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<document id="19DE92B1C37621106D5EA92818521909" ID-CLB-Dataset="71201" ID-DOI="10.11646/zootaxa.5154.1.2" ID-GBIF-Dataset="2fc6067e-600e-46ce-addc-c735e05bc789" ID-ISSN="1175-5326" ID-Zenodo-Dep="6637183" ID-ZooBank="209D4D88-BF8C-4124-95A1-31B6F43861D7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1655101583164" checkinUser="plazi" docAuthor="Bálint, Zsolt, Boyer, Pierre, Farfán, Jackie, Cerdeña, José &amp; Pyrcz, Tomasz W." docDate="2022" docId="547E756CFF99FFEF1EFD8BE6FE25F87D" docLanguage="en" docName="Zootaxa.5154.1.2.pdf" docOrigin="Zootaxa 5154 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Penaincisalia jadwigae Balint, Boyer &amp; Pyrcz 2022, sp. n." docType="treatment" docVersion="5" lastPageNumber="51" masterDocId="A8470D14FF98FFED1E6A8F4FFF90FF9E" masterDocTitle="A new high-altitude species of Penaincisalia Johnson, 1990 (Lepidoptera, Lycaenidae) from the Peruvian Andes" masterLastPageNumber="59" masterPageNumber="49" pageNumber="50" updateTime="1699339283048" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="55CC91BE1B1776780CE9D62B1C9E3B99" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D667AAA0C9F37EE10A9628DE3459D121">
<mods:title id="7670B903EAF792DEB7548EDB08B757C5">A new high-altitude species of Penaincisalia Johnson, 1990 (Lepidoptera, Lycaenidae) from the Peruvian Andes</mods:title>
</mods:titleInfo>
<mods:name id="EA2EBC202DD0D314B7E88F3C1C6727D5" type="personal">
<mods:role id="61DB54770B14A6B7FB2FF2496B91410F">
<mods:roleTerm id="E58AB19696E637AC028615E7882BB463">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CE8550AF0AD629C8DE884344C689142C">Bálint, Zsolt</mods:namePart>
</mods:name>
<mods:name id="805C6E31A8104DC5A47BAF86DA65CC40" type="personal">
<mods:role id="CA3A75A1E339BBDE8B0D8E6258CF6249">
<mods:roleTerm id="272B0C69E37C95464F52C2F1EFAD6647">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6A678D643EA0A6CFFD0A481DC7D24A36">Boyer, Pierre</mods:namePart>
<mods:nameIdentifier id="42DFEC4884414A81B7AC5F7E72C16F25" type="ORCID">0000-0001-7271-4246</mods:nameIdentifier>
<mods:affiliation id="678601479A6930F9ABC8F28AF9AE754D">Lotissement l’Horizon, 13610 Le Puy Sainte Réparade, France. pierdom @ aliceadsl. fr; https: // orcid. org / 0000 - 0001 - 7271 - 4246 Universidad Nacional de San Augustín de Arequipa, Museo de Historia Natural, Av Alcides Carrión s / n, Arequipa, Peru. Department of Evolution of Invertebrates, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 380 Kraków, Poland. tomasz. pyrcz @ uj. edu. pl; https: // orcid. org / 0000 - 0003 - 4822 - 0670</mods:affiliation>
<mods:nameIdentifier id="2C262E8926372EC6E2AB575E1F5543EE" type="email">pierdom@aliceadsl.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="B4B52A3F3B49C52D4C58D79DAAA34CFE" type="personal">
<mods:role id="FB7979925514E0F0F650ED0E9C6C47EE">
<mods:roleTerm id="646ED19F2D6A351A0FB5250A868AFE1D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A6C525395BCBAAE00DB300DED730CAAC">Farfán, Jackie</mods:namePart>
<mods:nameIdentifier id="6B639F34C94E9B1BFD56F6B0CF501EDC" type="ORCID">0000-0001-5913-0756</mods:nameIdentifier>
<mods:affiliation id="1DBC2A63ED33E8ECD221EB8A074FB31E">jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="127C71A27675A0894B1B1C9ED9780C7D" type="email">jjackie4u@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="55C373747EEC704CDA2072E8C5DDA515" type="personal">
<mods:role id="28FEDE28A4CA2D7AD3A7A9AFE79C55B3">
<mods:roleTerm id="768D6A916F91996A0F5785AADFB995DA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="47D154A835CDC6EF0C1497AF2DF8C6BF">Cerdeña, José</mods:namePart>
<mods:nameIdentifier id="984FE2BE5122D3369C5AAB34CC770503" type="ORCID">0000-0003-3930-6480</mods:nameIdentifier>
<mods:affiliation id="B6EEA50DCE8920801A150E1E06F0EC37">cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480</mods:affiliation>
<mods:nameIdentifier id="C4A2B8C8F0BFE46323545486F2D04E75" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
</mods:name>
<mods:name id="C5EDE301CB4AA678EB731887A1A6218C" type="personal">
<mods:role id="2B08C904F4993FA23EA27B39238DCAA2">
<mods:roleTerm id="A8B9E4F8CE1605606FF2D962B388803A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BA0A16E829390FC38C413B8FF404B1F8">Pyrcz, Tomasz W.</mods:namePart>
<mods:nameIdentifier id="296BDB8F20F0F40656A08693F5ED44B0" type="ORCID">0000-0003-4822-0670</mods:nameIdentifier>
<mods:nameIdentifier id="8E084372D6F4E0DBC530569FE10EF87D" type="email">tomasz.pyrcz@uj.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="C042BE3D64D3C4F3EF79BC7B78AB7DF1">text</mods:typeOfResource>
<mods:relatedItem id="E6A6AF62B8BD0B4EA1AEC988638396AD" type="host">
<mods:titleInfo id="E2C88A5F467F8455127FF49812B91CE0">
<mods:title id="93DEE64F6D2045D98B59E6C25F41F481">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9B0F918B3203A7632D2D2CC179D6B846">
<mods:date id="0BB52F8A7A9B038A5E399E8195A96BAB">2022</mods:date>
<mods:detail id="B82154463E07922AD7500BEECA2A9A89" type="pubDate">
<mods:number id="999661D7402482AEB9ABE32704B2160D">2022-06-13</mods:number>
</mods:detail>
<mods:detail id="2887A603C7E478D89043119C86F6EE1A" type="volume">
<mods:number id="585056F86B06B1AFA22E56D15B715D5B">5154</mods:number>
</mods:detail>
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<mods:number id="E7B89C13CAF44EED7516AEF4ED2900F9">1</mods:number>
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<mods:start id="0EED3BE19D68E8886F277B10CBE88707">49</mods:start>
<mods:end id="44F8E1248930BF8A73E0D32D419C8E38">59</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BC47BFB8A943A11847093B71F3984DA3">journal article</mods:classification>
<mods:identifier id="F71126CCEAEE544A0208665D63047E68" type="CLB-Dataset">71201</mods:identifier>
<mods:identifier id="13E1C46BC3B3DE659CB0F510AEA1B161" type="DOI">10.11646/zootaxa.5154.1.2</mods:identifier>
<mods:identifier id="42AD780B7B9AE03655DC91B0A81BA2BC" type="GBIF-Dataset">2fc6067e-600e-46ce-addc-c735e05bc789</mods:identifier>
<mods:identifier id="7DCCBFF9975747063E97AF7B9D0CFDF4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D6B49E8947096A76F5F180EFF3A59825" type="Zenodo-Dep">6637183</mods:identifier>
<mods:identifier id="E4E4D511C6B2B3FF85EA89D907251E0E" type="ZooBank">209D4D88-BF8C-4124-95A1-31B6F43861D7</mods:identifier>
</mods:mods>
<treatment id="547E756CFF99FFEF1EFD8BE6FE25F87D" ID-DOI="http://doi.org/10.5281/zenodo.6637185" ID-GBIF-Taxon="195864894" ID-Zenodo-Dep="6637185" LSID="urn:lsid:plazi:treatment:547E756CFF99FFEF1EFD8BE6FE25F87D" httpUri="http://treatment.plazi.org/id/547E756CFF99FFEF1EFD8BE6FE25F87D" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50">
<subSubSection id="94CD97F1FF99FFEC1EFD8BE6FC8BFB5A" box="[151,795,1193,1220]" pageId="1" pageNumber="50" type="nomenclature">
<paragraph id="DC68C47AFF99FFEC1EFD8BE6FC8BFB5A" blockId="1.[151,795,1193,1256]" box="[151,795,1193,1220]" pageId="1" pageNumber="50">
<heading id="87207316FF99FFEC1EFD8BE6FC8BFB5A" bold="true" box="[151,795,1193,1220]" fontSize="11" level="1" pageId="1" pageNumber="50" reason="1">
<emphasis id="EEA31868FF99FFEC1EFD8BE6FC8BFB5A" bold="true" box="[151,795,1193,1220]" pageId="1" pageNumber="50">
<taxonomicName id="1BD7BFF9FF99FFEC1EFD8BE6FD5CFB5A" ID-CoL="9FT69" authority="Balint, Boyer &amp; Pyrcz" authorityName="Balint, Boyer &amp; Pyrcz" authorityYear="2022" box="[151,716,1193,1220]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="jadwigae" status="sp. nov.">
<emphasis id="EEA31868FF99FFEC1EFD8BE6FE3FFB5A" bold="true" box="[151,431,1193,1220]" italics="true" pageId="1" pageNumber="50">Penaincisalia jadwigae</emphasis>
Bálint, Boyer &amp; Pyrcz
</taxonomicName>
,
<taxonomicNameLabel id="F590A513FF99FFEC1CB38BE5FC8BFB5A" box="[729,795,1194,1220]" pageId="1" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="94CD97F1FF99FFEC1EFD8B81FB66FACB" pageId="1" pageNumber="50" type="description">
<paragraph id="DC68C47AFF99FFEC1EFD8B81FEBFFB76" blockId="1.[151,795,1193,1256]" box="[151,303,1230,1256]" pageId="1" pageNumber="50">
(
<figureCitation id="44ECD8FFFF99FFEC1EF58B81FEB6FB76" box="[159,294,1230,1256]" captionStart-0="FIGURE 1–4" captionStart-1="FIGURE 5–6" captionStart-2="FIGURE 7–8" captionStartId-0="2.[151,250,1706,1730]" captionStartId-1="3.[151,250,830,854]" captionStartId-2="3.[151,250,1986,2010]" captionTargetBox-0="[151,1436,702,1680]" captionTargetBox-1="[223,1354,211,798]" captionTargetBox-2="[371,1213,922,1953]" captionTargetId-0="figure-270@2.[151,1436,702,1680]" captionTargetId-1="figure-20@3.[216,1370,181,805]" captionTargetId-2="figure-52@3.[360,1228,914,1962]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIGURE 1–4. Penaincisalia jadwigae sp. n. type material. 1 = holotype dorsum, 2 = ditto, ventrum, 3 = paratype no. 1. (allotype) female dorsum, 4 = ditto, ventrum. All specimens under same magnification. Scale: holotype fore wing costa length 11 mm." captionText-1="FIGURE 5–6. Male genitalia capsula and aedeagus of Penaincisalia jadwigae sp. n. in lateral (5) and ventral view (aedeagus excluded) (6). Scale bar: 1 mm." captionText-2="FIGURE 7–8. Female genitalia of Penaincisalia jadwigae sp. n. in lateral (7) and ventral view (8). Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6637187" figureDoi-1="http://doi.org/10.5281/zenodo.6637191" figureDoi-2="http://doi.org/10.5281/zenodo.6637193" httpUri-0="https://zenodo.org/record/6637187/files/figure.png" httpUri-1="https://zenodo.org/record/6637191/files/figure.png" httpUri-2="https://zenodo.org/record/6637193/files/figure.png" pageId="1" pageNumber="50">Figures 1–8</figureCitation>
)
</paragraph>
<paragraph id="DC68C47AFF99FFEC1EFD8A5AFB66FACB" blockId="1.[151,1437,1301,2012]" pageId="1" pageNumber="50">
<emphasis id="EEA31868FF99FFEC1EFD8A5AFE32FAAE" bold="true" box="[151,418,1301,1328]" pageId="1" pageNumber="50">Systematic placement.</emphasis>
Ordo:
<taxonomicName id="1BD7BFF9FF99FFEC1F988A59FDECFAAE" authorityName="Linnaeus" authorityYear="1758" box="[498,636,1302,1328]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="1" pageNumber="50" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
, family:
<taxonomicName id="1BD7BFF9FF99FFEC1C8A8A59FCF2FAAE" authorityName="Leach" authorityYear="1815" box="[736,866,1302,1328]" class="Insecta" family="Lycaenidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="family">Lycaenidae</taxonomicName>
, subfamily:
<taxonomicName id="1BD7BFF9FF99FFEC1D868A59FBCDFAAE" box="[1004,1117,1302,1328]" class="Insecta" family="Lycaenidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Theclinae">Theclinae</taxonomicName>
, tribe:
<taxonomicName id="1BD7BFF9FF99FFEC1AC18A59FA89FAAE" authorityName="E.Doubleday" authorityYear="1847" box="[1195,1305,1302,1328]" class="Insecta" family="Lycaenidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Eumaeini">Eumaeini</taxonomicName>
, genus:
<taxonomicName id="1BD7BFF9FF99FFEC1B1F8A58FE2CFACA" authority="Johnson, 1990" authorityName="Johnson" authorityYear="1990" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="EEA31868FF99FFEC1B1F8A58FE83FACA" italics="true" pageId="1" pageNumber="50">Penaincisalia</emphasis>
Johnson, 1990
</taxonomicName>
(
<typeStatus id="036C7AD8FF99FFEC1FA18A75FE6DFACA" box="[459,509,1338,1364]" pageId="1" pageNumber="50">type</typeStatus>
species:
<taxonomicName id="1BD7BFF9FF99FFEC1C0E8A75FB93FACA" authority="Staudinger, 1894" authorityName="Staudinger" authorityYear="1894" box="[612,1027,1338,1364]" class="Insecta" family="Lycaenidae" genus="Thecla" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="culminicola">
<emphasis id="EEA31868FF99FFEC1C0E8A75FCABFACA" box="[612,827,1338,1364]" italics="true" pageId="1" pageNumber="50">Thecla culminicola</emphasis>
Staudinger, 1894
</taxonomicName>
, original designation)
</paragraph>
</subSubSection>
<subSubSection id="94CD97F1FF99FFEC1EAD8A11FE12F997" pageId="1" pageNumber="50" type="materials_examined">
<paragraph id="DC68C47AFF99FFEC1EAD8A11FE12F997" blockId="1.[151,1437,1301,2012]" pageId="1" pageNumber="50">
<emphasis id="EEA31868FF99FFEC1EAD8A11FEE6FAE9" bold="true" box="[199,374,1373,1400]" pageId="1" pageNumber="50">Type material.</emphasis>
<materialsCitation id="6CBFCE27FF99FFEC1F158A11FCABFA5E" ID-GBIF-Occurrence="3807031302" collectingDate="2021-09-27" collectionCode="PBC, MUSM" country="Peru" elevation="4555" location="Pierre Boyer Leg" municipality="Route" pageId="1" pageNumber="50" specimenCount="1" stateProvince="Huancavelica" typeStatus="holotype">
<typeStatus id="036C7AD8FF99FFEC1F158A11FE77FAE6" box="[383,487,1374,1400]" pageId="1" pageNumber="50" type="holotype">Holotype</typeStatus>
male, set dorsally, in perfect condition (
<figureCitation id="44ECD8FFFF99FFEC1DD98A11FB87FAE6" box="[947,1047,1374,1400]" captionStart="FIGURE 1–4" captionStartId="2.[151,250,1706,1730]" captionTargetBox="[151,1436,702,1680]" captionTargetId="figure-270@2.[151,1436,702,1680]" captionTargetPageId="2" captionText="FIGURE 1–4. Penaincisalia jadwigae sp. n. type material. 1 = holotype dorsum, 2 = ditto, ventrum, 3 = paratype no. 1. (allotype) female dorsum, 4 = ditto, ventrum. All specimens under same magnification. Scale: holotype fore wing costa length 11 mm." figureDoi="http://doi.org/10.5281/zenodo.6637187" httpUri="https://zenodo.org/record/6637187/files/figure.png" pageId="1" pageNumber="50">Figs 1–2</figureCitation>
) with label “
<location id="D90892A1FF99FFEC1AC18A11FAE2FAE6" LSID="urn:lsid:plazi:treatment:547E756CFF99FFEF1EFD8BE6FE25F87D:D90892A1FF99FFEC1AC18A11FAE2FAE6" box="[1195,1394,1374,1400]" country="Peru" municipality="Route" name="Pierre Boyer Leg" pageId="1" pageNumber="50" stateProvince="Huancavelica">Pierre Boyer Leg</location>
[//]
<collectingMunicipality id="3C0C5E00FF99FFEC1EFD8ACDFF4AFA02" box="[151,218,1410,1436]" pageId="1" pageNumber="50">Route</collectingMunicipality>
26B pk112, [//]
<quantity id="1B2F699FFF99FFEC1FFF8ACDFDB9FA03" box="[405,553,1410,1437]" metricMagnitude="3" metricUnit="m" metricValue="4.555" metricValueMax="4.6" metricValueMin="4.51" pageId="1" pageNumber="50" unit="m" value="4555.0" valueMax="4600.0" valueMin="4510.0">
<elevation id="57FA2349FF99FFEC1FFF8ACDFDB9FA03" box="[405,553,1410,1437]" metricMagnitude="3" metricUnit="m" metricValue="4.555" metricValueMax="4.6" metricValueMin="4.51" pageId="1" pageNumber="50" unit="m" value="4555.0" valueMax="4600.0" valueMin="4510.0">4510–4600m</elevation>
</quantity>
[//] (
<collectingRegion id="1E130A98FF99FFEC1C0B8ACDFD6DFA03" box="[609,765,1410,1437]" country="Peru" name="Huancavelica" pageId="1" pageNumber="50">Huancavelica</collectingRegion>
) [//]
<collectingCountry id="A4C084EAFF99FFEC1D5C8ACDFC02FA02" box="[822,914,1410,1436]" name="Peru" pageId="1" pageNumber="50">PEROU</collectingCountry>
[//]
<date id="A869E2BAFF99FFEC1DA98ACDFBA7FA02" box="[963,1079,1410,1436]" pageId="1" pageNumber="50" value="2021-09-27">
<collectingDate id="B82D1B52FF99FFEC1DA98ACDFBA7FA02" box="[963,1079,1410,1436]" pageId="1" pageNumber="50" value="2021-09-27">27/9/2021</collectingDate>
</date>
” (white label, printed, letters in black frames), at present in
<collectionCode id="BAC65CBFFF99FFEC1FBA8AE9FD98FA5E" box="[464,520,1446,1472]" country="0" httpUri="http://grbio.org/cool/hvzk-w6r8" name="Pacific Bacterial Collection" pageId="1" pageNumber="50">PBC</collectionCode>
, to be deposited in
<collectionCode id="BAC65CBFFF99FFEC1C888AE9FCABFA5E" box="[738,827,1446,1472]" pageId="1" pageNumber="50">MUSM</collectionCode>
</materialsCitation>
.
<materialsCitation id="6CBFCE27FF99FFEC1D2C8AE9FEEEF997" ID-GBIF-Occurrence="3807031301" collectingDate="2021-09-27" collectionCode="PBC, T, HNHM" country="Peru" elevation="4555" location="Pierre Boyer Leg" municipality="Route" pageId="1" pageNumber="50" specimenCount="3" stateProvince="Huancavelica" typeStatus="paratype">
<typeStatus id="036C7AD8FF99FFEC1D2C8AE9FC23FA5E" box="[838,947,1446,1472]" pageId="1" pageNumber="50" type="paratype">Paratypes</typeStatus>
(n = 9): with
<typeStatus id="036C7AD8FF99FFEC1A3A8AE9FB22FA5E" box="[1104,1202,1446,1472]" pageId="1" pageNumber="50" type="holotype">holotype</typeStatus>
data (no. 1:
<collectionCode id="BAC65CBFFF99FFEC1B2B8AE9FAE6FA5E" box="[1345,1398,1446,1472]" country="0" httpUri="http://grbio.org/cool/hvzk-w6r8" name="Pacific Bacterial Collection" pageId="1" pageNumber="50">PBC</collectionCode>
female (“
<typeStatus id="036C7AD8FF99FFEC1E8D8A85FEDBFA7B" box="[231,331,1482,1509]" pageId="1" pageNumber="50" type="allotype">allotype</typeStatus>
”); nos 2–5:
<collectionCode id="BAC65CBFFF99FFEC1FA48A85FD93FA7A" box="[462,515,1482,1508]" country="0" httpUri="http://grbio.org/cool/hvzk-w6r8" name="Pacific Bacterial Collection" pageId="1" pageNumber="50">PBC</collectionCode>
males), same data as the holotype, but “leg.
<collectionCode id="BAC65CBFFF99FFEC1D9E8A85FB96FA7A" box="[1012,1030,1482,1508]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="50">T</collectionCode>
. Pyrcz” (nos 6–8: CEP-MZUJ male, no. 9:
<collectionCode id="BAC65CBFFF99FFEC1EB48AA1FEA4F996" box="[222,308,1518,1544]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="50">HNHM</collectionCode>
male)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="94CD97F1FF99FFEC1EAD895EFED8F97E" pageId="1" pageNumber="50" type="diagnosis">
<paragraph id="DC68C47AFF99FFEC1EAD895EFED8F97E" blockId="1.[151,1437,1301,2012]" pageId="1" pageNumber="50">
<emphasis id="EEA31868FF99FFEC1EAD895EFED2F9B5" bold="true" box="[199,322,1553,1579]" pageId="1" pageNumber="50">Diagnosis.</emphasis>
The new species is most similar to
<taxonomicName id="1BD7BFF9FF99FFEC1CB5895CFC48F9B2" authority="Balint, 2001" authorityName="Balint" authorityYear="2001" box="[735,984,1554,1580]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="perezi">
<emphasis id="EEA31868FF99FFEC1CB5895CFCAFF9B2" box="[735,831,1555,1580]" italics="true" pageId="1" pageNumber="50">P. perezi</emphasis>
Bálint, 2001
</taxonomicName>
, which has an almost identical size and orange dorsal wing surface colouration, but specimens can most readily be recognized by the following three characters: (1) male dorsal wing-surface shows slight violet iridescence, visible at a shallow angle (
<taxonomicName id="1BD7BFF9FF99FFEC1AD78914FA8BF9EA" authorityName="Balint" authorityYear="2001" box="[1213,1307,1627,1652]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="perezi">
<emphasis id="EEA31868FF99FFEC1AD78914FA8BF9EA" box="[1213,1307,1627,1652]" italics="true" pageId="1" pageNumber="50">P. perezi</emphasis>
</taxonomicName>
: has strong iridescence visible at a wider angle); (2) hindwing black border is obsolete in the tornal area (in
<emphasis id="EEA31868FF99FFEC1ABF8930FAADF906" box="[1237,1341,1663,1688]" italics="true" pageId="1" pageNumber="50">
<taxonomicName id="1BD7BFF9FF99FFEC1ABF8930FAA8F906" authorityName="Balint" authorityYear="2001" box="[1237,1336,1663,1688]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="perezi">P. perezi</taxonomicName>
:
</emphasis>
running parallel with outer margin and clearly visible in the tornal area), and (3) ventral forewing ground colour is grey (
<emphasis id="EEA31868FF99FFEC1BE289ECFF74F97E" italics="true" pageId="1" pageNumber="50">
<taxonomicName id="1BD7BFF9FF99FFEC1BE289ECFF4EF97E" authorityName="Balint" authorityYear="2001" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="perezi">P. perezi</taxonomicName>
:
</emphasis>
orange).
</paragraph>
</subSubSection>
<subSubSection id="94CD97F1FF99FFEF1EAD89A6FD1BFD37" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50" type="description">
<paragraph id="DC68C47AFF99FFEF1EAD89A6FD1BFD37" blockId="1.[151,1437,1301,2012]" lastBlockId="2.[151,1437,151,681]" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50">
<emphasis id="EEA31868FF99FFEC1EAD89A6FEC9F89A" bold="true" box="[199,345,1769,1796]" pageId="1" pageNumber="50">Description.</emphasis>
Male (
<figureCitation id="44ECD8FFFF99FFEC1FC289A5FD9DF89A" box="[424,525,1770,1796]" captionStart="FIGURE 1–4" captionStartId="2.[151,250,1706,1730]" captionTargetBox="[151,1436,702,1680]" captionTargetId="figure-270@2.[151,1436,702,1680]" captionTargetPageId="2" captionText="FIGURE 1–4. Penaincisalia jadwigae sp. n. type material. 1 = holotype dorsum, 2 = ditto, ventrum, 3 = paratype no. 1. (allotype) female dorsum, 4 = ditto, ventrum. All specimens under same magnification. Scale: holotype fore wing costa length 11 mm." figureDoi="http://doi.org/10.5281/zenodo.6637187" httpUri="https://zenodo.org/record/6637187/files/figure.png" pageId="1" pageNumber="50">Figs 1–2</figureCitation>
,
<figureCitation id="44ECD8FFFF99FFEC1C7389A5FDD7F89B" box="[537,583,1770,1797]" captionStart="FIGURE 5–6" captionStartId="3.[151,250,830,854]" captionTargetBox="[223,1354,211,798]" captionTargetId="figure-20@3.[216,1370,181,805]" captionTargetPageId="3" captionText="FIGURE 5–6. Male genitalia capsula and aedeagus of Penaincisalia jadwigae sp. n. in lateral (5) and ventral view (aedeagus excluded) (6). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6637191" httpUri="https://zenodo.org/record/6637191/files/figure.png" pageId="1" pageNumber="50">5–6</figureCitation>
). Head, thorax and abdomen black dorsally; thorax and abdomen with greyish pubescence ventrally; antennae 0.6 times the length of the forewing costa, clubs black. Forewing: length
<quantity id="1B2F699FFF99FFEC1B288841FF53F8D2" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="1" pageNumber="50" unit="mm" value="10.75" valueMax="11.0" valueMin="10.5">10,5–11 mm</quantity>
(n=6), measured from the base of cubital vein to vein R3 terminus, and triangular in shape. Ground colour on the dorsal surface golden orange with violet iridescence (depending on the direction of illuminating light); margins with very thin (&lt;
<quantity id="1B2F699FFF99FFEC1F338835FE0AF80A" box="[345,410,1914,1940]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="50" unit="mm" value="2.0">2 mm</quantity>
) black border running parallel with outer margin in both wings, becoming more extensive at the apex of the forewing and fading towards the hindwing tornus, fringes chequered. Forewing androconia present in the form of a light grey coloured, trapezoidal scent pad in the apical part of the discal area, and a minute light grey scent patch in the base of vein M3. Forewing ventral surface greyish brown with a faint discoidal patch and black, sharp and slightly ruptured postmedian line and a somewhat lighter ash grey patterned submarginal area; hindwing basal and medial band black, interrupted by lighter variegate scaling, but obsolete; the submarginal line appearing as continuous intercellular black arrowhead marks bordered distally by lighter ash grey crescent pattern, with some slight variation in their intensity. Genitalia: tegumen and uncus large when viewed seen from lateral and dorsal sides; gnathos straight with pointed apex when seen from lateral but curved when seen from ventral side; valve slender, as long as tegumen, with slightly bent upper and lower margins, distal apex conical-shaped extending half way along the valva; vinculum slender but well sclerotized; saccus membranous and short (shorter than valva conical apical part); aedeagus robust but slender, two times the length of the valva, with two apical cornuti. (
<figureCitation id="44ECD8FFFF9AFFEF1ECA8E94FE94FE6B" box="[160,260,474,501]" captionStart="FIGURE 5–6" captionStartId="3.[151,250,830,854]" captionTargetBox="[223,1354,211,798]" captionTargetId="figure-20@3.[216,1370,181,805]" captionTargetPageId="3" captionText="FIGURE 5–6. Male genitalia capsula and aedeagus of Penaincisalia jadwigae sp. n. in lateral (5) and ventral view (aedeagus excluded) (6). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6637191" httpUri="https://zenodo.org/record/6637191/files/figure.png" pageId="2" pageNumber="51">Fig. 5–6</figureCitation>
). Female (
<figureCitation id="44ECD8FFFF9AFFEF1F168E94FE73FE6B" box="[380,483,475,501]" captionStart="FIGURE 1–4" captionStartId="2.[151,250,1706,1730]" captionTargetBox="[151,1436,702,1680]" captionTargetId="figure-270@2.[151,1436,702,1680]" captionTargetPageId="2" captionText="FIGURE 1–4. Penaincisalia jadwigae sp. n. type material. 1 = holotype dorsum, 2 = ditto, ventrum, 3 = paratype no. 1. (allotype) female dorsum, 4 = ditto, ventrum. All specimens under same magnification. Scale: holotype fore wing costa length 11 mm." figureDoi="http://doi.org/10.5281/zenodo.6637187" httpUri="https://zenodo.org/record/6637187/files/figure.png" pageId="2" pageNumber="51">Figs 3–4</figureCitation>
,
<figureCitation id="44ECD8FFFF9AFFEF1F9A8E94FD8EFE6B" box="[496,542,475,501]" captionStart="FIGURE 7–8" captionStartId="3.[151,250,1986,2010]" captionTargetBox="[371,1213,922,1953]" captionTargetId="figure-52@3.[360,1228,914,1962]" captionTargetPageId="3" captionText="FIGURE 7–8. Female genitalia of Penaincisalia jadwigae sp. n. in lateral (7) and ventral view (8). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6637193" httpUri="https://zenodo.org/record/6637193/files/figure.png" pageId="2" pageNumber="51">7–8</figureCitation>
). Similar in markings to the male but without androconia and a more rounded wing-shape (
<figureCitation id="44ECD8FFFF9AFFEF1F428EB0FE00FD87" box="[296,400,511,537]" captionStart="FIGURE 1–4" captionStartId="2.[151,250,1706,1730]" captionTargetBox="[151,1436,702,1680]" captionTargetId="figure-270@2.[151,1436,702,1680]" captionTargetPageId="2" captionText="FIGURE 1–4. Penaincisalia jadwigae sp. n. type material. 1 = holotype dorsum, 2 = ditto, ventrum, 3 = paratype no. 1. (allotype) female dorsum, 4 = ditto, ventrum. All specimens under same magnification. Scale: holotype fore wing costa length 11 mm." figureDoi="http://doi.org/10.5281/zenodo.6637187" httpUri="https://zenodo.org/record/6637187/files/figure.png" pageId="2" pageNumber="51">Figs 3–4</figureCitation>
). Genitalia: ductus bursae composed of sclerotized anterior and posterior parts divided by a dorsally membranous region; anterior ductus bursae two times shorter than posterior ductus-bursae, with convergent edges in dorso-ventral view; posterior ductus bursae widely open ventrally from the middle and dorsally composed by two sclerotized lamellae with blunt apices divided by a membranous central area; bursa with ductus length and arrowhead signa typical of genus (
<figureCitation id="44ECD8FFFF9AFFEF1C728DC0FDEEFD37" box="[536,638,655,681]" captionStart="FIGURE 7–8" captionStartId="3.[151,250,1986,2010]" captionTargetBox="[371,1213,922,1953]" captionTargetId="figure-52@3.[360,1228,914,1962]" captionTargetPageId="3" captionText="FIGURE 7–8. Female genitalia of Penaincisalia jadwigae sp. n. in lateral (7) and ventral view (8). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6637193" httpUri="https://zenodo.org/record/6637193/files/figure.png" pageId="2" pageNumber="51">Figs 7–8</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="88A894F2FF9AFFEF1EFD89E5FF56F894" ID-DOI="http://doi.org/10.5281/zenodo.6637187" ID-Zenodo-Dep="6637187" httpUri="https://zenodo.org/record/6637187/files/figure.png" pageId="2" pageNumber="51" startId="2.[151,250,1706,1730]" targetBox="[151,1436,702,1680]" targetPageId="2">
<paragraph id="DC68C47AFF9AFFEF1EFD89E5FF56F894" blockId="2.[151,1437,1706,1802]" pageId="2" pageNumber="51">
<emphasis id="EEA31868FF9AFFEF1EFD89E5FEBEF95C" bold="true" box="[151,302,1706,1730]" pageId="2" pageNumber="51">FIGURE 1–4.</emphasis>
<taxonomicName id="1BD7BFF9FF9AFFEF1F5F89E4FDB5F95D" authority="Bálint &amp; Boyer &amp; Farfán &amp; Cerdeña &amp; Pyrcz, 2022" authorityName="Bálint &amp; Boyer &amp; Farfán &amp; Cerdeña &amp; Pyrcz" authorityYear="2022" box="[309,549,1707,1731]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="jadwigae" status="sp. nov.">
<emphasis id="EEA31868FF9AFFEF1F5F89E4FDB5F95D" box="[309,549,1707,1731]" italics="true" pageId="2" pageNumber="51">Penaincisalia jadwigae</emphasis>
</taxonomicName>
<emphasis id="EEA31868FF9AFFEF1C4689E4FDF7F95D" bold="true" box="[556,615,1707,1731]" pageId="2" pageNumber="51">
<taxonomicNameLabel id="F590A513FF9AFFEF1C4689E4FDF7F95D" box="[556,615,1707,1731]" pageId="2" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
type material. 1 = holotype dorsum, 2 =
<emphasis id="EEA31868FF9AFFEF1A7A89E4FBAEF95D" box="[1040,1086,1707,1731]" italics="true" pageId="2" pageNumber="51">ditto</emphasis>
, ventrum, 3 = paratype no. 1. (allotype) female dorsum, 4 =
<emphasis id="EEA31868FF9AFFEF1FD88980FE70F979" box="[434,480,1743,1767]" italics="true" pageId="2" pageNumber="51">ditto</emphasis>
, ventrum. All specimens under same magnification. Scale: holotype fore wing costa length 11 mm.
</paragraph>
</caption>
<subSubSection id="94CD97F1FF9AFFEF1EAD8877FE4DF8E9" pageId="2" pageNumber="51" type="distribution">
<paragraph id="DC68C47AFF9AFFEF1EAD8877FE4DF8E9" blockId="2.[151,1437,1848,2019]" pageId="2" pageNumber="51">
<emphasis id="EEA31868FF9AFFEF1EAD8877FEC9F8CC" bold="true" box="[199,345,1848,1874]" pageId="2" pageNumber="51">Distribution</emphasis>
(known only from
<typeStatus id="036C7AD8FF9AFFEF1C5A8876FDF0F8CD" box="[560,608,1849,1875]" pageId="2" pageNumber="51">type</typeStatus>
data). Geographic:
<collectingRegion id="1E130A98FF9AFFEF1D568876FC47F8CD" box="[828,983,1849,1875]" country="Peru" name="Huancavelica" pageId="2" pageNumber="51">Huancavelica</collectingRegion>
,
<collectingCountry id="A4C084EAFF9AFFEF1D888876FB85F8CD" box="[994,1045,1849,1875]" name="Peru" pageId="2" pageNumber="51">Peru</collectingCountry>
(
<figureCitation id="44ECD8FFFF9AFFEF1A4E8876FBF7F8CD" box="[1060,1127,1849,1875]" captionStart="FIGURE 9" captionStartId="4.[151,250,1718,1742]" captionTargetBox="[174,1414,181,1694]" captionTargetId="figure-23@4.[173,1414,181,1694]" captionTargetPageId="4" captionText="FIGURE 9. Type localities of Penaincisalia species with male orange dorsal colouration. Penaincisalia alina Bálint, 2019 (Apurímac: Abancay Pass), P. aurulenta Johnson, 1990 (Ancash: Caraz), P. jadwigae sp. n. (Huancavelica), P. perezi Bálint, 2001 (Ancash: Parque Nacional Huascarán, Quebrada Demanda) and P. sp. (Apurímac: Abancay Pass)." figureDoi="http://doi.org/10.5281/zenodo.6637195" httpUri="https://zenodo.org/record/6637195/files/figure.png" pageId="2" pageNumber="51">Fig. 9</figureCitation>
). Altitudinal:
<quantity id="1B2F699FFF9AFFEF1B6B8876FA09F8CA" box="[1281,1433,1849,1876]" metricMagnitude="3" metricUnit="m" metricValue="4.5649999999999995" metricValueMax="4.62" metricValueMin="4.51" pageId="2" pageNumber="51" unit="m" value="4565.0" valueMax="4620.0" valueMin="4510.0">4510–4620m</quantity>
. Temporal: end of September.
</paragraph>
</subSubSection>
<subSubSection id="94CD97F1FF9AFFEF1EAD88CEFE25F87D" pageId="2" pageNumber="51" type="etymology">
<paragraph id="DC68C47AFF9AFFEF1EAD88CEFE25F87D" blockId="2.[151,1437,1848,2019]" pageId="2" pageNumber="51">
<emphasis id="EEA31868FF9AFFEF1EAD88CEFEDEF805" bold="true" box="[199,334,1921,1947]" pageId="2" pageNumber="51">Etymology.</emphasis>
Named after Jadwiga Lorenc-Brudecka, an entomologist from the Nature Education Centre, Jagiellonian University (Kraków,
<collectingCountry id="A4C084EAFF9AFFEF1FBF88EAFDB7F85E" box="[469,551,1957,1984]" name="Poland" pageId="2" pageNumber="51">Poland</collectingCountry>
), who has taken part in many
<taxonomicName id="1BD7BFF9FF9AFFEF1D1488EAFB95F821" authorityName="Linnaeus" authorityYear="1758" box="[894,1029,1957,1983]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="2" pageNumber="51" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
projects, especially those dedicated to the family
<taxonomicName id="1BD7BFF9FF9AFFEF1F478886FE20F87D" authorityName="Leach" authorityYear="1815" box="[301,432,1993,2019]" class="Insecta" family="Lycaenidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="family">Lycaenidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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