emphasis>
</taxonomicName>
Crevost &amp; Lemarie (
<taxonomicName id="C625C8F74D0130A9AE04A8AD970F3149" class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Scitamineae" pageId="0" pageNumber="43" phylum="Angiospermae" rank="family">Zingiberaceae</taxonomicName>
). In the subpopulation in Biaoshuiyan, the plants were found beside the artificial trail in a scenic area. The main accompanying species were 
<taxonomicName id="582187D0BB112FD71EC7E6645A4A2556" class="Dicotyledoneae" family="Begoniaceae" genus="Begonia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Begonia" order="Violales" pageId="0" pageNumber="43" phylum="Angiospermae" rank="genus">
<emphasis id="99C0CE959B6FE6818B47D814C542DB32" italics="true" pageId="0" pageNumber="43">Begonia</emphasis>
</taxonomicName>
Linn. sp. (
<taxonomicName id="0185E8181FA9A9FACAED9ADAF946C0D5" class="Dicotyledoneae" family="Begoniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Violales" pageId="0" pageNumber="43" phylum="Angiospermae" rank="family">Begoniaceae</taxonomicName>
), 
<taxonomicName id="3ACA6C0B54D02EB9A4142A3ADBAA142C" class="Magnoliopsida" family="Gesneriaceae" genus="Didymocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Didymocarpus purpureobracteatus" order="Lamiales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="purpureobracteatus">
<emphasis id="E3F3049F3041152200C948BCD640AC36" italics="true" pageId="0" pageNumber="43">Didymocarpus purpureobracteatus</emphasis>
</taxonomicName>
W.W. Smith (
<taxonomicName id="938CE0CF151647321405150E76FA86D6" class="Magnoliopsida" family="Gesneriaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Lamiales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="family">Gesneriaceae</taxonomicName>
), 
<taxonomicName id="6FF59F9C5E2249EBAB1DD394863089B0" class="Magnoliopsida" family="Orchidaceae" genus="Epigeneium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Epigeneium amplum" order="Asparagales" pageId="0" pageNumber="43" phylum="Angiospermae" rank="species" species="amplum">
<emphasis id="99E89CCCEE0805FDBE689C9A10CC08A8" italics="true" pageId="0" pageNumber="43">Epigeneium amplum</emphasis>
</taxonomicName>
(Lindl.) Summerh. (
<taxonomicName id="8DB7D6133726FAECB7D1B79F141952F9" authorityName="Gagnepain" authorityYear="1932" class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asparagales" pageId="0" pageNumber="43" phylum="Angiospermae" rank="family">Orchidaceae</taxonomicName>
), 
<taxonomicName id="71CB076A8F5784812BCB90C3ECB40FF3" class="Monocotyledoneae" family="Zingiberaceae" genus="Hedychium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hedychium villosum" order="Scitamineae" pageId="0" pageNumber="43" phylum="Angiospermae" rank="species" species="villosum">
<emphasis id="1F7277D58F58E3BBB35E72542DA0BA74" italics="true" pageId="0" pageNumber="43">Hedychium villosum</emphasis>
</taxonomicName>
Wall. (
<taxonomicName id="9998FF8CBAC393EE8F81FEAA20F45168" class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Scitamineae" pageId="0" pageNumber="43" phylum="Angiospermae" rank="family">Zingiberaceae</taxonomicName>
), 
<taxonomicName id="F6F01BBD6BE921CAEB0B3112598929CE" class="Dicotyledoneae" family="Polygalaceae" genus="Polygala" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Polygala fallax" order="Rutales" pageId="0" pageNumber="43" phylum="Angiospermae" rank="species" species="fallax">
<emphasis id="23BC14D1DB1FBE4A46EC8CE42158D263" italics="true" pageId="0" pageNumber="43">Polygala fallax</emphasis>
</taxonomicName>
Hemsl. (
<taxonomicName id="C45D75B8986F59233524D23D0997D2EF" class="Dicotyledoneae" family="Polygalaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rutales" pageId="0" pageNumber="43" phylum="Angiospermae" rank="family">Polygalaceae</taxonomicName>
) and 
<taxonomicName id="DDA7DF4B9FE473AD9B910DC0BE7E0845" class="Dicotyledoneae" family="Ericaceae" genus="Vaccinium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Vaccinium" order="Ericales" pageId="0" pageNumber="43" phylum="Angiospermae" rank="genus">
<emphasis id="7BF3F1EC1EC718DF88F649C6DDFB329C" italics="true" pageId="0" pageNumber="43">Vaccinium</emphasis>
</taxonomicName>
Linn. sp. (
<taxonomicName id="F3ED9B02B820FEA4A5658B9349A4B78A" class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Ericales" pageId="0" pageNumber="43" phylum="Angiospermae" rank="family">Ericaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="E7CE8B2A819CC8714930F92D655CA81D" pageId="0" pageNumber="43" type="conservation status">
<paragraph id="9DD5D3A6F6028883FA376898509DDD6B" pageId="0" pageNumber="43">Conservation status.</paragraph>
<paragraph id="98CE66B3D6163EA889DD17C78144C7AB" pageId="0" pageNumber="43">
This species is currently known only from Jinping County, Yunnan, China with one type locality and a subpopulation. The Extent of Occurrence (EOO) is less than 100 km2 and the known Area of Occupancy (AOO) is less than 15 km2. The conservation status can be evaluated as Vulnerable (VU) D2 based on IUCN Red List Categories and Criteria (
<bibRefCitation id="C89AE15A559C0A66994AE0A7484187C5" author="IUCN" journalOrPublisher="Annales Botanici Fennici" pageId="0" pageNumber="43" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B12" refString="IUCN, 2019. Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Subcommittee. http://www.iucnredlist.org/documents/RedListGuidelines.pdf" title="Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Subcommittee." url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2019">IUCN 2019</bibRefCitation>
). About 350 and 200 individuals were known in the two subpopulations of Biaoshuiyan and Shidong, respectively, and both are exposed to human disturbance. Therefore, we assess this species as a Plant Species with Extremely Small Populations (PSESP) (
<bibRefCitation id="55435FC8897E0FD9312A5F6EA5DB62DF" DOI="https://doi.org/10.1016/j.biocon.2020.108535" author="Yang, J" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="43" refId="B24" refString="Yang, J, Cai, L, Liu, DT, Chen, G, Gratzfeld, J, Sun, WB, 2020. China's conservation program on Plant Species with Extremely Small Populations (PSESP): Progress and perspectives. Biological Conservation 244: e108535. https://doi.org/10.1016/j.biocon.2020.108535" title="China's conservation program on Plant Species with Extremely Small Populations (PSESP): Progress and perspectives. Biological Conservation 244: e 108535." url="https://doi.org/10.1016/j.biocon.2020.108535" year="2020">Yang et al. 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D59504F91BFE6E2634A6BCCD8BFD038E" pageId="0" pageNumber="43" type="etymology">
<paragraph id="268BB60CDD989B746D4EC464B976AE2F" pageId="0" pageNumber="43">Etymology.</paragraph>
<paragraph id="9C3EFFC58E324F7A72B5CDB570E6BA31" pageId="0" pageNumber="43">
The specific epithet ' 
<taxonomicName id="C8C03F5E70C75ED663E7072BCA16CD2E" authorityName="R. L. Liao &amp; Lei Cai" authorityYear="2021" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens wutaishanensis" order="Ericales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="wutaishanensis">
<emphasis id="DB6A962D5A5E61216B0ACB0E323F5154" italics="true" pageId="0" pageNumber="43">Impatiens wutaishanensis</emphasis>
</taxonomicName>
' refers to the type locality where the new species was found, located in the Wutaishan area of Jinping Fenshuiling National Nature Reserve, Jinping County, Southeast Yunnan, China. The Chinese name is given as 
<normalizedToken id="8285FB4CFF826B45E5368D2BAA3A37F7" originalValue="“五台山凤仙花”">&quot;五台山凤仙花&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="3F21CAB6288C3A8D2202E56AAA1FBE2C" pageId="0" pageNumber="43" type="materials_examined">
<paragraph id="00C22AF6B8804C823177EEC1E5819756" pageId="0" pageNumber="43">Additional specimens examined.</paragraph>
<paragraph id="3DB2B4713B24F515B956961910B788DE" pageId="0" pageNumber="43">
<materialsCitation id="B37CCE3DE44E091210D7A8C61D88A9C5" ID-GBIF-Occurrence="3079226302" collectingDate="2015-08-05" collectingDateMax="2020-08-02" collectingDateMin="2015-08-05" collectorName="Lei Cai, The, R. L. Liao, X. Y. Li" country="China" county="Jinping County" elevation="1650" latitude="22.772213" location="Maandi Town" longLatPrecision="1" longitude="103.47494" specimenCount="1" stateProvince="Yunnan Province" typeStatus="Paratypes">
<typeStatus id="3ADC6BF30AB39D88CC1CBA225D27E17E">Paratypes</typeStatus>
. 
<collectingCountry id="C93A5C2D0EB6B85852944887F7873753" name="China">China</collectingCountry>
. 
<collectingRegion id="AE31BFB24038088FCA7407488FE62D7D" country="China" name="Yunnan">Yunnan Province</collectingRegion>
: 
<collectingCounty id="2AA11EB4A85DF97593F39F283B21D85B">Jinping County</collectingCounty>
, 
<location id="2F95A83BE55E1FD4A1DCBB5FE80CB604" LSID="urn:lsid:plazi:treatment:7E11335ADD9554919F42A0A0D34D00F1:2F95A83BE55E1FD4A1DCBB5FE80CB604" country="China" county="Jinping County" latitude="22.772213" longLatPrecision="1" longitude="103.47494" name="Maandi Town" stateProvince="Yunnan Province">Maandi Town</location>
, Shidong. 
<geoCoordinate id="C2E001A98EE0CC4438E299CCB5E796EF" degrees="22" direction="north" minutes="46" orientation="latitude" precision="1" seconds="19.97" value="22.772213">22°46'19.97&quot;N</geoCoordinate>
, 
<geoCoordinate id="2D93E963AAA757E60138A2ED8FDB9471" degrees="103" direction="east" minutes="28" orientation="longitude" precision="1" seconds="29.78" value="103.47494">103°28'29.78&quot;E</geoCoordinate>
, 
<elevation id="F0ADC697A189D4F04683DBE2835A7FED" metricMagnitude="3" metricUnit="m" metricValue="1.65" unit="m" value="1650.0">
<quantity id="6265ABE5B31F5E46B1D6895363BE4209" metricMagnitude="3" metricUnit="m" metricValue="1.65" unit="m" value="1650.0">1650 m</quantity>
a.s.l.
</elevation>
, 
<collectingDate id="12C8CDFF69A1866235F1E446F4197EB9" value="2015-08-05">5 August 2015</collectingDate>
, 
<collectorName id="97349457B0C67A4EDF960A15266BE06C">Lei Cai</collectorName>
et al., CL155 (KUN!); 
<collectorName id="28F2BB806BE0C5DB793BCF5A1DBDE21E">The</collectorName>
same locality, 
<collectingDate id="D3CD808DFBC5435CB7060183206A5ABF" value="2020-08-02">2 August 2020</collectingDate>
, 
<collectorName id="0BA06FBF43C5B1DCCBBFE2466524AD7E">R.L. Liao</collectorName>
&amp; 
<collectorName id="30E76B6DABC4BC6887E106E485804693">X.Y. Li</collectorName>
, LRL202008003 (KUN!)
</materialsCitation>
; 
<materialsCitation id="6299D56537A7101AB11CC9F95B42F74F" ID-GBIF-Occurrence="3079226303" collectingDate="2020-08-02" collectorName="R. L. Liao, X. Y. Li" country="China" county="Jinping County" elevation="1311" latitude="22.732391" location="Biaoshuiyan" longLatPrecision="1" longitude="103.51016" municipality="Maandi Town" specimenCount="1" stateProvince="Yunnan Province" typeStatus="paratype">
<collectingRegion id="68FF71BC434D01D48C66DAB863835EB2" country="China" name="Yunnan">Yunnan Province</collectingRegion>
: 
<collectingCounty id="9E773FCBF46035E7D6536CE6BF896987">Jinping County</collectingCounty>
, 
<collectingMunicipality id="FB2430BA269E605270549A3CBBB0E1CF">Maandi Town</collectingMunicipality>
, 
<location id="2C3D063E5B6CFF48AF001882279AFF7D" LSID="urn:lsid:plazi:treatment:7E11335ADD9554919F42A0A0D34D00F1:2C3D063E5B6CFF48AF001882279AFF7D" country="China" county="Jinping County" latitude="22.732391" longLatPrecision="1" longitude="103.51016" municipality="Maandi Town" name="Biaoshuiyan" stateProvince="Yunnan Province">Biaoshuiyan</location>
, 
<geoCoordinate id="9A41A9E0F7ACE3683850157F219FEA05" degrees="22" direction="north" minutes="43" orientation="latitude" precision="1" seconds="56.61" value="22.732391">22°43'56.61&quot;N</geoCoordinate>
, 
<geoCoordinate id="454B12B6546D3DACB0549052A0CEEE0A" degrees="103" direction="east" minutes="30" orientation="longitude" precision="1" seconds="36.58" value="103.51016">103°30'36.58&quot;E</geoCoordinate>
, 
<elevation id="10B3216249EA917C8DFA6FC8A69AA018" metricMagnitude="3" metricUnit="m" metricValue="1.311" unit="m" value="1311.0">
<quantity id="E00B3AEFC5097E54B13D5E0629A21FDF" metricMagnitude="3" metricUnit="m" metricValue="1.311" unit="m" value="1311.0">1311 m</quantity>
a.s.l.
</elevation>
, 
<collectingDate id="F3C81038F50FC1065700E8E8101B26A7" value="2020-08-02">2 August 2020</collectingDate>
, 
<collectorName id="A434CBF7C619C0F39DFB3B4687C3778F">R.L. Liao</collectorName>
&amp; 
<collectorName id="A9D376893A177C7C7FC441E898033CA9">X.Y. Li</collectorName>
, LRL202008001 (KUN!)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="97BA681A92C5B880A4DD6462018DF2A1" pageId="0" pageNumber="43" type="discussion">
<paragraph id="B51CD2CED975734395B2B0A558454A61" pageId="0" pageNumber="43">Discussion.</paragraph>
<paragraph id="1D45182C5D8AD4E06083D82A0066705D" pageId="0" pageNumber="43">
The new species is most similar to 
<taxonomicName id="29FC2FB89D02FC756B42AE963FB750BF" authorityName="S.X. Yu &amp; Y.T. Hou" authorityYear="2011" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens parvisepala" order="Ericales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="parvisepala">
<emphasis id="D69AB1C7DCF96312497785217F6BAC0D" italics="true" pageId="0" pageNumber="43">Impatiens parvisepala</emphasis>
</taxonomicName>
in its racemose inflorescence, its yellow flowers with four lateral sepals. However, 
<taxonomicName id="B09428AC25C2503E981EFA9916133F00" lsidName="I. wutaishanensis" pageId="0" pageNumber="43" rank="species" species="wutaishanensis">
<emphasis id="22C5098501EFA8DB36F807C4C55EE89C" italics="true" pageId="0" pageNumber="43">I. wutaishanensis</emphasis>
</taxonomicName>
is usually a shorter plant and its petiolate leaf blades are smaller and elliptic, lanceolate, or lanceolate-oblong. Its nectar spur is slightly incurved or narrowing to an incurved spur, and each inflorescence may have up to 22 flowers (Figs 
<figureCitation id="6ED41700D8C83681A159C025CD3D3E2D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Impatiens wutaishanensis R. L. Liao &amp; Lei Cai A habit B whole plant C inflorescence D capsule E flower in front view F flower in lateral view G different parts of the flower." figureDoi="10.3897/phytokeys.176.58825.figure1" httpUri="https://binary.pensoft.net/fig/531547" pageId="0" pageNumber="43">1</figureCitation>
, 
<figureCitation id="C4C7F218F1030CE05ED1F255C6C8ADE1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Impatiens wutaishanensis R. L. Liao &amp; Lei Cai (drawn by R. M. Zhang) A habit B root C inner lateral sepal D outer lateral sepal E dorsal petal F lateral united petals (F 1: upper petal F 2: lower petal) G lower sepal H ovary and stamens I flower in front view J flower in lateral view K partial view of leaf margin." figureDoi="10.3897/phytokeys.176.58825.figure2" httpUri="https://binary.pensoft.net/fig/531548" pageId="0" pageNumber="43">2</figureCitation>
). In contrast, the plants of 
<taxonomicName id="F90D5BCB6965AD6D2C4F2F6172A2DE6C" lsidName="I. parvisepala" pageId="0" pageNumber="43" rank="species" species="parvisepala">
<emphasis id="7B8FB052BC0DBAE32496414209F8B24B" italics="true" pageId="0" pageNumber="43">I. parvisepala</emphasis>
</taxonomicName>
are taller, the sessile or subsessile leaf blades are larger and obovate or obovate-lanceolate, the spur is nearly straight and the number of flowers per inflorescence is 6-8 (
<bibRefCitation id="F66B51CF30BA41BD04D1F477462BF190" DOI="https://doi.org/10.5735/085.048.0108" author="Hou, YT" journalOrPublisher="Annales Botanici Fennici" pageId="0" pageNumber="43" pagination="57 - 62" refId="B11" refString="Hou, YT, Zhou, XR, Yu, SX, 2011. Impatiens parvisepala (Balsaminaceae), a new species from Guangxi, China. Annales Botanici Fennici 48 (1): 57 - 62, DOI: https://doi.org/10.5735/085.048.0108" title="Impatiens parvisepala (Balsaminaceae), a new species from Guangxi, China." url="https://doi.org/10.5735/085.048.0108" volume="48" year="2011">Hou et al. 2011</bibRefCitation>
; 
<bibRefCitation id="1447A4BC905236621A30767DEAE74A32" author="Son, HT" journalOrPublisher="Vietnam Journal of Forest Science" pageId="0" pageNumber="43" pagination="4018 - 4020" publicationUrl="http://vafs.gov.vn/" refId="B19" refString="Son, HT, Bon, Tn, Hung, NQ, Vinh, PV, Anh, TV, 2015. Impatiens parvisepala (Balsaminaceae): a newly recorded from Vietnam. Vietnam Journal of Forest Science 4: 4018 - 4020, http://vafs.gov.vn/" title="Impatiens parvisepala (Balsaminaceae): a newly recorded from Vietnam." url="http://vafs.gov.vn/" volume="4" year="2015">Son et al. 2015</bibRefCitation>
). The distribution of these two species is geographically isolated from each other: 
<taxonomicName id="903F7D67BB2006E44FA8A2D46FC2C786" authorityName="R. L. Liao &amp; Lei Cai" authorityYear="2021" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens wutaishanensis" order="Ericales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="wutaishanensis">
<emphasis id="8B043FF49254C5099E287B6D84AF3F3E" italics="true" pageId="0" pageNumber="43">Impatiens wutaishanensis</emphasis>
</taxonomicName>
, is confined to Southeast Yunnan Province with two subpopulations, where the 
<taxonomicName id="B8BE1A2062144C37A4DAAA93E99C7CF3" lsidName="I. parvisepala" pageId="0" pageNumber="43" rank="species" species="parvisepala">
<emphasis id="CFA57D39607BCD76E002E51EE5270717" italics="true" pageId="0" pageNumber="43">I. parvisepala</emphasis>
</taxonomicName>
is recorded in western Guangxi and northern Vietnam (Fig. 
