204 lines
32 KiB
XML
204 lines
32 KiB
XML
<document id="CF6860A324D7F18C839B1ED850F2C2C1" ID-DOI="10.5281/zenodo.186352" ID-GBIF-Dataset="c87b0ab2-a35e-4069-8c94-cb29159d4ec0" ID-ISSN="1175-5326" ID-Zenodo-Dep="186352" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461050678747" checkinUser="plazi" docAuthor="Raven, Robert J." docDate="2009" docId="774E87C4D666FF84FF3BFB78FB56FA06" docLanguage="en" docName="zt02035p040.pdf" docOrigin="Zootaxa 2035" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nuliodon Raven, 2009, gen. nov." docType="treatment" docVersion="9" lastPageNumber="32" masterDocId="8B77FFBCD678FF9BFFACFFF7FFD4FF81" masterDocTitle="Revisions of Australian ground-hunting spiders: IV. The spider subfamily Diaprograptinae subfam. nov. (Araneomorphae: Miturgidae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="31" updateTime="1698578758242" updateUser="plazi">
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<mods:titleInfo id="02C9DBE66F9E5D6467B4DA1997DECA67">
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<mods:title id="7F63684DC99A14EC535029676028BF85">Revisions of Australian ground-hunting spiders: IV. The spider subfamily Diaprograptinae subfam. nov. (Araneomorphae: Miturgidae)</mods:title>
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<mods:namePart id="9CE989BC0985585B2BED70C281AB319D">Raven, Robert J.</mods:namePart>
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<mods:title id="77A8B6E059ED6703275A36F94880D7CC">Zootaxa</mods:title>
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<mods:part id="00E6A5BB36A197D1451564EB1D79D3BE">
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<mods:date id="15B37CC449CE02A8ECB6F715D3633009">2009</mods:date>
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<mods:detail id="77A2B788D5A5AA670A411459FCB58A9D" type="volume">
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<mods:number id="8CB08847C14ECA08EB64A8E9B9F941C7">2035</mods:number>
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<mods:classification id="2E96F6736960550E5D422E5676FDB749">journal article</mods:classification>
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<mods:identifier id="463679C856271EEDDAA2CFD7C27B2FC3" type="DOI">10.5281/zenodo.186352</mods:identifier>
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<mods:identifier id="C56EBE4682F098AAA05BF98F64DE145A" type="GBIF-Dataset">c87b0ab2-a35e-4069-8c94-cb29159d4ec0</mods:identifier>
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<mods:identifier id="A2F87E9E53B8B0B7C210C7B72052683A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0C570777190CB1B26D1D0FAD9B9FC8A2" type="Zenodo-Dep">186352</mods:identifier>
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<treatment id="774E87C4D666FF84FF3BFB78FB56FA06" ID-DOI="http://doi.org/10.5281/zenodo.6216644" ID-GBIF-Taxon="119612061" ID-Zenodo-Dep="6216644" LSID="urn:lsid:plazi:treatment:774E87C4D666FF84FF3BFB78FB56FA06" httpUri="http://treatment.plazi.org/id/774E87C4D666FF84FF3BFB78FB56FA06" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
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<subSubSection id="B7FD6559D666FF85FF3BFB78FEAAFB28" box="[151,382,1167,1193]" pageId="30" pageNumber="31" type="nomenclature">
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<paragraph id="FF5836D2D666FF85FF3BFB78FEAAFB28" blockId="30.[151,382,1167,1193]" box="[151,382,1167,1193]" pageId="30" pageNumber="31">
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<heading id="A41081BED666FF85FF3BFB78FEAAFB28" bold="true" box="[151,382,1167,1193]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
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<emphasis id="CD93EAC0D666FF85FF3BFB78FEAAFB28" bold="true" box="[151,382,1167,1193]" pageId="30" pageNumber="31">
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<taxonomicName id="38E74D51D666FF85FF3BFB78FEDDFB28" ID-CoL="63GB7" ID-ENA="1956489" box="[151,265,1167,1193]" class="Arachnida" family="Miturgidae" genus="Nuliodon" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="CD93EAC0D666FF85FF3BFB78FEDDFB28" bold="true" box="[151,265,1167,1193]" italics="true" pageId="30" pageNumber="31">Nuliodon</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D6A057BBD666FF85FEBCFB78FEAAFB28" box="[272,382,1167,1193]" pageId="30" pageNumber="31" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B7FD6559D666FF85FF3BFB2FFD57FB73" box="[151,643,1240,1267]" pageId="30" pageNumber="31" type="materials_examined">
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<paragraph id="FF5836D2D666FF85FF3BFB2FFD57FB73" blockId="30.[151,1437,1240,2002]" box="[151,643,1240,1267]" pageId="30" pageNumber="31">
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<emphasis id="CD93EAC0D666FF85FF3BFB2FFEE6FB73" bold="true" box="[151,306,1240,1266]" pageId="30" pageNumber="31">
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<typeStatus id="205C8870D666FF85FF3BFB2FFF00FB73" box="[151,212,1240,1266]" pageId="30" pageNumber="31">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName id="38E74D51D666FF85FEECFB2EFDCBFB73" box="[320,543,1241,1266]" class="Arachnida" family="Miturgidae" genus="Nuliodon" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="fishburni" status="sp. nov.">
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<emphasis id="CD93EAC0D666FF85FEECFB2EFDCBFB73" box="[320,543,1241,1266]" italics="true" pageId="30" pageNumber="31">Nuliodon fishburni</emphasis>
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</taxonomicName>
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<emphasis id="CD93EAC0D666FF85FD8AFB2FFD57FB73" bold="true" box="[550,643,1240,1266]" pageId="30" pageNumber="31">
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<taxonomicNameLabel id="D6A057BBD666FF85FD8AFB2FFD57FB73" box="[550,643,1240,1266]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="B7FD6559D666FF85FF69FB08FD93FA5A" pageId="30" pageNumber="31" type="diagnosis">
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<paragraph id="FF5836D2D666FF85FF69FB08FD93FA5A" blockId="30.[151,1437,1240,2002]" pageId="30" pageNumber="31">
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<emphasis id="CD93EAC0D666FF85FF69FB08FE91FA98" bold="true" box="[197,325,1279,1305]" pageId="30" pageNumber="31">Diagnosis.</emphasis>
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Differs from
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<taxonomicName id="38E74D51D666FF85FE44FAF7FD93FA98" box="[488,583,1280,1305]" class="Arachnida" family="Miturgidae" genus="Miturga" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="CD93EAC0D666FF85FE44FAF7FD93FA98" box="[488,583,1280,1305]" italics="true" pageId="30" pageNumber="31">Miturga</emphasis>
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</taxonomicName>
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in the presence of claw tufts (
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<figureCitation id="67DC2A57D666FF85FC02FAF7FBD1FA9B" box="[942,1029,1280,1306]" captionStart="FIGURE 24" captionStartId="35.[151,257,1518,1542]" captionTargetBox="[151,1435,886,1494]" captionTargetId="figure@35.[151,1436,886,1494]" captionTargetPageId="35" captionText="FIGURE 24. Nuliodon fishburni sp. nov., female, scanning electron micrographs: a, tarsal trichobothria and cuticle, dorsal view; b, tarsal organ, dorsal view; c, claw tufts." httpUri="https://zenodo.org/record/186375/files/figure.png" pageId="30" pageNumber="31">Fig. 24</figureCitation>
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a), from
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<taxonomicName id="38E74D51D666FF85FBC6FAF7FB38FA98" box="[1130,1260,1280,1305]" class="Arachnida" family="Miturgidae" genus="Mituliodon" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="CD93EAC0D666FF85FBC6FAF7FB38FA98" box="[1130,1260,1280,1305]" italics="true" pageId="30" pageNumber="31">Mituliodon</emphasis>
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</taxonomicName>
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in the absence of strong scopulae on metatarsi and tarsi I–IV, the form of the male palp (
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<figureCitation id="67DC2A57D666FF85FBBEFAD1FBBEFAC1" box="[1042,1130,1318,1344]" captionStart="FIGURE 21" captionStartId="32.[151,255,1933,1957]" captionTargetBox="[151,1436,429,1906]" captionTargetId="figure@32.[151,1436,429,1906]" captionTargetPageId="32" captionText="FIGURE 21. Nuliodon fishburni sp. nov., male: a, palpal tibia, cymbium and bulb, ventral view; b, median apophysis, ventral view; c, d, RTA, ventral (c) and retrolateral view (d)." httpUri="https://zenodo.org/record/186372/files/figure.png" pageId="30" pageNumber="31">Fig. 21</figureCitation>
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a, b), viz., short embolus, absence of cymbial groove and small conductor. Resembles
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<taxonomicName id="38E74D51D666FF85FCC3FABAFC38FAE7" box="[879,1004,1357,1382]" class="Arachnida" family="Miturgidae" genus="Mitzoruga" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="CD93EAC0D666FF85FCC3FABAFC38FAE7" box="[879,1004,1357,1382]" italics="true" pageId="30" pageNumber="31">Mitzoruga</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D6A057BBD666FF85FC58FABAFBB6FAE6" box="[1012,1122,1357,1383]" pageId="30" pageNumber="31" rank="genus">gen. nov.</taxonomicNameLabel>
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in the distodorsal crest of thick curved setae on cymbium (
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<figureCitation id="67DC2A57D666FF85FDB8FA84FDBDFA0C" box="[532,617,1395,1421]" captionStart="FIGURE 21" captionStartId="32.[151,255,1933,1957]" captionTargetBox="[151,1436,429,1906]" captionTargetId="figure@32.[151,1436,429,1906]" captionTargetPageId="32" captionText="FIGURE 21. Nuliodon fishburni sp. nov., male: a, palpal tibia, cymbium and bulb, ventral view; b, median apophysis, ventral view; c, d, RTA, ventral (c) and retrolateral view (d)." httpUri="https://zenodo.org/record/186372/files/figure.png" pageId="30" pageNumber="31">Fig. 21</figureCitation>
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a), and the carapace shape but males differ in the stout RTA (
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<figureCitation id="67DC2A57D666FF85FA83FA84FA5DFA0C" box="[1327,1417,1395,1421]" captionStart="FIGURE 21" captionStartId="32.[151,255,1933,1957]" captionTargetBox="[151,1436,429,1906]" captionTargetId="figure@32.[151,1436,429,1906]" captionTargetPageId="32" captionText="FIGURE 21. Nuliodon fishburni sp. nov., male: a, palpal tibia, cymbium and bulb, ventral view; b, median apophysis, ventral view; c, d, RTA, ventral (c) and retrolateral view (d)." httpUri="https://zenodo.org/record/186372/files/figure.png" pageId="30" pageNumber="31">Figs 21</figureCitation>
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c, d), the absence of dark stripes on the carapace (
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<figureCitation id="67DC2A57D666FF85FD7EFA6DFCFDFA35" box="[722,809,1434,1460]" captionStart="FIGURE 22" captionStartId="33.[151,257,1302,1326]" captionTargetBox="[151,1435,197,1276]" captionTargetId="figure@33.[151,1436,197,1277]" captionTargetPageId="33" captionText="FIGURE 22. Nuliodon fishburni sp. nov., female: a, habitus; b, carapace, dorsal view; c, epigyne, external view; d, coxae IV, ventral view; e, carapace, abdomen and legs, dorsal view; f, sternum, maxillae and labium, ventral view; g, abdomen, ventral view. Scale lines: 5 mm for a, e; 1 mm for b, f; d, f, g to same scale; 0.2 mm for c." httpUri="https://zenodo.org/record/186373/files/figure.png" pageId="30" pageNumber="31">Fig. 22</figureCitation>
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e) and the large median apophysis is not in the same line as the embolus origin (
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<figureCitation id="67DC2A57D666FF85FE79FA36FDFFFA5A" box="[469,555,1473,1499]" captionStart="FIGURE 21" captionStartId="32.[151,255,1933,1957]" captionTargetBox="[151,1436,429,1906]" captionTargetId="figure@32.[151,1436,429,1906]" captionTargetPageId="32" captionText="FIGURE 21. Nuliodon fishburni sp. nov., male: a, palpal tibia, cymbium and bulb, ventral view; b, median apophysis, ventral view; c, d, RTA, ventral (c) and retrolateral view (d)." httpUri="https://zenodo.org/record/186372/files/figure.png" pageId="30" pageNumber="31">Fig. 21</figureCitation>
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a).
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</paragraph>
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</subSubSection>
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<subSubSection id="B7FD6559D666FF85FF69FA10FED3F9A9" pageId="30" pageNumber="31" type="etymology">
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<paragraph id="FF5836D2D666FF85FF69FA10FED3F9A9" blockId="30.[151,1437,1240,2002]" pageId="30" pageNumber="31">
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<emphasis id="CD93EAC0D666FF85FF69FA10FE9AF980" bold="true" box="[197,334,1511,1537]" pageId="30" pageNumber="31">Etymology</emphasis>
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. Based on the zoropsid genus name
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<taxonomicName id="38E74D51D666FF85FCA0FA1FFCBBF980" box="[780,879,1512,1537]" class="Arachnida" family="Zoropsidae" genus="Uliodon" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="CD93EAC0D666FF85FCA0FA1FFCBBF980" box="[780,879,1512,1537]" italics="true" pageId="30" pageNumber="31">Uliodon</emphasis>
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</taxonomicName>
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which it somatically resembles; the gender is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="B7FD6559D666FF84FF69F9C3FDFFFBD3" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
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<paragraph id="FF5836D2D666FF85FF69F9C3FA48F853" blockId="30.[151,1437,1240,2002]" pageId="30" pageNumber="31">
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<emphasis id="CD93EAC0D666FF85FF69F9C3FEB4F9CF" bold="true" box="[197,352,1588,1614]" pageId="30" pageNumber="31">Description.</emphasis>
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Carapace brown with irregular darker flecks along striae (
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<figureCitation id="67DC2A57D666FF85FB88F9C2FB55F9CE" box="[1060,1153,1589,1615]" captionStart="FIGURE 22" captionStartId="33.[151,257,1302,1326]" captionTargetBox="[151,1435,197,1276]" captionTargetId="figure@33.[151,1436,197,1277]" captionTargetPageId="33" captionText="FIGURE 22. Nuliodon fishburni sp. nov., female: a, habitus; b, carapace, dorsal view; c, epigyne, external view; d, coxae IV, ventral view; e, carapace, abdomen and legs, dorsal view; f, sternum, maxillae and labium, ventral view; g, abdomen, ventral view. Scale lines: 5 mm for a, e; 1 mm for b, f; d, f, g to same scale; 0.2 mm for c." httpUri="https://zenodo.org/record/186373/files/figure.png" pageId="30" pageNumber="31">Figs 22</figureCitation>
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b, e); abdomen dorsally fawn with irregular mottling forming weak internested “V”s; ventrally with irregular darker mottling; legs brown with weak paler bands (
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<figureCitation id="67DC2A57D666FF85FDAAF975FDB7F91D" box="[518,611,1666,1692]" captionStart="FIGURE 22" captionStartId="33.[151,257,1302,1326]" captionTargetBox="[151,1435,197,1276]" captionTargetId="figure@33.[151,1436,197,1277]" captionTargetPageId="33" captionText="FIGURE 22. Nuliodon fishburni sp. nov., female: a, habitus; b, carapace, dorsal view; c, epigyne, external view; d, coxae IV, ventral view; e, carapace, abdomen and legs, dorsal view; f, sternum, maxillae and labium, ventral view; g, abdomen, ventral view. Scale lines: 5 mm for a, e; 1 mm for b, f; d, f, g to same scale; 0.2 mm for c." httpUri="https://zenodo.org/record/186373/files/figure.png" pageId="30" pageNumber="31">Figs 22</figureCitation>
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a, e, g). Carapace pear-shaped with gentle anterior constriction; caput low; fovea present reaching to edge of posterior declivity; caput slightly elevated, distinct; other striae indistinct; lightly hirsute. Clypeus
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<emphasis id="CD93EAC0D666FF85FD97F927FD83F968" box="[571,599,1744,1769]" italics="true" pageId="30" pageNumber="31">ca</emphasis>
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. one AME diameter high. Eight eyes in two rows: anterior narrower, recurved; posterior more recurved (
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<figureCitation id="67DC2A57D666FF85FD93F901FD4CF891" box="[575,664,1782,1808]" captionStart="FIGURE 22" captionStartId="33.[151,257,1302,1326]" captionTargetBox="[151,1435,197,1276]" captionTargetId="figure@33.[151,1436,197,1277]" captionTargetPageId="33" captionText="FIGURE 22. Nuliodon fishburni sp. nov., female: a, habitus; b, carapace, dorsal view; c, epigyne, external view; d, coxae IV, ventral view; e, carapace, abdomen and legs, dorsal view; f, sternum, maxillae and labium, ventral view; g, abdomen, ventral view. Scale lines: 5 mm for a, e; 1 mm for b, f; d, f, g to same scale; 0.2 mm for c." httpUri="https://zenodo.org/record/186373/files/figure.png" pageId="30" pageNumber="31">Fig. 22</figureCitation>
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b). All eyes of similar size with anterior row eyes slightly larger. Group occupies two-thirds of headwidth. AME on common tubercle on clypeal edge, look forward, to side and up; ALE less than 1 diameter away look more up and to side, PME paler look up,
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<emphasis id="CD93EAC0D666FF85FB09F8B3FB15F8DC" box="[1189,1217,1860,1885]" italics="true" pageId="30" pageNumber="31">ca</emphasis>
