treatments-xml/data/03/FF/BE/03FFBE1DFFDE736332FAF973E192F832.xml
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<document id="1ED1470C93EC1F742EAC95595118A1AB" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFDE736332FAF973E192F832" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium pseudopriapus Wesener 2009, new species" docType="treatment" docUuid="3E4A0A52-336F-40A6-A654-79EDF9BE778D" docUuidSource="ZooBank" docVersion="5" lastPageNumber="52" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="46" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4FDFF54E94974454F1DD9912B875D19B">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
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<mods:namePart id="CC64B9B8ACE215B398A442AE15BA40AB">Wesener, Thomas</mods:namePart>
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<mods:title id="5DBAC6ECB09566A1A46427D5894C617A">Zootaxa</mods:title>
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<mods:date id="6EF21C02A7A7901B29E762D64D8E4A13">2009</mods:date>
<mods:detail id="A100106C8B69CFB1C2842039305B211D" type="pubDate">
<mods:number id="097A75D122B89051FC6D6E973D1B7D14">2009-05-11</mods:number>
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<mods:number id="BF97760863DFC00D9227A64BAA398DC4">2097</mods:number>
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<mods:classification id="4BF107A917F2C5E91895E834AB2FA9D5">journal article</mods:classification>
<mods:identifier id="D2850C8C358D9167F49A38DC03D47983" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
<mods:identifier id="CF01AEB0D25A2B5B75D5A077005B1CBB" type="ISSN">1175­5334</mods:identifier>
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<treatment id="03FFBE1DFFDE736332FAF973E192F832" ID-DOI="http://doi.org/10.5281/zenodo.5325461" ID-Zenodo-Dep="5325461" LSID="urn:lsid:zoobank.org:act:3E4A0A52-336F-40A6-A654-79EDF9BE778D" httpUri="http://treatment.plazi.org/id/03FFBE1DFFDE736332FAF973E192F832" lastPageId="51" lastPageNumber="52" pageId="45" pageNumber="46">
<subSubSection id="C34C5C80FFDE737D32FAF973E19BF93A" box="[151,668,1737,1763]" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8BE90F0BFFDE737D32FAF973E19BF93A" blockId="45.[151,668,1737,1763]" box="[151,668,1737,1763]" pageId="45" pageNumber="46">
<heading id="D0A1B867FFDE737D32FAF973E19BF93A" bold="true" box="[151,668,1737,1763]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
<emphasis id="B922D319FFDE737D32FAF973E19BF93A" bold="true" box="[151,668,1737,1763]" pageId="45" pageNumber="46">
<emphasis id="B922D319FFDE737D32FAF973E102F93A" bold="true" box="[151,517,1737,1763]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="4C567488FFDE737D32FAF973E2F8F93A" ID-CoL="5D7HT" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,511,1737,1763]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pseudopriapus" status="sp. nov.">Zoosphaerium pseudopriapus</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2116E62FFDE737D3061F973E19BF93A" box="[524,668,1737,1763]" pageId="45" pageNumber="46" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFDE737F32FAF8A8E14EFAC9" lastPageId="47" lastPageNumber="48" pageId="45" pageNumber="46" type="etymology">
<paragraph id="8BE90F0BFFDE737D32FAF8A8E2E0F88D" blockId="45.[151,1436,1810,2030]" pageId="45" pageNumber="46">
<emphasis id="B922D319FFDE737D32FAF8A8E281F8F5" bold="true" box="[151,390,1810,1836]" pageId="45" pageNumber="46">Derivatio nominis:</emphasis>
<taxonomicName id="4C567488FFDE737D33F9F8A9E146F8F5" authorityName="Wesener" authorityYear="2009" box="[404,577,1811,1836]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pseudopriapus">
<emphasis id="B922D319FFDE737D33F9F8A9E146F8F5" box="[404,577,1811,1836]" italics="true" pageId="45" pageNumber="46">pseudopriapus</emphasis>
</taxonomicName>
refers to the similarity of this species to the smaller
<taxonomicName id="4C567488FFDE737D3687F8A9E2E4F88D" authority="(de Saussure &amp; Zehntner, 1897)" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="priapus">
<emphasis id="B922D319FFDE737D3687F8A9E66EF8F5" box="[1258,1385,1811,1836]" italics="true" pageId="45" pageNumber="46">Z. priapus</emphasis>
(
<bibRefCitation id="EFC772FAFFDE737D3712F8A9E2DDF88D" author="de Saussure, H. &amp; Zehntner, L." pageId="45" pageNumber="46" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. &amp; Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure &amp; Zehntner, 1897</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BE90F0BFFDE737D32ABF8DAE14EF81E" blockId="45.[151,1436,1810,2030]" pageId="45" pageNumber="46">
<emphasis id="B922D319FFDE737D32ABF8DAE2A6F8A3" bold="true" box="[198,417,1888,1914]" pageId="45" pageNumber="46">Studied material:</emphasis>
<materialsCitation id="3B3E0556FFDE737D33C4F8DAE14EF81E" collectingDate="2003-12-07" collectionCode="CAS" collectorName="de Bekaraoka &amp; B. L. Fisher" country="Madagascar" county="Locus" elevation="150" latitude="-13.166667" location="Foret de Bekaraoka" longLatPrecision="21" longitude="49.71" municipality="Province" pageId="45" pageNumber="46" specimenCode="BLF 9871" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<emphasis id="B922D319FFDE737D33C4F8DAE11CF8A3" bold="true" box="[425,539,1888,1914]" pageId="45" pageNumber="46">
<typeStatus id="54EDB1A9FFDE737D33C4F8DAE11CF8A3" box="[425,539,1888,1914]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
</emphasis>
: 1 M (
<quantity id="4CAEA2EEFFDE737D3000F8DAE1C6F8A3" box="[621,705,1888,1914]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="45" pageNumber="46" unit="mm" value="32.0">32 mm</quantity>
long),
<emphasis id="B922D319FFDE737D317AF8DAE0D1F8A3" bold="true" box="[791,982,1888,1914]" pageId="45" pageNumber="46">
<collectionCode id="ED4797CEFFDE737D317AF8DAE055F8A3" box="[791,850,1888,1914]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="45" pageNumber="46" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFDE737D3136F8DAE0D1F8A3" box="[859,982,1888,1914]" collectionCode="BLF" pageId="45" pageNumber="46">BLF 9871</specimenCode>
</emphasis>
.
<emphasis id="B922D319FFDE737D318BF8DBE780F8A3" box="[998,1159,1889,1914]" italics="true" pageId="45" pageNumber="46">
<collectingCounty id="62887787FFDE737D318BF8DBE72BF8A3" box="[998,1068,1889,1914]" pageId="45" pageNumber="46">Locus</collectingCounty>
typicus
</emphasis>
:
<collectingCountry id="F3414F9BFFDE737D36F5F8DAE620F8A3" box="[1176,1319,1888,1914]" name="Madagascar" pageId="45" pageNumber="46">Madagascar</collectingCountry>
,
<collectingMunicipality id="6B8D9571FFDE737D3759F8DAE69BF8A3" box="[1332,1436,1888,1914]" pageId="45" pageNumber="46">Province</collectingMunicipality>
<collectingRegion id="4992C1E9FFDE737D32FAF83DE220F878" box="[151,295,1927,1953]" country="Madagascar" name="Antsiranana" pageId="45" pageNumber="46">Antsiranana</collectingRegion>
,
<location id="8E8959D0FFDE737D335BF83DE124F878" LSID="urn:lsid:plazi:treatment:03FFBE1DFFDE736332FAF973E192F832:8E8959D0FFDE737D335BF83DE124F878" box="[310,547,1927,1953]" country="Madagascar" county="Locus" latitude="-13.166667" longLatPrecision="21" longitude="49.71" municipality="Province" name="Foret de Bekaraoka" pageId="45" pageNumber="46" stateProvince="Antsiranana">
Forêt
<collectorName id="26A36ADDFFDE737D3313F83DE124F878" box="[382,547,1927,1953]" pageId="45" pageNumber="46">de Bekaraoka</collectorName>
</location>
,
<locationDeviation id="9A5B1629FFDE737D305FF83DE060F878" box="[562,871,1927,1953]" location="Daraina" pageId="45" pageNumber="46">
<quantity id="4CAEA2EEFFDE737D305FF83DE181F878" box="[562,646,1927,1953]" metricMagnitude="3" metricUnit="m" metricValue="6.8" pageId="45" pageNumber="46" unit="km" value="6.8">6.8 km</quantity>
60° ENE Daraina
</locationDeviation>
,
<geoCoordinate id="EE6269CCFFDE737D311BF83DE0FBF878" box="[886,1020,1927,1953]" degrees="13" direction="south" minutes="10" orientation="latitude" pageId="45" pageNumber="46" precision="15" seconds="00" value="-13.166667">13°10'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFFDE737D3661F83DE7A3F878" box="[1036,1188,1927,1953]" degrees="049" direction="east" minutes="42" orientation="longitude" pageId="45" pageNumber="46" precision="15" seconds="36" value="49.71">049°42'36&quot;E</geoCoordinate>
,
<quantity id="4CAEA2EEFFDE737D36DEF83DE605F878" box="[1203,1282,1927,1953]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="45" pageNumber="46" unit="m" value="150.0">
<elevation id="007BE838FFDE737D36DEF83DE605F878" box="[1203,1282,1927,1953]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="45" pageNumber="46" unit="m" value="150.0">150 m</elevation>
</quantity>
, tropical dry forest, coll.
<collectorName id="26A36ADDFFDE737D334FF817E2B7F81E" box="[290,432,1965,1991]" pageId="45" pageNumber="46">B. L. Fisher</collectorName>
,
<date id="FFE829CBFFDE737D33D1F817E143F81E" box="[444,580,1965,1991]" pageId="45" pageNumber="46" value="2003-12-07">
<collectingDate id="EFACD023FFDE737D33D1F817E143F81E" box="[444,580,1965,1991]" pageId="45" pageNumber="46" value="2003-12-07">07.xii.2003</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE90F0BFFDE737D32AAF86EE71FF837" blockId="45.[151,1436,1810,2030]" box="[199,1048,2004,2030]" pageId="45" pageNumber="46">
<materialsCitation id="3B3E0556FFDE737D32AAF86EE713F837" box="[199,1044,2004,2030]" collectingDate="2003-12-07" collectionCode="CAS" collectorName="de Bekaraoka &amp; B. L. Fisher" country="Madagascar" county="Locus" elevation="150" latitude="-13.166667" location="Foret de Bekaraoka" longLatPrecision="21" longitude="49.71" municipality="Province" pageId="45" pageNumber="46" specimenCode="BLF 9871" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<emphasis id="B922D319FFDE737D32AAF86EE249F837" bold="true" box="[199,334,2004,2030]" pageId="45" pageNumber="46">
<typeStatus id="54EDB1A9FFDE737D32AAF86EE24EF837" box="[199,329,2004,2030]" pageId="45" pageNumber="46" type="paratype">Paratypes</typeStatus>
:
</emphasis>
3 M, 4 F,
<emphasis id="B922D319FFDE737D33ABF86EE18AF837" bold="true" box="[454,653,2004,2030]" pageId="45" pageNumber="46">
<collectionCode id="ED4797CEFFDE737D33ABF86EE107F837" box="[454,512,2004,2030]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="45" pageNumber="46" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFDE737D3065F86EE18FF837" box="[520,648,2004,2030]" collectionCode="BLF" pageId="45" pageNumber="46">BLF 9871</specimenCode>
:
</emphasis>
same collection data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BE90F0BFFDD737E32FAF95DE22AF8BF" blockId="46.[151,1436,1767,1894]" pageId="46" pageNumber="47">
<emphasis id="B922D319FFDD737E32FAF95DE228F926" bold="true" box="[151,303,1767,1791]" pageId="46" pageNumber="47">FIGURE 030</emphasis>
.
<taxonomicName id="4C567488FFDD737E3353F95DE136F927" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[318,561,1767,1790]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="aureum" status="sp. nov.">
<emphasis id="B922D319FFDD737E3353F95DE136F927" box="[318,561,1767,1790]" italics="true" pageId="46" pageNumber="47">Zoosphaerium aureum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFDD737E3054F95DE170F926" bold="true" box="[569,631,1767,1791]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A2116E62FFDD737E3054F95DE170F926" box="[569,631,1767,1791]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, SEM,
<typeStatus id="54EDB1A9FFDD737E30A1F95DE02DF927" box="[716,810,1767,1790]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
,
<emphasis id="B922D319FFDD737E315BF95DE054F926" bold="true" box="[822,851,1767,1791]" pageId="46" pageNumber="47">A:</emphasis>
antenna, lateral view;
<emphasis id="B922D319FFDD737E3626F95DE75AF926" bold="true" box="[1099,1117,1767,1791]" pageId="46" pageNumber="47">B</emphasis>
: gnathochilarium, right inner palpus with rudimentary lateral palpus;
<emphasis id="B922D319FFDD737E3027F8B3E15AF8F8" bold="true" box="[586,605,1801,1825]" pageId="46" pageNumber="47">C</emphasis>
: gnathochilarium, details of sensory cells on endochilarium;
<emphasis id="B922D319FFDD737E3697F8B3E60AF8F8" bold="true" box="[1274,1293,1801,1825]" pageId="46" pageNumber="47">D</emphasis>
: epipharynx, apical view. Abbreviations:
<emphasis id="B922D319FFDD737E33A9F896E2DBF89D" bold="true" box="[452,476,1836,1860]" pageId="46" pageNumber="47">iP</emphasis>
= inner palpus;
<emphasis id="B922D319FFDD737E30E3F896E1B4F89D" bold="true" box="[654,691,1836,1860]" pageId="46" pageNumber="47">mp</emphasis>
= central pads or median palps on endochilarium;
<emphasis id="B922D319FFDD737E36B4F896E7FCF89D" bold="true" box="[1241,1275,1836,1860]" pageId="46" pageNumber="47">rlp</emphasis>
= rudimentary lateral palpus.
</paragraph>
<paragraph id="8BE90F0BFFDC737F32FAFBAFE7E1FB96" blockId="47.[151,1436,1045,1104]" pageId="47" pageNumber="48">
<emphasis id="B922D319FFDC737F32FAFBAFE228FBF4" bold="true" box="[151,303,1045,1069]" pageId="47" pageNumber="48">FIGURE 031</emphasis>
.
<taxonomicName id="4C567488FFDC737F3353FBACE136FBF4" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[318,561,1046,1069]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="aureum" status="sp. nov.">
<emphasis id="B922D319FFDC737F3353FBACE136FBF4" box="[318,561,1046,1069]" italics="true" pageId="47" pageNumber="48">Zoosphaerium aureum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFDC737F3056FBAFE17FFBF4" bold="true" box="[571,632,1045,1069]" pageId="47" pageNumber="48">
<taxonomicNameLabel id="A2116E62FFDC737F3056FBAFE17FFBF4" box="[571,632,1045,1069]" pageId="47" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, SEM,
<typeStatus id="54EDB1A9FFDC737F30A3FBACE02BFBF4" box="[718,812,1046,1069]" pageId="47" pageNumber="48" type="paratype">paratype</typeStatus>
, left mandible, mesal view. Abbreviations:
<emphasis id="B922D319FFDC737F3765FBAFE628FBF4" bold="true" box="[1288,1327,1045,1069]" pageId="47" pageNumber="48">3iT</emphasis>
= 3 inner teeth;
<emphasis id="B922D319FFDC737F32B4FB82E3FEFB89" bold="true" box="[217,249,1080,1104]" pageId="47" pageNumber="48">Co</emphasis>
= condylus;
<emphasis id="B922D319FFDC737F33E9FB82E2A5FB89" bold="true" box="[388,418,1080,1104]" pageId="47" pageNumber="48">eT</emphasis>
= external tooth;
<emphasis id="B922D319FFDC737F3032FB82E170FB89" bold="true" box="[607,631,1080,1104]" pageId="47" pageNumber="48">iP</emphasis>
= inner palpus;
<emphasis id="B922D319FFDC737F3149FB82E04EFB89" bold="true" box="[804,841,1080,1104]" pageId="47" pageNumber="48">mp</emphasis>
= molar plate;
<emphasis id="B922D319FFDC737F3181FB82E70BFB89" bold="true" box="[1004,1036,1080,1104]" pageId="47" pageNumber="48">pL</emphasis>
= pectinate lamella.
</paragraph>
<paragraph id="8BE90F0BFFDC737F32AAFB3BE14EFAC9" blockId="47.[151,1437,1153,2030]" pageId="47" pageNumber="48">
<emphasis id="B922D319FFDC737F32AAFB3BE292FB42" bold="true" box="[199,405,1153,1179]" pageId="47" pageNumber="48">Other material:</emphasis>
<materialsCitation id="3B3E0556FFDC737F33CBFB38E782FB1A" collectingDate="2001-11-03" collectingDateMax="2001-11-10" collectingDateMin="2001-11-03" collectionCode="FMMC" collectorName="S. Goodman" elevation="463" latitude="-13.255" location="Analamozava River" longLatPrecision="129" longitude="49.616665" pageId="47" pageNumber="48" specimenCode="FMMC 5488" specimenCount="1">
7 M,
<emphasis id="B922D319FFDC737F339AFB3BE1ADFB42" bold="true" box="[503,682,1153,1179]" pageId="47" pageNumber="48">
<specimenCode id="DBF0A770FFDC737F339AFB3BE1A2FB42" box="[503,677,1153,1179]" collectionCode="FMMC" pageId="47" pageNumber="48">FMMC 5488</specimenCode>
:
</emphasis>
near
<location id="8E8959D0FFDC737F3090FB38E0F5FB45" LSID="urn:lsid:plazi:treatment:03FFBE1DFFDE736332FAF973E192F832:8E8959D0FFDC737F3090FB38E0F5FB45" box="[765,1010,1154,1180]" latitude="-13.255" longLatPrecision="129" longitude="49.616665" name="Analamozava River" pageId="47" pageNumber="48">Analamozava River</location>
,
<locationDeviation id="9A5B1629FFDC737F366AFB38E609FB45" box="[1031,1294,1154,1180]" location="Daraina" pageId="47" pageNumber="48">
<quantity id="4CAEA2EEFFDC737F366AFB38E766FB45" box="[1031,1121,1154,1180]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="47" pageNumber="48" unit="km" value="7.5">7.5 km</quantity>
SW Daraina
</locationDeviation>
,
<geoCoordinate id="EE6269CCFFDC737F374EFB38E690FB45" box="[1315,1431,1154,1180]" degrees="13" direction="south" minutes="15.3" orientation="latitude" pageId="47" pageNumber="48" precision="92" value="-13.255">13°15.3'S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFFDC737F32FAFB13E21BFB1A" box="[151,284,1193,1219]" degrees="049" direction="east" minutes="37.0" orientation="longitude" pageId="47" pageNumber="48" precision="92" value="49.616665">049°37.0'E</geoCoordinate>
,
<quantity id="4CAEA2EEFFDC737F3344FB13E2ABFB1A" box="[297,428,1193,1219]" metricMagnitude="2" metricUnit="m" metricValue="4.625" metricValueMax="6.0" metricValueMin="3.25" pageId="47" pageNumber="48" unit="m" value="462.5" valueMax="600.0" valueMin="325.0">
<elevation id="007BE838FFDC737F3344FB13E2ABFB1A" box="[297,428,1193,1219]" metricMagnitude="2" metricUnit="m" metricValue="4.625" metricValueMax="6.0" metricValueMin="3.25" pageId="47" pageNumber="48" unit="m" value="462.5" valueMax="600.0" valueMin="325.0">325-600 m</elevation>
</quantity>
, undisturbed lowland forest, coll.
<collectorName id="26A36ADDFFDC737F3129FB13E0DDFB1A" box="[836,986,1193,1219]" pageId="47" pageNumber="48">S. Goodman</collectorName>
,
<date id="FFE829CBFFDC737F3185FB13E782FB1A" box="[1000,1157,1193,1219]" pageId="47" pageNumber="48" value="2001-11-03" valueMax="2001-11-10" valueMin="2001-11-03">
<collectingDate id="EFACD023FFDC737F3185FB13E782FB1A" box="[1000,1157,1193,1219]" pageId="47" pageNumber="48" value="2001-11-03" valueMax="2001-11-10" valueMin="2001-11-03">310.xi.2001</collectingDate>
</date>
</materialsCitation>
, pitfall trap. 5 M,
<materialsCitation id="3B3E0556FFDC737F370CFB12E142FAC9" collectingDate="2003-12-10" collectionCode="CAS" collectorName="B. L. Fisher" elevation="175" latitude="-12.969999" location="Foret d'Ampondrabe" longLatPrecision="21" longitude="49.7" pageId="47" pageNumber="48" specimenCode="BLF 9973" specimenCount="1">
<emphasis id="B922D319FFDC737F370CFB12E214FB30" bold="true" pageId="47" pageNumber="48">
<collectionCode id="ED4797CEFFDC737F370CFB12E69CFB1B" box="[1377,1435,1192,1218]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="47" pageNumber="48" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFDC737F32FAFB75E214FB30" box="[151,275,1231,1257]" collectionCode="BLF" pageId="47" pageNumber="48">BLF 9973</specimenCode>
</emphasis>
:
<location id="8E8959D0FFDC737F3348FB75E11BFB30" LSID="urn:lsid:plazi:treatment:03FFBE1DFFDE736332FAF973E192F832:8E8959D0FFDC737F3348FB75E11BFB30" box="[293,540,1231,1257]" latitude="-12.969999" longLatPrecision="21" longitude="49.7" name="Foret d'Ampondrabe" pageId="47" pageNumber="48">Forêt d'Ampondrabe</location>
,
<locationDeviation id="9A5B1629FFDC737F3047FB75E06AFB30" box="[554,877,1231,1257]" location="Daraina" pageId="47" pageNumber="48">
<quantity id="4CAEA2EEFFDC737F3047FB75E18CFB30" box="[554,651,1231,1257]" metricMagnitude="4" metricUnit="m" metricValue="2.63" pageId="47" pageNumber="48" unit="km" value="26.3">26.3 km</quantity>
10° NNE Daraina
</locationDeviation>
,
<geoCoordinate id="EE6269CCFFDC737F3117FB75E706FB30" box="[890,1025,1231,1257]" degrees="12" direction="south" minutes="58" orientation="latitude" pageId="47" pageNumber="48" precision="15" seconds="12" value="-12.969999">12°58'12&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFFDC737F367DFB75E7A0FB30" box="[1040,1191,1231,1257]" degrees="049" direction="east" minutes="42" orientation="longitude" pageId="47" pageNumber="48" precision="15" seconds="00" value="49.7">049°42'00&quot;E</geoCoordinate>
,
<quantity id="4CAEA2EEFFDC737F36DBFB75E602FB30" box="[1206,1285,1231,1257]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="47" pageNumber="48" unit="m" value="175.0">
<elevation id="007BE838FFDC737F36DBFB75E602FB30" box="[1206,1285,1231,1257]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="47" pageNumber="48" unit="m" value="175.0">175 m</elevation>
</quantity>
, tropical dry forest, coll.
<collectorName id="26A36ADDFFDC737F334EFB4CE2B7FAC9" box="[291,432,1270,1296]" pageId="47" pageNumber="48">B. L. Fisher</collectorName>
,
<date id="FFE829CBFFDC737F33D1FB4CE142FAC9" box="[444,581,1270,1296]" pageId="47" pageNumber="48" value="2003-12-10">
<collectingDate id="EFACD023FFDC737F33D1FB4CE142FAC9" box="[444,581,1270,1296]" pageId="47" pageNumber="48" value="2003-12-10">10.xii.2003</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFDC737F32ABFAA6E087FAEF" box="[198,896,1308,1334]" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8BE90F0BFFDC737F32ABFAA6E087FAEF" blockId="47.[151,1437,1153,2030]" box="[198,896,1308,1334]" pageId="47" pageNumber="48">
<emphasis id="B922D319FFDC737F32ABFAA6E26EFAEF" bold="true" box="[198,361,1308,1334]" pageId="47" pageNumber="48">Distribution:</emphasis>
Lowland dry forests around Daraina (Fig. 5).
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFDC736032ABFAF9E29DFAE7" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="8BE90F0BFFDC737F32ABFAF9E05AF8F4" blockId="47.[151,1437,1153,2030]" pageId="47" pageNumber="48">
<emphasis id="B922D319FFDC737F32ABFAF9E24FFA84" bold="true" box="[198,328,1347,1373]" pageId="47" pageNumber="48">Diagnosis:</emphasis>
Up to
<quantity id="4CAEA2EEFFDC737F33FAFAF9E2EFFA84" box="[407,488,1347,1373]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="47" pageNumber="48" unit="mm" value="44.0">44 mm</quantity>
long. Males smaller than females. Color light brown, posterior margin of tergites and anal shield laterally with black pattern. Head, collum and appendages dark olive green. Texture of tergites smooth and glabrous. Antennomeres 15 with sclerotized teeth (Fig. 33A), disc with 3585 apical cones (Fig. 33B), all antennomeres without groove (Fig. 33A). Spination of legs variable, in
<typeStatus id="54EDB1A9FFDC737F3629FA0DE7ADFA08" box="[1092,1194,1463,1489]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
first two pair of legs without apical spine. All stigmatic plates completely covered with numerous sclerotized spines. Movable finger of posterior telopods weakly curved with two large, non-sclerotized lobes and
<emphasis id="B922D319FFDC737F36F0F9BFE7DFF9C7" box="[1181,1240,1541,1566]" italics="true" pageId="47" pageNumber="48">circa</emphasis>
four sclerotized spines (Fig. 32C), posterior side with up to 20 sclerotized teeth (Fig. 32D). Fixed finger slightly shorter and more slender than movable finger, basally with single large, membranous lobe and sclerotized spine. First podomere of anterior telopods large, longer than third podomere and width up to 1.7 times larger than its height (Fig. 32A). Two stridulation ribs on each male harp (Fig. 32A). Anal shield with two locking carinae, shape and length variable, in
<typeStatus id="54EDB1A9FFDC737F3382F97CE152F939" box="[495,597,1734,1760]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
both carinae short, second even more weakly developed and as long as first. Female with 3 or 4 stridulation ribs on each half of washboard. Shape of operculum variable, not extending beyond coxa and apically at least weakly notched.
</paragraph>
<paragraph id="8BE90F0BFFDC737F32ABF883E2ADF837" blockId="47.[151,1437,1153,2030]" pageId="47" pageNumber="48">
<emphasis id="B922D319FFDC737F32ABF883E28CF88A" bold="true" box="[198,395,1849,1875]" pageId="47" pageNumber="48">Similar species:</emphasis>
Coloration, inconstant shape of the locking carinae of the anal shield, shape of anterior telopods and the absence of an apical spine on the third pair of legs are shared with the possible sister-species
<taxonomicName id="4C567488FFDC737F32FAF832E1A5F878" authority="(de Saussure &amp; Zehntner, 1897)" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[151,674,1927,1953]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="priapus">
<emphasis id="B922D319FFDC737F32FAF832E215F878" box="[151,274,1928,1953]" italics="true" pageId="47" pageNumber="48">Z. priapus</emphasis>
(
<bibRefCitation id="EFC772FAFFDC737F3348F83DE19EF878" author="de Saussure, H. &amp; Zehntner, L." box="[293,665,1927,1953]" pageId="47" pageNumber="48" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. &amp; Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure &amp; Zehntner, 1897</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C567488FFDC737F30DDF832E078F878" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[688,895,1928,1953]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pseudopriapus" status="sp. nov.">
<emphasis id="B922D319FFDC737F30DDF832E078F878" box="[688,895,1928,1953]" italics="true" pageId="47" pageNumber="48">Z. pseudopriapus</emphasis>
</taxonomicName>
<emphasis id="B922D319FFDC737F31E5F83DE0CAF878" bold="true" box="[904,973,1927,1953]" pageId="47" pageNumber="48">
<taxonomicNameLabel id="A2116E62FFDC737F31E5F83DE0CAF878" box="[904,973,1927,1953]" pageId="47" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C567488FFDC737F361CF832E7EAF878" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[1137,1261,1928,1953]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="priapus">
<emphasis id="B922D319FFDC737F361CF832E7EAF878" box="[1137,1261,1928,1953]" italics="true" pageId="47" pageNumber="48">Z. priapus</emphasis>
</taxonomicName>
in size, in the presence longer and more slender leg podomeres, far longer antenna (Fig. 33A), and different shaped anterior telopods (Figs 32A, B).
</paragraph>
<paragraph id="8BE90F0BFFC3736032FAFB04E29DFAE7" blockId="48.[151,1437,1214,1342]" pageId="48" pageNumber="49">
<emphasis id="B922D319FFC3736032FAFB04E22AFB0F" bold="true" box="[151,301,1214,1238]" pageId="48" pageNumber="49">FIGURE 032</emphasis>
.
<taxonomicName id="4C567488FFC373603357FB05E170FB0F" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[314,631,1215,1238]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="pseudopriapus" status="sp. nov.">
<emphasis id="B922D319FFC373603357FB05E170FB0F" box="[314,631,1215,1238]" italics="true" pageId="48" pageNumber="49">Zoosphaerium pseudopriapus</emphasis>
</taxonomicName>
<emphasis id="B922D319FFC373603010FB04E1BEFB0F" bold="true" box="[637,697,1214,1238]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A2116E62FFC373603010FB04E1BEFB0F" box="[637,697,1214,1238]" pageId="48" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFC3736030A8FB05E022FB0F" box="[709,805,1215,1238]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
,
<emphasis id="B922D319FFC37360315DFB04E044FB0F" bold="true" box="[816,835,1214,1238]" pageId="48" pageNumber="49">A</emphasis>
: right side of anterior telopod, anterior view;
<emphasis id="B922D319FFC373603745FB04E644FB0F" bold="true" box="[1320,1347,1214,1238]" pageId="48" pageNumber="49">B:</emphasis>
anterior telopod, last two podomeres, lateral view;
<emphasis id="B922D319FFC373603034FB5BE16BFB20" bold="true" box="[601,620,1249,1273]" pageId="48" pageNumber="49">C</emphasis>
: posterior telopod, anterior view;
<emphasis id="B922D319FFC3736031BBFB5BE0EEFB20" bold="true" box="[982,1001,1249,1273]" pageId="48" pageNumber="49">D</emphasis>
: posterior telopod, chela, posterior view. Abbreviations:
<emphasis id="B922D319FFC373603353FABEE263FAC5" bold="true" box="[318,356,1284,1308]" pageId="48" pageNumber="49">cav</emphasis>
= cavity;
<emphasis id="B922D319FFC3736033B8FABEE2F9FAC5" bold="true" box="[469,510,1284,1308]" pageId="48" pageNumber="49">cr-t</emphasis>
= crenulated teeth;
<emphasis id="B922D319FFC3736030B4FABEE1F0FAC5" bold="true" box="[729,759,1284,1308]" pageId="48" pageNumber="49">ml</emphasis>
= membranous lobe;
<emphasis id="B922D319FFC37360318BFABEE70FFAC5" bold="true" box="[998,1032,1284,1308]" pageId="48" pageNumber="49">s-p</emphasis>
= sclerotized spot;
<emphasis id="B922D319FFC37360368DFABEE605FAC5" bold="true" box="[1248,1282,1284,1308]" pageId="48" pageNumber="49">SR</emphasis>
= stridulation ribs. Scale bars =
<quantity id="4CAEA2EEFFC373603338FA9DE291FAE7" box="[341,406,1319,1342]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFC3736332ABFAD6E192F832" lastPageId="51" lastPageNumber="52" pageId="48" pageNumber="49" type="description">
<paragraph id="8BE90F0BFFC3736032ABFAD6E7C7FA0D" blockId="48.[151,1436,1388,2032]" pageId="48" pageNumber="49">
<emphasis id="B922D319FFC3736032ABFAD6E267FA5F" bold="true" box="[198,352,1388,1414]" pageId="48" pageNumber="49">Description:</emphasis>
<emphasis id="B922D319FFC373603305FAD7E2F2FA5F" box="[360,501,1389,1414]" italics="true" pageId="48" pageNumber="49">Body length</emphasis>
: Males (
<specimenCount id="9D50C482FFC373603031FAD6E005FA5F" box="[604,770,1388,1414]" pageId="48" pageNumber="49" type="generic">16 specimens</specimenCount>
): length up to
<quantity id="4CAEA2EEFFC3736031C5FAD6E0FAFA5F" box="[936,1021,1388,1414]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="48" pageNumber="49" unit="mm" value="33.0">33 mm</quantity>
, width of thoracic shield up to
<quantity id="4CAEA2EEFFC373603705FAD6E3CFFA74" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="48" pageNumber="49" unit="mm" value="18.4">18.4 mm</quantity>
, height of thoracic shield up to
<quantity id="4CAEA2EEFFC373603025FA29E1A0FA74" box="[584,679,1427,1453]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="48" pageNumber="49" unit="mm" value="9.3">9.3 mm</quantity>
. Females (
<specimenCount id="9D50C482FFC373603145FA29E0C6FA74" box="[808,961,1427,1453]" pageId="48" pageNumber="49" type="generic">4 specimens</specimenCount>
): length up to
<quantity id="4CAEA2EEFFC37360361DFA29E7C0FA74" box="[1136,1223,1427,1453]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="48" pageNumber="49" unit="mm" value="44.0">44 mm</quantity>
, width up to
<quantity id="4CAEA2EEFFC373603705FA29E3CFFA0D" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="48" pageNumber="49" unit="mm" value="21.1">21.1 mm</quantity>
, height up to
<quantity id="4CAEA2EEFFC37360330BFA00E2D6FA0D" box="[358,465,1466,1492]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="48" pageNumber="49" unit="mm" value="11.8">11.8 mm</quantity>
. Females more voluminous and
<emphasis id="B922D319FFC373603121FA00E080FA0A" box="[844,903,1466,1491]" italics="true" pageId="48" pageNumber="49">circa</emphasis>
<quantity id="4CAEA2EEFFC3736031E3FA00E0E6FA0D" box="[910,993,1466,1492]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="mm" value="10.0">10 mm</quantity>
longer than males.
</paragraph>
<paragraph id="8BE90F0BFFC3736032FAFA5BE75FFA23" blockId="48.[151,1436,1388,2032]" box="[151,1112,1504,1530]" pageId="48" pageNumber="49">
<emphasis id="B922D319FFC3736032FAFA5BE3F9FA23" box="[151,254,1505,1530]" italics="true" pageId="48" pageNumber="49">Habitus:</emphasis>
Width of specimens seems to be unusually great in relation to the length.
</paragraph>
<paragraph id="8BE90F0BFFC3736032AAF9BCE068F9B4" blockId="48.[151,1436,1388,2032]" pageId="48" pageNumber="49">
<emphasis id="B922D319FFC3736032AAF9BCE256F9C6" box="[199,337,1542,1567]" italics="true" pageId="48" pageNumber="49">Coloration:</emphasis>
Faded in alcohol. Tergites light brown, through posterior body end dark black coloration pattern, getting much thicker towards paratergite tips. Posterior end of anal shield also dark. Head and collum dark olive-green to blackish. Antenna and legs olive-greenish.
</paragraph>
<paragraph id="8BE90F0BFFC3736032AAF9C0E03DF8D1" blockId="48.[151,1436,1388,2032]" pageId="48" pageNumber="49">
<emphasis id="B922D319FFC3736032AAF9C0E238F94A" box="[199,319,1658,1683]" italics="true" pageId="48" pageNumber="49">Antennae:</emphasis>
Antenna long, reaching up to 4th tergite. Length of antennomeres: 1&lt;2&gt;3&gt;4=5&lt;6 (Fig. 33A), sixth antennomere longest, bearing disc with 3586 sensory cones (Fig. 33B). First antennomere remarkably broader than the others, but short and without groove, apically with row of
<emphasis id="B922D319FFC373603647F97DE60BF939" box="[1066,1292,1735,1760]" italics="true" pageId="48" pageNumber="49">sensilla basiconica</emphasis>
(Fig. 33A). Sclerotized teeth at base of antennomeres 15 (Fig. 33A).
</paragraph>
<paragraph id="8BE90F0BFFC3736032AAF8AFE055F88C" blockId="48.[151,1436,1388,2032]" pageId="48" pageNumber="49">
<emphasis id="B922D319FFC3736032AAF8AFE230F8F7" box="[199,311,1813,1838]" italics="true" pageId="48" pageNumber="49">Mandible</emphasis>
: Thin, five rows of pectinate lamellae, number of teeth declining apically to proximally (Fig. 34). Condylus with single step near apical margin (Fig. 34).
</paragraph>
<paragraph id="8BE90F0BFFC3736032AAF8D8E086F87B" blockId="48.[151,1436,1388,2032]" pageId="48" pageNumber="49">
<emphasis id="B922D319FFC3736032AAF8D8E296F8A2" box="[199,401,1890,1915]" italics="true" pageId="48" pageNumber="49">Gnathochilarium</emphasis>
: Lateral of palpi four sensory cones, located together (Fig. 33C). Depression on posterior side of palpi with numerous sensory cones (Fig. 33C).
</paragraph>
<paragraph id="8BE90F0BFFC3736032AAF815E799F810" blockId="48.[151,1436,1388,2032]" box="[199,1182,1967,1993]" pageId="48" pageNumber="49">
<emphasis id="B922D319FFC3736032AAF815E220F811" box="[199,295,1967,1992]" italics="true" pageId="48" pageNumber="49">Collum:</emphasis>
Median part of collum glabrous, in some specimens with 4 or 5 short hairs.
</paragraph>
<paragraph id="8BE90F0BFFC3736032AAF86CE06CF829" blockId="48.[151,1436,1388,2032]" box="[199,875,2006,2032]" pageId="48" pageNumber="49">
<emphasis id="B922D319FFC3736032AAF86CE279F836" box="[199,382,2006,2031]" italics="true" pageId="48" pageNumber="49">Thoracic shield</emphasis>
: Surface similar to those of other tergites.
</paragraph>
<paragraph id="8BE90F0BFFC2736132FAF8D3E2F4F831" blockId="49.[151,1436,1897,2025]" pageId="49" pageNumber="50">
<emphasis id="B922D319FFC2736132FAF8D3E228F858" bold="true" box="[151,303,1897,1921]" pageId="49" pageNumber="50">FIGURE 033</emphasis>
.
<taxonomicName id="4C567488FFC273613352F8D0E178F858" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[319,639,1898,1921]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pseudopriapus" status="sp. nov.">
<emphasis id="B922D319FFC273613352F8D0E178F858" box="[319,639,1898,1921]" italics="true" pageId="49" pageNumber="50">Zoosphaerium pseudopriapus</emphasis>
</taxonomicName>
<emphasis id="B922D319FFC2736130E5F8D3E1C1F858" bold="true" box="[648,710,1897,1921]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="A2116E62FFC2736130E5F8D3E1C1F858" box="[648,710,1897,1921]" pageId="49" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, SEM,
<typeStatus id="54EDB1A9FFC273613171F8D3E07DF859" box="[796,890,1897,1920]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
,
<emphasis id="B922D319FFC2736131EAF8D3E0A4F858" bold="true" box="[903,931,1897,1921]" pageId="49" pageNumber="50">A:</emphasis>
antenna, lateral view, arrow points to
<emphasis id="B922D319FFC273613727F8D0E20DF87A" italics="true" pageId="49" pageNumber="50">sensilla basiconica</emphasis>
;
<emphasis id="B922D319FFC273613376F831E22AF87A" bold="true" box="[283,301,1931,1955]" pageId="49" pageNumber="50">B</emphasis>
: antenna, apical view on disc;
<emphasis id="B922D319FFC2736130E9F831E190F87A" bold="true" box="[644,663,1931,1955]" pageId="49" pageNumber="50">C</emphasis>
: gnathochilarium, ventral view. Abbreviations:
<emphasis id="B922D319FFC2736136CEF831E7DDF87A" bold="true" box="[1187,1242,1931,1955]" pageId="49" pageNumber="50">endo</emphasis>
= endochilarium;
<emphasis id="B922D319FFC2736132FAF814E3B8F81F" bold="true" box="[151,191,1966,1990]" pageId="49" pageNumber="50">Epi</emphasis>
= tentorium?;
<emphasis id="B922D319FFC273613309F814E27BF81F" bold="true" box="[356,380,1966,1990]" pageId="49" pageNumber="50">iP</emphasis>
= inner palpus;
<emphasis id="B922D319FFC27361305EF814E15FF81F" bold="true" box="[563,600,1966,1990]" pageId="49" pageNumber="50">mp</emphasis>
= central pads or median palps on endochilarium;
<emphasis id="B922D319FFC2736136E6F814E7AAF81F" bold="true" box="[1163,1197,1966,1990]" pageId="49" pageNumber="50">rlp</emphasis>
= rudimentary lateral palpus;
<emphasis id="B922D319FFC273613287F86BE204F830" bold="true" box="[234,259,2001,2025]" pageId="49" pageNumber="50">sb</emphasis>
=
<emphasis id="B922D319FFC27361334DF868E2EBF830" box="[288,492,2002,2025]" italics="true" pageId="49" pageNumber="50">sensilla basiconica</emphasis>
.
</paragraph>
<paragraph id="8BE90F0BFFC1736232FAFB65E623FAC0" blockId="50.[151,1435,1247,1306]" pageId="50" pageNumber="51">
<emphasis id="B922D319FFC1736232FAFB65E22AFB2E" bold="true" box="[151,301,1247,1271]" pageId="50" pageNumber="51">FIGURE 034</emphasis>
.
<taxonomicName id="4C567488FFC173623356FB5AE17EFB2E" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[315,633,1248,1271]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="pseudopriapus" status="sp. nov.">
<emphasis id="B922D319FFC173623356FB5AE17EFB2E" box="[315,633,1248,1271]" italics="true" pageId="50" pageNumber="51">Zoosphaerium pseudopriapus</emphasis>
</taxonomicName>
<emphasis id="B922D319FFC1736230EDFB65E1BBFB2E" bold="true" box="[640,700,1247,1271]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="A2116E62FFC1736230EDFB65E1BBFB2E" box="[640,700,1247,1271]" pageId="50" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, SEM,
<typeStatus id="54EDB1A9FFC173623163FB65E06AFB2F" box="[782,877,1247,1270]" pageId="50" pageNumber="51" type="paratype">paratype</typeStatus>
, right mandible, mesal view. Abbreviations:
<emphasis id="B922D319FFC173623726FB65E675FB2E" bold="true" box="[1355,1394,1247,1271]" pageId="50" pageNumber="51">3iT</emphasis>
= 3 inner teeth;
<emphasis id="B922D319FFC17362337BFAB8E231FAC3" bold="true" box="[278,310,1282,1306]" pageId="50" pageNumber="51">Co</emphasis>
= condylus;
<emphasis id="B922D319FFC1736233ACFAB8E2D8FAC3" bold="true" box="[449,479,1282,1306]" pageId="50" pageNumber="51">eT</emphasis>
= external tooth;
<emphasis id="B922D319FFC1736230F0FAB8E1B2FAC3" bold="true" box="[669,693,1282,1306]" pageId="50" pageNumber="51">iP</emphasis>
= inner palpus;
<emphasis id="B922D319FFC17362310FFAB8E080FAC3" bold="true" box="[866,903,1282,1306]" pageId="50" pageNumber="51">mp</emphasis>
= molar plate;
<emphasis id="B922D319FFC173623647FAB8E74DFAC3" bold="true" box="[1066,1098,1282,1306]" pageId="50" pageNumber="51">pL</emphasis>
= pectinate lamella.
</paragraph>
<paragraph id="8BE90F0BFFC1736232AAFAF2E7CFFABB" blockId="50.[151,1436,1352,2035]" box="[199,1224,1352,1378]" pageId="50" pageNumber="51">
<emphasis id="B922D319FFC1736232AAFAF2E224FAB8" box="[199,291,1352,1377]" italics="true" pageId="50" pageNumber="51">Tergites</emphasis>
: Paratergites only slightly projecting posteriorly. Tergites smooth and glabrous.
</paragraph>
<paragraph id="8BE90F0BFFC1736232AAFAD5E06BFA0C" blockId="50.[151,1436,1352,2035]" pageId="50" pageNumber="51">
<emphasis id="B922D319FFC1736232AAFAD5E267FA51" box="[199,352,1391,1416]" italics="true" pageId="50" pageNumber="51">Endotergum:</emphasis>
Inner area with conical spines and long hair. Between inner area and marginal brim no cuticular impressions (Fig. 35D). Up to three rows of marginal bristles, standing closely to one another. Marginal bristles extending beyond tergite margin (Fig. 35D).
</paragraph>
<paragraph id="8BE90F0BFFC1736232AAFA58E2DCF993" blockId="50.[151,1436,1352,2035]" pageId="50" pageNumber="51">
<emphasis id="B922D319FFC1736232AAFA58E251FA22" box="[199,342,1506,1531]" italics="true" pageId="50" pageNumber="51">Anal shield:</emphasis>
Well-rounded, neither bell-shaped nor tapered, reaching its hindmost point slightly above its lower side. Underside carries two black locking carinae, anterior one small, posterior carina as large as first, but only weakly developed.
</paragraph>
<paragraph id="8BE90F0BFFC1736232AAF9EDE667F964" blockId="50.[151,1436,1352,2035]" pageId="50" pageNumber="51">
<emphasis id="B922D319FFC1736232AAF9EDE20FF9A9" box="[199,264,1623,1648]" italics="true" pageId="50" pageNumber="51">Legs:</emphasis>
Tarsi of first pair with five to seven, second with six to eight and third with eight to ten long ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 321 with curved claws, 1214 ventral spines and one apical spine. Femur 1.9 and tarsi 4.2 times longer than wide.
</paragraph>
<paragraph id="8BE90F0BFFC1736232AAF970E015F8D2" blockId="50.[151,1436,1352,2035]" pageId="50" pageNumber="51">
<emphasis id="B922D319FFC1736232AAF970E297F93A" box="[199,400,1738,1763]" italics="true" pageId="50" pageNumber="51">Stigmatic plates:</emphasis>
First lobe straight, only weakly curved, protruding above coxa. All stigmatic plates covered completely with numerous sclerotized spines.
</paragraph>
<paragraph id="8BE90F0BFFC1736232AAF8A2E641F8A6" blockId="50.[151,1436,1352,2035]" pageId="50" pageNumber="51">
<emphasis id="B922D319FFC1736232AAF8A2E2F8F8E8" box="[199,511,1816,1841]" italics="true" pageId="50" pageNumber="51">Female sexual characters</emphasis>
: Washboard with three or four ribs of medium length on each half. Vulva covering more than 2/3 of coxa surface. Operculum large, ending closely before apical coxal margin. Apical margin of operculum variable, always weakly notched. Mesal plate reaching up to operculum's height.
</paragraph>
<paragraph id="8BE90F0BFFC1736232AAF836E7EFF87C" blockId="50.[151,1436,1352,2035]" box="[199,1256,1931,1957]" pageId="50" pageNumber="51">
<emphasis id="B922D319FFC1736232AAF836E2DDF87C" box="[199,474,1932,1957]" italics="true" pageId="50" pageNumber="51">Male sexual characters</emphasis>
: Anal shield at hindmost point covered with numerous short hairs.
</paragraph>
<paragraph id="8BE90F0BFFC1736332AAF808E3C8FEFF" blockId="50.[151,1436,1352,2035]" lastBlockId="51.[151,1436,152,294]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
<emphasis id="B922D319FFC1736232AAF808E291F812" box="[199,406,1970,1995]" italics="true" pageId="50" pageNumber="51">Anterior telopods</emphasis>
(Figs 32A, B): First podomere remarkably broad, width 1.7 times greater than height, harp with two stridulation ribs, both straight and short. Second podomere process lobe-like and little curved with rounded edges, reaching almost half of third podomere height, apically with sclerotized spots. Third podomere cavity mesally with numerous sclerotized spots and four small spines, three of them located closely towards tip. Lateral of cavity in some specimens with 13 large, crenulated teeth. Tip with one sclerotized spot.
</paragraph>
<paragraph id="8BE90F0BFFC0736332FAFB0AE1B3FAD4" blockId="51.[151,1436,1200,1293]" pageId="51" pageNumber="52">
<emphasis id="B922D319FFC0736332FAFB0AE233FB11" bold="true" box="[151,308,1200,1224]" pageId="51" pageNumber="52">FIGURE 035</emphasis>
. Endoterga, underside of midbody tergites,
<typeStatus id="54EDB1A9FFC07363315EFB0AE09AFB1E" box="[819,925,1200,1223]" pageId="51" pageNumber="52" type="holotype">holotypes</typeStatus>
,
<emphasis id="B922D319FFC0736331C2FB0AE0C5FB11" bold="true" box="[943,962,1200,1224]" pageId="51" pageNumber="52">A</emphasis>
:
<taxonomicName id="4C567488FFC0736331B5FB0BE7FEFB11" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[984,1273,1201,1224]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="endemicum" status="sp. nov.">
<emphasis id="B922D319FFC0736331B5FB0BE7FEFB11" box="[984,1273,1201,1224]" italics="true" pageId="51" pageNumber="52">Zoosphaerium endemicum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFC07363376AFB0AE64DFB11" bold="true" box="[1287,1354,1200,1224]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A2116E62FFC07363376AFB0AE64DFB11" box="[1287,1354,1200,1224]" pageId="51" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B922D319FFC073633732FB0AE676FB11" bold="true" box="[1375,1393,1200,1224]" pageId="51" pageNumber="52">B</emphasis>
:
<taxonomicName id="4C567488FFC0736337EBFB0BE208FB33" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="xerophilum" status="sp. nov.">
<emphasis id="B922D319FFC0736337EBFB0BE208FB33" italics="true" pageId="51" pageNumber="52">Z. xerophilum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFC07363337BFB69E255FB32" bold="true" box="[278,338,1235,1259]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A2116E62FFC07363337BFB69E255FB32" box="[278,338,1235,1259]" pageId="51" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B922D319FFC07363330DFB69E274FB32" bold="true" box="[352,371,1235,1259]" pageId="51" pageNumber="52">C</emphasis>
:
<taxonomicName id="4C567488FFC0736333EFFB69E2E9FB33" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[386,494,1235,1258]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="aureum" status="sp. nov.">
<emphasis id="B922D319FFC0736333EFFB69E2E9FB33" box="[386,494,1235,1258]" italics="true" pageId="51" pageNumber="52">Z. aureum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFC073633398FB69E136FB32" bold="true" box="[501,561,1235,1259]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A2116E62FFC073633398FB69E136FB32" box="[501,561,1235,1259]" pageId="51" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B922D319FFC07363302DFB69E154FB32" bold="true" box="[576,595,1235,1259]" pageId="51" pageNumber="52">D</emphasis>
:
<taxonomicName id="4C567488FFC07363300CFB69E01DFB33" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[609,794,1235,1258]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pseudopriapus" status="sp. nov.">
<emphasis id="B922D319FFC07363300CFB69E01DFB33" box="[609,794,1235,1258]" italics="true" pageId="51" pageNumber="52">Z. pseudopriapus</emphasis>
</taxonomicName>
<emphasis id="B922D319FFC07363314CFB69E05AFB32" bold="true" box="[801,861,1235,1259]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A2116E62FFC07363314CFB69E05AFB32" box="[801,861,1235,1259]" pageId="51" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Abbreviations:
<emphasis id="B922D319FFC073633664FB69E71BFB32" bold="true" box="[1033,1052,1235,1259]" pageId="51" pageNumber="52">ci</emphasis>
= cuticular impressions;
<emphasis id="B922D319FFC073633746FB69E64FFB32" bold="true" box="[1323,1352,1235,1259]" pageId="51" pageNumber="52">IA</emphasis>
= inner area;
<emphasis id="B922D319FFC0736332BCFB4FE3F1FAD4" bold="true" box="[209,246,1269,1293]" pageId="51" pageNumber="52">mb</emphasis>
= marginal bristles;
<emphasis id="B922D319FFC0736333BEFB4FE2F2FAD4" bold="true" box="[467,501,1269,1293]" pageId="51" pageNumber="52">mr</emphasis>
= marginal ridge.
</paragraph>
<paragraph id="8BE90F0BFFC0736332AAFAFAE1DCF9C2" blockId="51.[151,1436,1343,2027]" pageId="51" pageNumber="52">
<emphasis id="B922D319FFC0736332AAFAFAE2A4FA80" box="[199,419,1344,1369]" italics="true" pageId="51" pageNumber="52">Posterior telopods</emphasis>
: Movable finger of chela weakly curved; hollowed-out inner margin with two nonsclerotized lobes and
<emphasis id="B922D319FFC0736333F1FADCE2D0FAA6" box="[412,471,1382,1407]" italics="true" pageId="51" pageNumber="52">circa</emphasis>
four smaller, sclerotized spines, posterior aspect with
<emphasis id="B922D319FFC073633606FADCE7A1FAA6" box="[1131,1190,1382,1407]" italics="true" pageId="51" pageNumber="52">circa</emphasis>
22 crenulated teeth. Fixed finger only half as wide as movable finger, with almost straight tip, basally with single membranous lobe and spine, on inner margin broadly covered with numerous sclerotized spots. First podomere on both sides almost glabrous, second podomere except for fixed finger on both sides covered with numerous hair. Third podomere only basally with numerous hair.
</paragraph>
<paragraph id="8BE90F0BFFC0736332ABF99DE0DAF8F0" blockId="51.[151,1436,1343,2027]" pageId="51" pageNumber="52">
<emphasis id="B922D319FFC0736332ABF99DE2EEF998" bold="true" box="[198,489,1575,1601]" pageId="51" pageNumber="52">Intraspecific variation:</emphasis>
In no other species of the genus
<taxonomicName id="4C567488FFC073633112F992E72FF998" authorityName="Pocock" authorityYear="1895" box="[895,1064,1576,1601]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B922D319FFC073633112F992E72FF998" box="[895,1064,1576,1601]" italics="true" pageId="51" pageNumber="52">Zoosphaerium</emphasis>
</taxonomicName>
was the observed intraspecific variation as strong as in
<taxonomicName id="4C567488FFC0736333D0F9F4E18DF9BE" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[445,650,1614,1639]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pseudopriapus" status="sp. nov.">
<emphasis id="B922D319FFC0736333D0F9F4E18DF9BE" box="[445,650,1614,1639]" italics="true" pageId="51" pageNumber="52">Z. pseudopriapus</emphasis>
</taxonomicName>
,
<emphasis id="B922D319FFC0736330F6F9F4E1D8F9B1" bold="true" box="[667,735,1614,1640]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A2116E62FFC0736330F6F9F4E1D8F9B1" box="[667,735,1614,1640]" pageId="51" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
. Even in the same specimen, the specificity of the locking carinae on the anal shield may vary. In some specimens, those locking carinae are barely visible at all, while in other those carinae are black and readily visible. Especially the second, more posterior carina is difficult to detect in some specimens. The length of the second podomere of posterior telopods varies to some degree in different males. Even more surprising, the number of sclerotized teeth on the third podomere of anterior telopods may even vary between none and three in the same individual.
</paragraph>
<paragraph id="8BE90F0BFFC0736332ABF88CE74DF8AF" blockId="51.[151,1436,1343,2027]" pageId="51" pageNumber="52">
The number of apical cones at the antenna in the
<typeStatus id="54EDB1A9FFC073633143F88CE092F889" box="[814,917,1846,1872]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
is 86 on the left, but only 62 of the right antenna, while other specimens of the type series have as few as 38 apical cones.
</paragraph>
<paragraph id="8BE90F0BFFC0736332ABF839E192F832" blockId="51.[151,1436,1343,2027]" pageId="51" pageNumber="52">
It is unclear if the differences found among specimens of
<taxonomicName id="4C567488FFC0736331F0F839E76AF845" authorityName="Wesener" authorityYear="2009" box="[925,1133,1923,1948]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pseudopriapus">
<emphasis id="B922D319FFC0736331F0F839E76AF845" box="[925,1133,1923,1948]" italics="true" pageId="51" pageNumber="52">Z. pseudopriapus</emphasis>
</taxonomicName>
from the three different localities Forêt Bekaraoka, Analamozava River and Forêt de Ampondrabe are present because of local intraspecific variation or cryptic speciation.
</paragraph>
</subSubSection>
</treatment>
</document>