<figureCitation id="36ADC90A7691E72BCB59069676B8AC38" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of Impatiens wutaishanensis R. L. Liao &amp; Lei Cai and I. parvisepala S. X. Yu &amp; Y. T. Hou." figureDoi="10.3897/phytokeys.176.58825.figure4" httpUri="https://binary.pensoft.net/fig/531550" pageId="0" pageNumber="43">4</figureCitation>
).
</paragraph>
<paragraph id="B75344B09B5A3B1A2C464AF616D9B266" pageId="0" pageNumber="43">
In order to illustrate the morphological circumscription of the new species, we compare the new species with four species with similar morphological characters in Table 
<tableCitation id="319CD3781A158326795F324F21103C1E" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of morphological characters in Impatiens wutaishanensis (n = 30 for the measurements), I. parvisepala (data from Hou et al. 2011 and Son et al. 2015), I. apalophylla (data from Hooker 1908 a and Chen et al. 2007), I. clavigera (data from Hooker 1909 b, Hou et al. 2011 and Tan et al. 2015,) and I. tianlinensis (data from Zeng et al. 2015)." httpUri="http://table.plazi.org/id/8CF1916B13D8048EDCAE8F69A94D95BE" pageId="0" pageNumber="43" tableUuid="8CF1916B13D8048EDCAE8F69A94D95BE">1</tableCitation>
: 
<taxonomicName id="EDD4F42B0E79C59F1955EC2231A10C81" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens apalophylla" order="Ericales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="apalophylla">
<emphasis id="1A4418D66FAF8D5BB706541630CD15DC" italics="true" pageId="0" pageNumber="43">Impatiens apalophylla</emphasis>
</taxonomicName>
Hook. f. (1908:243), 
<taxonomicName id="9DDFF8B8DB4BE2FAA79263222A696D9D" lsidName="I. clavigera" pageId="0" pageNumber="43" rank="species" species="clavigera">
<emphasis id="02B4825654B900AA75F8DCBF5E8E9A3B" italics="true" pageId="0" pageNumber="43">I. clavigera</emphasis>
</taxonomicName>
Hook. f. (1908:2863), 
<taxonomicName id="06E0CE3AB45F82BB5689B5C39C0CC04C" lsidName="I. parvisepala" pageId="0" pageNumber="43" rank="species" species="parvisepala">
<emphasis id="92B173286133CB57126249DF89EBA010" italics="true" pageId="0" pageNumber="43">I. parvisepala</emphasis>
</taxonomicName>
and 
<taxonomicName id="E57A52C631665160C0EBCBB8F33150B2" lsidName="I. tianlinensis" pageId="0" pageNumber="43" rank="species" species="tianlinensis">
<emphasis id="FE6DE72C7296F801548B909F24B6BF19" italics="true" pageId="0" pageNumber="43">I. tianlinensis</emphasis>
</taxonomicName>
S.X. Yu &amp; L.J. Zhang (2015: 253) (
<bibRefCitation id="EF859E6736E8F2137BE1AF83C0796733" DOI="https://doi.org/10.11646/phytotaxa.227.3.4" author="Zeng, L" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="43" pagination="253 - 260" refId="B27" refString="Zeng, L, Liu, YN, Gogoi, R, Zhang, LJ, Yu, SX, 2015. Impatiens tianlinensis (Balsaminaceae), a new species from Guangxi, China. Phytotaxa 227 (3): 253 - 260, DOI: https://doi.org/10.11646/phytotaxa.227.3.4" title="Impatiens tianlinensis (Balsaminaceae), a new species from Guangxi, China." url="https://doi.org/10.11646/phytotaxa.227.3.4" volume="227" year="2015">Zeng et al. 2015</bibRefCitation>
).
</paragraph>
<caption id="8CF1916B13D8048EDCAE8F69A94D95BE" ID-Table-UUID="8CF1916B13D8048EDCAE8F69A94D95BE" httpUri="http://table.plazi.org/id/8CF1916B13D8048EDCAE8F69A94D95BE" pageId="0" pageNumber="43" start="Table 1" startId="T1">
<paragraph id="A125C912B77521134C082FF35A5B639B" pageId="0" pageNumber="43">
<emphasis id="E33542F273E698152518039DE9D07C7B" bold="true" pageId="0" pageNumber="43">Table 1.</emphasis>
Comparison of morphological characters in 
<taxonomicName id="436C6B90771D4355112E7B64471C40C8" authorityName="R. L. Liao &amp; Lei Cai" authorityYear="2021" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens wutaishanensis" order="Ericales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="wutaishanensis">
<emphasis id="1E040EDC028C3C292626A3E32F665363" italics="true" pageId="0" pageNumber="43">Impatiens wutaishanensis</emphasis>
</taxonomicName>
(n=30 for the measurements), 
<taxonomicName id="3831948BDB7BFFA5200EA4AA88422A38" lsidName="I. parvisepala" pageId="0" pageNumber="43" rank="species" species="parvisepala">
<emphasis id="92634B68F712A37C6F88A463064A49E7" italics="true" pageId="0" pageNumber="43">I. parvisepala</emphasis>
</taxonomicName>
(data from 
<bibRefCitation id="108A5F9AFB0C704CCBCC2070FF81A935" DOI="https://doi.org/10.5735/085.048.0108" author="Hou, YT" journalOrPublisher="Annales Botanici Fennici" pageId="0" pageNumber="43" pagination="57 - 62" refId="B11" refString="Hou, YT, Zhou, XR, Yu, SX, 2011. Impatiens parvisepala (Balsaminaceae), a new species from Guangxi, China. Annales Botanici Fennici 48 (1): 57 - 62, DOI: https://doi.org/10.5735/085.048.0108" title="Impatiens parvisepala (Balsaminaceae), a new species from Guangxi, China." url="https://doi.org/10.5735/085.048.0108" volume="48" year="2011">Hou et al. 2011</bibRefCitation>