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. 1 diameter apart; PLE look to side. Chelicerae small, vertical with small boss; fangs short, diaxial; dentition 3 small P, 2 R, of short furrow; long pallid ovoid area prolaterally above fangs. Maxillae (
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<figureCitation id="67DC2A57D666FF85FC52F866FB81F82A" box="[1022,1109,1937,1963]" captionStart="FIGURE 22" captionStartId="33.[151,257,1302,1326]" captionTargetBox="[151,1435,197,1276]" captionTargetId="figure@33.[151,1436,197,1277]" captionTargetPageId="33" captionText="FIGURE 22. Nuliodon fishburni sp. nov., female: a, habitus; b, carapace, dorsal view; c, epigyne, external view; d, coxae IV, ventral view; e, carapace, abdomen and legs, dorsal view; f, sternum, maxillae and labium, ventral view; g, abdomen, ventral view. Scale lines: 5 mm for a, e; 1 mm for b, f; d, f, g to same scale; 0.2 mm for c." httpUri="https://zenodo.org/record/186373/files/figure.png" pageId="30" pageNumber="31">Fig. 22</figureCitation>
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f) roughly diamond-shaped with rounded corners and shallow basal diagonal groove; serrula long. Labium (
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<figureCitation id="67DC2A57D666FF85FBF5F84FFB7BF853" box="[1113,1199,1976,2002]" captionStart="FIGURE 22" captionStartId="33.[151,257,1302,1326]" captionTargetBox="[151,1435,197,1276]" captionTargetId="figure@33.[151,1436,197,1277]" captionTargetPageId="33" captionText="FIGURE 22. Nuliodon fishburni sp. nov., female: a, habitus; b, carapace, dorsal view; c, epigyne, external view; d, coxae IV, ventral view; e, carapace, abdomen and legs, dorsal view; f, sternum, maxillae and labium, ventral view; g, abdomen, ventral view. Scale lines: 5 mm for a, e; 1 mm for b, f; d, f, g to same scale; 0.2 mm for c." httpUri="https://zenodo.org/record/186373/files/figure.png" pageId="30" pageNumber="31">Fig. 22</figureCitation>
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f) short, shield-like.
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</paragraph>
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<paragraph id="FF5836D2D667FF84FF3BFF6FFDFFFBD3" blockId="31.[151,1437,152,1415]" pageId="31" pageNumber="32">
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Sternum (
|
||
<figureCitation id="67DC2A57D667FF84FEA7FF6FFEB4FF33" box="[267,352,152,178]" captionStart="FIGURE 22" captionStartId="33.[151,257,1302,1326]" captionTargetBox="[151,1435,197,1276]" captionTargetId="figure@33.[151,1436,197,1277]" captionTargetPageId="33" captionText="FIGURE 22. Nuliodon fishburni sp. nov., female: a, habitus; b, carapace, dorsal view; c, epigyne, external view; d, coxae IV, ventral view; e, carapace, abdomen and legs, dorsal view; f, sternum, maxillae and labium, ventral view; g, abdomen, ventral view. Scale lines: 5 mm for a, e; 1 mm for b, f; d, f, g to same scale; 0.2 mm for c." httpUri="https://zenodo.org/record/186373/files/figure.png" pageId="31" pageNumber="32">Fig. 22</figureCitation>
|
||
f) almost lenticular; converges in front, rounded margins, strong setae along marginal band, marginal setae outward, central setae radially directed inward.
|
||
<emphasis id="CD93EAC0D667FF84FCDFFF48FC7EFF59" box="[883,938,191,216]" italics="true" pageId="31" pageNumber="32">Legs</emphasis>
|
||
. Coxae with rounded inner edges, anterior corner of basal strial attachment triangular, smaller process on posterior corner. RCH raised, pallid. Coxae IV of male cluster of
|
||
<emphasis id="CD93EAC0D667FF84FEC2FEFBFE5EFEA4" box="[366,394,268,293]" italics="true" pageId="31" pageNumber="32">ca</emphasis>
|
||
. 10–20 long thick setae on ventral inner corner (
|
||
<figureCitation id="67DC2A57D667FF84FC64FEFBFBCBFEA7" box="[968,1055,268,294]" captionStart="FIGURE 22" captionStartId="33.[151,257,1302,1326]" captionTargetBox="[151,1435,197,1276]" captionTargetId="figure@33.[151,1436,197,1277]" captionTargetPageId="33" captionText="FIGURE 22. Nuliodon fishburni sp. nov., female: a, habitus; b, carapace, dorsal view; c, epigyne, external view; d, coxae IV, ventral view; e, carapace, abdomen and legs, dorsal view; f, sternum, maxillae and labium, ventral view; g, abdomen, ventral view. Scale lines: 5 mm for a, e; 1 mm for b, f; d, f, g to same scale; 0.2 mm for c." httpUri="https://zenodo.org/record/186373/files/figure.png" pageId="31" pageNumber="32">Fig. 22</figureCitation>
|
||
d). Trochanters short, with small but distinct deep notches
|
||
<emphasis id="CD93EAC0D667FF84FE6FFEC5FE0AFECA" box="[451,478,306,331]" italics="true" pageId="31" pageNumber="32">ca</emphasis>
|
||
. 1.5 times wider than deep (
|
||
<figureCitation id="67DC2A57D667FF84FC82FEC5FC52FECD" box="[814,902,306,332]" captionStart="FIGURE 22" captionStartId="33.[151,257,1302,1326]" captionTargetBox="[151,1435,197,1276]" captionTargetId="figure@33.[151,1436,197,1277]" captionTargetPageId="33" captionText="FIGURE 22. Nuliodon fishburni sp. nov., female: a, habitus; b, carapace, dorsal view; c, epigyne, external view; d, coxae IV, ventral view; e, carapace, abdomen and legs, dorsal view; f, sternum, maxillae and labium, ventral view; g, abdomen, ventral view. Scale lines: 5 mm for a, e; 1 mm for b, f; d, f, g to same scale; 0.2 mm for c." httpUri="https://zenodo.org/record/186373/files/figure.png" pageId="31" pageNumber="32">
|
||
Fig.
|
||
<quantity id="381F9B37D667FF84FCC9FEC5FC52FECD" box="[869,902,306,332]" metricMagnitude="-2" metricUnit="kg" metricValue="2.2" pageId="31" pageNumber="32" unit="g" value="22.0">22</quantity>
|
||
</figureCitation>
|
||
g). Tarsi slightly curved. Female palpal tarsus tapered conical, unmodified; claw clearly evident.
|
||
<taxonomicName id="38E74D51D667FF84FD56FEAEFC8EFEF3" box="[762,858,345,370]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CD93EAC0D667FF84FD56FEAEFC8EFEF3" box="[762,858,345,370]" italics="true" pageId="31" pageNumber="32">Scopula</emphasis>
|
||
</taxonomicName>
|
||
: on male weak, thin on tarsi I, II, not obscuring cuticle reduced to distal group of setae on III, IV, weak, thin, lateral group of scattered hairs on metatarsi I, II, absent on III, IV and tibiae.
|
||
<taxonomicName id="38E74D51D667FF84FE72FE51FDE8FE41" box="[478,572,422,448]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="37" phylum="Arthropoda" rank="genus">Scopula</taxonomicName>
|
||
slightly denser in females than males, for
|
||
<emphasis id="CD93EAC0D667FF84FB84FE50FB97FE41" box="[1064,1091,423,448]" italics="true" pageId="31" pageNumber="32">ca</emphasis>
|
||
. 0.8 of metatarsal length, full length of tarsus. Trichobothria in 2 rows on tibiae, an irregular line on metatarsi, two bands on tarsi, distal most trichobothria on each segment very long. Trichobothrial base collariform (
|
||
<figureCitation id="67DC2A57D667FF84FBE8FE03FB49FD8F" box="[1092,1181,500,526]" captionStart="FIGURE 24" captionStartId="35.[151,257,1518,1542]" captionTargetBox="[151,1435,886,1494]" captionTargetId="figure@35.[151,1436,886,1494]" captionTargetPageId="35" captionText="FIGURE 24. Nuliodon fishburni sp. nov., female, scanning electron micrographs: a, tarsal trichobothria and cuticle, dorsal view; b, tarsal organ, dorsal view; c, claw tufts." httpUri="https://zenodo.org/record/186375/files/figure.png" pageId="31" pageNumber="32">Fig. 24</figureCitation>
|
||
a). Tarsal organ ovoid with ovoid aperture (
|
||
<figureCitation id="67DC2A57D667FF84FE0DFDEDFDD4FDB5" box="[417,512,538,564]" captionStart="FIGURE 24" captionStartId="35.[151,257,1518,1542]" captionTargetBox="[151,1435,886,1494]" captionTargetId="figure@35.[151,1436,886,1494]" captionTargetPageId="35" captionText="FIGURE 24. Nuliodon fishburni sp. nov., female, scanning electron micrographs: a, tarsal trichobothria and cuticle, dorsal view; b, tarsal organ, dorsal view; c, claw tufts." httpUri="https://zenodo.org/record/186375/files/figure.png" pageId="31" pageNumber="32">Fig. 24</figureCitation>
|
||
b).
|
||
<emphasis id="CD93EAC0D667FF84FD8BFDEDFDADFDB2" box="[551,633,538,563]" italics="true" pageId="31" pageNumber="32">Spines</emphasis>
|
||
. Male (female): proventral spine on femora I, II weak; patellae aspinose; tibiae I, II with 3 pairs of weak spines (only 3–4 thicker bristles) ventrally, much weaker than on metatarsi; 2 (1) pairs on ventral metatarsi I, II. Female palp: fe p1; pa d1 elongate; ti p3d2 all strong; ta p3d1r
|
||
<date id="8B591012D667FF84FA21FD9FFF72FD29" pageId="31" pageNumber="32">2 v</date>
|
||
2 subdistally. Preening combs absent.
|
||
<emphasis id="CD93EAC0D667FF84FDD5FD78FD11FD29" box="[633,709,655,680]" italics="true" pageId="31" pageNumber="32">Claws</emphasis>
|
||
. 2 claws with 3–6 teeth; claw tufts dense, fused, as high as claws but not obscuring teeth. Female palpal claw with 3–4 teeth.
|
||
<emphasis id="CD93EAC0D667FF84FC7EFD42FB87FD4F" box="[978,1107,693,718]" italics="true" pageId="31" pageNumber="32">Spinnerets</emphasis>
|
||
(
|
||
<figureCitation id="67DC2A57D667FF84FBC9FD42FB17FD4E" box="[1125,1219,693,719]" captionStart="FIGURE 22" captionStartId="33.[151,257,1302,1326]" captionTargetBox="[151,1435,197,1276]" captionTargetId="figure@33.[151,1436,197,1277]" captionTargetPageId="33" captionText="FIGURE 22. Nuliodon fishburni sp. nov., female: a, habitus; b, carapace, dorsal view; c, epigyne, external view; d, coxae IV, ventral view; e, carapace, abdomen and legs, dorsal view; f, sternum, maxillae and labium, ventral view; g, abdomen, ventral view. Scale lines: 5 mm for a, e; 1 mm for b, f; d, f, g to same scale; 0.2 mm for c." httpUri="https://zenodo.org/record/186373/files/figure.png" pageId="31" pageNumber="32">
|
||
Fig.
|
||
<quantity id="381F9B37D667FF84FB0CFD42FB17FD4E" box="[1184,1219,693,719]" metricMagnitude="-2" metricUnit="kg" metricValue="2.2" pageId="31" pageNumber="32" unit="g" value="22.0">22</quantity>
|
||
</figureCitation>
|
||
g). Colulus small. Female: ALS
|
||
<emphasis id="CD93EAC0D667FF84FE90FD2BFE8CFD74" box="[316,344,732,757]" italics="true" pageId="31" pageNumber="32">ca</emphasis>
|
||
.
|
||
<date id="8B591012D667FF84FECBFD2BFE51FD77" box="[359,389,732,758]" pageId="31" pageNumber="32">2x</date>
|
||
diameter of PLS and longer, PLS slightly longer than PMS with conical apical segment; PMS long, cylindrical with no enlarged spigots on tip or dorsally.
|
||
<emphasis id="CD93EAC0D667FF84FC66FCF5FB9EFC9A" box="[970,1098,770,795]" italics="true" pageId="31" pageNumber="32">Male Palp</emphasis>
|
||
(
|
||
<figureCitation id="67DC2A57D667FF84FBF0FCF5FB6CFC9D" box="[1116,1208,770,796]" captionStart="FIGURE 21" captionStartId="32.[151,255,1933,1957]" captionTargetBox="[151,1436,429,1906]" captionTargetId="figure@32.[151,1436,429,1906]" captionTargetPageId="32" captionText="FIGURE 21. Nuliodon fishburni sp. nov., male: a, palpal tibia, cymbium and bulb, ventral view; b, median apophysis, ventral view; c, d, RTA, ventral (c) and retrolateral view (d)." httpUri="https://zenodo.org/record/186372/files/figure.png" pageId="31" pageNumber="32">Fig. 21</figureCitation>
|
||
). Tibia short, with retrodistal RTA with inner keel and outer flange with unsclerotised crescent between them. Cymbium short with ovoid apical
|
||
<taxonomicName id="38E74D51D667FF84FEDCFCA7FE1FFCEB" box="[368,459,848,874]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="37" phylum="Arthropoda" rank="genus">scopula</taxonomicName>
|
||
and long thick recurved setae apically; apical cone wide, short, retromargin with wide flattened area with medial groove for basal third; light cage of long bristles on promargin (
|
||
<figureCitation id="67DC2A57D667FF84FA8AFC81FA54FC11" box="[1318,1408,886,912]" captionStart="FIGURE 21" captionStartId="32.[151,255,1933,1957]" captionTargetBox="[151,1436,429,1906]" captionTargetId="figure@32.[151,1436,429,1906]" captionTargetPageId="32" captionText="FIGURE 21. Nuliodon fishburni sp. nov., male: a, palpal tibia, cymbium and bulb, ventral view; b, median apophysis, ventral view; c, d, RTA, ventral (c) and retrolateral view (d)." httpUri="https://zenodo.org/record/186372/files/figure.png" pageId="31" pageNumber="32">Fig. 21</figureCitation>
|
||
a). Bulb: tegulum with several processes and lobes apart from prominent distal bifid median apophysis and short embolus originating on mid-prolateral edge; small conductor below median apophysis.
|
||
<emphasis id="CD93EAC0D667FF84FB0CFC33FAD4FC5C" box="[1184,1280,964,989]" italics="true" pageId="31" pageNumber="32">Epigyne</emphasis>
|
||
(
|
||
<figureCitation id="67DC2A57D667FF84FABCFC33FAB8FC5F" box="[1296,1388,964,990]" captionStart="FIGURE 22" captionStartId="33.[151,257,1302,1326]" captionTargetBox="[151,1435,197,1276]" captionTargetId="figure@33.[151,1436,197,1277]" captionTargetPageId="33" captionText="FIGURE 22. Nuliodon fishburni sp. nov., female: a, habitus; b, carapace, dorsal view; c, epigyne, external view; d, coxae IV, ventral view; e, carapace, abdomen and legs, dorsal view; f, sternum, maxillae and labium, ventral view; g, abdomen, ventral view. Scale lines: 5 mm for a, e; 1 mm for b, f; d, f, g to same scale; 0.2 mm for c." httpUri="https://zenodo.org/record/186373/files/figure.png" pageId="31" pageNumber="32">Figs 22</figureCitation>
|
||
c, g, 23a, b). Externally a pair of anterior crescentic ridges set diagonally; medially forming shallow trough posteriorly; vulva simple, copulatory fossae funnel-shaped with short twisted duct leading to enlarged distal spermatheca and fertilisation duct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7FD6559D667FF84FF69FBA9FDF3FB47" pageId="31" pageNumber="32" type="distribution">
|
||
<paragraph id="FF5836D2D667FF84FF69FBA9FDF3FB47" blockId="31.[151,1437,152,1415]" pageId="31" pageNumber="32">
|
||
<emphasis id="CD93EAC0D667FF84FF69FBA9FE2CFBF9" bold="true" box="[197,504,1118,1144]" pageId="31" pageNumber="32">Distribution and habitat</emphasis>
|
||
(
|
||
<figureCitation id="67DC2A57D667FF84FDA5FBA9FD8DFBF9" box="[521,601,1118,1144]" captionStart="FIGURE 25" captionStartId="36.[151,255,1809,1833]" captionTargetBox="[158,1430,1193,1764]" captionTargetId="figure@36.[151,1436,1164,1784]" captionTargetPageId="36" captionText="FIGURE 25. Distribution records of Miturgidae in Australia: a, Diaprograpta, Eupograpta, Mitzoruga and Nuliodon; b, all records of Miturgidae for Australia." httpUri="https://zenodo.org/record/186376/files/figure.png" pageId="31" pageNumber="32">Fig 25</figureCitation>
|
||
a, 26b). Known only from leaf litter in open eucalypt, wet sclerophyll, semi-evergreen vine thicket, Blackbutt close forest and heath mixed with eucalypt forest from Rockhampton to Ipswich, southeast Queensland.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7FD6559D667FF84FF69FB25FB56FA06" pageId="31" pageNumber="32" type="discussion">
|
||
<paragraph id="FF5836D2D667FF84FF69FB25FB56FA06" blockId="31.[151,1437,152,1415]" pageId="31" pageNumber="32">
|
||
<emphasis id="CD93EAC0D667FF84FF69FB25FEEFFB6D" bold="true" box="[197,315,1234,1260]" pageId="31" pageNumber="32">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="38E74D51D667FF84FEE0FB24FE6BFB6D" box="[332,447,1235,1260]" class="Arachnida" family="Miturgidae" genus="Nuliodon" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CD93EAC0D667FF84FEE0FB24FE6BFB6D" box="[332,447,1235,1260]" italics="true" pageId="31" pageNumber="32">Nuliodon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D6A057BBD667FF84FE6BFB24FDECFB6C" box="[455,568,1235,1261]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
includes the smallest miturgids known (total length about
|
||
<quantity id="381F9B37D667FF84FAB8FB24FAB5FB6C" box="[1300,1377,1235,1261]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="mm" value="5.0">5 mm</quantity>
|
||
); the spiders are common in open forest, but not rainforest, leaf litter. Perhaps as a result of limited spider surveys in the Brisbane area prior to 1990, records of these spiders prior to 1978 are lacking. Despite this, these spiders were taken very close to long-occupied suburbs and in bushland on the Griffith University campus. Hence, this new species has literally long been in the backyards of Brisbane dwellers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |