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<document id="37F2B1294BC69E3D511D6C90E065E268" ID-DOI="10.5281/zenodo.5169575" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169575" ID-ZooBank="89FC75EA-2324-4361-B818-FBA7B7682A00" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628278715662" checkinUser="felipe" docAuthor="Larochelle, Larivière, Marie-Claude, Larochelle &amp; Larivière" docDate="2017" docId="556787D30246FF863FCAFBD7FC7CFB07" docLanguage="en" docName="InsectaMundi.0594.1-110.pdf" docOrigin="Insecta Mundi 2017 (594)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Oopterus marrisi Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière 2017, new species" docType="treatment" docUuid="7D2D572A-BDEC-4180-B779-F35366EBDADB" docUuidSource="ZooBank" docVersion="5" lastPageNumber="25" masterDocId="A95EFFAB025FFF9C3F0AFFF6FFFFFF9D" masterDocTitle="Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae)" masterLastPageNumber="110" masterPageNumber="1" pageNumber="24" updateTime="1699039365375" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="A3F9B5D1002FA62AD5308A7B66BB13CD">Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="D23B4AA3E9500F0027C01A709A36C642">Larochelle</mods:namePart>
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<mods:namePart id="82509F9C6832D1EE2CCEB084C422DF6C">Larivière, Marie-Claude</mods:namePart>
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<mods:namePart id="BE3DEAAF9B1B97DC87A0567873DB0670">Larochelle</mods:namePart>
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<mods:namePart id="08F3F200151067D2AF70521A225396DB">Larivière</mods:namePart>
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<mods:date id="9F9B59225A818BD2567C223D515B23F4">2017</mods:date>
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<treatment id="556787D30246FF863FCAFBD7FC7CFB07" ID-DOI="http://doi.org/10.5281/zenodo.5186063" ID-Zenodo-Dep="5186063" LSID="urn:lsid:zoobank.org:act:7D2D572A-BDEC-4180-B779-F35366EBDADB" httpUri="http://treatment.plazi.org/id/556787D30246FF863FCAFBD7FC7CFB07" lastPageId="26" lastPageNumber="25" pageId="25" pageNumber="24">
<subSubSection id="95D4654E0246FF853FCAFBD7FDA6FBA4" box="[192,601,1057,1082]" pageId="25" pageNumber="24" type="nomenclature">
<paragraph id="DD7136C50246FF853FCAFBD7FDA6FBA4" blockId="25.[192,601,1057,1114]" box="[192,601,1057,1082]" pageId="25" pageNumber="24">
<heading id="863981A90246FF853FCAFBD7FDA6FBA4" bold="true" box="[192,601,1057,1082]" fontSize="10" level="2" pageId="25" pageNumber="24" reason="3">
<emphasis id="EFBAEAD70246FF853FCAFBD7FDA6FBA4" bold="true" box="[192,601,1057,1082]" pageId="25" pageNumber="24">
<taxonomicName id="1ACE4D460246FF853FCAFBD7FE53FBA4" ID-CoL="8LY2R" authority="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière, 2017" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[192,428,1057,1081]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="marrisi" status="sp. nov.">
<emphasis id="EFBAEAD70246FF853FCAFBD7FE53FBA4" bold="true" box="[192,428,1057,1081]" italics="true" pageId="25" pageNumber="24">Oopterus marrisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F48957AC0246FF853EBEFBD4FDA6FBA4" box="[436,601,1057,1082]" pageId="25" pageNumber="24" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95D4654E0246FF863FCAFBB4FBACFE24" lastPageId="26" lastPageNumber="25" pageId="25" pageNumber="24" type="description">
<paragraph id="DD7136C50246FF853FCAFBB4FE83FBC7" blockId="25.[192,601,1057,1114]" box="[192,380,1089,1114]" pageId="25" pageNumber="24">
<figureCitation id="45F52A400246FF853FCAFBB4FEE8FBC7" box="[192,279,1090,1114]" captionStart="Figures 2730" captionStartId="83.[192,288,1832,1854]" captionTargetBox="[202,1427,270,1814]" captionTargetId="figure-43@83.[192,1435,192,1817]" captionTargetPageId="83" captionText="Figures 2730. Dorsal habitus. 27) Oopterus discoideus new species. 28) O. marrisi new species. 29) O. lewisi. 30) O. patulus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5169587" httpUri="https://zenodo.org/record/5169587/files/figure.png" pageId="25" pageNumber="24">Fig. 28</figureCitation>
,
<figureCitation id="45F52A400246FF853E2EFBB7FEBBFBC4" box="[292,324,1089,1113]" captionStart="Figures 7481" captionStartId="94.[192,288,1832,1854]" captionTargetBox="[254,1414,252,1806]" captionTargetId="figure-59@94.[192,1440,192,1819]" captionTargetPageId="94" captionText="Figures 7481. Aedeagi, lateral. 74) Oopterus discoideus new species. 75) O. marrisi new species. 76) O. lewisi. 77) O. patulus. 78) O. fulvipes. 79) O. sobrinus. 80) O. collaris. 81) O. laeviventris." figureDoi="http://doi.org/10.5281/zenodo.5169615" httpUri="https://zenodo.org/record/5169615/files/figure.png" pageId="25" pageNumber="24">75</figureCitation>
,
<figureCitation id="45F52A400246FF853E5AFBB4FE83FBC7" box="[336,380,1090,1114]" captionStart="Figures 130135" captionStartId="103.[192,288,1829,1851]" captionTargetBox="[192,1412,192,1818]" captionTargetId="figure-48@103.[192,1412,192,1818]" captionTargetPageId="103" captionText="Figures 130135. Species distribution maps. 130) Oopterus laeviventris. 131) O. lewisi. 132) O. marrineri. 133) O. marrisi new species. 134) O. mohiensis new species. 135) O. monticola new species." figureDoi="http://doi.org/10.5281/zenodo.5169635" httpUri="https://zenodo.org/record/5169635/files/figure.png" pageId="25" pageNumber="24">133</figureCitation>
</paragraph>
<paragraph id="DD7136C50246FF853FCAFB74FBDBFB67" blockId="25.[192,1440,1154,1274]" pageId="25" pageNumber="24">
<materialsCitation id="6DA63C980246FF853FCAFB74FBCAFB47" collectionCode="NZAC, FD" country="New Zealand" elevation="701" location="NEW ZEALAND" pageId="25" pageNumber="24" specimenCount="701" typeStatus="holotype">
<typeStatus id="027588670246FF853FCAFB74FECDFB07" box="[192,306,1154,1178]" pageId="25" pageNumber="24" type="holotype">Holotype</typeStatus>
: male (
<collectionCode id="BBDFAE000246FF853E89FB74FE2DFB07" box="[387,466,1154,1178]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="25" pageNumber="24" type="Other institutional type">NZAC</collectionCode>
) labeled: “[
<collectingCountry id="A5D976550246FF853D5EFB74FCDBFB07" box="[596,804,1154,1178]" name="New Zealand" pageId="25" pageNumber="24">NEW ZEALAND</collectingCountry>
<collectionCode id="BBDFAE000246FF853C23FB74FCAEFB04" box="[809,849,1154,1177]" pageId="25" pageNumber="24">FD</collectionCode>
] Wilmot Pass.
<specimenCount id="CBC8FD4C0246FF853B09FB74FBB1FB07" box="[1027,1102,1154,1178]" pageId="25" pageNumber="24" type="generic">
<quantity id="1A369B200246FF853B09FB74FBB5FB07" box="[1027,1098,1154,1178]" metricMagnitude="2" metricUnit="m" metricValue="7.01" pageId="25" pageNumber="24" unit="m" value="701.0">
<elevation id="56E3D1F60246FF853B09FB74FBB5FB07" box="[1027,1098,1154,1178]" metricMagnitude="2" metricUnit="m" metricValue="7.01" pageId="25" pageNumber="24" unit="m" value="701.0">701m</elevation>
</quantity>
.
</specimenCount>
(hand-written) / Manapouri Exp. Jan 70 (typed) / at night (typed) on moss. (hand-written) / [male symbol] /
<typeStatus id="027588670246FF853BA7FB54FAB7FB27" box="[1197,1352,1186,1210]" pageId="25" pageNumber="24" type="holotype">HOLOTYPE</typeStatus>
[male symbol]
<taxonomicName id="1ACE4D460246FF853E2FFB34FCACFB47" authority="Larochelle &amp; Lariviere, 2017" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[293,851,1218,1242]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="marrisi">
<emphasis id="EFBAEAD70246FF853E2FFB34FE0BFB44" box="[293,500,1218,1241]" italics="true" pageId="25" pageNumber="24">Oopterus marrisi</emphasis>
Larochelle &amp; Larivière, 2017
</taxonomicName>
(red label; typed).”
</materialsCitation>
<typeStatus id="027588670246FF853B31FB34FB44FB47" box="[1083,1211,1218,1242]" pageId="25" pageNumber="24" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="CBC8FD4C0246FF853BCEFB34FAC2FB47" box="[1220,1341,1218,1242]" pageId="25" pageNumber="24" type="male">two males</specimenCount>
(LUNZ, NZAC) from Murchison Mountains (FD), bearing blue
<typeStatus id="027588670246FF853C55FB14FC32FB67" box="[863,973,1250,1274]" pageId="25" pageNumber="24" type="paratype">paratype</typeStatus>
labels.
</paragraph>
<paragraph id="DD7136C50246FF863FCAFAD7FBACFE24" blockId="25.[192,1440,1313,1914]" lastBlockId="26.[192,1441,193,442]" lastPageId="26" lastPageNumber="25" pageId="25" pageNumber="24">
<emphasis id="EFBAEAD70246FF853FCAFAD7FE9FFAA4" bold="true" box="[192,352,1313,1337]" pageId="25" pageNumber="24">Description</emphasis>
. Body length
<quantity id="1A369B200246FF853D08FAD7FD6DFAA4" box="[514,658,1313,1337]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.6" metricValueMin="5.7" pageId="25" pageNumber="24" unit="mm" value="6.15" valueMax="6.6" valueMin="5.7">5.76.6 mm</quantity>
. Head, pronotum, elytra, and abdomen mostly reddish black; base of pronotum reddish medially; lateral margins, suture, and extreme apex of elytra reddish; abdomen bright reddish laterally and apically; antennae, palpi, and legs reddish; apical half of femora yellow. Microsculpture absent. Iridescence absent. Very shiny, with metallic luster (aeneous) on pronotum and elytra.
<emphasis id="EFBAEAD70246FF853E1FFA54FEA2FA27" bold="true" box="[277,349,1442,1466]" pageId="25" pageNumber="24">Head</emphasis>
. Labrum moderately transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula.
<emphasis id="EFBAEAD70246FF853A3EF9D4FA67F9A7" bold="true" box="[1332,1432,1570,1594]" pageId="25" pageNumber="24">Thorax</emphasis>
. Pronotum moderately convex, finely punctate between laterobasal foveae, wrinkled discally, strongly transverse, widest about middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, slightly sinuate posteriorly; lateral grooves absent; setiferous punctures absent on each side; posterolateral angles subrectangular; posterolateral carinae obsolete; laterobasal foveae well defined, virtually impunctate, very deep and wide, oblong, prolonged forward, double; posterior bead absent; base emarginate, wider than pronotal apex, about as wide as elytral base.
<emphasis id="EFBAEAD70246FF853D63F8F4FD56F887" bold="true" box="[617,681,1794,1818]" pageId="25" pageNumber="24">Legs</emphasis>
. Moderately long.
<emphasis id="EFBAEAD70246FF853C8DF8F4FC21F887" bold="true" box="[903,990,1794,1818]" pageId="25" pageNumber="24">Elytra</emphasis>
. Strongly convex, ovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Striae 14 complete; striae 57 incomplete; stria 1 deep; striae 24 shallow, finely punctate; striae 57 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals subdepressed. Sutural apices rounded.
<emphasis id="EFBAEAD70245FF863FCAFF17FEBCFF64" bold="true" box="[192,323,225,249]" pageId="26" pageNumber="25">Abdomen</emphasis>
. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae.
<emphasis id="EFBAEAD70245FF863D38FEF7FD48FE84" bold="true" box="[562,695,257,281]" pageId="26" pageNumber="25">Aedeagus</emphasis>
. Lateral view (
<figureCitation id="45F52A400245FF863C7EFEF4FC32FE84" box="[884,973,257,282]" captionStart="Figures 7481" captionStartId="94.[192,288,1832,1854]" captionTargetBox="[254,1414,252,1806]" captionTargetId="figure-59@94.[192,1440,192,1819]" captionTargetPageId="94" captionText="Figures 7481. Aedeagi, lateral. 74) Oopterus discoideus new species. 75) O. marrisi new species. 76) O. lewisi. 77) O. patulus. 78) O. fulvipes. 79) O. sobrinus. 80) O. collaris. 81) O. laeviventris." figureDoi="http://doi.org/10.5281/zenodo.5169615" httpUri="https://zenodo.org/record/5169615/files/figure.png" pageId="26" pageNumber="25">Fig. 75</figureCitation>
): moderately arcuate, widened in apical half; base slightly convex dorsally; middle slightly convex dorsally, strongly convex ventrally, with dorsal membranous area moderately wide and very long; apex subtriangular, slightly convex dorsally, straight ventrally, with extreme tip narrow and rather long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with five to seven apical setae.
</paragraph>
</subSubSection>
<subSubSection id="95D4654E0245FF863FCAFE17FBB3FDC7" pageId="26" pageNumber="25" type="materials_examined">
<paragraph id="DD7136C50245FF863FCAFE17FCCCFE67" blockId="26.[192,819,481,506]" box="[192,819,481,506]" pageId="26" pageNumber="25">
<emphasis id="EFBAEAD70245FF863FCAFE17FE3AFE64" bold="true" box="[192,453,481,505]" pageId="26" pageNumber="25">Material examined</emphasis>
.
<specimenCount id="CBC8FD4C0245FF863EDEFE17FD96FE67" box="[468,617,481,506]" pageId="26" pageNumber="25" type="generic">6 specimens</specimenCount>
(
<collectionCode id="BBDFAE000245FF863D72FE14FD35FE67" box="[632,714,482,506]" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="26" pageNumber="25">LUNZ</collectionCode>
,
<collectionCode id="BBDFAE000245FF863DDCFE14FCD7FE67" box="[726,808,482,506]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="26" pageNumber="25" type="Other institutional type">NZAC</collectionCode>
).
</paragraph>
<paragraph id="DD7136C50245FF863FCAFDD7FBB3FDC7" blockId="26.[192,1440,545,602]" pageId="26" pageNumber="25">
<emphasis id="EFBAEAD70245FF863FCAFDD7FDEAFDA4" bold="true" box="[192,533,545,569]" pageId="26" pageNumber="25">Geographic distribution</emphasis>
(
<figureCitation id="45F52A400245FF863D22FDD4FD6BFDA7" box="[552,660,546,570]" captionStart="Figures 130135" captionStartId="103.[192,288,1829,1851]" captionTargetBox="[192,1412,192,1818]" captionTargetId="figure-48@103.[192,1412,192,1818]" captionTargetPageId="103" captionText="Figures 130135. Species distribution maps. 130) Oopterus laeviventris. 131) O. lewisi. 132) O. marrineri. 133) O. marrisi new species. 134) O. mohiensis new species. 135) O. monticola new species." figureDoi="http://doi.org/10.5281/zenodo.5169635" httpUri="https://zenodo.org/record/5169635/files/figure.png" pageId="26" pageNumber="25">Fig. 133</figureCitation>
). South Island:
<collectionCode id="BBDFAE000245FF863C6BFDD4FC77FDA7" box="[865,904,546,570]" pageId="26" pageNumber="25">FD</collectionCode>
Fiordland National Park (Borland Valley Bivouac (South of); McKenzie Burn; Murchison Mountains; Wilmot Pass).
</paragraph>
</subSubSection>
<subSubSection id="95D4654E0245FF863FCAFD74FC17FC07" pageId="26" pageNumber="25" type="biology_ecology">
<paragraph id="DD7136C50245FF863FCAFD74FC3CFD27" blockId="26.[192,1440,642,698]" pageId="26" pageNumber="25">
<emphasis id="EFBAEAD70245FF863FCAFD74FED1FD07" bold="true" box="[192,302,642,666]" pageId="26" pageNumber="25">Ecology</emphasis>
. Montane. Endogean. Wet beech forests. Shaded ground. Nocturnal; active at night on moss; hides during the day in thick moss, leaf litter, and under bark.
</paragraph>
<paragraph id="DD7136C50245FF863FCAFD17FE9EFC87" blockId="26.[192,1440,737,794]" pageId="26" pageNumber="25">
<emphasis id="EFBAEAD70245FF863FCAFD17FED5FD64" bold="true" box="[192,298,737,761]" pageId="26" pageNumber="25">Biology</emphasis>
. Seasonality: NovemberJanuary. Tenerals: November. Predacious (based on mouthpart morphology).
</paragraph>
<paragraph id="DD7136C50245FF863FCAFCB7FBD3FCC4" blockId="26.[192,1068,833,858]" box="[192,1068,833,858]" pageId="26" pageNumber="25">
<emphasis id="EFBAEAD70245FF863FCAFCB7FE5DFCC7" bold="true" box="[192,418,833,858]" pageId="26" pageNumber="25">Dispersal power</emphasis>
. Subapterous (incapable of flight). Moderate runner.
</paragraph>
<paragraph id="DD7136C50245FF863FCAFC77FC17FC07" blockId="26.[192,1000,897,922]" box="[192,1000,897,922]" pageId="26" pageNumber="25">
<emphasis id="EFBAEAD70245FF863FCAFC77FE12FC04" bold="true" box="[192,493,897,921]" pageId="26" pageNumber="25">Collecting techniques</emphasis>
. Sifting moss and leaf litter; lifting bark.
</paragraph>
</subSubSection>
<subSubSection id="95D4654E0245FF863FCAFC34FC7CFB07" pageId="26" pageNumber="25" type="discussion">
<paragraph id="DD7136C50245FF863FCAFC34FC7CFB07" blockId="26.[192,1441,962,1178]" pageId="26" pageNumber="25">
<emphasis id="EFBAEAD70245FF863FCAFC34FEC3FC47" bold="true" box="[192,316,962,986]" pageId="26" pageNumber="25">Remarks</emphasis>
. This species is named after our friend and colleague John M. W. Marris (Lincoln University, Lincoln, Christchurch) for his special help and encouragement in our entomological studies.
<taxonomicName id="1ACE4D460245FF863A3BFC14FEE3FB84" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="marrisi">
<emphasis id="EFBAEAD70245FF863A3BFC14FEE3FB84" italics="true" pageId="26" pageNumber="25">Oopterus marrisi</emphasis>
</taxonomicName>
is morphologically close to
<taxonomicName id="1ACE4D460245FF863D60FBF4FCF3FB87" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[618,780,1026,1050]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="discoideus">
<emphasis id="EFBAEAD70245FF863D60FBF4FCF3FB87" box="[618,780,1026,1050]" italics="true" pageId="26" pageNumber="25">O. discoideus</emphasis>
</taxonomicName>
. In addition to diagnostic characters of the male genitalia,
<taxonomicName id="1ACE4D460245FF863E02FBD4FE76FBA4" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[264,393,1058,1081]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="marrisi">
<emphasis id="EFBAEAD70245FF863E02FBD4FE76FBA4" box="[264,393,1058,1081]" italics="true" pageId="26" pageNumber="25">O. marrisi</emphasis>
</taxonomicName>
has the following distinguishing features: body length
<quantity id="1A369B200245FF863B3FFBD7FB37FBA4" box="[1077,1224,1057,1081]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.6" metricValueMin="5.7" pageId="26" pageNumber="25" unit="mm" value="6.15" valueMax="6.6" valueMin="5.7">5.76.6 mm</quantity>
; head, pronotum, and elytra mostly reddish black; eyes moderately convex; tempora not inflated; pronotum moderately convex, sides moderately rounded, slightly sinuate posteriorly, laterobasal foveae oblong. Both species are known only from the Fiordland region (South Island).
</paragraph>
</subSubSection>
</treatment>
</document>