and 
<bibRefCitation id="56F0FB3FEA47A72A9A19B4D75510C073" author="Son, HT" journalOrPublisher="Vietnam Journal of Forest Science" pageId="0" pageNumber="43" pagination="4018 - 4020" publicationUrl="http://vafs.gov.vn/" refId="B19" refString="Son, HT, Bon, Tn, Hung, NQ, Vinh, PV, Anh, TV, 2015. Impatiens parvisepala (Balsaminaceae): a newly recorded from Vietnam. Vietnam Journal of Forest Science 4: 4018 - 4020, http://vafs.gov.vn/" title="Impatiens parvisepala (Balsaminaceae): a newly recorded from Vietnam." url="http://vafs.gov.vn/" volume="4" year="2015">Son et al. 2015</bibRefCitation>
), 
<taxonomicName id="BC1E4D404EF9B272311FE7B6F1308976" lsidName="I. apalophylla" pageId="0" pageNumber="43" rank="species" species="apalophylla">
<emphasis id="0EE7DABD2F73C9C8C49CEF850C27E054" italics="true" pageId="0" pageNumber="43">I. apalophylla</emphasis>
</taxonomicName>
(data from 
<bibRefCitation id="91C9C6BD66E59750E256B5E9DE5E69E5" author="Hooker, JD" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="43" refId="B9" refString="Hooker, JD, 1908a. Les Especes Du Genre &quot;Impatiens&quot; dans l'Herbier du Museum de Paris. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, ser. 4, 10: 233-272." title="Les Especes Du Genre &quot; Impatiens &quot; dans l'Herbier du Museum de Paris. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, ser. 4, 10: 233 - 272." year="1908 a">Hooker 1908a</bibRefCitation>
and 
<bibRefCitation id="47FE9C7FEF90B85ADFAFCD974F09D06B" author="Chen, YL" editor="Wu, ZY" journalOrPublisher="12). Science Press, Beijing &amp; Missouri Botanical Garden Press, St. Louis" pageId="0" pageNumber="43" pagination="43 - 113" refId="B3" refString="Chen, YL, Akiyama, S, Ohba, H, 2007. Balsaminaceae. In: Wu, ZY, Raven, PH, Hong, DY, Eds., Flora of China (Vol. 12). Science Press, Beijing &amp; Missouri Botanical Garden Press, St. Louis: 43 - 113" title="Balsaminaceae." volumeTitle="Flora of China (Vol." year="2007">Chen et al. 2007</bibRefCitation>
), 
<taxonomicName id="C0D0310FFA8FEDBE419B09A9DB11A2D2" lsidName="I. clavigera" pageId="0" pageNumber="43" rank="species" species="clavigera">
<emphasis id="841272C12A99D00167AAC4AD76BFFE8A" italics="true" pageId="0" pageNumber="43">I. clavigera</emphasis>
</taxonomicName>
(data from Hooker 1909b, 
<bibRefCitation id="AB8904FC20B7E44104DAA8E8EDC2D611" DOI="https://doi.org/10.5735/085.048.0108" author="Hou, YT" journalOrPublisher="Annales Botanici Fennici" pageId="0" pageNumber="43" pagination="57 - 62" refId="B11" refString="Hou, YT, Zhou, XR, Yu, SX, 2011. Impatiens parvisepala (Balsaminaceae), a new species from Guangxi, China. Annales Botanici Fennici 48 (1): 57 - 62, DOI: https://doi.org/10.5735/085.048.0108" title="Impatiens parvisepala (Balsaminaceae), a new species from Guangxi, China." url="https://doi.org/10.5735/085.048.0108" volume="48" year="2011">Hou et al. 2011</bibRefCitation>
and 
<bibRefCitation id="26378B429B33A665DF389D21A3C38549" DOI="https://doi.org/10.1186/s40529-015-0108-4" author="Tan, YH" journalOrPublisher="Caryologia" pageId="0" pageNumber="43" refId="B21" refString="Tan, YH, Liu, YN, Jiang, H, Zhu, XX, Zhang, W, Yu, SX, 2015. Impatiens pandurata (Balsaminaceae), a new species from Yunnan, China. Botanical Studies 56: 29. https://doi.org/10.1186/s40529-015-0108-4" title="Impatiens pandurata (Balsaminaceae), a new species from Yunnan, China. Botanical Studies 56: 29." url="https://doi.org/10.1186/s40529-015-0108-4" year="2015">Tan et al. 2015</bibRefCitation>
,) and 
<taxonomicName id="10CFA40D79FD90726B5D0892E19C9CE7" lsidName="I. tianlinensis" pageId="0" pageNumber="43" rank="species" species="tianlinensis">
<emphasis id="9F079C169A61F1A91975C8901E035771" italics="true" pageId="0" pageNumber="43">I. tianlinensis</emphasis>
</taxonomicName>
(data from 
<bibRefCitation id="DCE0E660013495D23126B3940F772097" DOI="https://doi.org/10.11646/phytotaxa.227.3.4" author="Zeng, L" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="43" pagination="253 - 260" refId="B27" refString="Zeng, L, Liu, YN, Gogoi, R, Zhang, LJ, Yu, SX, 2015. Impatiens tianlinensis (Balsaminaceae), a new species from Guangxi, China. Phytotaxa 227 (3): 253 - 260, DOI: https://doi.org/10.11646/phytotaxa.227.3.4" title="Impatiens tianlinensis (Balsaminaceae), a new species from Guangxi, China." url="https://doi.org/10.11646/phytotaxa.227.3.4" volume="227" year="2015">Zeng et al. 2015</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="56360FEB20DF42AF593B1B35735CCCDD" pageId="0" pageNumber="43">
<table id="70982BEAC669456D1AC3E4A2E491CA0B" pageId="0" pageNumber="43">
<tr id="D44BB4E536E19B0E0B2AE65B5FC2F9A1" pageId="0" pageNumber="43">
<th id="5465A7C66EA4F34A206E0FD1BA217939" colspan="1" pageId="0" pageNumber="43" rowspan="1">Character</th>
<th id="431D090FDEB42A0C8A187FBA5B2FE281" colspan="1" pageId="0" pageNumber="43" rowspan="1">
<taxonomicName id="8E6C09ED982F852F6CAAC0F4E732F940" lsidName="I. wutaishanensis" pageId="0" pageNumber="43" rank="species" species="wutaishanensis">
<emphasis id="741652DB17E6130BE29941BFD8558640" italics="true" pageId="0" pageNumber="43">I. wutaishanensis</emphasis>
</taxonomicName>
</th>
<th id="EE74F88AE870AC60DBD232C70C4C6462" colspan="1" pageId="0" pageNumber="43" rowspan="1">
<taxonomicName id="7F5BD01801DE75684E8499C4FB66C9A5" lsidName="I. parvisepala" pageId="0" pageNumber="43" rank="species" species="parvisepala">
<emphasis id="A5467AB9276F368BCFC0F7A8A5958E8A" italics="true" pageId="0" pageNumber="43">I. parvisepala</emphasis>
</taxonomicName>
</th>
<th id="BB3FDDD6A4C8F6E3911B8CF4CDB4CC8B" colspan="1" pageId="0" pageNumber="43" rowspan="1">
<taxonomicName id="55990DB86B58A19B89C395F2CD49FEAE" lsidName="I. apalophylla" pageId="0" pageNumber="43" rank="species" species="apalophylla">
<emphasis id="03C8A1ED55F277CA052842DF35A1C06D" italics="true" pageId="0" pageNumber="43">I. apalophylla</emphasis>
</taxonomicName>
</th>
<th id="44CBB6B0404DAB99CD6618F0F4C0EBA3" colspan="1" pageId="0" pageNumber="43" rowspan="1">
<taxonomicName id="DC2AB3ADE6F0B548690F2DC4EAB80E0D" lsidName="I. clavigera" pageId="0" pageNumber="43" rank="species" species="clavigera">
<emphasis id="61C9C762F9B4E6D577A74577D88D2C11" italics="true" pageId="0" pageNumber="43">I. clavigera</emphasis>
</taxonomicName>
</th>
<th id="92F8906BDFAC3F717A95150B89D3B795" colspan="1" pageId="0" pageNumber="43" rowspan="1">
<taxonomicName id="172F03B02D978DD5EA813FE844F572D8" lsidName="I. tianlinensis" pageId="0" pageNumber="43" rank="species" species="tianlinensis">
<emphasis id="114F15A32F9EC22E8ACC8991CB60D4DA" italics="true" pageId="0" pageNumber="43">I. tianlinensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="B9A37787220090700EE3581D918A669C" pageId="0" pageNumber="43">
<td id="77F51A28C53A8E082C68C9C9D3900CFF" colspan="1" pageId="0" pageNumber="43" rowspan="1">Plant height</td>
<td id="365E9A7CBA0137DF97D3989AC57CCBB9" colspan="1" pageId="0" pageNumber="43" rowspan="1">
10-35 (16.71 
<normalizedToken id="8DA8161A0D1AA5BF92AC554CCC3DCADA" originalValue="±">+/-</normalizedToken>
5.30) cm
</td>
<td id="71083BC21A88ADA76CD9FD37C015E375" colspan="1" pageId="0" pageNumber="43" rowspan="1">35-60 cm</td>
<td id="2CDA54101B37F73C6886532FF0059218" colspan="1" pageId="0" pageNumber="43" rowspan="1">30-60 cm</td>
<td id="8E4D7BB124C5954635C7DEFC5CCA2C2F" colspan="1" pageId="0" pageNumber="43" rowspan="1">50-60 cm</td>
<td id="BD742F38383A48314ECCF8ED91512FD7" colspan="1" pageId="0" pageNumber="43" rowspan="1">50-80 cm</td>
</tr>
<tr id="E71CA5EB6774799E3D329E2687F515F3" pageId="0" pageNumber="43">
<td id="A0DF77542CFA4FB59D07962B3FCFB5A6" colspan="1" pageId="0" pageNumber="43" rowspan="1">Length of petiole</td>
<td id="F48E6C1936363509E90AF0BC6D89C578" colspan="1" pageId="0" pageNumber="43" rowspan="1">
0.5-2.4 (1.02 
<normalizedToken id="597F6D6C3C30AF063B443819E39E75CE" originalValue="±">+/-</normalizedToken>
0.44) cm
</td>
<td id="DD5C7F72C3608AF2C0DB9CED0E296491" colspan="1" pageId="0" pageNumber="43" rowspan="1">sub-sessile or sessile</td>
<td id="0F9EA7DD18B04F840BE53FC33E839DEB" colspan="1" pageId="0" pageNumber="43" rowspan="1">2-4 cm</td>
<td id="7A1376DF9CADBB9C16363DC89E43867C" colspan="1" pageId="0" pageNumber="43" rowspan="1">1-2 cm</td>
<td id="0DFF00F427A7409B86A22052B1C7EEA1" colspan="1" pageId="0" pageNumber="43" rowspan="1">(0.5-) 1-2 cm</td>
</tr>
<tr id="0199C80BB8A4D8C0D7C71C712A349E56" pageId="0" pageNumber="43">
<td id="EE8B94203C6F5053CFEC86C9B77FBB06" colspan="1" pageId="0" pageNumber="43" rowspan="1">Shape of leaf blade</td>
<td id="E48E3ECC537308FD4DCA6CCC0C5044FA" colspan="1" pageId="0" pageNumber="43" rowspan="1">elliptic, lanceolate, or lanceolate-oblong</td>
<td id="C65977F40B97BC02A2A1610A080491B6" colspan="1" pageId="0" pageNumber="43" rowspan="1">obovate or obovate-lanceolate</td>
<td id="0D6A72E2535EEB3D9C4BC59C8F2B47C7" colspan="1" pageId="0" pageNumber="43" rowspan="1">oblong-ovate or oblong-oblanceolate</td>
<td id="5F763C6783D70419541B84D57746A7D4" colspan="1" pageId="0" pageNumber="43" rowspan="1">obovate or oblanceolate</td>
<td id="366633E3BF3BB2D7D1BC35C30EA8DDF1" colspan="1" pageId="0" pageNumber="43" rowspan="1">obovate to oblanceolate</td>
</tr>
<tr id="3A46C2460A4687E993611F2E35B9AD8D" pageId="0" pageNumber="43">
<td id="D28BF36DF69EDC3016F96914B305097A" colspan="1" pageId="0" pageNumber="43" rowspan="1">Size of leaf</td>
<td id="E284076F6CD3D45A923A11DE67C810D7" colspan="1" pageId="0" pageNumber="43" rowspan="1">
3.5-12 (7.53 
<normalizedToken id="64BD1819D9B88BEB6FBB707EF8DCC079" originalValue="±">+/-</normalizedToken>
1.86) 
<normalizedToken id="4E355C66D12F1DD182DB0EB874C5F1AE" originalValue="×">x</normalizedToken>
1.5-4 (2.90 
<normalizedToken id="5FDEF4C2F6BAD388CDA6B6C16501A281" originalValue="±">+/-</normalizedToken>
0.79) cm
</td>
<td id="9AEB6AB2A4D97EC521AEE9BE4EF278EF" colspan="1" pageId="0" pageNumber="43" rowspan="1">
12-20 
<normalizedToken id="85B1F79A412AA4B8749E0EC1EBF9D7DD" originalValue="×">x</normalizedToken>
3.5-6 cm
</td>
<td id="C8E8C4EBA3FF4A8ED922EC02BCCAA357" colspan="1" pageId="0" pageNumber="43" rowspan="1">
10-22 
<normalizedToken id="37AF456E0203819727BF983C79677CF0" originalValue="×">x</normalizedToken>
4-8 cm
</td>
<td id="10F4F857A7BC81B34A1BCB4229A77EC7" colspan="1" pageId="0" pageNumber="43" rowspan="1">
8-15 (-18) 
<normalizedToken id="C482E98219C92BD1609F8C031CED57F8" originalValue="×">x</normalizedToken>
3.5-5 cm
</td>
<td id="1271FBD3BCE3038347296AB2EE79D88C" colspan="1" pageId="0" pageNumber="43" rowspan="1">
10-15 (-18) 
<normalizedToken id="D8854AB7DF2D60C4B4BD9619406C7A30" originalValue="×">x</normalizedToken>
5-8 cm
</td>
</tr>
<tr id="86EE73A5631337ABE54A4C09BDFFF389" pageId="0" pageNumber="43">
<td id="0E56CE10846E15A4FEAFAEBC51F51121" colspan="1" pageId="0" pageNumber="43" rowspan="1">Length of peduncle</td>
<td id="7F9C721339F0275DC4953A85B7981792" colspan="1" pageId="0" pageNumber="43" rowspan="1">
4-28 (13.00 
<normalizedToken id="1335637CEFEF5886EC03FC075C12BAF0" originalValue="±">+/-</normalizedToken>
6.71) cm
</td>
<td id="43933FE9CCF3A14D40059470E341A878" colspan="1" pageId="0" pageNumber="43" rowspan="1">15-17 cm</td>
<td id="76003C2026B4055BFBA52AE8B6F8C9A3" colspan="1" pageId="0" pageNumber="43" rowspan="1">7-15 cm</td>
<td id="4723BF6A66F1494572FBDA817449B968" colspan="1" pageId="0" pageNumber="43" rowspan="1">8-10 cm</td>
<td id="5E40EE7EAADF24694DC6CA72A5B1A713" colspan="1" pageId="0" pageNumber="43" rowspan="1">10-15 cm,</td>
</tr>
<tr id="CBD98E9D2628FC2BE04504FEE86ECF2F" pageId="0" pageNumber="43">
<td id="70B5849C8FA1E146E3A3DD4DA392EABE" colspan="1" pageId="0" pageNumber="43" rowspan="1">No. of flowers in an inflorescence</td>
<td id="9CDC6D928BE933D7DBCC295AA593F79B" colspan="1" pageId="0" pageNumber="43" rowspan="1">up to 22 flowers</td>
<td id="101CC973302FCEE62A6CB408D5CF1FA3" colspan="1" pageId="0" pageNumber="43" rowspan="1">6-8 flowers</td>
<td id="73CE2147EFB40FB260E11089A3ABEB5F" colspan="1" pageId="0" pageNumber="43" rowspan="1">4-10 flowers</td>
<td id="DBE66BBD46E4A821FD562596A13FFC3D" colspan="1" pageId="0" pageNumber="43" rowspan="1">5-9 flowers</td>
<td id="DB8BB4583F0FEEF961555C33C3D341A8" colspan="1" pageId="0" pageNumber="43" rowspan="1">3-5 (-7) flowers</td>
</tr>
<tr id="3FB5B26876184BD87A865AF11888FB84" pageId="0" pageNumber="43">
<td id="909B8E89ADAA700448709BF957EAE3D9" colspan="1" pageId="0" pageNumber="43" rowspan="1">Shape and length of spur</td>
<td id="10F38E7005C7488E2460F944A76288B2" colspan="1" pageId="0" pageNumber="43" rowspan="1">
slightly incurved or incurved, gradually constricted, 1.2-3 (2.06 
<normalizedToken id="FB4778D56A80E054ADF1CD5AC247054A" originalValue="±">+/-</normalizedToken>
0.45) cm long
</td>
<td id="103AC025EEC3F96C72D97FD003AE3686" colspan="1" pageId="0" pageNumber="43" rowspan="1">nearly straight, gradually constricted, 2-2.5 cm long</td>
<td id="0B9607058EF7BC6D303477B643D7D2AB" colspan="1" pageId="0" pageNumber="43" rowspan="1">incurved, abruptly elongated, 2-2.5 cm long</td>
<td id="470C1E2AD0CC323A9C4D946F605EAC9D" colspan="1" pageId="0" pageNumber="43" rowspan="1">incurved, abruptly narrowed, 5-6 mm long</td>
<td id="3F043D038DE08ECB2358ADB8AE819C28" colspan="1" pageId="0" pageNumber="43" rowspan="1">involuted, abruptly constricted, 1-1.5 cm long</td>
</tr>
<tr id="90740C51545D0DFA845136CE0BDEB9C1" pageId="0" pageNumber="43">
<td id="BC1782C313DE481C1F9BE6FC8AD69D8B" colspan="1" pageId="0" pageNumber="43" rowspan="1">Bracts</td>
<td id="778C11E9976051CD4AB5ABBE0C6658E2" colspan="1" pageId="0" pageNumber="43" rowspan="1">lanceolate to subulate, persistent</td>
<td id="154D453E12230D58802809EA58724D6D" colspan="1" pageId="0" pageNumber="43" rowspan="1">lanceolate or subulate, persistent</td>
<td id="EAC7967D2D85B765450B8895B1A79BFF" colspan="1" pageId="0" pageNumber="43" rowspan="1">invisible</td>
<td id="8218C382F62A83D7A7C59DBD966924DE" colspan="1" pageId="0" pageNumber="43" rowspan="1">ovate, caducous</td>
<td id="D32E635EBF0013014DF0D9E65C85FA23" colspan="1" pageId="0" pageNumber="43" rowspan="1">ovate, deciduous</td>
</tr>
<tr id="B64FD17E90F7A2A22175BD249995B45A" pageId="0" pageNumber="43">
<td id="4048C6EF067E214FCF4CB558D6465090" colspan="1" pageId="0" pageNumber="43" rowspan="1">Lower sepal</td>
<td id="BEC923228211B63374854283DEF31E70" colspan="1" pageId="0" pageNumber="43" rowspan="1">
obliquely infundibuliform, with reddish patches, 1.2-2.8 (1.94 
<normalizedToken id="A7E9F1ED1FB8C259C20B3B4B75F01EAA" originalValue="±">+/-</normalizedToken>
0.38) 
<normalizedToken id="03F502BE2437BFFC126619BD19BF1444" originalValue="×">x</normalizedToken>
0.8-2.2 (1.38 
<normalizedToken id="A60BD5ACC93B1F17D984D9166B7BAC60" originalValue="±">+/-</normalizedToken>
0.37) cm
</td>
<td id="99A876D2876DE250086A2389204FCC25" colspan="1" pageId="0" pageNumber="43" rowspan="1">
obliquely infundibuliform, with reddish patches, 1.8-2 
<normalizedToken id="D6DEC12C6D03ADBCA6D8FDB36A6E0FBD" originalValue="×">x</normalizedToken>
2.5-3 cm
</td>
<td id="7418DCD09F471BAE4CF8BABEE2FECD1F" colspan="1" pageId="0" pageNumber="43" rowspan="1">narrowly funnel-shaped, with reddish patches, 3 cm</td>
<td id="87138DE7957CA62350F58F0CE5632FF2" colspan="1" pageId="0" pageNumber="43" rowspan="1">
narrowly funnel-shaped, absent patch, 2 
<normalizedToken id="8538DFD2C3112B882F9336D83C7C5EB1" originalValue="×">x</normalizedToken>
3 cm
</td>
<td id="546A66231235788FD958EBA1C70A76E7" colspan="1" pageId="0" pageNumber="43" rowspan="1">slightly narrowly funnel-shaped, with reddish patches, 2.5-3.5 cm</td>
</tr>
<tr id="805A4318866430D059DD256EF5602E03" pageId="0" pageNumber="43">
<td id="D58ABC0A249273C27EE9FAD54DADC88D" colspan="1" pageId="0" pageNumber="43" rowspan="1">Dorsal petal</td>
<td id="35CB360126F699FEAD1FD7627487C75F" colspan="1" pageId="0" pageNumber="43" rowspan="1">
ovate, 1.3-2.3 (1.80 
<normalizedToken id="956B12662CD53A98339FBC74217E0CB2" originalValue="±">+/-</normalizedToken>
0.28) 
<normalizedToken id="30400CCD7BA0558F18EE882148EDAC65" originalValue="×">x</normalizedToken>
0.7-1.6 (1.15 
<normalizedToken id="73988FEAFCE7CB5E4173531C29F04D7D" originalValue="±">+/-</normalizedToken>
0.26) cm
</td>
<td id="800CBF37F43835404547826EF7244891" colspan="1" pageId="0" pageNumber="43" rowspan="1">
obovate, 1.8-2.2 
<normalizedToken id="2D4E5BD202286CB78A7500331264ACE5" originalValue="×">x</normalizedToken>
1.5-2 cm
</td>
<td id="424E6E68CC69CDA2ED6EE4F2A748E83A" colspan="1" pageId="0" pageNumber="43" rowspan="1">ellipticum, 1-4 cm</td>
<td id="64973BE487FF310F9A5C200746BE495F" colspan="1" pageId="0" pageNumber="43" rowspan="1">obovate, ca. 2 cm</td>
<td id="D8216B889AF4D29E96C322C5838AB558" colspan="1" pageId="0" pageNumber="43" rowspan="1">
ovate, 10-12 
<normalizedToken id="170FB6E32E0F6A49D6163DE3FD663F0C" originalValue="×">x</normalizedToken>
7-9 mm
</td>
</tr>
<tr id="BC54A7E4FE4C02BE4FC94B7C7A4DFF2A" pageId="0" pageNumber="43">
<td id="593BE036DAA03C5940D1658670833FAD" colspan="1" pageId="0" pageNumber="43" rowspan="1">Lateral united petals</td>
<td id="C587D414CE705E5AF2A4822C3C990E84" colspan="1" pageId="0" pageNumber="43" rowspan="1">1.6-3 cm long, with reddish patches</td>
<td id="0C53F8D373686C46A45AC04EDE079329" colspan="1" pageId="0" pageNumber="43" rowspan="1">2.8-3 cm long, with reddish patches</td>
<td id="2D3F0E1755C55BC9DEA613D71BD828A6" colspan="1" pageId="0" pageNumber="43" rowspan="1">2-3 cm long, with apparent reddish patches</td>
<td id="5F874813E9998864E46B832C9C6655B8" colspan="1" pageId="0" pageNumber="43" rowspan="1">2-6 cm long, absent patch</td>
<td id="97396F672991A1A9ADB51E10CD278A6B" colspan="1" pageId="0" pageNumber="43" rowspan="1">2-2.5 cm long, with reddish patches</td>
</tr>
<tr id="69AEB79F73784BACDC7C26C01DCA6B7B" pageId="0" pageNumber="43">
<td id="14DE56C2E6AC3496E4D7833B86107300" colspan="1" pageId="0" pageNumber="43" rowspan="1">Pollen grains</td>
<td id="9B4619800128157A747391EA0A165531" colspan="1" pageId="0" pageNumber="43" rowspan="1">
30.00 
<normalizedToken id="60369FA2B098DE40FA2DBA1A667D8B64" originalValue="±">+/-</normalizedToken>
0.98 (28.35-32.11) 
<normalizedToken id="30E4E07744569EA8F1E33664BA6F6742" originalValue="×">x</normalizedToken>
16.56 
<normalizedToken id="37E1459D2F22419AB9A128663B4D5DC8" originalValue="±">+/-</normalizedToken>
1.78 (14.51-21.73) 
<normalizedToken id="42F826E8C0CCAF71CF4457FEA95CF992" originalValue="μm">μm</normalizedToken>
</td>
<td id="9C6548DC77ECEFEBFF86EA2DF504E74A" colspan="1" pageId="0" pageNumber="43" rowspan="1">
28.84 (27.98-30.17) 
<normalizedToken id="57E4502FCA973A52EAAA196DC6D0B435" originalValue="×">x</normalizedToken>
20.57 (19.70-21.63) 
<normalizedToken id="C1FE148752A5F5CD25055747D89006D2" originalValue="μm">μm</normalizedToken>
</td>
<td id="D00F37684FD8DCE23B886686CE4A3DD7" colspan="1" pageId="0" pageNumber="43" rowspan="1">
31.85 (30.96-32.73) 
<normalizedToken id="0F39018E5686F4756F949D2F9A66765A" originalValue="×">x</normalizedToken>
5.57 (5.13-6.75) 
<normalizedToken id="0528397FC89E2492EC94B98CB9607D36" originalValue="μm">μm</normalizedToken>
</td>
<td id="D11FB5750C50ECF73A691350A1EB56FD" colspan="1" pageId="0" pageNumber="43" rowspan="1">Missing data</td>
<td id="32407CC96133ED02CB15424A1074EC78" colspan="1" pageId="0" pageNumber="43" rowspan="1">
30.13 (29.62-30.47) 
<normalizedToken id="2D18F3CE141CC06A77AE59BCCB49E36D" originalValue="×">x</normalizedToken>
12.95 (12.68-13.54) 
<normalizedToken id="B47205735B9DA9B56D1B3970E66B6D8C" originalValue="μm">μm</normalizedToken>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>