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<document id="75A3E05EA622A95C1405596C3C6D0752" ID-DOI="10.1051/acarologia/20162252" ID-ISSN="2107-7207" ID-Zenodo-Dep="5403994" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630342510265" checkinUser="marcus" docAuthor="Ermilov, S. G. &amp; Friedrich, S." docDate="2016" docId="1A38721C3451FFA934ABFD73FECF32AF" docLanguage="en" docName="Acarologia.2016.56.3.379-391.pdf" docOrigin="Acarologia 56 (3)" docSource="http://dx.doi.org/10.1051/acarologia/20162252" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Hammerella (Woasella) huanucoensis Ermilov 2016, n. sp." docType="treatment" docVersion="3" lastPageNumber="389" masterDocId="E6010A643456FFA23446FFDCFF91343E" masterDocTitle="Additions to the oppioid oribatid mite fauna of Peru (Acari, Oribatida, Oppioidea)" masterLastPageNumber="391" masterPageNumber="379" pageNumber="385" updateTime="1699236003866" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:title id="61684E07092AC6DB6DC958A607F1FB74">Additions to the oppioid oribatid mite fauna of Peru (Acari, Oribatida, Oppioidea)</mods:title>
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<paragraph id="922EC30A3451FFA534ABFD73FDB336D4" blockId="7.[237,683,687,781]" pageId="7" pageNumber="385">
<heading id="C96674663451FFA534ABFD73FDB336D4" bold="true" centered="true" fontSize="10" level="2" pageId="7" pageNumber="385" reason="2">
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<taxonomicName id="5591B8893451FFA534ABFD73FE4936D4" ID-CoL="8TBG4" authority="Ermilov" authorityName="Ermilov" authorityYear="2016" class="Arachnida" family="Granuloppiidae" genus="Hammerella" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="385" phylum="Arthropoda" rank="species" species="huanucoensis" status="sp. nov." subGenus="Woasella">
<emphasis id="A0E51F183451FFA534ABFD73FEEE36F9" bold="true" box="[237,383,687,711]" italics="true" pageId="7" pageNumber="385">Hammerella</emphasis>
(
<emphasis id="A0E51F183451FFA535C8FD73FE6C36F9" bold="true" box="[398,509,687,711]" italics="true" pageId="7" pageNumber="385">Woasella</emphasis>
)
<emphasis id="A0E51F183451FFA5364AFD73FD3A36F9" bold="true" box="[524,683,687,711]" italics="true" pageId="7" pageNumber="385">huanucoensis</emphasis>
Ermilov
</taxonomicName>
<taxonomicNameLabel id="BBD6A2633451FFA53599FD0EFDB336D4" box="[479,546,722,746]" pageId="7" pageNumber="385" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="DA8B90813451FFA93539FD29FECF32AF" lastPageId="11" lastPageNumber="389" pageId="7" pageNumber="385" type="description">
<paragraph id="922EC30A3451FFA53539FD29FD8B3733" blockId="7.[237,683,687,781]" box="[383,538,757,781]" pageId="7" pageNumber="385">
<emphasis id="A0E51F183451FFA53539FD29FD8B3733" bold="true" box="[383,538,757,781]" pageId="7" pageNumber="385">
(
<figureCitation id="0AAADF8F3451FFA535C0FD29FD833733" box="[390,530,757,781]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="8.[159,171,1869,1888]" captionStartId-1="9.[159,171,1752,1771]" captionStartId-2="10.[159,171,1738,1757]" captionTargetBox-1="[288,1311,497,1616]" captionTargetBox-2="[382,1274,550,1627]" captionTargetId-1="figure-6@9.[242,1346,415,1720]" captionTargetId-2="figure-5@10.[242,1346,457,1706]" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 4: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A dorsal view (legs except basal parts of legs II and III not illustrated); B ventral view (palps and legs except basal parts of legs IV not illustrated). Scale bar 100 µm." captionText-1="FIGURE 5: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A lateral view, right side (gnathosoma and legs except basal part of legs I, III and IV not illustrated); B tibia (solenidion not illustrated) and basal part of tarsus of leg I, right, antiaxial view; C tibia (solenidion except basal part not illustrated) and tarsus of leg IV, right, paraxial view. Scale bar (A) 100 µm, scale bar (B, C) 20 µm." captionText-2="FIGURE 6: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A subcapitulum, ventral view, and palptrochanter; B palp (trochanter not illustrated), left, ventral view; C chelicera, left, paraxial view. Scale bar (A, C; B) 16 µm." figureDoi-1="http://doi.org/10.5281/zenodo.5404000" figureDoi-2="http://doi.org/10.5281/zenodo.5404002" httpUri-1="https://zenodo.org/record/5404000/files/figure.png" httpUri-2="https://zenodo.org/record/5404002/files/figure.png" pageId="7" pageNumber="385">Figures 46</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="922EC30A3451FFA534D9FCF1FDB5309A" blockId="7.[159,762,813,1948]" pageId="7" pageNumber="385">
Diagnosis — Body size: 298 332 × 166 182. Rostrum with median indentation. Costulae slightly developed. Transcostula absent. Lamellar setae shorter and thinner than rostral and interlamellar setae, all barbed. Exobothridial setae shortest. Bothridial setae with long stalk and shorter head, having five to six long cilia and some short barbs unilaterally. Ten pairs of notogastral setae present, setae
<emphasis id="A0E51F183451FFA5348DFB9AFF473063" box="[203,214,1094,1117]" italics="true" pageId="7" pageNumber="385">c</emphasis>
minute, other setae of medium size, smooth. Epimeral setae setiform, slightly barbed. Discidia absent. Anogenital setae smooth.
</paragraph>
<paragraph id="922EC30A3451FFA53483FB6BFD6B3165" blockId="7.[159,762,813,1948]" pageId="7" pageNumber="385">
Description — Measurements Body length: 332 (
<typeStatus id="4D2A7DA83451FFA5349DFB06FED630CC" box="[219,327,1242,1266]" pageId="7" pageNumber="385" type="holotype">holotype</typeStatus>
: female), 298 332 (
<specimenCount id="849708833451FFA53611FB06FD6730CC" box="[599,758,1242,1266]" pageId="7" pageNumber="385" type="generic" typeStatus="paratypes">16 paratypes</specimenCount>
:
<specimenCount id="849708833451FFA534D9FB21FED6312B" box="[159,327,1277,1301]" pageId="7" pageNumber="385" type="female">seven females</specimenCount>
and
<specimenCount id="849708833451FFA535CDFB21FD80312B" box="[395,529,1277,1301]" pageId="7" pageNumber="385" type="male">nine males</specimenCount>
); notogaster width 182 (
<typeStatus id="4D2A7DA83451FFA53490FAFCFED33106" box="[214,322,1312,1336]" pageId="7" pageNumber="385" type="holotype">holotype</typeStatus>
), 166 182 (
<specimenCount id="849708833451FFA53595FAFCFDFF3106" box="[467,622,1312,1336]" pageId="7" pageNumber="385" type="generic" typeStatus="paratypes">16 paratypes</specimenCount>
). No differences between females and males in the body sizes.
</paragraph>
<paragraph id="922EC30A3451FFA53483FAB3FD1731CE" blockId="7.[159,762,813,1948]" pageId="7" pageNumber="385">
Integument (
<figureCitation id="0AAADF8F3451FFA53527FAB2FE5531B8" box="[353,452,1390,1414]" captionStart="FIGURE" captionStartId="8.[159,171,1869,1888]" captionText="FIGURE 4: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A dorsal view (legs except basal parts of legs II and III not illustrated); B ventral view (palps and legs except basal parts of legs IV not illustrated). Scale bar 100 µm." pageId="7" pageNumber="385">Figs 4A</figureCitation>
,
<figureCitation id="0AAADF8F3451FFA53592FAB2FE6631B8" box="[468,503,1390,1414]" captionStart="FIGURE" captionStartId="9.[159,171,1752,1771]" captionTargetBox="[288,1311,497,1616]" captionTargetId="figure-6@9.[242,1346,415,1720]" captionTargetPageId="9" captionText="FIGURE 5: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A lateral view, right side (gnathosoma and legs except basal part of legs I, III and IV not illustrated); B tibia (solenidion not illustrated) and basal part of tarsus of leg I, right, antiaxial view; C tibia (solenidion except basal part not illustrated) and tarsus of leg IV, right, paraxial view. Scale bar (A) 100 µm, scale bar (B, C) 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5404000" httpUri="https://zenodo.org/record/5404000/files/figure.png" pageId="7" pageNumber="385">5A</figureCitation>
) — Body color light brownish. Body surface punctate, lateral side and podosomal region with sparse tuberculate cerotegument (diameter tubercle up to 4).
</paragraph>
<paragraph id="922EC30A3451FFA53483F9DFFB643589" blockId="7.[159,762,813,1948]" lastBlockId="7.[826,1429,311,1949]" pageId="7" pageNumber="385">
Prodorsum (
<figureCitation id="0AAADF8F3451FFA53511F9DFFE253225" box="[343,436,1539,1563]" captionStart="FIGURE" captionStartId="8.[159,171,1869,1888]" captionText="FIGURE 4: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A dorsal view (legs except basal parts of legs II and III not illustrated); B ventral view (palps and legs except basal parts of legs IV not illustrated). Scale bar 100 µm." pageId="7" pageNumber="385">Figs 4A</figureCitation>
,
<figureCitation id="0AAADF8F3451FFA535F9F9DFFE703225" box="[447,481,1539,1563]" captionStart="FIGURE" captionStartId="9.[159,171,1752,1771]" captionTargetBox="[288,1311,497,1616]" captionTargetId="figure-6@9.[242,1346,415,1720]" captionTargetPageId="9" captionText="FIGURE 5: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A lateral view, right side (gnathosoma and legs except basal part of legs I, III and IV not illustrated); B tibia (solenidion not illustrated) and basal part of tarsus of leg I, right, antiaxial view; C tibia (solenidion except basal part not illustrated) and tarsus of leg IV, right, paraxial view. Scale bar (A) 100 µm, scale bar (B, C) 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5404000" httpUri="https://zenodo.org/record/5404000/files/figure.png" pageId="7" pageNumber="385">5A</figureCitation>
) — Rostrum with small median indentation (
<emphasis id="A0E51F183451FFA535EEF9FAFE5A3200" box="[424,459,1574,1598]" italics="true" pageId="7" pageNumber="385">ind</emphasis>
; well visible in frontal view). Costulae short, thin, parallel. Transcostula absent. Rostral (28), lamellar (18 20), interlamellar (32) and exobothridial (16 20) setae setiform, barbed;
<emphasis id="A0E51F183451FFA53545F96EFE8432F4" box="[259,277,1714,1738]" italics="true" pageId="7" pageNumber="385">le</emphasis>
and
<emphasis id="A0E51F183451FFA53510F96FFEFF32F4" box="[342,366,1715,1738]" italics="true" pageId="7" pageNumber="385">ex</emphasis>
thinnest,
<emphasis id="A0E51F183451FFA535AFF96EFE6A32F4" box="[489,507,1714,1738]" italics="true" pageId="7" pageNumber="385">le</emphasis>
located nearer to interlamellar setae, than to rostral setae. Transverse thin ridge (r) located posteriorly to insertions of rostral setae. Bothridial setae (53 57) with long stalks and shorter, elongate, rounded distally heads, having five to six of long (45 53) cilia and three to five of short barbs unilaterally. Longitudinal row of muscle sigilla present anteriad to bothridia and lateral to costulae. Interbothridial region with two pairs of muscle sigilla. Interbothridial tubercles absent. Postbothridial tubercles present.
</paragraph>
<paragraph id="922EC30A3451FFA53726FE17FCC03720" blockId="7.[826,1429,311,1949]" pageId="7" pageNumber="385">
Notogaster (
<figureCitation id="0AAADF8F3451FFA537BFFE16FBCF35DC" box="[1017,1118,458,482]" captionStart="FIGURE" captionStartId="8.[159,171,1869,1888]" captionText="FIGURE 4: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A dorsal view (legs except basal parts of legs II and III not illustrated); B ventral view (palps and legs except basal parts of legs IV not illustrated). Scale bar 100 µm." pageId="7" pageNumber="385">Figs 4A</figureCitation>
,
<figureCitation id="0AAADF8F3451FFA53036FE16FB0235DC" box="[1136,1171,458,482]" captionStart="FIGURE" captionStartId="9.[159,171,1752,1771]" captionTargetBox="[288,1311,497,1616]" captionTargetId="figure-6@9.[242,1346,415,1720]" captionTargetPageId="9" captionText="FIGURE 5: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A lateral view, right side (gnathosoma and legs except basal part of legs I, III and IV not illustrated); B tibia (solenidion not illustrated) and basal part of tarsus of leg I, right, antiaxial view; C tibia (solenidion except basal part not illustrated) and tarsus of leg IV, right, paraxial view. Scale bar (A) 100 µm, scale bar (B, C) 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5404000" httpUri="https://zenodo.org/record/5404000/files/figure.png" pageId="7" pageNumber="385">5A</figureCitation>
) — Anterior border straight. Humeral tubercles indistinct, directed to postbothridial tubercles. Ten pairs of notogastral setae present, setae
<emphasis id="A0E51F183451FFA5304DFDE8FB873675" box="[1035,1046,564,587]" italics="true" pageId="7" pageNumber="385">c</emphasis>
minute (2), other setae setiform, smooth, inserted in four parallel rows (except posterior setae). Setae
<emphasis id="A0E51F183451FFA5306FFDA6FBA736AF" box="[1065,1078,634,657]" italics="true" pageId="7" pageNumber="385">p</emphasis>
<subScript id="0E15C14F3451FFA53070FD5FFBAE36AA" attach="left" box="[1078,1087,643,660]" fontSize="7" pageId="7" pageNumber="385">2</subScript>
and
<emphasis id="A0E51F183451FFA530C4FDA6FB1E36AF" box="[1154,1167,634,657]" italics="true" pageId="7" pageNumber="385">p</emphasis>
<subScript id="0E15C14F3451FFA530C9FD5FFB0936AA" attach="left" box="[1167,1176,643,660]" fontSize="7" pageId="7" pageNumber="385">3</subScript>
(2024) shorter than
<emphasis id="A0E51F183451FFA5377CFD41FCD6368A" box="[826,839,669,692]" italics="true" pageId="7" pageNumber="385">p</emphasis>
<subScript id="0E15C14F3451FFA53701FD7AFCC13689" attach="left" box="[839,848,678,695]" fontSize="7" pageId="7" pageNumber="385">1</subScript>
(2428), other setae longest (3241) and thickest. All lyrifissures clearly visible. Opisthonotal gland openings located posterior to
<emphasis id="A0E51F183451FFA530D7FD3FFB3C36C4" box="[1169,1197,739,762]" italics="true" pageId="7" pageNumber="385">im</emphasis>
and distanced from it.
</paragraph>
<paragraph id="922EC30A3451FFA53726FCECFCFA3168" blockId="7.[826,1429,311,1949]" pageId="7" pageNumber="385">
Gnathosoma (
<figureCitation id="0AAADF8F3451FFA5304FFCECFBF73776" box="[1033,1126,816,840]" captionStart="FIGURE" captionStartId="8.[159,171,1869,1888]" captionText="FIGURE 4: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A dorsal view (legs except basal parts of legs II and III not illustrated); B ventral view (palps and legs except basal parts of legs IV not illustrated). Scale bar 100 µm." pageId="7" pageNumber="385">Figs 4B</figureCitation>
,
<figureCitation id="0AAADF8F3451FFA53033FCECFB083776" box="[1141,1177,816,840]" captionStart="FIGURE" captionStartId="9.[159,171,1752,1771]" captionTargetBox="[288,1311,497,1616]" captionTargetId="figure-6@9.[242,1346,415,1720]" captionTargetPageId="9" captionText="FIGURE 5: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A lateral view, right side (gnathosoma and legs except basal part of legs I, III and IV not illustrated); B tibia (solenidion not illustrated) and basal part of tarsus of leg I, right, antiaxial view; C tibia (solenidion except basal part not illustrated) and tarsus of leg IV, right, paraxial view. Scale bar (A) 100 µm, scale bar (B, C) 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5404000" httpUri="https://zenodo.org/record/5404000/files/figure.png" pageId="7" pageNumber="385">5A</figureCitation>
,
<figureCitation id="0AAADF8F3451FFA530EEFCECFB7A3776" box="[1192,1259,816,840]" captionStart="FIGURE" captionStartId="10.[159,171,1738,1757]" captionTargetBox="[382,1274,550,1627]" captionTargetId="figure-5@10.[242,1346,457,1706]" captionTargetPageId="10" captionText="FIGURE 6: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A subcapitulum, ventral view, and palptrochanter; B palp (trochanter not illustrated), left, ventral view; C chelicera, left, paraxial view. Scale bar (A, C; B) 16 µm." figureDoi="http://doi.org/10.5281/zenodo.5404002" httpUri="https://zenodo.org/record/5404002/files/figure.png" pageId="7" pageNumber="385">6AC</figureCitation>
) — Morphology of subcapitulum, palp and chelicera typical for
<taxonomicName id="5591B8893451FFA5377CFCAAFC2C37B0" authorityName="Balogh" authorityYear="1983" box="[826,957,886,910]" class="Arachnida" family="Granuloppiidae" genus="Hammerella" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="A0E51F183451FFA5377CFCAAFC2C37B0" box="[826,957,886,910]" italics="true" pageId="7" pageNumber="385">Hammerella</emphasis>
</taxonomicName>
(e.g.
<bibRefCitation id="F600BEFB3451FFA53040FCAAFAB937B0" author="Franklin E. &amp; Woas S." box="[1030,1320,886,910]" pageId="7" pageNumber="385" pagination="5 - 56" refId="ref5961" refString="Franklin E., Woas S. 1992 - Some oribatid mites of the family Oppiidae (Acari, Oribatei) from Amazonia - Andrias, 9: 5 - 56." type="journal article" year="1992">Franklin and Woas 1992</bibRefCitation>
; Ermilov
<emphasis id="A0E51F183451FFA5377CFC46FCE0378F" box="[826,881,921,945]" italics="true" pageId="7" pageNumber="385">et al.</emphasis>
2012). Subcapitulum longer than wide (73 77 × 53 57). Antero-medial parts of rutelli with thin tooth. Three pairs of subcapitular setae setiform;
<emphasis id="A0E51F183451FFA537C2FBDFFC083024" box="[900,921,1027,1050]" italics="true" pageId="7" pageNumber="385">m</emphasis>
longest (28) barbed,
<emphasis id="A0E51F183451FFA530DEFBDEFB343024" box="[1176,1189,1026,1050]" italics="true" pageId="7" pageNumber="385">h</emphasis>
(24) sparsely ciliate on dorsal side,
<emphasis id="A0E51F183451FFA537B1FBFAFB923003" box="[1015,1027,1062,1085]" italics="true" pageId="7" pageNumber="385">a</emphasis>
shortest (16), barbed. Two pairs of adoral setae (6) thin, smooth. Palps (41 45) with setation 02138(+ω); solenidion of palptarsi as long as one third of tarsi, thick, expanded distally, pressed to the surface, attached in mediodistal parts to seta
<emphasis id="A0E51F183451FFA53055FB09FBA730D3" box="[1043,1078,1237,1261]" italics="true" pageId="7" pageNumber="385">ul&quot;</emphasis>
. Chelicerae (69 73) with two setiform, barbed setae;
<emphasis id="A0E51F183451FFA5300BFB24FBE0312E" box="[1101,1137,1272,1296]" italics="true" pageId="7" pageNumber="385">cha</emphasis>
(20) longer than
<emphasis id="A0E51F183451FFA5317AFB24FAF1312E" box="[1340,1376,1272,1296]" italics="true" pageId="7" pageNumber="385">chb</emphasis>
(12) barbed. Trägårdhs organ tapered, slightly granulate.
</paragraph>
<paragraph id="922EC30A3451FFA53726FAB4FB003248" blockId="7.[826,1429,311,1949]" pageId="7" pageNumber="385">
Epimeral and lateral podosomal regions (
<figureCitation id="0AAADF8F3451FFA53125FAB4FCCB319D" captionStart="FIGURE" captionStartId="8.[159,171,1869,1888]" captionText="FIGURE 4: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A dorsal view (legs except basal parts of legs II and III not illustrated); B ventral view (palps and legs except basal parts of legs IV not illustrated). Scale bar 100 µm." pageId="7" pageNumber="385">Figs 4B</figureCitation>
,
<figureCitation id="0AAADF8F3451FFA53722FA57FC16319D" box="[868,903,1419,1443]" captionStart="FIGURE" captionStartId="9.[159,171,1752,1771]" captionTargetBox="[288,1311,497,1616]" captionTargetId="figure-6@9.[242,1346,415,1720]" captionTargetPageId="9" captionText="FIGURE 5: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A lateral view, right side (gnathosoma and legs except basal part of legs I, III and IV not illustrated); B tibia (solenidion not illustrated) and basal part of tarsus of leg I, right, antiaxial view; C tibia (solenidion except basal part not illustrated) and tarsus of leg IV, right, paraxial view. Scale bar (A) 100 µm, scale bar (B, C) 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5404000" httpUri="https://zenodo.org/record/5404000/files/figure.png" pageId="7" pageNumber="385">5A</figureCitation>
) — Sejugal apodemes and apodemes 2 short, similar in length, apodemes 4 absent, substituted by muscle sigilla. Epimeral setal formula: 3133; all setae setiform, slightly barbed,
<emphasis id="A0E51F183451FFA530FCFA29FB423233" box="[1210,1235,1525,1549]" italics="true" pageId="7" pageNumber="385">1a</emphasis>
,
<emphasis id="A0E51F183451FFA530A3FA29FB6F3233" box="[1253,1278,1525,1549]" italics="true" pageId="7" pageNumber="385">2a</emphasis>
and
<emphasis id="A0E51F183451FFA53106FA29FACB3233" box="[1344,1370,1525,1549]" italics="true" pageId="7" pageNumber="385">3a</emphasis>
(16 20) shorter than
<emphasis id="A0E51F183451FFA537BDF9C4FB84320E" box="[1019,1045,1560,1584]" italics="true" pageId="7" pageNumber="385">1b</emphasis>
,
<emphasis id="A0E51F183451FFA53065F9C4FBAC320E" box="[1059,1085,1560,1584]" italics="true" pageId="7" pageNumber="385">3b</emphasis>
,
<emphasis id="A0E51F183451FFA5300DF9C4FB18320E" box="[1099,1161,1560,1584]" italics="true" pageId="7" pageNumber="385">3c, 4a</emphasis>
,
<emphasis id="A0E51F183451FFA530DEF9C4FB20320E" box="[1176,1201,1560,1584]" italics="true" pageId="7" pageNumber="385">4b</emphasis>
and
<emphasis id="A0E51F183451FFA530ABF9C4FA94320E" box="[1261,1285,1560,1584]" italics="true" pageId="7" pageNumber="385">4c</emphasis>
(32 41),
<emphasis id="A0E51F183451FFA5313BF9C4FA04320E" box="[1405,1429,1560,1584]" italics="true" pageId="7" pageNumber="385">1c</emphasis>
longest (45 53), inserted on pedotecta I,
<emphasis id="A0E51F183451FFA5315EF9E7FABE326D" box="[1304,1327,1595,1619]" italics="true" pageId="7" pageNumber="385">3c</emphasis>
inserted on tubercles. Discidia absent.
</paragraph>
<paragraph id="922EC30A3451FFA53726F954FC4C3371" blockId="7.[826,1429,311,1949]" pageId="7" pageNumber="385">
Anogenital region (
<figureCitation id="0AAADF8F3451FFA5300FF954FB34329E" box="[1097,1189,1672,1696]" captionStart="FIGURE" captionStartId="8.[159,171,1869,1888]" captionText="FIGURE 4: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A dorsal view (legs except basal parts of legs II and III not illustrated); B ventral view (palps and legs except basal parts of legs IV not illustrated). Scale bar 100 µm." pageId="7" pageNumber="385">Figs 4B</figureCitation>
,
<figureCitation id="0AAADF8F3451FFA530F5F954FB44329E" box="[1203,1237,1672,1696]" captionStart="FIGURE" captionStartId="9.[159,171,1752,1771]" captionTargetBox="[288,1311,497,1616]" captionTargetId="figure-6@9.[242,1346,415,1720]" captionTargetPageId="9" captionText="FIGURE 5: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A lateral view, right side (gnathosoma and legs except basal part of legs I, III and IV not illustrated); B tibia (solenidion not illustrated) and basal part of tarsus of leg I, right, antiaxial view; C tibia (solenidion except basal part not illustrated) and tarsus of leg IV, right, paraxial view. Scale bar (A) 100 µm, scale bar (B, C) 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5404000" httpUri="https://zenodo.org/record/5404000/files/figure.png" pageId="7" pageNumber="385">5A</figureCitation>
) — Six pairs of genital setae minute (
<emphasis id="A0E51F183451FFA5307FF970FBD732FD" box="[1081,1094,1708,1731]" italics="true" pageId="7" pageNumber="385">g</emphasis>
<subScript id="0E15C14F3451FFA53001F968FBC132FB" attach="left" box="[1095,1104,1716,1733]" fontSize="7" pageId="7" pageNumber="385">1</subScript>
<emphasis id="A0E51F183451FFA53018F970FBFA32FD" box="[1118,1131,1708,1731]" italics="true" pageId="7" pageNumber="385">g</emphasis>
<subScript id="0E15C14F3451FFA5302AF968FBE432FB" attach="left" box="[1132,1141,1716,1733]" fontSize="7" pageId="7" pageNumber="385">6</subScript>
, 8). One pair of aggenital (32), three pairs of adanal (32) and two pairs of anal (16 20) setae setiform, smooth. Adanal lyrifissures distinct, inverse apoanal, distanced from the anal aperture.
</paragraph>
<paragraph id="922EC30A3451FFA93726F8BEFD273659" blockId="7.[826,1429,311,1949]" lastBlockId="11.[159,762,311,1681]" lastPageId="11" lastPageNumber="389" pageId="7" pageNumber="385">
Legs (
<figureCitation id="0AAADF8F3451FFA537EFF8BDFBB83347" box="[937,1065,1889,1914]" captionStart="FIGURE" captionStartId="8.[159,171,1869,1888]" captionText="FIGURE 4: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A dorsal view (legs except basal parts of legs II and III not illustrated); B ventral view (palps and legs except basal parts of legs IV not illustrated). Scale bar 100 µm." pageId="7" pageNumber="385">Figs 4A, B</figureCitation>
,
<figureCitation id="0AAADF8F3451FFA5307EF8BDFBEA3347" box="[1080,1147,1889,1913]" captionStart="FIGURE" captionStartId="9.[159,171,1752,1771]" captionTargetBox="[288,1311,497,1616]" captionTargetId="figure-6@9.[242,1346,415,1720]" captionTargetPageId="9" captionText="FIGURE 5: Hammerella (Woasella) huanucoensis Ermilov n. sp.: A lateral view, right side (gnathosoma and legs except basal part of legs I, III and IV not illustrated); B tibia (solenidion not illustrated) and basal part of tarsus of leg I, right, antiaxial view; C tibia (solenidion except basal part not illustrated) and tarsus of leg IV, right, paraxial view. Scale bar (A) 100 µm, scale bar (B, C) 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5404000" httpUri="https://zenodo.org/record/5404000/files/figure.png" pageId="7" pageNumber="385">5AC</figureCitation>
) — Morphology of leg segments, setae and solenidia typical for
<taxonomicName id="5591B8893451FFA53154F858FA0433A2" authorityName="Balogh" authorityYear="1983" box="[1298,1429,1924,1948]" class="Arachnida" family="Granuloppiidae" genus="Hammerella" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="A0E51F183451FFA53154F858FA0433A2" box="[1298,1429,1924,1948]" italics="true" pageId="7" pageNumber="385">Hammerella</emphasis>
</taxonomicName>
(e.g.
<bibRefCitation id="F600BEFB345DFFA9349BFEEBFE6F3571" author="Franklin E. &amp; Woas S." box="[221,510,311,335]" pageId="11" pageNumber="389" pagination="5 - 56" refId="ref5961" refString="Franklin E., Woas S. 1992 - Some oribatid mites of the family Oppiidae (Acari, Oribatei) from Amazonia - Andrias, 9: 5 - 56." type="journal article" year="1992">Franklin and Woas 1992</bibRefCitation>
; Ermilov
<emphasis id="A0E51F18345DFFA93634FEE4FD393571" box="[626,680,311,335]" italics="true" pageId="11" pageNumber="389">et al.</emphasis>
2012). Claw of each leg indistinctly serrate on dorsal side. Formulas of leg setation and solenidia: I (1524 20) [122], II (152416) [112], III (231315) [110], IV (122312) [010]; homology of setae and solenidia indicated in
<tableCitation id="DF13F6B1345DFFA9359CFE3AFDBC35C0" box="[474,557,486,510]" captionStart="TABLE" captionStartId="6.[159,172,308,327]" captionText="TABLE 1: Leg setation and solenidia of adult Lasiobelba (Antennoppia) parachistyakovi Ermilov n. sp. and Hammerella (Woasella) huanucoen-" pageId="11" pageNumber="389">Table 1</tableCitation>
. Setae
<emphasis id="A0E51F18345DFFA936C5FE3BFD0135C0" box="[643,656,487,510]" italics="true" pageId="11" pageNumber="389">p</emphasis>
setiform on tarsi I, and very short, conical on tarsi IIIV. Famuli (Ɛ) of tarsi I short, thin, swollen and truncated distally, inserted posteriorly to solenidion ω
<subScript id="0E15C14F345DFFA936E2FD85FD3C3654" attach="left" box="[676,685,601,618]" fontSize="7" pageId="11" pageNumber="389">1</subScript>
.
</paragraph>
<caption id="C6EE9382345EFFAA34D9F891FBB43342" pageId="8" pageNumber="386" startId="8.[159,171,1869,1888]">
<paragraph id="922EC30A345EFFAA34D9F891FBB43342" blockId="8.[159,1429,1869,1916]" pageId="8" pageNumber="386">
FIGURE 4:
<taxonomicName id="5591B889345EFFAA3543F891FD09335E" authority="Ermilov" authorityName="Ermilov" authorityYear="2016" box="[261,664,1869,1888]" class="Arachnida" family="Granuloppiidae" genus="Hammerella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="huanucoensis" status="sp. nov." subGenus="Woasella">
<emphasis id="A0E51F18345EFFAA3543F891FEFF335E" box="[261,366,1869,1888]" italics="true" pageId="8" pageNumber="386">Hammerella</emphasis>
(
<emphasis id="A0E51F18345EFFAA353CF891FE57335E" box="[378,454,1869,1888]" italics="true" pageId="8" pageNumber="386">Woasella</emphasis>
)
<emphasis id="A0E51F18345EFFAA3595F891FDD6335E" box="[467,583,1869,1888]" italics="true" pageId="8" pageNumber="386">huanucoensis</emphasis>
Ermilov
</taxonomicName>
<emphasis id="A0E51F18345EFFAA36D8F891FD43335E" bold="true" box="[670,722,1869,1888]" pageId="8" pageNumber="386">
<taxonomicNameLabel id="BBD6A263345EFFAA36D8F891FD43335E" box="[670,722,1869,1888]" pageId="8" pageNumber="386" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A dorsal view (legs except basal parts of legs II and III not illustrated); B ventral view (palps and legs except basal parts of legs IV not illustrated). Scale bar 100 µm.
</paragraph>
</caption>
<caption id="C6EE9382345FFFAB34D9F904FA12331D" ID-DOI="http://doi.org/10.5281/zenodo.5404000" ID-Zenodo-Dep="5404000" httpUri="https://zenodo.org/record/5404000/files/figure.png" pageId="9" pageNumber="387" startId="9.[159,171,1752,1771]" targetBox="[288,1311,497,1616]" targetPageId="9">
<paragraph id="922EC30A345FFFAB34D9F904FA12331D" blockId="9.[159,1429,1752,1827]" pageId="9" pageNumber="387">
FIGURE 5:
<taxonomicName id="5591B889345FFFAB354DF904FD3532D5" authority="Ermilov" authorityName="Ermilov" authorityYear="2016" box="[267,676,1752,1771]" class="Arachnida" family="Granuloppiidae" genus="Hammerella" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="387" phylum="Arthropoda" rank="species" species="huanucoensis" status="sp. nov." subGenus="Woasella">
<emphasis id="A0E51F18345FFFAB354DF904FEE532D5" box="[267,372,1752,1771]" italics="true" pageId="9" pageNumber="387">Hammerella</emphasis>
(
<emphasis id="A0E51F18345FFFAB35C4F904FE5F32D5" box="[386,462,1752,1771]" italics="true" pageId="9" pageNumber="387">Woasella</emphasis>
)
<emphasis id="A0E51F18345FFFAB359AF904FDC132D5" box="[476,592,1752,1771]" italics="true" pageId="9" pageNumber="387">huanucoensis</emphasis>
Ermilov
</taxonomicName>
<emphasis id="A0E51F18345FFFAB36EDF904FD7732D5" bold="true" box="[683,742,1752,1771]" pageId="9" pageNumber="387">
<taxonomicNameLabel id="BBD6A263345FFFAB36EDF904FD7732D5" box="[683,742,1752,1771]" pageId="9" pageNumber="387" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A lateral view, right side (gnathosoma and legs except basal part of legs I, III and IV not illustrated); B tibia (solenidion not illustrated) and basal part of tarsus of leg I, right, antiaxial view; C tibia (solenidion except basal part not illustrated) and tarsus of leg IV, right, paraxial view. Scale bar (A) 100 µm, scale bar (B, C) 20 µm.
</paragraph>
</caption>
<caption id="C6EE9382345CFFA834D9F916FBB232C6" ID-DOI="http://doi.org/10.5281/zenodo.5404002" ID-Zenodo-Dep="5404002" httpUri="https://zenodo.org/record/5404002/files/figure.png" pageId="10" pageNumber="388" startId="10.[159,171,1738,1757]" targetBox="[382,1274,550,1627]" targetPageId="10">
<paragraph id="922EC30A345CFFA834D9F916FBB232C6" blockId="10.[159,1429,1737,1784]" pageId="10" pageNumber="388">
FIGURE 6:
<taxonomicName id="5591B889345CFFA83540F915FD0A32E2" authority="Ermilov" authorityName="Ermilov" authorityYear="2016" box="[262,667,1737,1756]" class="Arachnida" family="Granuloppiidae" genus="Hammerella" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="388" phylum="Arthropoda" rank="species" species="huanucoensis" status="sp. nov." subGenus="Woasella">
<emphasis id="A0E51F18345CFFA83540F915FEFE32E2" box="[262,367,1737,1756]" italics="true" pageId="10" pageNumber="388">Hammerella</emphasis>
(
<emphasis id="A0E51F18345CFFA8353AF915FE5932E2" box="[380,456,1737,1756]" italics="true" pageId="10" pageNumber="388">Woasella</emphasis>
)
<emphasis id="A0E51F18345CFFA83593F915FDD832E2" box="[469,585,1737,1756]" italics="true" pageId="10" pageNumber="388">huanucoensis</emphasis>
Ermilov
</taxonomicName>
<emphasis id="A0E51F18345CFFA836E7F915FD4632E2" bold="true" box="[673,727,1737,1756]" pageId="10" pageNumber="388">
<taxonomicNameLabel id="BBD6A263345CFFA836E7F915FD4632E2" box="[673,727,1737,1756]" pageId="10" pageNumber="388" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A subcapitulum, ventral view, and palptrochanter; B palp (trochanter not illustrated), left, ventral view; C chelicera, left, paraxial view. Scale bar (A, C; B) 16 µm.
</paragraph>
</caption>
<paragraph id="922EC30A345DFFA93483FDA3FE0637EC" blockId="11.[159,762,311,1681]" pageId="11" pageNumber="389">
<materialsCitation id="22F9C957345DFFA93483FDA3FF4936C1" country="Peru" location="Museo de Historia Natural" municipality="Universidad Nacional Mayor de San Marcos" pageId="11" pageNumber="389" specimenCount="1" stateProvince="Lima" typeStatus="holotype">
Type deposition — The
<typeStatus id="4D2A7DA8345DFFA935B4FDA3FDCB36A9" box="[498,602,639,663]" pageId="11" pageNumber="389" type="holotype">holotype</typeStatus>
is deposited in the collection of the
<location id="974E95D1345DFFA935F4FD7EFD643684" LSID="urn:lsid:plazi:treatment:1A38721C3451FFA934ABFD73FECF32AF:974E95D1345DFFA935F4FD7EFD643684" box="[434,757,674,698]" country="Peru" municipality="Universidad Nacional Mayor de San Marcos" name="Museo de Historia Natural" pageId="11" pageNumber="389" stateProvince="Lima">Museo de Historia Natural</location>
,
<collectingMunicipality id="724A5970345DFFA934D9FD19FD3A36E2" box="[159,683,709,733]" pageId="11" pageNumber="389">Universidad Nacional Mayor de San Marcos</collectingMunicipality>
,
<collectingRegion id="50550DE8345DFFA936F0FD19FD6436E3" box="[694,757,709,733]" country="Peru" name="Lima" pageId="11" pageNumber="389">Lima</collectingRegion>
,
<collectingCountry id="EA86839A345DFFA934D9FD34FF4936C1" box="[159,216,744,767]" name="Peru" pageId="11" pageNumber="389">Peru</collectingCountry>
</materialsCitation>
;
<materialsCitation id="22F9C957345DFFA934A3FD34FE813778" country="Germany" location="Collection of Zoology" municipality="Munich" pageId="11" pageNumber="389" specimenCount="1" stateProvince="Bavarian State" typeStatus="paratype">
<specimenCount id="84970883345DFFA934A3FD34FE1F373E" box="[229,398,744,768]" pageId="11" pageNumber="389" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
are deposited in the collection of the
<collectingRegion id="50550DE8345DFFA934A3FCD7FE1D371D" box="[229,396,779,803]" inferredBySuffix="true" pageId="11" pageNumber="389">Bavarian State</collectingRegion>
<location id="974E95D1345DFFA935D4FCD7FD1F371D" LSID="urn:lsid:plazi:treatment:1A38721C3451FFA934ABFD73FECF32AF:974E95D1345DFFA935D4FCD7FD1F371D" box="[402,654,779,803]" country="Germany" municipality="Munich" name="Collection of Zoology" pageId="11" pageNumber="389" stateProvince="Bavarian State">Collection of Zoology</location>
,
<collectingMunicipality id="724A5970345DFFA936D1FCD7FD64371D" box="[663,757,779,803]" pageId="11" pageNumber="389">Munich</collectingMunicipality>
,
<collectingCountry id="EA86839A345DFFA934D9FCF2FE813778" box="[159,272,814,838]" name="Germany" pageId="11" pageNumber="389">Germany</collectingCountry>
</materialsCitation>
;
<materialsCitation id="22F9C957345DFFA9355CFCF2FF7537B3" country="Germany" location="Senckenberg Institution Frankfurt" pageId="11" pageNumber="389" specimenCount="1" typeStatus="paratype">
<specimenCount id="84970883345DFFA9355CFCF2FE513778" box="[282,448,814,838]" pageId="11" pageNumber="389" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
are deposited in the collection of the
<location id="974E95D1345DFFA93565FC8DFD293757" LSID="urn:lsid:plazi:treatment:1A38721C3451FFA934ABFD73FECF32AF:974E95D1345DFFA93565FC8DFD293757" box="[291,696,849,873]" country="Germany" name="Senckenberg Institution Frankfurt" pageId="11" pageNumber="389">Senckenberg Institution Frankfurt</location>
,
<collectingCountry id="EA86839A345DFFA93680FC8DFF7537B3" name="Germany" pageId="11" pageNumber="389">Germany</collectingCountry>
</materialsCitation>
;
<materialsCitation id="22F9C957345DFFA934B4FCA8FE0237EC" country="Russia" location="University Museum of Zoology" pageId="11" pageNumber="389" specimenCount="12" stateProvince="Tyumen'" typeStatus="paratype">
<specimenCount id="84970883345DFFA934B4FCA8FE1837B3" box="[242,393,884,909]" pageId="11" pageNumber="389" type="generic" typeStatus="paratypes">12 paratypes</specimenCount>
are deposited in the collection of the
<collectingRegion id="50550DE8345DFFA934B6FC44FE053791" box="[240,404,919,944]" country="Russia" name="Tyumen'" pageId="11" pageNumber="389">Tyumen State</collectingRegion>
<location id="974E95D1345DFFA935D8FC4BFF5B37ED" LSID="urn:lsid:plazi:treatment:1A38721C3451FFA934ABFD73FECF32AF:974E95D1345DFFA935D8FC4BFF5B37ED" country="Russia" name="University Museum of Zoology" pageId="11" pageNumber="389" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="50550DE8345DFFA93493FC67FEA937ED" box="[213,312,955,979]" country="Russia" name="Tyumen'" pageId="11" pageNumber="389">Tyumen</collectingRegion>
,
<collectingCountry id="EA86839A345DFFA93505FC66FE0237EC" box="[323,403,954,978]" name="Russia" pageId="11" pageNumber="389">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="922EC30A345DFFA93483FC35FD933076" blockId="11.[159,762,311,1681]" pageId="11" pageNumber="389">
Etymology — The specific name
<taxonomicName id="5591B889345DFFA9362EFC35FD68303F" authorityName="Ermilov" authorityYear="2016" box="[616,761,1001,1025]" class="Arachnida" family="Granuloppiidae" genus="Hammerella" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="389" phylum="Arthropoda" rank="species" species="huanucoensis">
<emphasis id="A0E51F18345DFFA9362EFC35FD68303F" box="[616,761,1001,1025]" italics="true" pageId="11" pageNumber="389">huanucoensis</emphasis>
</taxonomicName>
refers to the Peruvian Department, HuAEnuco, where the new species was collected.
</paragraph>
<paragraph id="922EC30A345DFFA93483FB83FE9E3177" blockId="11.[159,762,311,1681]" pageId="11" pageNumber="389">
Remarks — The genus
<taxonomicName id="5591B889345DFFA935B7FB83FDE53049" authorityName="Balogh" authorityYear="1983" box="[497,628,1119,1143]" class="Arachnida" family="Granuloppiidae" genus="Hammerella" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="A0E51F18345DFFA935B7FB83FDE53049" box="[497,628,1119,1143]" italics="true" pageId="11" pageNumber="389">Hammerella</emphasis>
</taxonomicName>
comprises five subgenera and 13 species, and is distributed in the tropics (
<bibRefCitation id="F600BEFB345DFFA9356BFB79FE2D3083" author="Subias L. S." box="[301,444,1189,1213]" pageId="11" pageNumber="389" refId="ref6228" refString="Subias L. S. 2004 - Listado sistemAEtico, sinonimico y biogeogrAEfico de los AEcaros oribAEtidos (Acariformes: Oribatida) del mundo (excepto fosiles) - Graellsia, 60 (noemero extraordinario): 3 - 305. Online version accessed in February 2016, 593 pp." type="book" year="2004">Subías 2004</bibRefCitation>
, updated 2016). The main morphological traits for these genus and subgenera and an identification key for all species (known to 2012) of
<taxonomicName id="5591B889345DFFA9354EFAD2FE1A3118" authorityName="Balogh" authorityYear="1983" box="[264,395,1294,1318]" class="Arachnida" family="Granuloppiidae" genus="Hammerella" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="A0E51F18345DFFA9354EFAD2FE1A3118" box="[264,395,1294,1318]" italics="true" pageId="11" pageNumber="389">Hammerella</emphasis>
</taxonomicName>
were presented by Ermilov
<emphasis id="A0E51F18345DFFA936A0FAD3FF283177" italics="true" pageId="11" pageNumber="389">et al.</emphasis>
(2012).
</paragraph>
<paragraph id="922EC30A345DFFA93483FABCFECF32AF" blockId="11.[159,762,311,1681]" pageId="11" pageNumber="389">
The new species is morphologically most similar to
<taxonomicName id="5591B889345DFFA934F8FA58FD8F3181" authority="Ermilov, Sandmann, Marian and Maraun, 2013" authorityName="Ermilov, Sandmann, Marian and Maraun" authorityYear="2013" class="Arachnida" family="Granuloppiidae" genus="Hammerella" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="389" phylum="Arthropoda" rank="species" species="parasufflata" subGenus="Woasella">
<emphasis id="A0E51F18345DFFA934F8FA58FED031A2" box="[190,321,1412,1436]" italics="true" pageId="11" pageNumber="389">Hammerella</emphasis>
(
<emphasis id="A0E51F18345DFFA93514FA58FE2031A2" box="[338,433,1412,1436]" italics="true" pageId="11" pageNumber="389">Woasella</emphasis>
)
<emphasis id="A0E51F18345DFFA93585FA58FDD331A2" box="[451,578,1412,1436]" italics="true" pageId="11" pageNumber="389">parasufflata</emphasis>
<bibRefCitation id="F600BEFB345DFFA9360CFA58FD8F3181" author="Ermilov S. G. &amp; Sandmann D. &amp; Marian F. &amp; Maraun M." pageId="11" pageNumber="389" pagination="218 - 224" refId="ref5788" refString="Ermilov S. G., Sandmann D., Marian F., Maraun M. 2013 - Oribatid mites of the superfamily Oppioidea from Ecuador (Acari: Oribatida) - Systematic and Applied Acarology, 18 (3): 218 - 224. doi: 10.11158 / saa. 18.3.3" type="journal article" year="2013">Ermilov, Sandmann, Marian and Maraun, 2013</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="EA86839A345DFFA93625FA7BFD573181" box="[611,710,1447,1471]" name="Ecuador" pageId="11" pageNumber="389">Ecuador</collectingCountry>
(see
<bibRefCitation id="F600BEFB345DFFA934D9FA16FEEA31DC" author="Ermilov S. G. &amp; Sandmann D. &amp; Marian F. &amp; Maraun M." box="[159,379,1482,1506]" pageId="11" pageNumber="389" pagination="218 - 224" refId="ref5788" refString="Ermilov S. G., Sandmann D., Marian F., Maraun M. 2013 - Oribatid mites of the superfamily Oppioidea from Ecuador (Acari: Oribatida) - Systematic and Applied Acarology, 18 (3): 218 - 224. doi: 10.11158 / saa. 18.3.3" type="journal article" year="2013">
Ermilov
<emphasis id="A0E51F18345DFFA93543FA16FEAB31DC" box="[261,314,1482,1506]" italics="true" pageId="11" pageNumber="389">et al.</emphasis>
2013
</bibRefCitation>
) in having an indentate rostrum and ciliate unilaterally bothridial seta. However, it differs from the latter by the thin costulae (
<emphasis id="A0E51F18345DFFA936CCF9CDFD333216" box="[650,674,1553,1576]" italics="true" pageId="11" pageNumber="389">vs</emphasis>
. thick), shorter notogastral setae (
<emphasis id="A0E51F18345DFFA9358CF9E8FE733275" box="[458,482,1588,1611]" italics="true" pageId="11" pageNumber="389">vs</emphasis>
. comparatively longer), and the absence of tubercles anterior to lamellar setae (
<emphasis id="A0E51F18345DFFA93497F9A6FF7832AF" box="[209,233,1658,1681]" italics="true" pageId="11" pageNumber="389">vs</emphasis>
. present).
</paragraph>
</subSubSection>
</treatment>
</document>