254 lines
28 KiB
XML
254 lines
28 KiB
XML
<document id="730743B5AD3B1CC14C8ED90B9C47E622" ID-DOI="10.5281/zenodo.5179307" ID-ISSN="1942-1354" ID-Zenodo-Dep="5179307" ID-ZooBank="1BC176E9-DD01-4279-8CCE-BFAC23948502" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628323712886" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2014" docId="2C5F8794FFC4720CFF0722AB5212FC3D" docLanguage="en" docName="InsectaMundi.0369.1-6.pdf" docOrigin="Insecta Mundi 2014 (369)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Parochodaeus perdidus Paulsen 2014, new species" docType="treatment" docVersion="4" lastPageNumber="5" masterDocId="D066FFECFFC1720AFFC7226A5171FFA7" masterDocTitle="Correction of the misidentifications and confusion surrounding Ochodaeus grandiceps Fairmaire, 1897 (Coleoptera: Ochodaeidae), and the description of a new species of ochodaeid from Cuba" masterLastPageNumber="6" masterPageNumber="1" pageNumber="4" updateTime="1699043076186" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="B87143A436E4F496A4F4DBE50F28EE02" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BC5BAF44F034B41B68A78C6C693E567E">
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<mods:title id="D077365CF9F39458219BE5D5DA160D85">Correction of the misidentifications and confusion surrounding Ochodaeus grandiceps Fairmaire, 1897 (Coleoptera: Ochodaeidae), and the description of a new species of ochodaeid from Cuba</mods:title>
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</mods:titleInfo>
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<mods:name id="CFB4E134B5C53FF5F8E413E3FEA261F6" type="personal">
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<mods:role id="D7DE8E1141943CAAAC9EB555BE28C7DA">
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<mods:roleTerm id="6AE795E3D0D5620D92D63C257A32C8C3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1B58B5E2DB58264B3B9A636D28CB9FF7">Paulsen, M. J.</mods:namePart>
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<mods:typeOfResource id="0EC1922AB27049A59FA30D1EDD7D8663">text</mods:typeOfResource>
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<mods:titleInfo id="6390A90490B69610A81D480B02ED8488">
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<mods:title id="2BE883EB844837174131DF98A4CF6402">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="F9CEBB5667233A040C46DDF02D691702">
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<mods:date id="4B994269DF19EB0FA5BC872CBD9580E6">2014</mods:date>
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<mods:detail id="DC0C82E5CD1C18B2EB4D3F82FF8789AA" type="pubDate">
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<mods:number id="DDA09BAF1FC17262BB5668605D5F1B9A">2014-08-29</mods:number>
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</mods:detail>
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<mods:detail id="A7083967DFE5559E40FEA768E753495B" type="volume">
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<mods:number id="050C2D4433A07C62F7F47DAF0D1DEE7A">2014</mods:number>
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</mods:detail>
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<mods:detail id="1CFF6CBCBCD690427DA2BFE14BAE70E9" type="issue">
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<mods:number id="2A5532B349FBFDE158F9321F5567E62B">369</mods:number>
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</mods:detail>
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<mods:start id="2ED5E158CD1BE550B023B1D5BA98696F">1</mods:start>
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<mods:end id="8887237442D5C86503E9834C04E55AA1">6</mods:end>
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</mods:extent>
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<mods:classification id="B7FE93CF1C3E612A4F74E0A132FB9390">journal article</mods:classification>
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<mods:identifier id="8C9E6956DBDCA1E0A1F3D732F5F29931" type="DOI">10.5281/zenodo.5179307</mods:identifier>
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<mods:identifier id="E738F612E35DD19FB90B3760C40D0A3F" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="ADC4257455CA2AA528B1889F91B61E76" type="Zenodo-Dep">5179307</mods:identifier>
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<mods:identifier id="137FA32C2DC2BE8637F0094933D15803" type="ZooBank">1BC176E9-DD01-4279-8CCE-BFAC23948502</mods:identifier>
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</mods:mods>
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<treatment id="2C5F8794FFC4720CFF0722AB5212FC3D" ID-DOI="http://doi.org/10.5281/zenodo.5189973" ID-Zenodo-Dep="5189973" LSID="urn:lsid:plazi:treatment:2C5F8794FFC4720CFF0722AB5212FC3D" httpUri="http://treatment.plazi.org/id/2C5F8794FFC4720CFF0722AB5212FC3D" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
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<subSubSection id="ECEC6509FFC4720FFF0722AB5241FF7E" box="[192,816,193,218]" pageId="5" pageNumber="4" type="nomenclature">
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||
<paragraph id="A4493682FFC4720FFF0722AB5241FF7E" blockId="5.[192,816,193,218]" box="[192,816,193,218]" pageId="5" pageNumber="4">
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||
<heading id="FF0181EEFFC4720FFF0722AB5241FF7E" bold="true" box="[192,816,193,218]" fontSize="10" level="2" pageId="5" pageNumber="4" reason="3">
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||
<emphasis id="9682EA90FFC4720FFF0722AB5241FF7E" bold="true" box="[192,816,193,218]" pageId="5" pageNumber="4">
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||
<taxonomicName id="63F64D01FFC4720FFF0722AB530DFF7D" ID-CoL="75RFQ" authority="Paulsen" authorityName="Paulsen" authorityYear="2014" box="[192,636,193,218]" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perdidus" status="sp. nov.">
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<emphasis id="9682EA90FFC4720FFF0722AB5374FF7D" bold="true" box="[192,517,193,218]" italics="true" pageId="5" pageNumber="4">Parochodaeus perdidus</emphasis>
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Paulsen
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</taxonomicName>
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,
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<taxonomicNameLabel id="8DB157EBFFC4720FFD4C22A85241FF7E" box="[651,816,193,218]" pageId="5" pageNumber="4" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="ECEC6509FFC4720FFF07236854D1FE9D" pageId="5" pageNumber="4" type="materials_examined">
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<paragraph id="A4493682FFC4720FFF07236854D1FE9D" blockId="5.[192,1440,258,314]" pageId="5" pageNumber="4">
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<materialsCitation id="149E3CDFFFC4720FFF07236854D1FE9D" collectingDate="1944-10-18" collectionCode="USNM, T" country="Cuba" county="Paulsen" location="Vista Alegre" municipality="Ramsden" pageId="5" pageNumber="4" specimenCount="2" typeStatus="holotype">
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<emphasis id="9682EA90FFC4720FFF07236850F7FEBD" bold="true" box="[192,390,258,282]" pageId="5" pageNumber="4">
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<typeStatus id="7B4D8820FFC4720FFF072368504FFEBD" box="[192,318,258,282]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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(
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<collectionCode id="C2E7AE47FFC4720FFE532368509FFEBD" box="[404,494,258,282]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="4" type="Museum">USNM</collectionCode>
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), labeled: a) “[
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<date id="D0481042FFC4720FFD532368525CFEBD" box="[660,813,258,282]" pageId="5" pageNumber="4" value="1944-10-18">
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<collectingDate id="C00CE9AAFFC4720FFD532368525CFEBD" box="[660,813,258,282]" pageId="5" pageNumber="4" value="1944-10-18">18-Oct-1944</collectingDate>
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</date>
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] /
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<location id="A1296059FFC4720FFC81236852ACFEBD" LSID="urn:lsid:plazi:treatment:2C5F8794FFC4720CFF0722AB5212FC3D:A1296059FFC4720FFC81236852ACFEBD" box="[838,989,258,282]" country="Cuba" county="Paulsen" municipality="Ramsden" name="Vista Alegre" pageId="5" pageNumber="4">Vista Alegre</location>
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/ Stgo.
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<collectingCountry id="DCE17612FFC4720FFBF423685506FEBD" box="[1075,1143,258,282]" name="Cuba" pageId="5" pageNumber="4">Cuba</collectingCountry>
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, at light / C.
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<collectionCode id="C2E7AE47FFC4720FFAC02368546DFEBD" box="[1287,1308,258,282]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="4">T</collectionCode>
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.
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<collectingMunicipality id="442DACF8FFC4720FFAE1236854EDFEBD" box="[1318,1436,258,282]" pageId="5" pageNumber="4">Ramsden</collectingMunicipality>
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, Coll.”; b) male symbol; c) on red paper, “
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<typeStatus id="7B4D8820FFC4720FFD642348524FFE9D" box="[675,830,290,314]" pageId="5" pageNumber="4" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="63F64D01FFC4720FFC9523485500FE9D" authorityName="Paulsen" authorityYear="2014" box="[850,1137,290,314]" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perdidus">
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<emphasis id="9682EA90FFC4720FFC9523485500FE9D" box="[850,1137,290,314]" italics="true" pageId="5" pageNumber="4">Parochodaeus perdidus</emphasis>
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</taxonomicName>
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/
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<collectingCounty id="4D284E0EFFC4720FFB422348559AFE9D" box="[1157,1259,290,314]" pageId="5" pageNumber="4">Paulsen</collectingCounty>
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, 2014” (
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<figureCitation id="3CCD2A07FFC4720FFA8C234854E0FE9D" box="[1355,1425,290,314]" captionStart="Figures 2–4" captionStartId="4.[192,288,1768,1790]" captionTargetBox="[208,1440,661,1756]" captionTargetId="figure-224@4.[192,1440,661,1757]" captionTargetPageId="4" captionText="Figures 2–4. Parochodaeus perdidus, n.sp. 2) Dorsal habitus of holotype. 3) Holotype labels. 4) Oblique lateral view of holotype, showing the ventrally projected pollex of the right protibia in lateral view (arrow), and bituberculate clypeus." figureDoi="http://doi.org/10.5281/zenodo.5179311" httpUri="https://zenodo.org/record/5179311/files/figure.png" pageId="5" pageNumber="4">Fig. 3</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="ECEC6509FFC4720FFF07230B52EFFD5D" pageId="5" pageNumber="4" type="description">
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<paragraph id="A4493682FFC4720FFF07230B52EFFD5D" blockId="5.[192,1441,353,762]" pageId="5" pageNumber="4">
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<emphasis id="9682EA90FFC4720FFF07230B50A5FEDD" bold="true" box="[192,468,353,378]" pageId="5" pageNumber="4">
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Description.
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<emphasis id="9682EA90FFC4720FFEA9230B50BCFEDD" bold="true" box="[366,461,353,378]" italics="true" pageId="5" pageNumber="4">Length</emphasis>
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:
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</emphasis>
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7.5 mm.
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<emphasis id="9682EA90FFC4720FFDF8230B53E9FEDD" bold="true" box="[575,664,353,378]" pageId="5" pageNumber="4">
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<emphasis id="9682EA90FFC4720FFDF8230B53E0FEDD" bold="true" box="[575,657,353,378]" italics="true" pageId="5" pageNumber="4">Width</emphasis>
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:
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</emphasis>
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4.0 mm.
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<emphasis id="9682EA90FFC4720FFCC4230B5223FEDD" bold="true" box="[771,850,353,378]" pageId="5" pageNumber="4">
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<emphasis id="9682EA90FFC4720FFCC4230B523AFEDD" bold="true" box="[771,843,353,378]" italics="true" pageId="5" pageNumber="4">Head</emphasis>
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:
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</emphasis>
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Surface shiny, everywhere sparsely, setigerously punctate. Frons with vertex tuberculate. Clypeus trapezoidal, long (length equal to 1/2 width), anterior margin with an erect, horn-like tubercle on each end (
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<figureCitation id="3CCD2A07FFC4720FFC8923C852E7FE1D" box="[846,918,418,442]" captionStart="Figures 2–4" captionStartId="4.[192,288,1768,1790]" captionTargetBox="[208,1440,661,1756]" captionTargetId="figure-224@4.[192,1440,661,1757]" captionTargetPageId="4" captionText="Figures 2–4. Parochodaeus perdidus, n.sp. 2) Dorsal habitus of holotype. 3) Holotype labels. 4) Oblique lateral view of holotype, showing the ventrally projected pollex of the right protibia in lateral view (arrow), and bituberculate clypeus." figureDoi="http://doi.org/10.5281/zenodo.5179311" httpUri="https://zenodo.org/record/5179311/files/figure.png" pageId="5" pageNumber="4">Fig. 4</figureCitation>
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), margin between tubercles distinct, thickened. Labrum emarginate. Mandibles rounded externally; left mandible with bicuspid tooth just behind apex and 1 strong basal tooth; right mandible with acute apex and 1 strong subapical tooth. Mentum with distinct longitudinal furrow in anterior half.
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<emphasis id="9682EA90FFC4720FFCD4206852D3FDBD" bold="true" box="[787,930,514,538]" pageId="5" pageNumber="4">
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<emphasis id="9682EA90FFC4720FFCD4206852EAFDBE" bold="true" box="[787,923,514,537]" italics="true" pageId="5" pageNumber="4">Pronotum</emphasis>
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:
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</emphasis>
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Form evenly convex. Surface with tile-like tubercles; tubercles large, setigerous with short setae; surface between tubercles punctate; punctures moderate, lacking setae.
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<emphasis id="9682EA90FFC4720FFE2A202B533BFDFD" bold="true" box="[493,586,577,602]" pageId="5" pageNumber="4">
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<emphasis id="9682EA90FFC4720FFE2A202B5332FDFD" bold="true" box="[493,579,577,602]" italics="true" pageId="5" pageNumber="4">Elytra</emphasis>
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:
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</emphasis>
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Striae moderately impressed, punctate with moderate punctures lacking setae. Interstriae tuberculate, intervals 1–2 with tubercles larger than strial punctures, elsewhere tubercles moderate.
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<emphasis id="9682EA90FFC4720FFE7820E85375FD3D" bold="true" box="[447,516,642,666]" pageId="5" pageNumber="4">
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<emphasis id="9682EA90FFC4720FFE7820E8508CFD3E" bold="true" box="[447,509,642,665]" italics="true" pageId="5" pageNumber="4">Legs</emphasis>
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:
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</emphasis>
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Protibia tridentate with basal tooth strongly developed, median; internal apical tooth (pollex) large, obtuse, directed ventrally (
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<figureCitation id="3CCD2A07FFC4720FFC9520C852EBFD1D" box="[850,922,674,698]" captionStart="Figures 2–4" captionStartId="4.[192,288,1768,1790]" captionTargetBox="[208,1440,661,1756]" captionTargetId="figure-224@4.[192,1440,661,1757]" captionTargetPageId="4" captionText="Figures 2–4. Parochodaeus perdidus, n.sp. 2) Dorsal habitus of holotype. 3) Holotype labels. 4) Oblique lateral view of holotype, showing the ventrally projected pollex of the right protibia in lateral view (arrow), and bituberculate clypeus." figureDoi="http://doi.org/10.5281/zenodo.5179311" httpUri="https://zenodo.org/record/5179311/files/figure.png" pageId="5" pageNumber="4">Fig. 4</figureCitation>
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). All femora unarmed. Metatibia straight, narrow (>4x longer than wide) expanding gradually to apex. Metatarsomere 1 not greatly enlarged.
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<emphasis id="9682EA90FFC4720FFF07208B5034FD5D" bold="true" box="[192,325,737,762]" pageId="5" pageNumber="4">
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<emphasis id="9682EA90FFC4720FFF07208B504FFD5D" bold="true" box="[192,318,737,762]" italics="true" pageId="5" pageNumber="4">Abdomen</emphasis>
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:
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</emphasis>
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Stridulatory peg present. Pygidium tuberculate.
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</paragraph>
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</subSubSection>
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<subSubSection id="ECEC6509FFC4720FFF07214B521BFCDD" pageId="5" pageNumber="4" type="diagnosis">
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<paragraph id="A4493682FFC4720FFF07214B521BFCDD" blockId="5.[192,1441,801,890]" pageId="5" pageNumber="4">
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<emphasis id="9682EA90FFC4720FFF07214B5021FC9E" bold="true" box="[192,336,801,825]" pageId="5" pageNumber="4">Diagnosis.</emphasis>
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The protibial pollex (“thumb”) of this species is large and obtuse (
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<figureCitation id="3CCD2A07FFC4720FFB44214855BCFC9D" box="[1155,1229,802,826]" captionStart="Figures 2–4" captionStartId="4.[192,288,1768,1790]" captionTargetBox="[208,1440,661,1756]" captionTargetId="figure-224@4.[192,1440,661,1757]" captionTargetPageId="4" captionText="Figures 2–4. Parochodaeus perdidus, n.sp. 2) Dorsal habitus of holotype. 3) Holotype labels. 4) Oblique lateral view of holotype, showing the ventrally projected pollex of the right protibia in lateral view (arrow), and bituberculate clypeus." figureDoi="http://doi.org/10.5281/zenodo.5179311" httpUri="https://zenodo.org/record/5179311/files/figure.png" pageId="5" pageNumber="4">Fig. 4</figureCitation>
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), which is unlike any other
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<taxonomicName id="63F64D01FFC4720FFEFB2128533CFCFD" box="[316,589,834,858]" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="9682EA90FFC4720FFEFB21285096FCFD" box="[316,487,834,858]" italics="true" pageId="5" pageNumber="4">Parochodaeus</emphasis>
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species.
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</taxonomicName>
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Some members of the genus from South America display pollices, but they are generally much narrower and acute or conical.
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</paragraph>
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</subSubSection>
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<subSubSection id="ECEC6509FFC4720FFF0721C85549FB9D" pageId="5" pageNumber="4" type="etymology">
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<paragraph id="A4493682FFC4720FFF0721C85549FB9D" blockId="5.[192,1440,930,1082]" pageId="5" pageNumber="4">
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<emphasis id="9682EA90FFC4720FFF0721C8502DFC1D" bold="true" box="[192,348,930,954]" pageId="5" pageNumber="4">Etymology.</emphasis>
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The species is named ‘
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<taxonomicName id="63F64D01FFC4720FFDB621C853AFFC1D" authorityName="Paulsen" authorityYear="2014" box="[625,734,930,954]" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perdidus">
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<emphasis id="9682EA90FFC4720FFDB621C853AFFC1D" box="[625,734,930,954]" italics="true" pageId="5" pageNumber="4">perdidus</emphasis>
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</taxonomicName>
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’, a Latinized form of the Spanish word ‘
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<emphasis id="9682EA90FFC4720FFB0421C85457FC1D" box="[1219,1318,930,954]" italics="true" pageId="5" pageNumber="4">perdido’</emphasis>
|
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, meaning lost. I chose this name primarily to allude to the isolation achieved by the founding individuals, however they arrived in the West Indies. No species of
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<taxonomicName id="63F64D01FFC4720FFCF4218852AFFC5D" authorityName="Nikolajev" authorityYear="1995" box="[819,990,994,1018]" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="9682EA90FFC4720FFCF4218852AFFC5D" box="[819,990,994,1018]" italics="true" pageId="5" pageNumber="4">Parochodaeus</emphasis>
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</taxonomicName>
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are known from the mainland areas nearest to
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<collectingCountry id="DCE17612FFC4720FFE84266850F9FBBD" box="[323,392,1026,1050]" name="Cuba" pageId="5" pageNumber="4">Cuba</collectingCountry>
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. Secondarily, the name denotes the confusion surrounding the identity of the species. It is an adjective in the nominative singular and is masculine in gender.
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</paragraph>
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</subSubSection>
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<subSubSection id="ECEC6509FFC4720CFF07260B50E6FEBD" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="distribution">
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<paragraph id="A4493682FFC4720FFF07260B5533FB3D" blockId="5.[192,1440,1121,1178]" pageId="5" pageNumber="4">
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<emphasis id="9682EA90FFC4720FFF07260B5005FBDE" bold="true" box="[192,372,1121,1145]" pageId="5" pageNumber="4">Distribution.</emphasis>
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(
|
||
<figureCitation id="3CCD2A07FFC4720FFE42260850BCFBDE" box="[389,461,1121,1146]" captionStart="Figure 5" captionStartId="5.[192,276,1838,1860]" captionTargetBox="[192,1440,1213,1819]" captionTargetId="figure-412@5.[192,1440,1213,1819]" captionTargetPageId="5" captionText="Figure 5. Distribution of Parochodaeus specimens from Cuba (circles) with the type locality of P. perdidus, n.sp., indicated in orange. Provincial abbreviations shown: CF = Cienfuegos, CM = Camagüey, GU = Guantánamo, and SC = Santiago de Cuba." figureDoi="http://doi.org/10.5281/zenodo.5179313" httpUri="https://zenodo.org/record/5179313/files/figure.png" pageId="5" pageNumber="4">Fig. 5</figureCitation>
|
||
)
|
||
<emphasis id="9682EA90FFC4720FFE19260B5348FBDE" bold="true" box="[478,569,1121,1145]" pageId="5" pageNumber="4">
|
||
<collectingCountry id="DCE17612FFC4720FFE19260B5345FBDE" box="[478,564,1121,1145]" name="Cuba" pageId="5" pageNumber="4">CUBA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<collectingRegion id="6632F860FFC4720FFD85260B539BFBDD" box="[578,746,1121,1146]" country="Cuba" name="Camaguey" pageId="5" pageNumber="4">CAMAGÜEY</collectingRegion>
|
||
: Galbis (1);
|
||
<collectingRegion id="6632F860FFC4720FFCB92608554FFBDD" box="[894,1086,1122,1146]" country="Cuba" name="Cienfuegos" pageId="5" pageNumber="4">CIENFUEGOS</collectingRegion>
|
||
: Soledad, nr.
|
||
<collectingRegion id="6632F860FFC4720FFB2F26085401FBDD" box="[1256,1392,1122,1146]" country="Cuba" name="Cienfuegos" pageId="5" pageNumber="4">Cienfuegos</collectingRegion>
|
||
(1);
|
||
<collectingRegion id="6632F860FFC4720FFF0726EB50E5FB3D" box="[192,404,1153,1178]" country="Cuba" name="Guantanamo" pageId="5" pageNumber="4">GUANTÁNAMO</collectingRegion>
|
||
: San Carlos (1);
|
||
<collectingRegion id="6632F860FFC4720FFD9B26E8521FFB3D" box="[604,878,1154,1178]" country="Cuba" name="Santiago de Cuba" pageId="5" pageNumber="4">SANTIAGO de CUBA</collectingRegion>
|
||
: Vista Alegre (4).
|
||
</paragraph>
|
||
<caption id="F089660AFFC4720FFF07254450B9F8D9" ID-DOI="http://doi.org/10.5281/zenodo.5179313" ID-Zenodo-Dep="5179313" httpUri="https://zenodo.org/record/5179313/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,276,1838,1860]" targetBox="[192,1440,1213,1819]" targetPageId="5">
|
||
<paragraph id="A4493682FFC4720FFF07254450B9F8D9" blockId="5.[192,1440,1838,1918]" pageId="5" pageNumber="4">
|
||
<emphasis id="9682EA90FFC4720FFF072544505EF8E3" bold="true" box="[192,303,1838,1860]" pageId="5" pageNumber="4">Figure 5.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="63F64D01FFC4720FFE1925455309F8E3" authorityName="Nikolajev" authorityYear="1995" box="[478,632,1839,1860]" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9682EA90FFC4720FFE1925455309F8E3" box="[478,632,1839,1860]" italics="true" pageId="5" pageNumber="4">Parochodaeus</emphasis>
|
||
</taxonomicName>
|
||
specimens from Cuba (circles) with the type locality of
|
||
<taxonomicName id="63F64D01FFC4720FFB1E25455429F8E3" authority="Paulsen, 2014" authorityName="Paulsen" authorityYear="2014" box="[1241,1368,1839,1860]" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perdidus" status="sp. nov.">
|
||
<emphasis id="9682EA90FFC4720FFB1E25455429F8E3" box="[1241,1368,1839,1860]" italics="true" pageId="5" pageNumber="4">P. perdidus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="8DB157EBFFC4720FFAA3254554EAF8E3" box="[1380,1435,1839,1860]" pageId="5" pageNumber="4" rank="species">n.sp.</taxonomicNameLabel>
|
||
, indicated in orange. Provincial abbreviations shown: CF = Cienfuegos, CM = Camagüey, GU = Guantánamo, and SC = Santiago de Cuba.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A4493682FFC7720CFF0722A850E6FEBD" blockId="6.[192,1440,193,282]" pageId="6" pageNumber="5">
|
||
<emphasis id="9682EA90FFC7720CFF0722A85370FF7E" bold="true" box="[192,513,193,218]" pageId="6" pageNumber="5">Temporal Distribution.</emphasis>
|
||
May (2), July (1), October (1), November (3). As with other species of
|
||
<taxonomicName id="63F64D01FFC7720CFA9A22A85040FF5D" authorityName="Nikolajev" authorityYear="1995" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9682EA90FFC7720CFA9A22A85040FF5D" italics="true" pageId="6" pageNumber="5">Parochodaeus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="C0674B73FFC7720CFE86228853E3FF5D" author="Ratcliffe, B. C. & M. J. Paulsen" box="[321,658,226,250]" pageId="6" pageNumber="5" pagination="1 - 570" refId="ref3115" refString="Ratcliffe, B. C., and M. J. Paulsen. 2008. The scarabaeoid beetles of Nebraska. Bulletin of the University of Nebraska State Museum 22: 1 - 570." type="journal article" year="2008">Ratcliffe and Paulsen 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="C0674B73FFC7720CFD592288529AFF5D" author="Paulsen, M. J. & F. C. Ocampo" box="[670,1003,226,250]" pageId="6" pageNumber="5" pagination="7 - 30" refId="ref3042" refString="Paulsen, M. J., and F. C. Ocampo. 2012. The Ochodaeidae of Argentina (Coleoptera, Scarabaeoidea). ZooKeys 174: 7 - 30." type="journal article" year="2012">Paulsen and Ocampo 2012</bibRefCitation>
|
||
), the Cuban species displays a long period of activity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ECEC6509FFC7720CFF0723285212FC3D" pageId="6" pageNumber="5" type="discussion">
|
||
<paragraph id="A4493682FFC7720CFF0723285212FC3D" blockId="6.[192,1441,322,922]" pageId="6" pageNumber="5">
|
||
<emphasis id="9682EA90FFC7720CFF0723285035FEFD" bold="true" box="[192,324,322,346]" pageId="6" pageNumber="5">Remarks.</emphasis>
|
||
The species is based solely on the largest male from Vista Alegre, although all specimens examined are tentatively and conservatively assigned to this single species. Unfortunately, there are not enough specimens available for study to be confident that the variation present in the remaining material is intraspecific. Although the armature of the internal sac of the male genitalia can be useful for species determination the
|
||
<specimenCount id="B2F0FD0BFFC7720CFD9E23A8539AFE7D" box="[601,747,450,474]" pageId="6" pageNumber="5" type="male">three males</specimenCount>
|
||
available did not display any obvious differences in this structure. Many species of
|
||
<taxonomicName id="63F64D01FFC7720CFDCB238853C6FE5D" authorityName="Nikolajev" authorityYear="1995" box="[524,695,482,506]" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9682EA90FFC7720CFDCB238853C6FE5D" box="[524,695,482,506]" italics="true" pageId="6" pageNumber="5">Parochodaeus</emphasis>
|
||
</taxonomicName>
|
||
demonstrate a large range in body size, and this can lead to differences in armature and punctation. The three smaller specimens from the type locality (
|
||
<specimenCount id="B2F0FD0BFFC7720CFAE2206854EDFDBD" box="[1317,1436,514,538]" pageId="6" pageNumber="5" type="female">2 females</specimenCount>
|
||
,
|
||
<specimenCount id="B2F0FD0BFFC7720CFF0720485063FD9D" box="[192,274,546,570]" pageId="6" pageNumber="5" type="male">1 male</specimenCount>
|
||
) have much smoother pronota and less well-developed pollices, but these differences may be related to their smaller size (
|
||
<quantity id="630E9B67FFC7720CFDC3202B5304FDFD" box="[516,629,577,602]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="5" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5 – 6 mm</quantity>
|
||
). The large female specimen from
|
||
<collectingRegion id="6632F860FFC7720CFBD5202855E4FDFD" box="[1042,1173,578,602]" country="Cuba" name="Camaguey" pageId="6" pageNumber="5">Camagüey</collectingRegion>
|
||
(7.6 mm) most closely approximates the
|
||
<typeStatus id="7B4D8820FFC7720CFE642008537EFDDD" box="[419,527,610,634]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
, but has the mentum more weakly furrowed and the elytra less strongly tuberculate. Because the form of the mentum usually is a reliable character for distinguishing species of
|
||
<taxonomicName id="63F64D01FFC7720CFF2720C850FAFD1D" authorityName="Nikolajev" authorityYear="1995" box="[224,395,674,698]" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9682EA90FFC7720CFF2720C850FAFD1D" box="[224,395,674,698]" italics="true" pageId="6" pageNumber="5">Parochodaeus</emphasis>
|
||
</taxonomicName>
|
||
and the specimen is from quite far away (
|
||
<figureCitation id="3CCD2A07FFC7720CFC6520C8529DFD1E" box="[930,1004,673,698]" captionStart="Figure 5" captionStartId="5.[192,276,1838,1860]" captionTargetBox="[192,1440,1213,1819]" captionTargetId="figure-412@5.[192,1440,1213,1819]" captionTargetPageId="5" captionText="Figure 5. Distribution of Parochodaeus specimens from Cuba (circles) with the type locality of P. perdidus, n.sp., indicated in orange. Provincial abbreviations shown: CF = Cienfuegos, CM = Camagüey, GU = Guantánamo, and SC = Santiago de Cuba." figureDoi="http://doi.org/10.5281/zenodo.5179313" httpUri="https://zenodo.org/record/5179313/files/figure.png" pageId="6" pageNumber="5">Fig. 5</figureCitation>
|
||
), this female specimen may not be conspecific. The partial specimen from
|
||
<collectingRegion id="6632F860FFC7720CFD5E20A8524AFD7D" box="[665,827,706,730]" country="Cuba" name="Guantanamo" pageId="6" pageNumber="5">Guantánamo</collectingRegion>
|
||
lacks a head and thorax but does display a similar stridulatory peg and large elytral tubercles. In the female specimen from
|
||
<collectingRegion id="6632F860FFC7720CFB9E20885590FD5D" box="[1113,1249,738,762]" country="Cuba" name="Cienfuegos" pageId="6" pageNumber="5">Cienfuegos</collectingRegion>
|
||
the mentum is weakly notched anteriorly and not distinctly longitudinally furrowed, with no evidence of a pollex; this specimen is the most likely to belong to another species, however no males from this locality are known. More specimens are necessary, and especially males, to discern if there may be additional species of
|
||
<taxonomicName id="63F64D01FFC7720CFF072108501AFCDD" authorityName="Nikolajev" authorityYear="1995" box="[192,363,866,890]" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9682EA90FFC7720CFF072108501AFCDD" box="[192,363,866,890]" italics="true" pageId="6" pageNumber="5">Parochodaeus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="DCE17612FFC7720CFE53210850A8FCDD" box="[404,473,866,890]" name="Cuba" pageId="6" pageNumber="5">Cuba</collectingCountry>
|
||
. Because it is possible that more than one species is present, specimens of the remaining material are not included in the type series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |