311 lines
36 KiB
XML
311 lines
36 KiB
XML
<document id="AE076B939C345246161C55FBC0A4297D" ID-DOI="10.11646/phytotaxa.348.2.7" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725593459450" checkinUser="felipe" docAuthor="Szeląg, Zbigniew" docDate="2018" docId="5507123A816CFFABFF5A36A5FDC0D8E2" docLanguage="en" docName="phytotaxa.348.2.7.pdf" docOrigin="Phytotaxa 348 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.348.2.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Hieracium gussevii Szelag 2018, sp. nov." docType="treatment" docVersion="1" lastPageNumber="126" masterDocId="A93E6A42816CFFAAFFD23151FFDDDF78" masterDocTitle="Hieracia balcanica XIV. Two new species in Hieracium sect. Cernua (Asteraceae) from Bulgaria" masterLastPageNumber="132" masterPageNumber="125" pageNumber="125" updateTime="1725602298533" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="12647A2F129E00CC6D4AA2B1D0FC6C04">Hieracia balcanica XIV. Two new species in Hieracium sect. Cernua (Asteraceae) from Bulgaria</mods:title>
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</mods:titleInfo>
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<mods:name id="F129E7E79468C7848D2B620700BF1029" type="personal">
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<mods:role id="AF25BDEB37E19948D3A8F07F82EA6798">
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<mods:roleTerm id="19DFD8316687F81941C7B42D17754ACD">Author</mods:roleTerm>
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<mods:namePart id="3E0E9965D9B3EA4DD72E723D5A1CB8D0">Szeląg, Zbigniew</mods:namePart>
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<mods:typeOfResource id="30B90FCA3F917C786700A8C702435EF3">text</mods:typeOfResource>
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<mods:title id="44923596A8F7DB26BB27DC6B1577BE68">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1EDAFB7DFEDA67C5E6A66080CD15E613">
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<mods:date id="766E4ED5D957967F8C7BAF7D257333ED">2018</mods:date>
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<mods:detail id="0B24AF5486898E61D951AC436FDD666D" type="pubDate">
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<mods:number id="D7B3ADB93A8B76F38118989BCA34086C">2018-04-27</mods:number>
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<mods:number id="3ECEC5085C7EB5635EDB0168A51477D8">348</mods:number>
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<mods:number id="31F0310CF610347D54952087E421C195">2</mods:number>
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<mods:start id="986DB2263C6E0312AB20D88A6317F6FB">125</mods:start>
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<mods:location id="BB3B31B4183F62E113C572784263A180">
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<mods:url id="31C64EEA3FF8694B1BA9EB4B263F2C2C">http://dx.doi.org/10.11646/phytotaxa.348.2.7</mods:url>
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<mods:classification id="A5940E4E318E58D5B51E40DF7D2295A0">journal article</mods:classification>
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<mods:identifier id="D9C5687495680935FED1978B1CF1447A" type="DOI">10.11646/phytotaxa.348.2.7</mods:identifier>
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<mods:identifier id="36C22C530B760BB957427884472C8776" type="ISSN">1179-3163</mods:identifier>
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</mods:mods>
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<treatment id="5507123A816CFFABFF5A36A5FDC0D8E2" LSID="urn:lsid:plazi:treatment:5507123A816CFFABFF5A36A5FDC0D8E2" httpUri="http://treatment.plazi.org/id/5507123A816CFFABFF5A36A5FDC0D8E2" lastPageId="1" lastPageNumber="126" pageId="0" pageNumber="125">
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<subSubSection id="95B4F0A7816CFFAAFF5A36A5FD7ED777" box="[136,675,2036,2063]" pageId="0" pageNumber="125" type="nomenclature">
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<paragraph id="DD11A32C816CFFAAFF5A36A5FD7ED777" blockId="0.[136,675,2036,2063]" box="[136,675,2036,2063]" pageId="0" pageNumber="125">
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<heading id="86591440816CFFAAFF5A36A5FD7ED777" box="[136,675,2036,2063]" fontSize="11" level="2" pageId="0" pageNumber="125" reason="3">
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<taxonomicName id="1AAED8AF816CFFAAFF5A36A5FE66D777" ID-CoL="3LPBQ" authority="Szelag" authorityName="Szelag" authorityYear="2018" box="[136,443,2036,2063]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="0" pageNumber="125" phylum="Tracheophyta" rank="species" species="gussevii" status="sp. nov.">
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<emphasis id="EFDA7F3E816CFFAAFF5A36A5FEB5D776" bold="true" box="[136,360,2036,2062]" italics="true" pageId="0" pageNumber="125">Hieracium gussevii</emphasis>
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Szeląg
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</taxonomicName>
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,
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<emphasis id="EFDA7F3E816CFFAAFE1536A7FDFDD777" bold="true" box="[455,544,2038,2063]" pageId="0" pageNumber="125">
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<taxonomicNameLabel id="F4E9C245816CFFAAFE1536A7FDFDD777" box="[455,544,2038,2063]" pageId="0" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
(
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<figureCitation id="4595BFA9816CFFAAFDE236A7FD47D777" box="[560,666,2038,2063]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[136,229,1953,1974]" captionStartId-1="3.[136,229,1184,1205]" captionTargetBox-1="[151,1436,190,1159]" captionTargetId-1="figure-14@3.[151,1437,190,1159]" captionTargetPageId-1="3" captionText-0="FIGURE 1. Holotype of Hieracium gussevii (SOM 175982)." captionText-1="FIGURE 2. Holotype of Hieracium gussevii: involucres (left) and indumentum of the lower part of the stem (right)." pageId="0" pageNumber="125">Figs. 1‒2</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="95B4F0A7816DFFABFF5A31F1FA77DF85" pageId="1" pageNumber="126" type="materials_examined">
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<paragraph id="DD11A32C816DFFABFF5A31F1FA77DF85" blockId="1.[136,1453,160,253]" pageId="1" pageNumber="126">
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<emphasis id="EFDA7F3E816DFFABFF5A31F1FF1FDFCD" bold="true" box="[136,194,160,181]" pageId="1" pageNumber="126">Type:</emphasis>
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―
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<materialsCitation id="6DC6A971816DFFABFF0831F1FA7BDF85" collectingDate="2008-07-20" collectingDateMax="2017-07-20" collectingDateMin="2008-07-20" collectionCode="Z, KRAM" collectorName="Vitosha Mountains, SE" country="Bulgaria" county="Szelag" elevation="1800" location="Specimens" municipality="Aleko" pageId="1" pageNumber="126" specimenCode="SOM 175982, SOM 175983" specimenCount="2" typeStatus="holotype">
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<location id="D871F5F7816DFFABFF0831F1FE9FDFCD" LSID="urn:lsid:plazi:treatment:5507123A816CFFABFF5A36A5FDC0D8E2:D871F5F7816DFFABFF0831F1FE9FDFCD" box="[218,322,160,181]" country="Bulgaria" county="Szelag" municipality="Aleko" name="Specimens" pageId="1" pageNumber="126">Specimens</location>
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from plants cultivated in the author’s garden, pressed on
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<date id="A91085EC816DFFABFC5031F1FBD7DFCD" box="[898,1034,160,181]" pageId="1" pageNumber="126" value="2017-07-20">
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<collectingDate id="B9547C04816DFFABFC5031F1FBD7DFCD" box="[898,1034,160,181]" pageId="1" pageNumber="126" value="2017-07-20">July 20, 2017</collectingDate>
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</date>
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, from the seeds collected in
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<collectingCountry id="A5B9E3BC816DFFABFAF831F1FA74DFCD" box="[1322,1449,160,181]" name="Bulgaria" pageId="1" pageNumber="126">BULGARIA</collectingCountry>
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.
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<collectorName id="705BC6FA816DFFABFF6F3195FE46DFA1" box="[189,411,196,217]" pageId="1" pageNumber="126">
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<location id="D871F5F7816DFFABFF6F3195FEA8DFA1" LSID="urn:lsid:plazi:treatment:5507123A816CFFABFF5A36A5FDC0D8E2:D871F5F7816DFFABFF6F3195FEA8DFA1" box="[189,373,196,217]" country="Bulgaria" county="Szelag" municipality="Aleko" name="Vitosha Mountains" pageId="1" pageNumber="126">Vitosha Mountains</location>
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, SE
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</collectorName>
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slope of
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<location id="D871F5F7816DFFABFE213195FDA1DFA1" LSID="urn:lsid:plazi:treatment:5507123A816CFFABFF5A36A5FDC0D8E2:D871F5F7816DFFABFE213195FDA1DFA1" box="[499,636,196,217]" country="Bulgaria" county="Szelag" municipality="Aleko" name="Mt. Goli Vrah" pageId="1" pageNumber="126">Mt. Goli Vrah</location>
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, open places in
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<taxonomicName id="1AAED8AF816DFFABFCC63195FC59DFA0" box="[788,900,195,217]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="abies">
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<emphasis id="EFDA7F3E816DFFABFCC63195FC59DFA0" box="[788,900,195,217]" italics="true" pageId="1" pageNumber="126">Picea abies</emphasis>
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</taxonomicName>
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forest, along tourist route leading to the
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<collectingMunicipality id="3D753956816DFFABFADA3195FA9EDFA1" box="[1288,1347,196,217]" pageId="1" pageNumber="126">Aleko</collectingMunicipality>
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ski centre,
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<elevation id="5683441F816DFFABFF6F31B9FEE7DF85" box="[189,314,232,253]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="126" unit="m" value="1800.0">
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<quantity id="1A560EC9816DFFABFF6F31B9FEDBDF85" box="[189,262,232,253]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="126" unit="m" value="1800.0">1800 m</quantity>
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a.s.l.
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</elevation>
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,
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<date id="A91085EC816DFFABFE9731B9FE1EDF85" box="[325,451,232,253]" pageId="1" pageNumber="126" value="2008-07-20">
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<collectingDate id="B9547C04816DFFABFE9731B9FE1EDF85" box="[325,451,232,253]" pageId="1" pageNumber="126" value="2008-07-20">20 July 2008</collectingDate>
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</date>
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,
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<emphasis id="EFDA7F3E816DFFABFE1F31B9FDF8DF85" box="[461,549,231,253]" italics="true" pageId="1" pageNumber="126">
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<collectionCode id="BBBF3BE9816DFFABFE1F31B9FE00DF85" box="[461,477,232,253]" country="Switzerland" name="Universität Zürich" pageId="1" pageNumber="126" type="Herbarium">Z</collectionCode>
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.
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<collectingCounty id="3470DBA0816DFFABFE3431B6FDF8DF85" box="[486,549,231,253]" pageId="1" pageNumber="126">Szeląg</collectingCounty>
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</emphasis>
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(
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<typeStatus id="02151D8E816DFFABFDE331B9FD5ADF85" box="[561,647,232,253]" pageId="1" pageNumber="126" type="holotype">holotype</typeStatus>
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<specimenCode id="8D080B57816DFFABFD5F31B9FCCFDF85" box="[653,786,232,253]" collectionCode="SOM" country="Bulgaria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12731" lsid="urn:lsid:biocol.org:col:12731" name="Bulgarian Academy of Sciences" pageId="1" pageNumber="126" type="Herbarium">SOM 175982</specimenCode>
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;
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<typeStatus id="02151D8E816DFFABFCC931B9FCB7DF85" box="[795,874,232,253]" pageId="1" pageNumber="126" type="isotype">isotypes</typeStatus>
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<specimenCode id="8D080B57816DFFABFCA231B9FC2FDF85" box="[880,1010,232,253]" collectionCode="SOM" country="Bulgaria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12731" lsid="urn:lsid:biocol.org:col:12731" name="Bulgarian Academy of Sciences" pageId="1" pageNumber="126" type="Herbarium">SOM 175983</specimenCode>
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& 175984,
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<collectionCode id="BBBF3BE9816DFFABFBB731B9FB72DF85" box="[1125,1199,232,253]" country="Poland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15714" lsid="urn:lsid:biocol.org:col:15714" name="Polish Academy of Sciences" pageId="1" pageNumber="126" type="Herbarium">KRAM</collectionCode>
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, Herb. Hierac.
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<collectionCode id="BBBF3BE9816DFFABFA9331B9FA8FDF85" box="[1345,1362,232,253]" country="Switzerland" name="Universität Zürich" pageId="1" pageNumber="126" type="Herbarium">Z</collectionCode>
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. Szeląg)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="95B4F0A7816DFFABFF5A307CFC71D94A" pageId="1" pageNumber="126" type="description">
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<paragraph id="DD11A32C816DFFABFF5A307CFE2ADCD2" blockId="1.[136,1452,301,1946]" pageId="1" pageNumber="126">
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<emphasis id="EFDA7F3E816DFFABFF5A307CFEC1DE3E" bold="true" box="[136,284,301,326]" pageId="1" pageNumber="126">Description:</emphasis>
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―Aphyllopodous. Stem
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<quantity id="1A560EC9816DFFABFDE4307CFD7BDE3E" box="[566,678,301,326]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="1" pageNumber="126" unit="cm" value="70.0" valueMax="90.0" valueMin="50.0">50–90 cm</quantity>
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high, purplish at base, in lower half with dense, stiff, pale,
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<quantity id="1A560EC9816DFFABFA9C307CFA71DE3E" box="[1358,1452,301,326]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="1" pageNumber="126" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
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long simple hairs, and few stellate hairs; in upper half with sparse, pale,
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<quantity id="1A560EC9816DFFABFC663000FBCCDE12" box="[948,1041,337,362]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="126" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
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long simple hairs and sparse stellate hairs; within synflorescence with numerous stellate hairs. Synflorescence branches
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<quantity id="1A560EC9816DFFABFB9F3024FB7BDEF6" box="[1101,1190,373,398]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="3.81" metricValueMin="1.27" pageId="1" pageNumber="126" unit="in" value="10.0" valueMax="15.0" valueMin="5.0">5–15 in</quantity>
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axils of upper cauline leaves, up to
|
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<quantity id="1A560EC9816DFFABFECE30C8FEB2DECA" box="[284,367,409,434]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="1" pageNumber="126" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 cm</quantity>
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||
long, with 2–4 capitula (and usually 1–3 capitula aborted), covered by numerous to rather dense stellate hairs. Acladium
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<quantity id="1A560EC9816DFFABFE4A30ECFE31DEAE" box="[408,492,445,470]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="126" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
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long. Rosette leaves (withering at anthesis) 8–12, up to
|
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<quantity id="1A560EC9816DFFABFBB530ECFB70DEAE" box="[1127,1197,445,470]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="126" unit="cm" value="15.0">15 cm</quantity>
|
||
long and up to
|
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<quantity id="1A560EC9816DFFABFA8D30ECFA76DEAE" box="[1375,1451,445,470]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="126" unit="cm" value="1.5">1.5 cm</quantity>
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wide, in shape and indumentum similar to lowest cauline leaves. Cauline leaves 15–30, somewhat coriaceous and glaucescent, sessile, ±semi-amplexicaul, gradually reduced in size upwards, lanceolate, acute at apex, sharply, remotely denticulate, with ±reticulate and protruding veins on lower surface. Lower cauline leaves up to
|
||
<quantity id="1A560EC9816DFFABFB7A3378FB31DD3A" box="[1192,1260,553,578]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="126" unit="cm" value="18.0">18 cm</quantity>
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||
long, at base and along midrib with dense, stiff, pale,
|
||
<quantity id="1A560EC9816DFFABFDCE331CFDA4DD1E" box="[540,633,589,614]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="1" pageNumber="126" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
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||
long simple hairs, on margins with numerous
|
||
<quantity id="1A560EC9816DFFABFB52331CFB00DD1E" box="[1152,1245,589,614]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="126" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||
long, simple hairs mixed with a few microglands and sparse stellate hairs; on lower surface with ±numerous stellate hairs and scattered simple hairs. Upper cauline leaves on both surfaces glabrous or almost so, on margins and along midrib with a few
|
||
<quantity id="1A560EC9816DFFABFF5A33E8FF3BDDAA" box="[136,230,697,722]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="126" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
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||
long, simple hairs, sparse stellate hairs and ±numerous microglands. Synflorescence with 20–40(–50), erect capitula. Peduncles thin, green, with dense stellate hairs. Bracteoles 2–4, green with scattered stellate hairs. Involucres
|
||
<quantity id="1A560EC9816DFFABFF5A3250FF28DC62" box="[136,245,769,794]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="1" pageNumber="126" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
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||
long, campanulate, with very sparse indumentum. Involucral bracts in two rows, obtuse at apex, up to
|
||
<quantity id="1A560EC9816DFFABFA4C3250FF69DC46" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="126" unit="mm" value="2.0">2 mm</quantity>
|
||
wide at base, dark green (in nature to blackish green), with a few, patent, dark glandular hairs
|
||
<quantity id="1A560EC9816DFFABFB013274FA84DC46" box="[1235,1369,805,830]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="1" pageNumber="126" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.2–0.4 mm</quantity>
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||
long, a few stellate hairs, and a tuft at apex. Ligules yellow, glabrous at apex. Styles yellow with dense, dark microtrichomes. Achenes dark brown to blackish, 3.7–4.0 mm long. Pappus pale-grey. Pollen in anthers numerous, spherical and of varying size. Flowering: August.
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||
</paragraph>
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<paragraph id="DD11A32C816DFFABFF6F32E4FDF1DB26" blockId="1.[136,1452,301,1946]" pageId="1" pageNumber="126">
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||
<emphasis id="EFDA7F3E816DFFABFF6F32E4FEF9DCB6" bold="true" box="[189,292,949,974]" pageId="1" pageNumber="126">Affinity:</emphasis>
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―
|
||
<taxonomicName id="1AAED8AF816DFFABFE9232E5FDC6DCB5" authorityName="Szelag" authorityYear="2018" box="[320,539,948,973]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="gussevii">
|
||
<emphasis id="EFDA7F3E816DFFABFE9232E5FDC6DCB5" box="[320,539,948,973]" italics="true" pageId="1" pageNumber="126">Hieracium gussevii</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="1AAED8AF816DFFABFD6432E5FCECDCB5" authorityName="Szelag" authorityYear="2006" box="[694,817,948,973]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="werneri">
|
||
<emphasis id="EFDA7F3E816DFFABFD6432E5FCECDCB5" box="[694,817,948,973]" italics="true" pageId="1" pageNumber="126">H. werneri</emphasis>
|
||
</taxonomicName>
|
||
from the Rhodopes in southern
|
||
<collectingCountry id="A5B9E3BC816DFFABFB7532E4FAD7DCB6" box="[1191,1290,949,974]" name="Bulgaria" pageId="1" pageNumber="126">Bulgaria</collectingCountry>
|
||
, but differs in its dense, simple hairs of the lower half of stem and the lower-cauline leaves, more numerous and glaucous cauline leaves, numerous stellate hairs on the stem within synflorescence, peduncles without microglands, involucral bracts with glandular hairs, and longer, dark brown to blackish achenes. The morphological differences between both species are stable in garden-cultivated plants.
|
||
</paragraph>
|
||
<paragraph id="DD11A32C816DFFABFF6F3538FA76DB96" blockId="1.[136,1452,301,1946]" pageId="1" pageNumber="126">
|
||
The following morphologic features of
|
||
<taxonomicName id="1AAED8AF816DFFABFDAE3539FD24DBF9" authorityName="Szelag" authorityYear="2018" box="[636,761,1128,1153]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="gussevii">
|
||
<emphasis id="EFDA7F3E816DFFABFDAE3539FD24DBF9" box="[636,761,1128,1153]" italics="true" pageId="1" pageNumber="126">H. gussevii</emphasis>
|
||
</taxonomicName>
|
||
seem to suggest a relationship with
|
||
<taxonomicName id="1AAED8AF816DFFABFB443539FECFDBDE" authority="Linnaeus (1753: 804)" authorityName="Linnaeus" authorityPageNumber="804" authorityYear="1753" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="umbellatum">
|
||
<emphasis id="EFDA7F3E816DFFABFB443539FAE0DBF9" box="[1174,1341,1127,1153]" italics="true" pageId="1" pageNumber="126">H. umbellatum</emphasis>
|
||
<bibRefCitation id="B93FDEDD816DFFABFA973538FECFDBDE" author="Linnaeus, C." pageId="1" pageNumber="126" refId="ref3236" refString="Linnaeus, C. (1753) Species plantarum. L. Salvius, Holmia, 1200 pp. http: // dx. doi. org / 10.5962 / bhl. title. 669" type="book" year="1753">Linnaeus (1753: 804)</bibRefCitation>
|
||
</taxonomicName>
|
||
: (
|
||
<emphasis id="EFDA7F3E816DFFABFEFB35DDFEEADBDE" box="[297,311,1164,1190]" italics="true" pageId="1" pageNumber="126">1</emphasis>
|
||
) the lack of the rosette leaves by anthesis, (
|
||
<emphasis id="EFDA7F3E816DFFABFCEB35DDFC9ADBDE" box="[825,839,1164,1190]" italics="true" pageId="1" pageNumber="126">2</emphasis>
|
||
) the unbranched stem up to
|
||
<quantity id="1A560EC9816DFFABFB4435DCFB03DBDE" box="[1174,1246,1165,1190]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="126" unit="cm" value="90.0">90 cm</quantity>
|
||
high, with 20–30 cauline leaves, (
|
||
<emphasis id="EFDA7F3E816DFFABFEEE35E1FE97DBB2" box="[316,330,1200,1226]" italics="true" pageId="1" pageNumber="126">3</emphasis>
|
||
) short synflorescence branches confined to the upper part of the stem, (
|
||
<emphasis id="EFDA7F3E816DFFABFBB535E1FBA8DBB2" box="[1127,1141,1200,1226]" italics="true" pageId="1" pageNumber="126">4</emphasis>
|
||
) few, patent glandular hairs on the involucral bracts, and (
|
||
<emphasis id="EFDA7F3E816DFFABFE043585FE39DB96" box="[470,484,1236,1262]" italics="true" pageId="1" pageNumber="126">5</emphasis>
|
||
) numerous stellate hairs in the upper part of the stem and especially on the peduncles.
|
||
</paragraph>
|
||
<paragraph id="DD11A32C816DFFABFF6F35A8FCD7DA22" blockId="1.[136,1452,301,1946]" pageId="1" pageNumber="126">
|
||
However, (
|
||
<emphasis id="EFDA7F3E816DFFABFEE835A9FE95DA6A" box="[314,328,1272,1298]" italics="true" pageId="1" pageNumber="126">1</emphasis>
|
||
) overwintering rosette leaves, (
|
||
<emphasis id="EFDA7F3E816DFFABFD7E35A9FD67DA6A" box="[684,698,1272,1298]" italics="true" pageId="1" pageNumber="126">2</emphasis>
|
||
) the stem purplish at the base, (
|
||
<emphasis id="EFDA7F3E816DFFABFBF235A9FBF3DA6A" box="[1056,1070,1272,1298]" italics="true" pageId="1" pageNumber="126">3</emphasis>
|
||
) the lower part of stem and lower cauline leaves covered by rough, dense, simple, hairs (
|
||
<emphasis id="EFDA7F3E816DFFABFD3D344DFD20DA4E" box="[751,765,1308,1334]" italics="true" pageId="1" pageNumber="126">4</emphasis>
|
||
) short involucral bracts (
|
||
<emphasis id="EFDA7F3E816DFFABFBC8344DFBF5DA4E" box="[1050,1064,1308,1334]" italics="true" pageId="1" pageNumber="126">5</emphasis>
|
||
) capitula spherical in bud, and (
|
||
<emphasis id="EFDA7F3E816DFFABFA46344AFA7FDA4D" box="[1428,1442,1307,1333]" italics="true" pageId="1" pageNumber="126">6</emphasis>
|
||
) glaucous leaves, resemble
|
||
<taxonomicName id="1AAED8AF816DFFABFE613411FCDBDA22" authority="Pallas (1771: 501)" authorityName="Pallas" authorityPageNumber="501" authorityYear="1771" box="[435,774,1344,1370]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="virosum">
|
||
<emphasis id="EFDA7F3E816DFFABFE613411FDF3DA21" box="[435,558,1344,1369]" italics="true" pageId="1" pageNumber="126">H. virosum</emphasis>
|
||
<bibRefCitation id="B93FDEDD816DFFABFDE73410FCDBDA22" author="Pallas, P. S." box="[565,774,1345,1370]" pageId="1" pageNumber="126" refId="ref3326" refString="Pallas, P. S. (1771) Reise durch verschiedene Provinzen des russischen Reichs, vol. 1. Kayserliche Academie der Wissenschaften, St. Petersburg, 504 pp. + 11 tab." type="book" year="1771">Pallas (1771: 501)</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DD11A32C816DFFABFF6F3434FC71D94A" blockId="1.[136,1452,301,1946]" pageId="1" pageNumber="126">
|
||
In southeastern Europe,
|
||
<taxonomicName id="1AAED8AF816DFFABFE033435FD90DA05" authorityName="Pallas" authorityYear="1771" box="[465,589,1380,1405]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="virosum">
|
||
<emphasis id="EFDA7F3E816DFFABFE033435FD90DA05" box="[465,589,1380,1405]" italics="true" pageId="1" pageNumber="126">H. virosum</emphasis>
|
||
</taxonomicName>
|
||
reaches the western limit of its geographical range and is represented by
|
||
<taxonomicName id="1AAED8AF816DFFABFA423435FDDADADA" authority="Willdenow (1803: 1589)" authorityName="Willdenow" authorityPageNumber="1589" authorityYear="1803" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="foliosum">
|
||
<emphasis id="EFDA7F3E816DFFABFA423435FF35DAD9" italics="true" pageId="1" pageNumber="126">H. foliosum</emphasis>
|
||
<bibRefCitation id="B93FDEDD816DFFABFF2234D8FDDADADA" author="Willdenow, C. L." box="[240,519,1417,1442]" pageId="1" pageNumber="126" pagination="1476 - 2409" refId="ref3758" refString="Willdenow, C. L. (1803) Species Plantarum. Ed. 4, vol. 3 (3). G. C. Nauk, Berlin, pp. 1476 - 2409. https: // doi. org / 10.5962 / bhl / title. 727" type="book chapter" year="1803">Willdenow (1803: 1589)</bibRefCitation>
|
||
</taxonomicName>
|
||
, currently treated as conspecific with
|
||
<taxonomicName id="1AAED8AF816DFFABFC6634D9FB2ADADA" authority="(Sennikov 2008)" baseAuthorityName="Sennikov" baseAuthorityYear="2008" box="[948,1271,1416,1442]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="virosum">
|
||
<emphasis id="EFDA7F3E816DFFABFC6634D9FBF2DAD9" box="[948,1071,1416,1441]" italics="true" pageId="1" pageNumber="126">H. virosum</emphasis>
|
||
(
|
||
<bibRefCitation id="B93FDEDD816DFFABFBED34D8FB32DADA" author="Sennikov, A. N." box="[1087,1263,1417,1442]" pageId="1" pageNumber="126" pagination="19 - 32" refId="ref3363" refString="Sennikov, A. N. (2008) Hieracium L., Pilosella Hill. In: Grubov, V. I. (Ed.) Rastenija tsentralnoj Asii [Plants of Central Asia], vol. 14 b. KMK, Moskva, pp. 19 - 32." type="book chapter" year="2008">Sennikov 2008</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. In
|
||
<collectingCountry id="A5B9E3BC816DFFABFAF334D8FA59DADA" box="[1313,1412,1417,1442]" name="Bulgaria" pageId="1" pageNumber="126">Bulgaria</collectingCountry>
|
||
,
|
||
<taxonomicName id="1AAED8AF816DFFABFA4234D9FF3CDABD" baseAuthorityName="Sennikov" baseAuthorityYear="2008" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="virosum">
|
||
<emphasis id="EFDA7F3E816DFFABFA4234D9FF3CDABD" italics="true" pageId="1" pageNumber="126">H. virosum</emphasis>
|
||
</taxonomicName>
|
||
is a rare species and has been classified as ‘Endangered’ (
|
||
<bibRefCitation id="B93FDEDD816DFFABFCA734FCFBABDABE" author="Vladimirov, V." box="[885,1142,1453,1478]" pageId="1" pageNumber="126" refId="ref3646" refString="Vladimirov, V. (2015) Hieracium virosum Pall. In: Peev, D., Petrova, A. S., Anchev, M., Temniskova, D., Denchev, C. M., Ganeva, A., Gussev, Ch. & Vladimirov, V. (Eds.) Red Data Book of the Republic of Bulgaria. Vol. 1. Plants and Fungi. Bulgarian Academy of Sciences & Ministry of Environment and Water of Bulgaria, Sofia, 521 pp. [digital edition: http: // e-ecodb. bas. bg / rdb / en]" type="book" year="2015">Vladimirov 2015: 521</bibRefCitation>
|
||
); it was also recorded from the Vitosha region (
|
||
<bibRefCitation id="B93FDEDD816DFFABFEB03480FD9CDA92" author="Stojanov, N. & Stefanov, B. & Kitanov, B." box="[354,577,1487,1514]" pageId="1" pageNumber="126" refId="ref3463" refString="Stojanov, N., Stefanov, B. & Kitanov, B. (1967) Flora Bulgarica. Ed. 4. Nauka & Izkustvo, Sofia, 1325 pp." type="book" year="1967">
|
||
Stojanov
|
||
<emphasis id="EFDA7F3E816DFFABFE1C3481FE22DA91" box="[462,511,1487,1513]" italics="true" pageId="1" pageNumber="126">et al</emphasis>
|
||
1967
|
||
</bibRefCitation>
|
||
). In my opinion,
|
||
<taxonomicName id="1AAED8AF816DFFABFCD23481FCA6DA91" authorityName="Szelag" authorityYear="2018" box="[768,891,1488,1513]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="gussevii">
|
||
<emphasis id="EFDA7F3E816DFFABFCD23481FCA6DA91" box="[768,891,1488,1513]" italics="true" pageId="1" pageNumber="126">H. gussevii</emphasis>
|
||
</taxonomicName>
|
||
may have originated as the result of hybridization between
|
||
<taxonomicName id="1AAED8AF816DFFABFF3C34A5FEAFD975" authorityName="Frivaldszky" authorityYear="1836" box="[238,370,1524,1549]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="sparsum">
|
||
<emphasis id="EFDA7F3E816DFFABFF3C34A5FEAFD975" box="[238,370,1524,1549]" italics="true" pageId="1" pageNumber="126">H. sparsum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1AAED8AF816DFFABFE7E34A5FCD6D976" authority="Fries (1848: 193)" authorityName="Fries" authorityPageNumber="193" authorityYear="1848" box="[428,779,1523,1550]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="robustum">
|
||
<emphasis id="EFDA7F3E816DFFABFE7E34A5FDE4D975" box="[428,569,1523,1549]" italics="true" pageId="1" pageNumber="126">H. robustum</emphasis>
|
||
<bibRefCitation id="B93FDEDD816DFFABFD9034A4FCD6D976" author="Fries, E." box="[578,779,1525,1550]" pageId="1" pageNumber="126" pagination="1 - 220" refId="ref3103" refString="Fries, E. (1848) Symbolae ad historiam Hieraciorum. Leffler et Sebell, Upsaliae, i - xxxiv, 1 - 220 pp." type="book chapter" year="1848">Fries (1848: 193)</bibRefCitation>
|
||
</taxonomicName>
|
||
, a hybridogenous species of the morphological formula
|
||
<emphasis id="EFDA7F3E816DFFABFA4234A5FE7ED949" italics="true" pageId="1" pageNumber="126">
|
||
<taxonomicName id="1AAED8AF816DFFABFA4234A5FF3CD949" baseAuthorityName="Sennikov" baseAuthorityYear="2008" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="virosum">H. virosum</taxonomicName>
|
||
—
|
||
<taxonomicName id="1AAED8AF816DFFABFF2F3749FE7ED949" authorityName="Linnaeus" authorityYear="1753" box="[253,419,1559,1585]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="umbellatum">H. umbellatum</taxonomicName>
|
||
</emphasis>
|
||
(cf.
|
||
<bibRefCitation id="B93FDEDD816DFFABFE0A3748FD59D94A" author="Sennikov, A. N." box="[472,644,1561,1586]" pageId="1" pageNumber="126" pagination="85 - 88" refId="ref3416" refString="Sennikov, A. N. (2012) Critical notes on the genera Hieracium and Pilosella (Asteraceae, Cichorieae) in the Himalayas. Willdenowia 42: 85 - 88. http: // dx. doi. org / 10.3372 / wi. 42.42110" type="journal article" year="2012">Sennikov 2012</bibRefCitation>
|
||
, see also
|
||
<bibRefCitation id="B93FDEDD816DFFABFD3F3748FC7DD94A" author="Tupitsyna, N. N." box="[749,928,1561,1586]" pageId="1" pageNumber="126" refId="ref3557" refString="Tupitsyna, N. N. (2004) Yastrebinki Sibiri [Hawkweeds of Siberia]. Nauka, Novosibirsk, 208 pp." type="book" year="2004">Tupitsyna 2004</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95B4F0A7816DFFABFF6F376CFE69D856" pageId="1" pageNumber="126" type="distribution">
|
||
<paragraph id="DD11A32C816DFFABFF6F376CFD28D99E" blockId="1.[136,1452,301,1946]" pageId="1" pageNumber="126">
|
||
<emphasis id="EFDA7F3E816DFFABFF6F376CFE2ED92E" bold="true" box="[189,499,1597,1622]" pageId="1" pageNumber="126">Distribution and habitats:</emphasis>
|
||
—Endemic to the Vitosha Mountains in western
|
||
<collectingCountry id="A5B9E3BC816DFFABFBDD376CFBAED92E" box="[1039,1139,1597,1622]" name="Bulgaria" pageId="1" pageNumber="126">Bulgaria</collectingCountry>
|
||
.
|
||
<taxonomicName id="1AAED8AF816DFFABFBAE376DFA89D92D" authorityName="Szelag" authorityYear="2018" box="[1148,1364,1596,1621]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="gussevii">
|
||
<emphasis id="EFDA7F3E816DFFABFBAE376DFA89D92D" box="[1148,1364,1596,1621]" italics="true" pageId="1" pageNumber="126">Hieracium gussevii</emphasis>
|
||
</taxonomicName>
|
||
is quite frequent in the area between the Aleko chalet and Mt. Goli Vrah, at
|
||
<quantity id="1A560EC9816DFFABFCA43730FBCDD902" box="[886,1040,1633,1658]" metricMagnitude="3" metricUnit="m" metricValue="1.815" metricValueMax="1.83" metricValueMin="1.8" pageId="1" pageNumber="126" unit="m" value="1815.0" valueMax="1830.0" valueMin="1800.0">1800–1830 m</quantity>
|
||
a.s.l., but it has been overlooked due to the morphological similarity with ubiquitous
|
||
<taxonomicName id="1AAED8AF816DFFABFD4937D5FC58D9E6" authority="Freyn" authorityName="Freyn" box="[667,901,1667,1694]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="velenovskyi">
|
||
<emphasis id="EFDA7F3E816DFFABFD4937D5FCE0D9E5" box="[667,829,1667,1693]" italics="true" pageId="1" pageNumber="126">H. velenovskyi</emphasis>
|
||
Freyn
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="B93FDEDD816DFFABFC7437D4FB69D9E6" author="Velenovsky, J." box="[934,1204,1669,1694]" pageId="1" pageNumber="126" refId="ref3615" refString="Velenovsky, J. (1891) Flora Bulgarica. Descriptio et enumeratio systematica plantarum vascularium in Principatu Bulgariae sponte nascentium. F. Rivnac, Pragae, 676 pp." type="book" year="1891">Velenovský (1891: 346)</bibRefCitation>
|
||
, growing side by side. The populations consisted of numerous plants growing in small clusters on the
|
||
<taxonomicName id="1AAED8AF816DFFABFC3D37F9FBB2D9B9" box="[1007,1135,1703,1729]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="abies">
|
||
<emphasis id="EFDA7F3E816DFFABFC3D37F9FBB2D9B9" box="[1007,1135,1703,1729]" italics="true" pageId="1" pageNumber="126">Picea abies</emphasis>
|
||
</taxonomicName>
|
||
forest margin, both in sunny and in partly overshadowed places, on silicate bedrock.
|
||
</paragraph>
|
||
<paragraph id="DD11A32C816DFFABFF6F37A0FE69D856" blockId="1.[136,1452,301,1946]" pageId="1" pageNumber="126">
|
||
<emphasis id="EFDA7F3E816DFFABFF6F37A0FCDED872" bold="true" box="[189,771,1777,1802]" pageId="1" pageNumber="126">Chromosome number and mode of reproduction:</emphasis>
|
||
—2n = 3x = 27, agamospermous (
|
||
<bibRefCitation id="B93FDEDD816DFFABFBA837A0FA7BD872" author="Ilnicki, T. & Szelag, Z." box="[1146,1446,1777,1802]" pageId="1" pageNumber="126" pagination="102 - 110" refId="ref3176" refString="Ilnicki, T. & Szelag, Z. (2011) Chromosome numbers in Hieracium and Pilosella (Asteraceae) from Central and Southeastern Europe. Acta Biologica Cracoviensia Series Botanica 53 (1): 102 - 110. http: // dx. doi. org / 10.2478 / v 10182 - 011 - 0014 - 3" type="journal article" year="2011">Ilnicki & Szeląg 2011: 107</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="1AAED8AF816DFFABFF743645FE98D856" box="[166,325,1812,1838]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="EFDA7F3E816DFFABFF743645FEC3D855" box="[166,286,1812,1837]" italics="true" pageId="1" pageNumber="126">Hieracium</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
‘Aleko’).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95B4F0A7816DFFABFF6F3668FDC0D8E2" pageId="1" pageNumber="126" type="etymology">
|
||
<paragraph id="DD11A32C816DFFABFF6F3668FDC0D8E2" blockId="1.[136,1452,301,1946]" pageId="1" pageNumber="126">
|
||
<emphasis id="EFDA7F3E816DFFABFF6F3668FE95D82A" bold="true" box="[189,328,1849,1874]" pageId="1" pageNumber="126">Etymology:</emphasis>
|
||
―The new species is named in honour of my colleague Dr. Chavdar Gussev (Чавдар Гусев), Institute of Biodiversity and Ecosystem Research Bulgarian Academy of Sciences,
|
||
<collectingRegion id="1F6A6DCE816DFFABFC1E360CFBD7D80E" box="[972,1034,1885,1910]" country="Bulgaria" name="Sofiya-Grad" pageId="1" pageNumber="126">Sofia</collectingRegion>
|
||
, for his contribution to knowledge of the flora and vegetation in
|
||
<collectingCountry id="A5B9E3BC816DFFABFE6736D0FDC4D8E2" box="[437,537,1921,1946]" name="Bulgaria" pageId="1" pageNumber="126">Bulgaria</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |